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<document id="105020CB7EAD47CDDA09BC7C8377F530" ID-DOI="10.5852/ejt.2024.935.2547" ID-ISSN="2118-9773" ID-Zenodo-Dep="11284619" ID-ZooBank="C6FAFB3A-1B4B-4245-A424-0E4EDDA8AF05" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1716568146815" checkinUser="plazi" docAuthor="Jaume-Schinkel, Santiago &amp; Kvifte, Gunnar Mikalsen" docDate="2024" docId="BB4F87F6FFCDF524FDFBFD8DFC85FD97" docLanguage="en" docName="EJT.2024.935.81-100.pdf" docOrigin="European Journal of Taxonomy 935" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2547/11497" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Eugenys clavellata Quate 1996" docType="treatment" docVersion="4" lastPageNumber="85" masterDocId="4776FF8EFFCEF520FFA0FFCBFF93FFB1" masterDocTitle="Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species" masterLastPageNumber="100" masterPageNumber="81" pageNumber="84" updateTime="1717032848221" updateUser="jonas" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5ED2A7350EDEE3945BCA80D54340DF4B">Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species</mods:title>
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<mods:namePart id="6C6D75D663143C9E54BF2068BAB055DA">Jaume-Schinkel, Santiago</mods:namePart>
<mods:nameIdentifier id="85AEBBD045ED9A7A39A865C7B93AA67D" type="LSID">CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096</mods:nameIdentifier>
<mods:affiliation id="916E3BED806EFB79B8BFBDCFAE140F89">Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="F336EC782EA12BF542F17AC1087925A6" type="email">santijaumes@hotmail.com &amp; s.jaume@leibniz-lib.de</mods:nameIdentifier>
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<mods:namePart id="454FF0D15C5C76D9813D6B545BDE095D">Kvifte, Gunnar Mikalsen</mods:namePart>
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<mods:affiliation id="BB95B29F7ECD7809F554C6BB0336AD62">Department of Biosciences and Aquaculture, Nord University, P. O. Box 2501, 7729 Steinkjer, Norway.</mods:affiliation>
<mods:nameIdentifier id="ADBA5BA5B6C71E8169DB5E81528C2798" type="email">gunnar.mikalsen-kvifte@nord.no</mods:nameIdentifier>
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<mods:title id="9D4BB20C16BF83DDF386C00F65830281">European Journal of Taxonomy</mods:title>
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<mods:part id="6891A7B86F5504472F637F30422F0A13">
<mods:date id="90BA36C4FA9DFF2C173C1FD6BA71F1F7">2024</mods:date>
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<mods:number id="A90A5812312B1CC5D83CEF5117E54F0B">2024-05-23</mods:number>
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<mods:number id="1002DDA734E5CEE097073B562EBB5E06">935</mods:number>
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<mods:start id="2E8F55ED1F3C887E56773E8F8168F428">81</mods:start>
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<treatment id="BB4F87F6FFCDF524FDFBFD8DFC85FD97" ID-DOI="http://doi.org/10.5281/zenodo.11284641" ID-Zenodo-Dep="11284641" LSID="urn:lsid:plazi:treatment:BB4F87F6FFCDF524FDFBFD8DFC85FD97" httpUri="http://treatment.plazi.org/id/BB4F87F6FFCDF524FDFBFD8DFC85FD97" lastPageId="4" lastPageNumber="85" pageId="3" pageNumber="84">
<subSubSection id="7BFC656BFFCDF523FDFBFD8DFC4AFDEE" box="[603,985,581,608]" pageId="3" pageNumber="84" type="nomenclature">
<paragraph id="335936E0FFCDF523FDFBFD8DFC4AFDEE" blockId="3.[603,985,581,643]" box="[603,985,581,608]" pageId="3" pageNumber="84">
<heading id="6811818CFFCDF523FDFBFD8DFC4AFDEE" box="[603,985,581,608]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="84" reason="2">
<taxonomicName id="F4E64D63FFCDF523FDFBFD8DFC4AFDEE" ID-CoL="3C6Z4" authority="Quate, 1996" authorityName="Quate" authorityYear="1996" box="[603,985,581,608]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="clavellata">
<emphasis id="0192EAF2FFCDF523FDFBFD8DFCD2FDD1" bold="true" box="[603,833,581,608]" italics="true" pageId="3" pageNumber="84">Eugenys clavellata</emphasis>
<bibRefCitation id="57774B11FFCDF523FCE9FD8DFC4AFDEE" author="Quate L. W." box="[841,985,581,608]" pageId="3" pageNumber="84" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFCDF523FD64FDA2FCE3FD32" box="[708,880,617,643]" pageId="3" pageNumber="84" type="description">
<paragraph id="335936E0FFCDF523FD64FDA2FCE3FD32" blockId="3.[603,985,581,643]" box="[708,880,617,643]" pageId="3" pageNumber="84">
<figureCitation id="ABDD2A65FFCDF523FD64FDA2FC9EFD32" box="[708,781,617,643]" captionStart="Fig" captionStartId="3.[187,230,1998,2024]" captionTargetBox="[189,1398,1352,1957]" captionTargetId="figure-251@3.[189,1398,1352,1957]" captionTargetPageId="3" captionText="Fig. 1. Distribution map of the species of Eugenys Quate, 1996." figureDoi="http://doi.org/10.5281/zenodo.11284621" httpUri="https://zenodo.org/record/11284621/files/figure.png" pageId="3" pageNumber="84">Figs 1</figureCitation>
,
<figureCitation id="ABDD2A65FFCDF523FCBBFDA2FCD2FD32" box="[795,833,617,643]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[409,1206,265,1842]" captionTargetId="figure-12@5.[361,1227,265,1852]" captionTargetPageId="5" captionText="Fig. 2. A. Eugenys clavellata Quate, 1996, ♂, paratype (USNMENT00857092), genitalia (ventral view). B. Eugenys cymosa Quate, 1999, ♂, paratype (BMNH(E)2001-8), genitalia (ventral view). Photos A by David Pecor, Smithsonian Institute Museum, B by Santiago Jaume-Schinkel. Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284623" httpUri="https://zenodo.org/record/11284623/files/figure.png" pageId="3" pageNumber="84">2A</figureCitation>
,
<figureCitation id="ABDD2A65FFCDF523FCEDFDA2FCE3FD32" box="[845,880,617,643]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="3" pageNumber="84">3C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFCDF523FF1DFD60FDE2FD75" box="[189,625,682,708]" pageId="3" pageNumber="84" type="reference_group">
<paragraph id="335936E0FFCDF523FF1DFD60FDE2FD75" blockId="3.[189,1399,682,743]" box="[189,625,682,708]" pageId="3" pageNumber="84">
<treatmentCitationGroup id="13F611CEFFCDF523FF1DFD60FDE2FD75" box="[189,625,682,708]" pageId="3" pageNumber="84">
<treatmentCitation id="B24710F1FFCDF523FF1DFD60FDFEFD75" author="Quate L. W." box="[189,621,682,708]" page="43" pageId="3" pageNumber="84" year="1996">
<taxonomicName id="F4E64D63FFCDF523FF1DFD60FDFEFD75" ID-CoL="3C6Z4" authority="Quate, 1996: 43" authorityName="Quate" authorityPageNumber="43" authorityYear="1996" box="[189,621,682,708]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="clavellata">
<emphasis id="0192EAF2FFCDF523FF1DFD60FE33FD75" box="[189,416,682,708]" italics="true" pageId="3" pageNumber="84">Eugenys clavellata</emphasis>
<bibRefCitation id="57774B11FFCDF523FE09FD61FDFEFD75" author="Quate L. W." box="[425,621,682,708]" pageId="3" pageNumber="84" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate, 1996: 43</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFCDF523FDD9FD60FAE4FD75" box="[633,1399,682,709]" pageId="3" pageNumber="84" type="materials_examined">
<paragraph id="335936E0FFCDF523FDD9FD60FAE4FD75" blockId="3.[189,1399,682,743]" box="[633,1399,682,709]" pageId="3" pageNumber="84">
<typeStatus id="EC5D8842FFCDF523FDD9FD60FD20FD74" box="[633,691,683,709]" pageId="3" pageNumber="84">Type</typeStatus>
locality:
<collectingCountry id="4BF17670FFCDF523FC86FD61FC38FD75" box="[806,939,682,708]" name="Costa Rica" pageId="3" pageNumber="84">Costa Rica</collectingCountry>
,
<collectingRegion id="F122F802FFCDF523FC19FD61FB8AFD75" box="[953,1049,682,708]" country="Costa Rica" name="Heredia" pageId="3" pageNumber="84">Heredia</collectingRegion>
, Estacion Biologica la Selva.
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFCDF523FF4CFD06FC31FD56" box="[236,930,717,743]" pageId="3" pageNumber="84" type="reference_group">
<paragraph id="335936E0FFCDF523FF4CFD06FC31FD56" blockId="3.[189,1399,682,743]" box="[236,930,717,743]" pageId="3" pageNumber="84">
Additional reference:
<bibRefCitation id="57774B11FFCDF523FE4EFD06FC0CFD56" author="Collantes F. &amp; Martinez-Ortega E." box="[494,927,717,743]" pageId="3" pageNumber="84" pagination="17 - 27" refId="ref6673" refString="Collantes F. &amp; Martinez-Ortega E. 1999. Nuevas citas de especies conocidas de Psychodinae (Diptera: Psychodidae) en Nicaragua. Revista Nicaraguense de Entomologia 46: 17 - 27." type="journal article" year="1999">Collantes &amp; Martinez-Ortega (1999)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFCDF523FF1DFCC5FEA7FBB6" pageId="3" pageNumber="84" type="diagnosis">
<paragraph id="335936E0FFCDF523FF1DFCC5FE5FFC99" blockId="3.[189,460,782,808]" box="[189,460,782,808]" pageId="3" pageNumber="84">
<heading id="6811818CFFCDF523FF1DFCC5FE5FFC99" bold="true" box="[189,460,782,808]" fontSize="11" level="3" pageId="3" pageNumber="84" reason="3">
<emphasis id="0192EAF2FFCDF523FF1DFCC5FE5FFC99" bold="true" box="[189,460,782,808]" pageId="3" pageNumber="84">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="335936E0FFCDF523FF1DFCF6FEA7FBB6" blockId="3.[188,1399,829,1031]" pageId="3" pageNumber="84">
This species can be easily differentiated from all other species of
<taxonomicName id="F4E64D63FFCDF523FC61FCF5FBB6FCE6" authorityName="Quate" authorityYear="1996" box="[961,1061,830,855]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="0192EAF2FFCDF523FC61FCF5FBB6FCE6" box="[961,1061,830,855]" italics="true" pageId="3" pageNumber="84">Eugenys</emphasis>
</taxonomicName>
by the following characters: hypopods with singular apical tenaculum (
<quantity id="F41E9B05FFCDF523FD08FCAAFC9CFCCB" box="[680,783,864,891]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.556" metricValueMin="2.54" pageId="3" pageNumber="84" unit="in" value="12.0" valueMax="14.0" valueMin="10.0">10–14 in</quantity>
<taxonomicName id="F4E64D63FFCDF523FCB3FCAAFC38FCCB" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[787,939,864,890]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="0192EAF2FFCDF523FCB3FCAAFC38FCCB" box="[787,939,864,890]" italics="true" pageId="3" pageNumber="84">E. singularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA15789FFCDF523FC10FCAAFB9AFCCA" box="[944,1033,865,891]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="F4E64D63FFCDF523FB9EFCAAFB2BFCCB" authorityName="Quate" authorityYear="1999" box="[1086,1208,865,890]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="cymosa">
<emphasis id="0192EAF2FFCDF523FB9EFCAAFB2BFCCB" box="[1086,1208,865,890]" italics="true" pageId="3" pageNumber="84">E. cymosa</emphasis>
</taxonomicName>
, one in
<taxonomicName id="F4E64D63FFCDF523FAB0FCAAFAE5FCCB" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[1296,1398,865,890]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="micra" status="sp. nov.">
<emphasis id="0192EAF2FFCDF523FAB0FCAAFAE5FCCB" box="[1296,1398,865,890]" italics="true" pageId="3" pageNumber="84">E. micra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA15789FFCDF523FF1CFC4FFE8DFC2F" box="[188,286,900,926]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="F4E64D63FFCDF523FE8DFC4EFE4DFC2F" authorityName="Quate" authorityYear="1999" box="[301,478,901,926]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="0192EAF2FFCDF523FE8DFC4EFE4DFC2F" box="[301,478,901,926]" italics="true" pageId="3" pageNumber="84">E. panamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="F4E64D63FFCDF523FDBFFC4EFD09FC2F" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[543,666,900,926]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="upsilon" status="sp. nov.">
<emphasis id="0192EAF2FFCDF523FDBFFC4EFD09FC2F" box="[543,666,900,926]" italics="true" pageId="3" pageNumber="84">E. upsilon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA15789FFCDF523FD06FC4FFC9BFC2F" box="[678,776,900,926]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
); antennal flagellomeres without patch of darkened sensilla (present in
<taxonomicName id="F4E64D63FFCDF523FE3FFC63FD84FC70" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[415,535,935,961]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="upsilon" status="sp. nov.">
<emphasis id="0192EAF2FFCDF523FE3FFC63FD84FC70" box="[415,535,935,961]" italics="true" pageId="3" pageNumber="84">E. upsilon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA15789FFCDF523FDBFFC6CFDEAFC70" box="[543,633,935,961]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="F4E64D63FFCDF523FD11FC63FCCDFC70" authorityName="Quate" authorityYear="1999" box="[689,862,936,961]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="0192EAF2FFCDF523FD11FC63FCCDFC70" box="[689,862,936,961]" italics="true" pageId="3" pageNumber="84">E. panamensis</emphasis>
</taxonomicName>
, absent in remaining species); aedeagus long, more than half length of ejaculatory apodeme (aedeagus less than half length of ejaculatory apodeme in
<taxonomicName id="F4E64D63FFCDF523FF1DFC25FEB0FBB6" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[189,291,1006,1031]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="micra">
<emphasis id="0192EAF2FFCDF523FF1DFC25FEB0FBB6" box="[189,291,1006,1031]" italics="true" pageId="3" pageNumber="84">E. micra</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFCDF524FF1DFBE5FDAAFEF6" lastPageId="4" lastPageNumber="85" pageId="3" pageNumber="84" type="materials_examined">
<paragraph id="335936E0FFCDF523FF1DFBE5FE3EFBF9" blockId="3.[189,429,1070,1096]" box="[189,429,1070,1096]" pageId="3" pageNumber="84">
<heading id="6811818CFFCDF523FF1DFBE5FE3EFBF9" bold="true" box="[189,429,1070,1096]" fontSize="11" level="3" pageId="3" pageNumber="84" reason="3">
<emphasis id="0192EAF2FFCDF523FF1DFBE5FE3EFBF9" bold="true" box="[189,429,1070,1096]" pageId="3" pageNumber="84">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="335936E0FFCDF524FF4AFB96FDAAFEF6" blockId="3.[188,1399,1117,1250]" lastBlockId="4.[189,1398,265,327]" lastPageId="4" lastPageNumber="85" pageId="3" pageNumber="84">
<materialsCitation id="838E3CBDFFCDF523FF4AFB96FCCEFB0F" collectingDate="1993-07-01" collectingMethod="Malaise trap" collectionCode="USNM" country="Costa Rica" elevation="75" location="Puerto viejo de Sarapaqui" pageId="3" pageNumber="84" specimenCode="USNMENT00857092" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<emphasis id="0192EAF2FFCDF523FF4AFB96FEFAFBC6" bold="true" box="[234,361,1117,1143]" pageId="3" pageNumber="84">
<typeStatus id="EC5D8842FFCDF523FF4AFB96FEFAFBC6" box="[234,361,1117,1143]" pageId="3" pageNumber="84" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="4BF17670FFCDF523FF1DFB4AFEF7FB2A" box="[189,356,1153,1179]" name="Costa Rica" pageId="3" pageNumber="84">COSTA RICA</collectingCountry>
–
<emphasis id="0192EAF2FFCDF523FE20FB4BFE76FB2B" bold="true" box="[384,485,1152,1178]" pageId="3" pageNumber="84">
<collectingRegion id="F122F802FFCDF523FE20FB4BFE76FB2B" box="[384,485,1152,1178]" country="Costa Rica" name="Heredia" pageId="3" pageNumber="84">Heredia</collectingRegion>
</emphasis>
•
<specimenCount id="25E0FD69FFCDF523FE5EFB4AFDBFFB2A" box="[510,556,1153,1179]" count="1" pageId="3" pageNumber="84" type="male">1 ♂</specimenCount>
;
<location id="3639603BFFCDF523FD98FB4AFCF4FB2B" LSID="urn:lsid:plazi:treatment:BB4F87F6FFCDF524FDFBFD8DFC85FD97:3639603BFFCDF523FD98FB4AFCF4FB2B" box="[568,871,1152,1179]" country="Costa Rica" name="Puerto viejo de Sarapaqui" pageId="3" pageNumber="84" stateProvince="Heredia">Puerto viejo de Sarapaqui</location>
,
<location id="3639603BFFCDF523FCD3FB4BFB28FB2B" LSID="urn:lsid:plazi:treatment:BB4F87F6FFCDF524FDFBFD8DFC85FD97:3639603BFFCDF523FCD3FB4BFB28FB2B" box="[883,1211,1152,1178]" country="Costa Rica" name="Estacion Biologica LaSelva" pageId="3" pageNumber="84" stateProvince="Heredia">Estacion Biologica LaSelva</location>
; alt.
<quantity id="F41E9B05FFCDF523FB51FB4BFAFEFB2A" box="[1265,1389,1152,1179]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="3" pageNumber="84" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
<elevation id="B8CBD1D3FFCDF523FB51FB4BFAFEFB2A" box="[1265,1389,1152,1179]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="3" pageNumber="84" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50–100 m</elevation>
</quantity>
.;
<date id="47581020FFCDF523FF1CFB6FFED7FB0F" box="[188,324,1188,1214]" pageId="3" pageNumber="84" value="1993-07-01">
<collectingDate id="571CE9C8FFCDF523FF1CFB6FFED7FB0F" box="[188,324,1188,1214]" pageId="3" pageNumber="84" value="1993-07-01">1 Jul. 1993</collectingDate>
</date>
;
<collectingMethod id="EAA74EF7FFCDF523FEF1FB6FFE74FB0F" box="[337,487,1188,1214]" pageId="3" pageNumber="84">Malaise trap</collectingMethod>
;
<specimenCode id="63409E9BFFCDF523FE54FB68FD68FB0F" box="[500,763,1187,1214]" pageId="3" pageNumber="84">USNMENT00857092</specimenCode>
;
<collectionCode id="55F7AE25FFCDF523FCA8FB6FFCCEFB0F" box="[776,861,1188,1214]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="84" type="Museum">USNM</collectionCode>
</materialsCitation>
•
<materialsCitation id="838E3CBDFFCDF523FCD8FB6FFDB7FB50" collectingDate="1993-07-01" collectingMethod="Malaise trap" collectionCode="USNM" country="Costa Rica" elevation="75" location="Puerto viejo de Sarapaqui" pageId="3" pageNumber="84" specimenCode="USNMENT00857112" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="25E0FD69FFCDF523FCD8FB6FFC3BFB0F" box="[888,936,1188,1214]" count="1" pageId="3" pageNumber="84" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<specimenCode id="63409E9BFFCDF523FF1EFB0CFE57FB53" box="[190,452,1223,1250]" pageId="3" pageNumber="84">USNMENT00857112</specimenCode>
;
<collectionCode id="55F7AE25FFCDF523FE6FFB0CFDB7FB50" box="[463,548,1223,1249]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="84" type="Museum">USNM</collectionCode>
</materialsCitation>
•
<materialsCitation id="838E3CBDFFCDF523FD9AFB0CFBFDFB50" box="[570,1134,1223,1249]" collectingDate="1993-07-01" collectingMethod="Malaise trap" collectionCode="CAS" country="Costa Rica" elevation="75" location="Puerto viejo de Sarapaqui" pageId="3" pageNumber="84" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="25E0FD69FFCDF523FD9AFB0CFDFAFB50" box="[570,617,1223,1249]" count="1" pageId="3" pageNumber="84" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="55F7AE25FFCDF523FB95FB0CFBFDFB50" box="[1077,1134,1223,1249]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="84" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="838E3CBDFFCDF524FB25FB0CFCD4FE92" collectingDate="1993-07-01" collectingMethod="Malaise trap" collectionCode="INBIO, NHM" country="Costa Rica" elevation="75" lastPageId="4" lastPageNumber="85" location="Puerto viejo de Sarapaqui" pageId="3" pageNumber="84" specimenCode="INBIO-CRI0Q1-470625" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="25E0FD69FFCDF523FB25FB0CFB27FB50" box="[1157,1204,1223,1249]" count="1" pageId="3" pageNumber="84" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<specimenCode id="63409E9BFFCAF524FE6FFEC2FD60FE95" box="[463,755,265,292]" pageId="4" pageNumber="85">INBIO-CRI0Q1-470625</specimenCode>
;
<collectionCode id="55F7AE25FFCAF524FCA3FEC2FCD4FE92" box="[771,839,265,291]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="4" pageNumber="85">NHM</collectionCode>
</materialsCitation>
•
<materialsCitation id="838E3CBDFFCAF524FCC7FEC2FDA6FEF6" collectingDate="1993-07-01" collectingMethod="Malaise trap" collectionCode="INBIO, NHM" country="Costa Rica" elevation="75" location="Puerto viejo de Sarapaqui" pageId="4" pageNumber="85" specimenCode="INBIO-CRI0Q1-470323" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<specimenCount id="25E0FD69FFCAF524FCC7FEC2FC0AFE92" box="[871,921,265,291]" count="1" pageId="4" pageNumber="85" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<specimenCode id="63409E9BFFCAF524FF1DFEE7FE72FEF7" box="[189,481,300,326]" pageId="4" pageNumber="85">INBIO-CRI0Q1-470323</specimenCode>
;
<collectionCode id="55F7AE25FFCAF524FE4DFEE6FDA6FEF6" box="[493,565,301,327]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="4" pageNumber="85">NHM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="67996668FFCDF523FF1BF805FC39F859" ID-DOI="http://doi.org/10.5281/zenodo.11284621" ID-Zenodo-Dep="11284621" box="[187,938,1998,2024]" httpUri="https://zenodo.org/record/11284621/files/figure.png" pageId="3" pageNumber="84" startId="3.[187,230,1998,2024]" targetBox="[189,1398,1352,1957]" targetPageId="3">
<paragraph id="335936E0FFCDF523FF1BF805FC39F859" blockId="3.[187,938,1998,2024]" box="[187,938,1998,2024]" pageId="3" pageNumber="84">
<emphasis id="0192EAF2FFCDF523FF1BF805FE9AF859" bold="true" box="[187,265,1998,2024]" pageId="3" pageNumber="84">Fig. 1.</emphasis>
Distribution map of the species of
<taxonomicName id="F4E64D63FFCDF523FD07F804FC35F859" authority="Quate, 1996" authorityName="Quate" authorityYear="1996" box="[679,934,1998,2024]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="0192EAF2FFCDF523FD07F804FC98F859" box="[679,779,1999,2024]" italics="true" pageId="3" pageNumber="84">Eugenys</emphasis>
<bibRefCitation id="57774B11FFCDF523FCB3F805FC35F859" author="Quate L. W." box="[787,934,1998,2024]" pageId="3" pageNumber="84" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="7BFC656BFFCAF524FF1DFEA6FB8BFE07" box="[189,1048,365,438]" pageId="4" pageNumber="85" type="distribution">
<paragraph id="335936E0FFCAF524FF1DFEA6FECBFE36" blockId="4.[189,344,365,391]" box="[189,344,365,391]" pageId="4" pageNumber="85">
<heading id="6811818CFFCAF524FF1DFEA6FECBFE36" bold="true" box="[189,344,365,391]" fontSize="11" level="3" pageId="4" pageNumber="85" reason="3">
<emphasis id="0192EAF2FFCAF524FF1DFEA6FECBFE36" bold="true" box="[189,344,365,391]" pageId="4" pageNumber="85">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="335936E0FFCAF524FF1DFE57FB8BFE07" blockId="4.[189,1048,412,438]" box="[189,1048,412,438]" pageId="4" pageNumber="85">
<collectingCountry id="4BF17670FFCAF524FF1DFE57FED3FE07" box="[189,320,412,438]" name="Costa Rica" pageId="4" pageNumber="85">Costa Rica</collectingCountry>
,
<collectingCountry id="4BF17670FFCAF524FEECFE57FE56FE07" box="[332,453,412,438]" name="Nicaragua" pageId="4" pageNumber="85">Nicaragua</collectingCountry>
(
<bibRefCitation id="57774B11FFCAF524FE73FE57FDF2FE07" author="Quate L. W." box="[467,609,412,438]" pageId="4" pageNumber="85" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate 1996</bibRefCitation>
;
<bibRefCitation id="57774B11FFCAF524FDCDFE57FB98FE07" author="Collantes F. &amp; Martinez-Ortega E." box="[621,1035,412,438]" pageId="4" pageNumber="85" pagination="17 - 27" refId="ref6673" refString="Collantes F. &amp; Martinez-Ortega E. 1999. Nuevas citas de especies conocidas de Psychodinae (Diptera: Psychodidae) en Nicaragua. Revista Nicaraguense de Entomologia 46: 17 - 27." type="journal article" year="1999">Collantes &amp; Martinez-Ortega 1999</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFCAF524FF1DFE16FC85FD97" box="[189,790,477,551]" pageId="4" pageNumber="85" type="biology_ecology">
<paragraph id="335936E0FFCAF524FF1DFE16FEB9FE46" blockId="4.[189,298,477,503]" box="[189,298,477,503]" pageId="4" pageNumber="85">
<heading id="6811818CFFCAF524FF1DFE16FEB9FE46" bold="true" box="[189,298,477,503]" fontSize="11" level="3" pageId="4" pageNumber="85" reason="3">
<emphasis id="0192EAF2FFCAF524FF1DFE16FEB9FE46" bold="true" box="[189,298,477,503]" pageId="4" pageNumber="85">Genetics</emphasis>
</heading>
</paragraph>
<paragraph id="335936E0FFCAF524FF1DFDC6FC85FD97" blockId="4.[189,790,524,551]" box="[189,790,524,551]" pageId="4" pageNumber="85">No specimens were available for DNA sequencing.</paragraph>
</subSubSection>
</treatment>
</document>
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