256 lines
30 KiB
XML
256 lines
30 KiB
XML
<document id="D7E65A2C7757A0C1E19F82D4B587CFBB" ID-DOI="10.5852/ejt.2024.935.2547" ID-ISSN="2118-9773" ID-Zenodo-Dep="11284619" ID-ZooBank="C6FAFB3A-1B4B-4245-A424-0E4EDDA8AF05" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1716568146815" checkinUser="plazi" docAuthor="Jaume-Schinkel, Santiago & Kvifte, Gunnar Mikalsen" docDate="2024" docId="BB4F87F6FFCCF523FDD1FCD4FB0AFDB9" docLanguage="en" docName="EJT.2024.935.81-100.pdf" docOrigin="European Journal of Taxonomy 935" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2547/11497" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Eugenys Quate 1996" docType="treatment" docVersion="4" lastPageNumber="84" masterDocId="4776FF8EFFCEF520FFA0FFCBFF93FFB1" masterDocTitle="Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species" masterLastPageNumber="100" masterPageNumber="81" pageNumber="83" updateTime="1717032848221" updateUser="jonas" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="DEC3E28AD4F2F4E3764F8D25AEC974AB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D74D42DEF0ED7F1B99F12456B34860E7">
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<mods:title id="73214155749CFE83ABC9B723E21A6F57">Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species</mods:title>
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<mods:name id="9B9D92A687FB433AA2AF235244FAABE6" type="personal">
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<mods:roleTerm id="D181E54CE5DA139B37CDBC5E3E8C7E06">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="242A642C7D6E326B8027C9E2D6ABAFC0">Jaume-Schinkel, Santiago</mods:namePart>
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<mods:nameIdentifier id="978952015DAF5E79EF37146381BF2BEC" type="LSID">CB8722CA-9B1E-4BB6-AF0C-1A8BD57CC096</mods:nameIdentifier>
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<mods:affiliation id="1FAE70B57FF16F61212FF7E68A92B4B1">Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="C22E181022B0D22FD4D95E27FA24D30A" type="email">santijaumes@hotmail.com & s.jaume@leibniz-lib.de</mods:nameIdentifier>
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</mods:name>
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<mods:name id="29F765D4E8A050604D62020DDDD66A81" type="personal">
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<mods:roleTerm id="42BA18E118C21400DD27E798C381532F">Author</mods:roleTerm>
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<mods:namePart id="F35E2CE5E94C7F75B16CDC7F2E40FC8A">Kvifte, Gunnar Mikalsen</mods:namePart>
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<mods:nameIdentifier id="0798302C96A0340FEF4B9C55BB94F5AD" type="LSID">A9E0868C-34C9-474C-BFB6-2D35CCBD8371</mods:nameIdentifier>
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<mods:affiliation id="522C0363845566A5AEE4F5941602E7BF">Department of Biosciences and Aquaculture, Nord University, P. O. Box 2501, 7729 Steinkjer, Norway.</mods:affiliation>
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<mods:nameIdentifier id="FC7A556F75C1AA34D604934E34F1DF2C" type="email">gunnar.mikalsen-kvifte@nord.no</mods:nameIdentifier>
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<mods:typeOfResource id="E81CFFD383160D7526E97B17561436AB">text</mods:typeOfResource>
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<mods:title id="D9C7C56B955F73859225F5234D8BB1E3">European Journal of Taxonomy</mods:title>
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<mods:part id="CC63D9D14BFCE41AEC471944A0767AB3">
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<mods:date id="6D282851551DB9322809EFB27B842228">2024</mods:date>
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<mods:number id="A1628B31A917245A6E6D4DCE24D3C102">2024-05-23</mods:number>
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<mods:number id="3841236FF47DBF00CE76D3D8F91586AD">935</mods:number>
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<mods:start id="529976CE2D281A2B9D8E53291483C603">81</mods:start>
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<mods:url id="99A3A9F19B21BF428996A878A4167917">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2547/11497</mods:url>
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<mods:identifier id="8E4EA0F5E124A08BC853471382381CD5" type="DOI">10.5852/ejt.2024.935.2547</mods:identifier>
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<mods:identifier id="58FDE3A9E02D48F1E230A161E3C0592C" type="ISSN">2118-9773</mods:identifier>
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<subSubSection id="7BFC656BFFCCF522FDD1FCD4FC50FC89" box="[625,963,798,825]" pageId="2" pageNumber="83" type="nomenclature">
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<paragraph id="335936E0FFCCF522FDD1FCD4FC50FC89" blockId="2.[189,963,798,860]" box="[625,963,798,825]" pageId="2" pageNumber="83">
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<heading id="6811818CFFCCF522FDD1FCD4FC50FC89" box="[625,963,798,825]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="83" reason="2">
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Genus
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<taxonomicName id="F4E64D63FFCCF522FD63FCD4FC50FC89" ID-CoL="4FLT" authority="Quate, 1996" authorityName="Quate" authorityYear="1996" box="[707,963,798,825]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="0192EAF2FFCCF522FD63FCD4FCBFFC88" bold="true" box="[707,812,799,825]" italics="true" pageId="2" pageNumber="83">Eugenys</emphasis>
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<bibRefCitation id="57774B11FFCCF522FC93FCD4FC50FC89" author="Quate L. W." box="[819,963,798,825]" pageId="2" pageNumber="83" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7BFC656BFFCCF522FF1DFC88FADCFC2C" pageId="2" pageNumber="83" type="reference_group">
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<paragraph id="335936E0FFCCF522FF1DFC88FE7FFCED" blockId="2.[189,963,798,860]" box="[189,492,834,860]" pageId="2" pageNumber="83">
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<treatmentCitationGroup id="13F611CEFFCCF522FF1DFC88FE7FFCED" box="[189,492,834,860]" pageId="2" pageNumber="83">
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<treatmentCitation id="B24710F1FFCCF522FF1DFC88FE7AFCED" author="Quate L. W." box="[189,489,834,860]" page="43" pageId="2" pageNumber="83" year="1996">
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||
<taxonomicName id="F4E64D63FFCCF522FF1DFC88FE7AFCED" ID-CoL="4FLT" authority="Quate, 1996: 43" authorityName="Quate" authorityPageNumber="43" authorityYear="1996" box="[189,489,834,860]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="0192EAF2FFCCF522FF1DFC88FEB2FCED" box="[189,289,835,860]" italics="true" pageId="2" pageNumber="83">Eugenys</emphasis>
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||
<bibRefCitation id="57774B11FFCCF522FE89FC89FE7AFCED" author="Quate L. W." box="[297,489,834,860]" pageId="2" pageNumber="83" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate, 1996: 43</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="335936E0FFCCF522FF1DFC4FFADCFC2C" blockId="2.[189,1359,899,926]" box="[189,1359,899,926]" pageId="2" pageNumber="83">
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<treatmentCitationGroup id="13F611CEFFCCF522FF1DFC4FFADCFC2C" box="[189,1359,899,926]" pageId="2" pageNumber="83">
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<taxonomicName id="F4E64D63FFCCF522FF1DFC4FFEB3FC2C" ID-CoL="4FLT" authorityName="Quate" authorityYear="1996" box="[189,288,900,925]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">
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||
<emphasis id="0192EAF2FFCCF522FF1DFC4FFEB3FC2C" box="[189,288,900,925]" italics="true" pageId="2" pageNumber="83">Eugenys</emphasis>
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||
</taxonomicName>
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||
–
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||
<treatmentCitation id="B24710F1FFCCF522FE9EFC48FD92FC2C" author="Quate L. W." box="[318,513,899,925]" page="432" pageId="2" pageNumber="83" year="1999">
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<bibRefCitation id="57774B11FFCCF522FE9EFC48FD92FC2C" author="Quate L. W." box="[318,513,899,925]" pageId="2" pageNumber="83" pagination="409 - 441" refId="ref7259" refString="Quate L. W. 1999. Taxonomy of neotropical Psychodidae. (Diptera) 3. Psychodines of Barro Colorado Island and San Blas, Panama. Memoirs on Entomology International 14: 409 - 441." type="journal article" year="1999">Quate 1999: 432</bibRefCitation>
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||
</treatmentCitation>
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||
(description of new species). —
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||
<treatmentCitation id="B24710F1FFCCF522FC26FC48FBDDFC2C" author="Kvifte G. M." box="[902,1102,899,925]" page="603" pageId="2" pageNumber="83" year="2018">
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||
<bibRefCitation id="57774B11FFCCF522FC26FC48FBDDFC2C" author="Kvifte G. M." box="[902,1102,899,925]" pageId="2" pageNumber="83" pagination="596 - 605" refId="ref7116" refString="Kvifte G. M. 2018. Molecular phylogeny of moth flies (Diptera, Psychodidae, Psychodinae) revisited, with a revised tribal classification. Systematic Entomology 43 (3): 596 - 605. https: // doi. org / 10.1111 / syen. 12288" type="journal article" year="2018">Kvifte 2018: 603</bibRefCitation>
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||
</treatmentCitation>
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||
(tribal classification).
|
||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7BFC656BFFCCF522FF1DFC0EFC31FBBC" pageId="2" pageNumber="83" type="type_taxon">
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||
<paragraph id="335936E0FFCCF522FF1DFC0EFECBFC6F" blockId="2.[189,344,964,991]" box="[189,344,964,991]" pageId="2" pageNumber="83">
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||
<heading id="6811818CFFCCF522FF1DFC0EFECBFC6F" bold="true" box="[189,344,964,991]" fontSize="11" level="3" pageId="2" pageNumber="83" reason="3">
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||
<emphasis id="0192EAF2FFCCF522FF1DFC0EFECBFC6F" bold="true" box="[189,344,964,991]" pageId="2" pageNumber="83">
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||
<typeStatus id="EC5D8842FFCCF522FF1DFC0EFF69FC6E" box="[189,250,965,991]" pageId="2" pageNumber="83">Type</typeStatus>
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species
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="335936E0FFCCF522FF1DFC3FFC31FBBC" blockId="2.[189,930,1011,1037]" box="[189,930,1011,1037]" pageId="2" pageNumber="83">
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||
<taxonomicName id="F4E64D63FFCCF522FF1DFC3FFDE3FBBC" ID-CoL="3C6Z4" authority="Quate, 1996: 432" authorityName="Quate" authorityPageNumber="432" authorityYear="1996" box="[189,624,1011,1037]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="clavellata">
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<emphasis id="0192EAF2FFCCF522FF1DFC3FFE0DFBBC" box="[189,414,1011,1037]" italics="true" pageId="2" pageNumber="83">Eugenys clavellata</emphasis>
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<bibRefCitation id="57774B11FFCCF522FE06FC38FDE3FBBC" author="Quate L. W." box="[422,624,1011,1037]" pageId="2" pageNumber="83" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate, 1996: 432</bibRefCitation>
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</taxonomicName>
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(by original designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="7BFC656BFFCCF522FF1DFBFFFAE4FA26" pageId="2" pageNumber="83" type="diagnosis">
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||
<paragraph id="335936E0FFCCF522FF1DFBFFFE5FFBFF" blockId="2.[189,460,1076,1102]" box="[189,460,1076,1102]" pageId="2" pageNumber="83">
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||
<heading id="6811818CFFCCF522FF1DFBFFFE5FFBFF" bold="true" box="[189,460,1076,1102]" fontSize="11" level="3" pageId="2" pageNumber="83" reason="3">
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<emphasis id="0192EAF2FFCCF522FF1DFBFFFE5FFBFF" bold="true" box="[189,460,1076,1102]" pageId="2" pageNumber="83">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="335936E0FFCCF522FF1DFBAFFAE4FA26" blockId="2.[189,1399,1123,1431]" pageId="2" pageNumber="83">
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||
<taxonomicName id="F4E64D63FFCCF522FF1DFBAFFEB2FBCC" authorityName="Quate" authorityYear="1996" box="[189,289,1124,1149]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">
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||
<emphasis id="0192EAF2FFCCF522FF1DFBAFFEB2FBCC" box="[189,289,1124,1149]" italics="true" pageId="2" pageNumber="83">Eugenys</emphasis>
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</taxonomicName>
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is a very distinctive genus and it can be easily differentiated from other genera of
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<taxonomicName id="F4E64D63FFCCF522FB42FBA8FAE5FBCC" box="[1250,1398,1123,1149]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="subFamily" subFamily="Psychodinae">Psychodinae</taxonomicName>
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by the following characters: eyes contiguous, eye bridge with three facet rows; antenna with 14 symmetric fusiform flagellomeres, except flagellomere 14 which is elongated and longer than the previous flagellomere, with a terminal apiculus; ascoids very long (the length of 3–4 flagellomeres), wavy-zigzag and rod-shaped (unknown in
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<taxonomicName id="F4E64D63FFCCF522FD65FB3AFCD0FABB" authorityName="Quate" authorityYear="1999" box="[709,835,1265,1290]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="cymosa">
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||
<emphasis id="0192EAF2FFCCF522FD65FB3AFCD0FABB" box="[709,835,1265,1290]" italics="true" pageId="2" pageNumber="83">E. cymosa</emphasis>
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||
</taxonomicName>
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||
); mouthparts atrophied or very reduced; palpi short, not extending beyond flagellomere 3–4; thorax without allurement organs, with alveoli on the central part of anepisternum, posteriorly to anterior spiracle; broad wings, vein R
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<subScript id="AF6234A5FFCCF522FBDDFA82FB0BFAE6" attach="left" box="[1149,1176,1353,1367]" fontSize="6" pageId="2" pageNumber="83">2+3</subScript>
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||
very short, arising from R
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||
<subScript id="AF6234A5FFCCF522FEB1FAA7FE89FACB" attach="left" box="[273,282,1388,1402]" fontSize="6" pageId="2" pageNumber="83">4</subScript>
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||
(Rs pectinate), R
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||
<subScript id="AF6234A5FFCCF522FE46FAA7FE7CFACB" attach="left" box="[486,495,1388,1402]" fontSize="6" pageId="2" pageNumber="83">5</subScript>
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ending in wing apex or slightly after apex, CuA
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||
<subScript id="AF6234A5FFCCF522FB83FAA7FBBFFACB" attach="left" box="[1059,1068,1388,1402]" fontSize="6" pageId="2" pageNumber="83">2</subScript>
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ending beyond medial fork; male genitalia with asymmetrical aedeagal complex, hypopods with one tenaculum or multiple tenacula.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7BFC656BFFCCF522FF1DFA75FE8AF8D6" pageId="2" pageNumber="83" type="discussion">
|
||
<paragraph id="335936E0FFCCF522FF1DFA75FEA3FA69" blockId="2.[189,304,1470,1496]" box="[189,304,1470,1496]" pageId="2" pageNumber="83">
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||
<heading id="6811818CFFCCF522FF1DFA75FEA3FA69" bold="true" box="[189,304,1470,1496]" fontSize="11" level="3" pageId="2" pageNumber="83" reason="3">
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||
<emphasis id="0192EAF2FFCCF522FF1DFA75FEA3FA69" bold="true" box="[189,304,1470,1496]" pageId="2" pageNumber="83">Remarks</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="335936E0FFCCF522FF1DFA26FE8AF8D6" blockId="2.[188,1400,1517,1895]" pageId="2" pageNumber="83">
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||
<bibRefCitation id="57774B11FFCCF522FF1DFA26FEFFF9B6" author="Quate L. W." box="[189,364,1517,1543]" pageId="2" pageNumber="83" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate’s (1996)</bibRefCitation>
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||
original description of the genus was based only on the
|
||
<typeStatus id="EC5D8842FFCCF522FBA3FA26FBA5F9B6" box="[1027,1078,1517,1543]" pageId="2" pageNumber="83">type</typeStatus>
|
||
species; however, when he later described two additional species in this genus (
|
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<bibRefCitation id="57774B11FFCCF522FCBDF9DBFC38F99B" author="Quate L. W." box="[797,939,1552,1578]" pageId="2" pageNumber="83" pagination="409 - 441" refId="ref7259" refString="Quate L. W. 1999. Taxonomy of neotropical Psychodidae. (Diptera) 3. Psychodines of Barro Colorado Island and San Blas, Panama. Memoirs on Entomology International 14: 409 - 441." type="journal article" year="1999">Quate 1999</bibRefCitation>
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||
:
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||
<taxonomicName id="F4E64D63FFCCF522FC16F9DAFBA3F99B" authorityName="Quate" authorityYear="1999" box="[950,1072,1553,1578]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="cymosa">
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||
<emphasis id="0192EAF2FFCCF522FC16F9DAFBA3F99B" box="[950,1072,1553,1578]" italics="true" pageId="2" pageNumber="83">E. cymosa</emphasis>
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||
</taxonomicName>
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and
|
||
<taxonomicName id="F4E64D63FFCCF522FBC8F9DAFA86F99B" authorityName="Quate" authorityYear="1999" box="[1128,1301,1553,1578]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="0192EAF2FFCCF522FBC8F9DAFA86F99B" box="[1128,1301,1553,1578]" italics="true" pageId="2" pageNumber="83">E. panamensis</emphasis>
|
||
</taxonomicName>
|
||
), not all the characters and structures were available for comparison, e.g., in
|
||
<typeStatus id="EC5D8842FFCCF522FC49F9FFFB8FF9FF" box="[1001,1052,1588,1614]" pageId="2" pageNumber="83">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="F4E64D63FFCCF522FB6AF9FFFAEBF9FC" authorityName="Quate" authorityYear="1999" box="[1226,1400,1588,1613]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="0192EAF2FFCCF522FB6AF9FFFAEBF9FC" box="[1226,1400,1588,1613]" italics="true" pageId="2" pageNumber="83">E. panamensis</emphasis>
|
||
</taxonomicName>
|
||
the antennae are incomplete; therefore, the last flagellomere is unknown. In
|
||
<typeStatus id="EC5D8842FFCCF522FB32F99CFB56F9C0" box="[1170,1221,1623,1649]" pageId="2" pageNumber="83">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="F4E64D63FFCCF522FF1DF9B1FEABF922" authorityName="Quate" authorityYear="1999" box="[189,312,1658,1683]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="cymosa">
|
||
<emphasis id="0192EAF2FFCCF522FF1DF9B1FEABF922" box="[189,312,1658,1683]" italics="true" pageId="2" pageNumber="83">E. cymosa</emphasis>
|
||
</taxonomicName>
|
||
the ascoids are missing; consequently, there is no comparison of this character between the species. Furthermore, the last flagellomere in
|
||
<taxonomicName id="F4E64D63FFCCF522FD6FF956FCF7F906" authorityName="Quate" authorityYear="1996" box="[719,868,1693,1719]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="clavellata">
|
||
<emphasis id="0192EAF2FFCCF522FD6FF956FCF7F906" box="[719,868,1693,1719]" italics="true" pageId="2" pageNumber="83">E. clavellata</emphasis>
|
||
</taxonomicName>
|
||
is considerably longer than the one present in
|
||
<taxonomicName id="F4E64D63FFCCF522FF1CF90AFEA5F96B" authorityName="Quate" authorityYear="1999" box="[188,310,1729,1754]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="cymosa">
|
||
<emphasis id="0192EAF2FFCCF522FF1CF90AFEA5F96B" box="[188,310,1729,1754]" italics="true" pageId="2" pageNumber="83">E. cymosa</emphasis>
|
||
</taxonomicName>
|
||
. Lastly, the number of tenacula present on the hypopods varies from one in
|
||
<taxonomicName id="F4E64D63FFCCF522FB0FF90AFAD7F96B" authorityName="Quate" authorityYear="1996" box="[1199,1348,1728,1754]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="clavellata">
|
||
<emphasis id="0192EAF2FFCCF522FB0FF90AFAD7F96B" box="[1199,1348,1728,1754]" italics="true" pageId="2" pageNumber="83">E. clavellata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F4E64D63FFCCF522FF1DF92FFEFAF94C" authorityName="Quate" authorityYear="1999" box="[189,361,1764,1789]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="0192EAF2FFCCF522FF1DF92FFEFAF94C" box="[189,361,1764,1789]" italics="true" pageId="2" pageNumber="83">E. panamensis</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<quantity id="F41E9B05FFCCF522FE2EF928FE66F94C" box="[398,501,1763,1789]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.556" pageId="2" pageNumber="83" unit="in" value="15.0" valueMax="16.0" valueMin="14.0">14–16 in</quantity>
|
||
<taxonomicName id="F4E64D63FFCCF522FE5BF92FFDE7F94C" authorityName="Quate" authorityYear="1999" box="[507,628,1764,1789]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="cymosa">
|
||
<emphasis id="0192EAF2FFCCF522FE5BF92FFDE7F94C" box="[507,628,1764,1789]" italics="true" pageId="2" pageNumber="83">E. cymosa</emphasis>
|
||
</taxonomicName>
|
||
which makes
|
||
<taxonomicName id="F4E64D63FFCCF522FCBBF92FFC06F94C" authorityName="Quate" authorityYear="1999" box="[795,917,1764,1789]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="cymosa">
|
||
<emphasis id="0192EAF2FFCCF522FCBBF92FFC06F94C" box="[795,917,1764,1789]" italics="true" pageId="2" pageNumber="83">E. cymosa</emphasis>
|
||
</taxonomicName>
|
||
hard to fit in the original diagnosis of the genus by
|
||
<bibRefCitation id="57774B11FFCCF522FE8DF8CDFE5BF891" author="Quate L. W." box="[301,456,1798,1824]" pageId="2" pageNumber="83" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate (1996)</bibRefCitation>
|
||
based solely on the number of tenacula and terminal flagellomeres. The updated diagnosis proposed above fits all of the previously described species, as well as the herein described species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BFC656BFFCDF523FF1DFEC3FAE4FEC5" pageId="3" pageNumber="84" type="biology_ecology">
|
||
<paragraph id="335936E0FFCDF523FF1DFEC3FE8FFE93" blockId="3.[189,284,264,290]" box="[189,284,264,290]" pageId="3" pageNumber="84">
|
||
<heading id="6811818CFFCDF523FF1DFEC3FE8FFE93" bold="true" box="[189,284,264,290]" fontSize="11" level="3" pageId="3" pageNumber="84" reason="3">
|
||
<emphasis id="0192EAF2FFCDF523FF1DFEC3FE8FFE93" bold="true" box="[189,284,264,290]" pageId="3" pageNumber="84">Biology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="335936E0FFCDF523FF1DFEF3FAE4FEC5" blockId="3.[189,1399,311,373]" pageId="3" pageNumber="84">
|
||
To date, nothing is known about the immature stages and the biology of the species of
|
||
<emphasis id="0192EAF2FFCDF523FB15FEF3FAB2FEE0" box="[1205,1313,312,337]" italics="true" pageId="3" pageNumber="84">
|
||
<taxonomicName id="F4E64D63FFCDF523FB15FEF3FA8EFEE0" authorityName="Quate" authorityYear="1996" box="[1205,1309,312,337]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">Eugenys</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Adults have atrophied (or very reduced) mouthparts, and it is believed that adults do not feed (see Discussion).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BFC656BFFCDF523FF1DFE50FB0AFDB9" pageId="3" pageNumber="84" type="taxon_list">
|
||
<paragraph id="335936E0FFCDF523FF1DFE50FE1DFE04" blockId="3.[189,398,411,437]" box="[189,398,411,437]" pageId="3" pageNumber="84">
|
||
<heading id="6811818CFFCDF523FF1DFE50FE1DFE04" bold="true" box="[189,398,411,437]" fontSize="11" level="3" pageId="3" pageNumber="84" reason="3">
|
||
<emphasis id="0192EAF2FFCDF523FF1DFE50FE1DFE04" bold="true" box="[189,398,411,437]" pageId="3" pageNumber="84">Species included</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="335936E0FFCDF523FF1DFE00FB0AFDB9" blockId="3.[188,1398,458,520]" pageId="3" pageNumber="84">
|
||
<taxonomicName id="F4E64D63FFCDF523FF1DFE00FDA7FE55" authority="Quate, 1996" authorityName="Quate" authorityYear="1996" box="[189,564,458,484]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="clavellata">
|
||
<emphasis id="0192EAF2FFCDF523FF1DFE00FE0EFE55" box="[189,413,458,484]" italics="true" pageId="3" pageNumber="84">Eugenys clavellata</emphasis>
|
||
<bibRefCitation id="57774B11FFCDF523FE03FE01FDA7FE55" author="Quate L. W." box="[419,564,458,484]" pageId="3" pageNumber="84" pagination="1 - 81" refId="ref7225" refString="Quate L. W. 1996. Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical 44 (1): 1 - 81." type="journal article" year="1996">Quate, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F4E64D63FFCDF523FD9FFE00FCDCFE55" authority="Quate, 1999" authorityName="Quate" authorityYear="1999" box="[575,847,458,484]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="cymosa">
|
||
<emphasis id="0192EAF2FFCDF523FD9FFE00FD2BFE55" box="[575,696,459,484]" italics="true" pageId="3" pageNumber="84">E. cymosa</emphasis>
|
||
<bibRefCitation id="57774B11FFCDF523FD1EFE01FCDCFE55" author="Quate L. W." box="[702,847,458,484]" pageId="3" pageNumber="84" pagination="409 - 441" refId="ref7259" refString="Quate L. W. 1999. Taxonomy of neotropical Psychodidae. (Diptera) 3. Psychodines of Barro Colorado Island and San Blas, Panama. Memoirs on Entomology International 14: 409 - 441." type="journal article" year="1999">Quate, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F4E64D63FFCDF523FCFAFE00FC2CFE55" authority="Jaume-Schinkel & Kvifte, 2024" authorityName="Jaume-Schinkel & Kvifte" authorityYear="2024" box="[858,959,459,484]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="micra" status="sp. nov.">
|
||
<emphasis id="0192EAF2FFCDF523FCFAFE00FC2CFE55" box="[858,959,459,484]" italics="true" pageId="3" pageNumber="84">E. micra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1AA15789FFCDF523FC66FE00FBB1FE54" box="[966,1058,459,485]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="F4E64D63FFCDF523FB8DFE00FAE2FE55" authority="Quate, 1999" authorityName="Quate" authorityYear="1999" box="[1069,1393,458,484]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="panamensis">
|
||
<emphasis id="0192EAF2FFCDF523FB8DFE00FB4AFE55" box="[1069,1241,459,484]" italics="true" pageId="3" pageNumber="84">E. panamensis</emphasis>
|
||
<bibRefCitation id="57774B11FFCDF523FB40FE01FAE2FE55" author="Quate L. W." box="[1248,1393,458,484]" pageId="3" pageNumber="84" pagination="409 - 441" refId="ref7259" refString="Quate L. W. 1999. Taxonomy of neotropical Psychodidae. (Diptera) 3. Psychodines of Barro Colorado Island and San Blas, Panama. Memoirs on Entomology International 14: 409 - 441." type="journal article" year="1999">Quate, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F4E64D63FFCDF523FF1CFE25FEC7FDB6" authority="Jaume-Schinkel & Kvifte, 2024" authorityName="Jaume-Schinkel & Kvifte" authorityYear="2024" box="[188,340,493,519]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
|
||
<emphasis id="0192EAF2FFCDF523FF1CFE25FEC7FDB6" box="[188,340,493,519]" italics="true" pageId="3" pageNumber="84">E. singularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1AA15789FFCDF523FEFCFE25FE24FDB9" box="[348,439,494,520]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="F4E64D63FFCDF523FE50FE25FDFBFDB6" authority="Jaume-Schinkel & Kvifte, 2024" authorityName="Jaume-Schinkel & Kvifte" authorityYear="2024" box="[496,616,493,519]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="upsilon" status="sp. nov.">
|
||
<emphasis id="0192EAF2FFCDF523FE50FE25FDFBFDB6" box="[496,616,493,519]" italics="true" pageId="3" pageNumber="84">E. upsilon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1AA15789FFCDF523FDD0FE25FD59FDB9" box="[624,714,494,520]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Species distribution is shown in
|
||
<figureCitation id="ABDD2A65FFCDF523FBEEFE26FB00FDB9" box="[1102,1171,493,520]" captionStart="Fig" captionStartId="3.[187,230,1998,2024]" captionTargetBox="[189,1398,1352,1957]" captionTargetId="figure-251@3.[189,1398,1352,1957]" captionTargetPageId="3" captionText="Fig. 1. Distribution map of the species of Eugenys Quate, 1996." figureDoi="http://doi.org/10.5281/zenodo.11284621" httpUri="https://zenodo.org/record/11284621/files/figure.png" pageId="3" pageNumber="84">Fig. 1</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |