treatments-xml/data/BB/4F/87/BB4F87F6FFC7F52CFDD3FBC7FD42FAD5.xml
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<document id="2A9308BACE42F0F7DE067131DCA125BB" ID-DOI="10.5852/ejt.2024.935.2547" ID-ISSN="2118-9773" ID-Zenodo-Dep="11284619" ID-ZooBank="C6FAFB3A-1B4B-4245-A424-0E4EDDA8AF05" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1716568146815" checkinUser="plazi" docAuthor="Jaume-Schinkel, Santiago &amp; Kvifte, Gunnar Mikalsen" docDate="2024" docId="BB4F87F6FFC7F52CFDD3FBC7FD42FAD5" docLanguage="en" docName="EJT.2024.935.81-100.pdf" docOrigin="European Journal of Taxonomy 935" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2547/11497" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Eugenys singularis Jaume-Schinkel &amp; Kvifte 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="93" masterDocId="4776FF8EFFCEF520FFA0FFCBFF93FFB1" masterDocTitle="Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species" masterLastPageNumber="100" masterPageNumber="81" pageNumber="90" updateTime="1717032848221" updateUser="jonas" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0006327DBF1A7C6E408D3095F1FAA723">Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species</mods:title>
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<mods:namePart id="40E4794FA2F9CD7845C0DCA7CEC8BF10">Jaume-Schinkel, Santiago</mods:namePart>
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<mods:affiliation id="9C962AE45035C66C78B2482BE4AB6A6F">Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="AA7120EACE23773CE6609A2E3D4BA0E0" type="email">santijaumes@hotmail.com &amp; s.jaume@leibniz-lib.de</mods:nameIdentifier>
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<mods:namePart id="7C760667754215E137AA3614BF5210BE">Kvifte, Gunnar Mikalsen</mods:namePart>
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<mods:affiliation id="47F488E7D8A102C405FF28B5BC23E69B">Department of Biosciences and Aquaculture, Nord University, P. O. Box 2501, 7729 Steinkjer, Norway.</mods:affiliation>
<mods:nameIdentifier id="D9903F5E51AA66FD496836025D4F76A1" type="email">gunnar.mikalsen-kvifte@nord.no</mods:nameIdentifier>
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<mods:date id="DCAEC0BE0F0CA4FB0CDD98B71F97E096">2024</mods:date>
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<treatment id="BB4F87F6FFC7F52CFDD3FBC7FD42FAD5" LSID="urn:lsid:plazi:treatment:BB4F87F6FFC7F52CFDD3FBC7FD42FAD5" httpUri="http://treatment.plazi.org/id/BB4F87F6FFC7F52CFDD3FBC7FD42FAD5" lastPageId="12" lastPageNumber="93" pageId="9" pageNumber="90">
<subSubSection id="7BFC656BFFC7F529FDD3FBC7FC52FB96" box="[627,961,1035,1063]" pageId="9" pageNumber="90" type="nomenclature">
<paragraph id="335936E0FFC7F529FDD3FBC7FC52FB96" blockId="9.[366,1221,1035,1133]" box="[627,961,1035,1063]" pageId="9" pageNumber="90">
<heading id="6811818CFFC7F529FDD3FBC7FC52FB96" box="[627,961,1035,1063]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="90" reason="2">
<taxonomicName id="F4E64D63FFC7F529FDD3FBC7FCCDFB97" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[627,862,1035,1062]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="0192EAF2FFC7F529FDD3FBC7FCCDFB97" bold="true" box="[627,862,1035,1062]" italics="true" pageId="9" pageNumber="90">Eugenys singularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA15789FFC7F529FCC6FBC6FC52FB96" box="[870,961,1037,1063]" pageId="9" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFC7F529FECEFBE4FCF0FBDD" pageId="9" pageNumber="90" type="description">
<paragraph id="335936E0FFC7F529FECEFBE4FB56FBFB" blockId="9.[366,1221,1035,1133]" box="[366,1221,1071,1098]" pageId="9" pageNumber="90">
<uri id="47773AE2FFC7F529FECEFBE4FB56FBFB" box="[366,1221,1071,1098]" pageId="9" pageNumber="90">
urn:lsid:zoobank.org:act:
<uuid id="47400C35FFC7F529FD2CFBE4FB56FBFB" box="[652,1221,1071,1098]" pageId="9" pageNumber="90">AA14FE8F-5A54-430F-9D08-56AC089FFB68</uuid>
</uri>
</paragraph>
<paragraph id="335936E0FFC7F529FD71FB99FCF0FBDD" blockId="9.[366,1221,1035,1133]" box="[721,867,1106,1133]" pageId="9" pageNumber="90">
<figureCitation id="ABDD2A65FFC7F529FD71FB99FC89FBDC" box="[721,794,1106,1133]" captionStart="Fig" captionStartId="3.[187,230,1998,2024]" captionTargetBox="[189,1398,1352,1957]" captionTargetId="figure-251@3.[189,1398,1352,1957]" captionTargetPageId="3" captionText="Fig. 1. Distribution map of the species of Eugenys Quate, 1996." figureDoi="http://doi.org/10.5281/zenodo.11284621" httpUri="https://zenodo.org/record/11284621/files/figure.png" pageId="9" pageNumber="90">Figs 1</figureCitation>
,
<figureCitation id="ABDD2A65FFC7F529FC88FB98FCDBFBDC" box="[808,840,1107,1133]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: AB, DF by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="9" pageNumber="90">3F</figureCitation>
,
<figureCitation id="ABDD2A65FFC7F529FCF4FB99FCF0FBDD" box="[852,867,1106,1132]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[300,1338,296,1820]" captionTargetId="figure-12@11.[220,1367,260,1850]" captionTargetPageId="11" captionText="Fig. 6. Eugenys singularis sp. nov., ♂, holotype (INBIO-CRI001470242). A. Head. B. Aedeagus, gonocoxites and gonostyli (ventral view). C. Hypopods (ventral view). D. Aedeagus, gonocoxites and gonostyli (ventral view). E. Hypopods. Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284645" httpUri="https://zenodo.org/record/11284645/files/figure.png" pageId="9" pageNumber="90">6</figureCitation>
</paragraph>
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<subSubSection id="7BFC656BFFC7F529FF1DFB5FFBFDFAB1" pageId="9" pageNumber="90" type="diagnosis">
<paragraph id="335936E0FFC7F529FF1DFB5FFE5FFB1F" blockId="9.[189,460,1172,1198]" box="[189,460,1172,1198]" pageId="9" pageNumber="90">
<heading id="6811818CFFC7F529FF1DFB5FFE5FFB1F" bold="true" box="[189,460,1172,1198]" fontSize="11" level="3" pageId="9" pageNumber="90" reason="3">
<emphasis id="0192EAF2FFC7F529FF1DFB5FFE5FFB1F" bold="true" box="[189,460,1172,1198]" pageId="9" pageNumber="90">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="335936E0FFC7F529FF1DFB09FBFDFAB1" blockId="9.[188,1398,1218,1280]" pageId="9" pageNumber="90">
This species can be easily differentiated from all other species of
<taxonomicName id="F4E64D63FFC7F529FC0EFB08FB81FB6D" authorityName="Quate" authorityYear="1996" box="[942,1042,1219,1244]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="0192EAF2FFC7F529FC0EFB08FB81FB6D" box="[942,1042,1219,1244]" italics="true" pageId="9" pageNumber="90">Eugenys</emphasis>
</taxonomicName>
by the hypopods with a lateral digitiform projection (hypopods without lateral projection in remaining species).
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFC7F529FF1DFAECFD92FA22" pageId="9" pageNumber="90" type="etymology">
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<heading id="6811818CFFC7F529FF1DFAECFED6FAF0" bold="true" box="[189,325,1319,1345]" fontSize="11" level="3" pageId="9" pageNumber="90" reason="3">
<emphasis id="0192EAF2FFC7F529FF1DFAECFED6FAF0" bold="true" box="[189,325,1319,1345]" pageId="9" pageNumber="90">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="335936E0FFC7F529FF1DFA9EFD92FA22" blockId="9.[189,1398,1365,1427]" pageId="9" pageNumber="90">
The species epithet is derived from the Latin
<taxonomicName id="F4E64D63FFC7F529FD7BFA9DFCC8FADE" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[731,859,1365,1392]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="0192EAF2FFC7F529FD44FA9EFCC8FADE" box="[740,859,1365,1391]" italics="true" pageId="9" pageNumber="90">singulāris</emphasis>
</taxonomicName>
, referring to the unusual shape of the lateral projection of the hypopods.
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFC7F529FF1DFA71FB7FF91F" pageId="9" pageNumber="90" type="materials_examined">
<paragraph id="335936E0FFC7F529FF1DFA71FE3EFA65" blockId="9.[189,429,1466,1492]" box="[189,429,1466,1492]" pageId="9" pageNumber="90">
<heading id="6811818CFFC7F529FF1DFA71FE3EFA65" bold="true" box="[189,429,1466,1492]" fontSize="11" level="3" pageId="9" pageNumber="90" reason="3">
<emphasis id="0192EAF2FFC7F529FF1DFA71FE3EFA65" bold="true" box="[189,429,1466,1492]" pageId="9" pageNumber="90">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="335936E0FFC7F529FF4AFA22FC33F9F8" blockId="9.[189,1399,1513,1610]" pageId="9" pageNumber="90">
<materialsCitation id="838E3CBDFFC7F529FF4AFA22FC33F9F8" collectingDate="1993-12-01" collectingMethod="Malaise trap" collectionCode="INBIO, LACM" country="Costa Rica" elevation="75" location="Puerto Viejo de Sarapaqui" pageId="9" pageNumber="90" specimenCode="INBIO-CRI001470242" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
<emphasis id="0192EAF2FFC7F529FF4AFA22FECEF9B2" bold="true" box="[234,349,1513,1539]" pageId="9" pageNumber="90">
<typeStatus id="EC5D8842FFC7F529FF4AFA22FECEF9B2" box="[234,349,1513,1539]" pageId="9" pageNumber="90" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="4BF17670FFC7F529FF1DF9C7FEF4F997" box="[189,359,1548,1574]" name="Costa Rica" pageId="9" pageNumber="90">COSTA RICA</collectingCountry>
<emphasis id="0192EAF2FFC7F529FE2AF9C7FE7CF997" bold="true" box="[394,495,1548,1574]" pageId="9" pageNumber="90">
<collectingRegion id="F122F802FFC7F529FE2AF9C7FE7CF997" box="[394,495,1548,1574]" country="Costa Rica" name="Heredia" pageId="9" pageNumber="90">Heredia</collectingRegion>
</emphasis>
<specimenCount id="25E0FD69FFC7F529FDAEF9C4FDB7F994" box="[526,548,1551,1573]" count="1" pageId="9" pageNumber="90" type="male"></specimenCount>
;
<location id="3639603BFFC7F529FD96F9C7FCE0F997" LSID="urn:lsid:plazi:treatment:BB4F87F6FFC7F52CFDD3FBC7FD42FAD5:3639603BFFC7F529FD96F9C7FCE0F997" box="[566,883,1548,1574]" country="Costa Rica" name="Puerto Viejo de Sarapaqui" pageId="9" pageNumber="90" stateProvince="Heredia">Puerto Viejo de Sarapaquí</location>
,
<location id="3639603BFFC7F529FC21F9C7FBF3F997" LSID="urn:lsid:plazi:treatment:BB4F87F6FFC7F52CFDD3FBC7FD42FAD5:3639603BFFC7F529FC21F9C7FBF3F997" box="[897,1120,1548,1574]" country="Costa Rica" name="Est. Biol. LaSelva" pageId="9" pageNumber="90" stateProvince="Heredia">Est. Biol. LaSelva</location>
; alt.
<quantity id="F41E9B05FFC7F529FB3DF9C7FA8BF997" box="[1181,1304,1548,1575]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="9" pageNumber="90" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
<elevation id="B8CBD1D3FFC7F529FB3DF9C7FA8BF997" box="[1181,1304,1548,1575]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="9" pageNumber="90" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50100 m</elevation>
</quantity>
;
<date id="47581020FFC7F529FA87F9C7FF68F9F8" pageId="9" pageNumber="90" value="1993-12-01">
<collectingDate id="571CE9C8FFC7F529FA87F9C7FF68F9F8" pageId="9" pageNumber="90" value="1993-12-01">1 Dec. 1993</collectingDate>
</date>
;
<collectingMethod id="EAA74EF7FFC7F529FEA7F9E4FE0AF9FB" box="[263,409,1583,1610]" pageId="9" pageNumber="90">Malaise trap</collectingMethod>
M/00/009;
<specimenCode id="63409E9BFFC7F529FD83F9E4FCA4F9F8" box="[547,823,1583,1609]" collectionCode="INBIO-CRI" pageId="9" pageNumber="90">INBIO-CRI001470242</specimenCode>
;
<collectionCode id="55F7AE25FFC7F529FCE3F9E4FC0FF9F8" box="[835,924,1583,1609]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="9" pageNumber="90" type="Museum">LACM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="335936E0FFC7F529FF4AF9BAFB7FF91F" blockId="9.[189,1260,1649,1710]" pageId="9" pageNumber="90">
<materialsCitation id="838E3CBDFFC7F529FF4AF9BAFB7AF91F" collectingDate="1993-12-01" collectionCode="INBIO, LACM" country="Costa Rica" elevation="75" location="COSTA RICA" pageId="9" pageNumber="90" specimenCode="INBIO-CRI001470896" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<emphasis id="0192EAF2FFC7F529FF4AF9BAFEFAF93A" bold="true" box="[234,361,1649,1675]" pageId="9" pageNumber="90">
<typeStatus id="EC5D8842FFC7F529FF4AF9BAFEFAF93A" box="[234,361,1649,1675]" pageId="9" pageNumber="90" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="4BF17670FFC7F529FF1DF95FFEF6F91F" box="[189,357,1684,1710]" name="Costa Rica" pageId="9" pageNumber="90">COSTA RICA</collectingCountry>
<specimenCount id="25E0FD69FFC7F529FEDEF95FFE39F91C" box="[382,426,1684,1710]" count="1" pageId="9" pageNumber="90" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<specimenCode id="63409E9BFFC7F529FCD0F95FFB17F91F" box="[880,1156,1684,1710]" collectionCode="INBIO-CRI" pageId="9" pageNumber="90">INBIO-CRI001470896</specimenCode>
;
<collectionCode id="55F7AE25FFC7F529FB30F95FFB7AF91F" box="[1168,1257,1684,1710]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="9" pageNumber="90" type="Museum">LACM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFC7F52CFF1DF91EFEAAFB35" lastPageId="12" lastPageNumber="93" pageId="9" pageNumber="90" type="description">
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<heading id="6811818CFFC7F529FF1DF91EFEDCF95E" bold="true" box="[189,335,1749,1775]" fontSize="11" level="3" pageId="9" pageNumber="90" reason="3">
<emphasis id="0192EAF2FFC7F529FF1DF91EFEDCF95E" bold="true" box="[189,335,1749,1775]" pageId="9" pageNumber="90">Description</emphasis>
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</paragraph>
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<emphasis id="0192EAF2FFC7F529FF4AF8CFFEB9F8AF" bold="true" box="[234,298,1796,1822]" pageId="9" pageNumber="90">Male</emphasis>
</paragraph>
<paragraph id="335936E0FFC7F529FF1DF8ECFBCAF81A" blockId="9.[189,1399,1796,1963]" pageId="9" pageNumber="90">
<smallCapsWord id="35BFA03CFFC7F529FF1DF8ECFEFDF88E" baselines="1850,1850" box="[189,366,1831,1857]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="9" pageNumber="90">MEASUREMENTS</smallCapsWord>
(in mm; n = 2). Wing length 1.8 (1.81.8), width 0.89 (0.890.90); head length 0.35 (0.320.38), width 0.40 (0.400.40); antennal segments, scape 0.09 (0.080.10), pedicel 0.05 (0.05 0.06), flagellomeres 1: 0.09 (0.090.10), 213: 0.08 (0.080.09), flagellomere 14: 0.09; palpomeres 1: 0.04 (0.040.05), 2: 0.06 (0.060.06), 3: 0.05 (0.050.05), 4: 0.04 (0.040.04).
</paragraph>
<caption id="67996668FFC4F52AFF1FF843FD39F859" ID-DOI="http://doi.org/10.5281/zenodo.11284643" ID-Zenodo-Dep="11284643" httpUri="https://zenodo.org/record/11284643/files/figure.png" pageId="10" pageNumber="91" startId="10.[191,234,1928,1954]" targetBox="[434,1189,339,1846]" targetPageId="10">
<paragraph id="335936E0FFC4F52AFF1FF843FD39F859" blockId="10.[191,1402,1928,2025]" pageId="10" pageNumber="91">
<emphasis id="0192EAF2FFC4F52AFF1FF843FE81F813" bold="true" box="[191,274,1928,1954]" pageId="10" pageNumber="91">Fig. 5.</emphasis>
Male genitalia, ventral view.
<emphasis id="0192EAF2FFC4F52AFD28F843FD0EF813" bold="true" box="[648,669,1928,1954]" pageId="10" pageNumber="91">A</emphasis>
.
<taxonomicName id="F4E64D63FFC4F52AFD10F842FCF6F813" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[688,869,1929,1954]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="micra" status="sp. nov.">
<emphasis id="0192EAF2FFC4F52AFD10F842FCF6F813" box="[688,869,1929,1954]" italics="true" pageId="10" pageNumber="91">Eugenys micra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA15789FFC4F52AFCD1F843FC41F813" box="[881,978,1928,1954]" pageId="10" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (ZFMK-DIP-00097917).
<emphasis id="0192EAF2FFC4F52AFF1FF860FF40F874" bold="true" box="[191,211,1963,1989]" pageId="10" pageNumber="91">B</emphasis>
.
<taxonomicName id="F4E64D63FFC4F52AFF49F867FD1CF874" authority="Quate, 1999" authorityName="Quate" authorityYear="1999" box="[233,655,1963,1989]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="0192EAF2FFC4F52AFF49F867FE7AF874" box="[233,489,1964,1989]" italics="true" pageId="10" pageNumber="91">Eugenys panamensis</emphasis>
<bibRefCitation id="57774B11FFC4F52AFE58F860FD1CF874" author="Quate L. W." box="[504,655,1963,1989]" pageId="10" pageNumber="91" pagination="409 - 441" refId="ref7259" refString="Quate L. W. 1999. Taxonomy of neotropical Psychodidae. (Diptera) 3. Psychodines of Barro Colorado Island and San Blas, Panama. Memoirs on Entomology International 14: 409 - 441." type="journal article" year="1999">Quate, 1999</bibRefCitation>
</taxonomicName>
, paratype (BMNH(E)-2001-8). Scales in millimeters (mm). Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="67996668FFC5F52BFF1DF8A8FEBBF856" ID-DOI="http://doi.org/10.5281/zenodo.11284645" ID-Zenodo-Dep="11284645" httpUri="https://zenodo.org/record/11284645/files/figure.png" pageId="11" pageNumber="92" startId="11.[189,232,1891,1917]" targetBox="[300,1338,296,1820]" targetPageId="11">
<paragraph id="335936E0FFC5F52BFF1DF8A8FEBBF856" blockId="11.[189,1398,1891,2023]" pageId="11" pageNumber="92">
<emphasis id="0192EAF2FFC5F52BFF1DF8A8FE83F8CC" bold="true" box="[189,272,1891,1917]" pageId="11" pageNumber="92">Fig. 6.</emphasis>
<taxonomicName id="F4E64D63FFC5F52BFEBDF8AFFD96F8CC" authority="Jaume-Schinkel &amp; Kvifte, 2024" authorityName="Jaume-Schinkel &amp; Kvifte" authorityYear="2024" box="[285,517,1891,1917]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="92" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
<emphasis id="0192EAF2FFC5F52BFEBDF8AFFD96F8CC" box="[285,517,1891,1917]" italics="true" pageId="11" pageNumber="92">Eugenys singularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AA15789FFC5F52BFDB2F8AFFDE7F8CF" box="[530,628,1892,1918]" pageId="11" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (INBIO-CRI001470242).
<emphasis id="0192EAF2FFC5F52BFBF9F8A8FBFDF8CC" bold="true" box="[1113,1134,1891,1917]" pageId="11" pageNumber="92">A</emphasis>
. Head.
<emphasis id="0192EAF2FFC5F52BFB74F8AFFB7BF8CF" bold="true" box="[1236,1256,1892,1918]" pageId="11" pageNumber="92">B</emphasis>
. Aedeagus, gonocoxites and gonostyli (ventral view).
<emphasis id="0192EAF2FFC5F52BFD12F84DFD55F811" bold="true" box="[690,710,1926,1952]" pageId="11" pageNumber="92">C</emphasis>
. Hypopods (ventral view).
<emphasis id="0192EAF2FFC5F52BFBAFF84CFBB0F810" bold="true" box="[1039,1059,1927,1953]" pageId="11" pageNumber="92">D</emphasis>
. Aedeagus, gonocoxites and gonostyli (ventral view).
<emphasis id="0192EAF2FFC5F52BFE4FF861FD90F875" bold="true" box="[495,515,1962,1988]" pageId="11" pageNumber="92">E</emphasis>
. Hypopods. Scales in millimeters (mm). Abbreviations: see Material and methods.
</paragraph>
</caption>
<paragraph id="335936E0FFC2F52CFF1DFEC2FA98FDA8" blockId="12.[188,1400,265,538]" pageId="12" pageNumber="93">
<smallCapsWord id="35BFA03CFFC2F52CFF1DFEC2FF6FFE93" baselines="284,285" box="[189,252,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="12" pageNumber="93">HEAD</smallCapsWord>
. Slightly wider than long; eye bridge contiguous, eye bridge with three facet rows; no enlarged alveoli on postocular area; interocular suture non-discernible in examined material (
<figureCitation id="ABDD2A65FFC2F52CFB01FEE7FA91FEF7" box="[1185,1282,300,326]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[300,1338,296,1820]" captionTargetId="figure-12@11.[220,1367,260,1850]" captionTargetPageId="11" captionText="Fig. 6. Eugenys singularis sp. nov., ♂, holotype (INBIO-CRI001470242). A. Head. B. Aedeagus, gonocoxites and gonostyli (ventral view). C. Hypopods (ventral view). D. Aedeagus, gonocoxites and gonostyli (ventral view). E. Hypopods. Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284645" httpUri="https://zenodo.org/record/11284645/files/figure.png" pageId="12" pageNumber="93">Fig. 6A</figureCitation>
). Antenna with scape cylindrical, about 2 × as long as pedicel; pedicel spherical; 14 flagellomeres, 113 fusiform, flagellomere 14 elongated, slightly longer than flagellomere 13, ending in digitiform apiculus, flagellomeres without patch of darkened sensilla; ascoids lanceolate, about 2 × as long as flagellomere carrying them; frontal alveoli patch square, lower margin with concavity in middle; mouthparts reduced, not extending beyond margin of head. Palpal segments short and sclerotized, apical segment not corrugated; not extending more than half of flagellomere 4, palpal proportions 1.0:1.4:1.1:1.0.
</paragraph>
<paragraph id="335936E0FFC2F52CFF1DFD8AFD58FD13" blockId="12.[189,1399,577,674]" pageId="12" pageNumber="93">
<smallCapsWord id="35BFA03CFFC2F52CFF1DFD8AFE8AFDE8" baselines="596,596" box="[189,281,577,603]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="12" pageNumber="93">THORAX</smallCapsWord>
. Without allurement organs; wing length about 2 × its width, hyaline, with alveoli on entire surface; Sc vein almost straight, ending at about origin of R
<subScript id="AF6234A5FFC2F52CFCD9FDBCFC11FD34" attach="left" box="[889,898,631,645]" fontSize="6" pageId="12" pageNumber="93">5</subScript>
, fork of R
<subScript id="AF6234A5FFC2F52CFC5CFDBCFB84FD34" attach="left" box="[1020,1047,631,645]" fontSize="6" pageId="12" pageNumber="93">2+3</subScript>
basal to M
<subScript id="AF6234A5FFC2F52CFB39FDBCFB26FD34" attach="left" box="[1177,1205,631,645]" fontSize="6" pageId="12" pageNumber="93">1+2</subScript>
, R
<subScript id="AF6234A5FFC2F52CFB76FDBCFB62FD34" attach="left" box="[1238,1265,631,645]" fontSize="6" pageId="12" pageNumber="93">2+3</subScript>
joining R
<subScript id="AF6234A5FFC2F52CFAC7FDBCFAE3FD34" attach="left" box="[1383,1392,631,645]" fontSize="6" pageId="12" pageNumber="93">4</subScript>
, wing apex rounded, R5 ending at wing apex.
</paragraph>
<paragraph id="335936E0FFC2F52CFF1DFD02FCFDFBAE" blockId="12.[187,1398,713,1056]" pageId="12" pageNumber="93">
<smallCapsWord id="35BFA03CFFC2F52CFF1DFD02FED0FD50" baselines="732,732" box="[189,323,713,739]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Terminalia" pageId="12" pageNumber="93">TERMINALIA</smallCapsWord>
. Hypandrium narrow, plate-like; gonocoxites cylindrical, about two-thirds length of gonostyli; gonocoxites joining in middle, and together giving origin to semi-sclerotized parameral sheath protruding beyond aedeagus, covering aedeagus only on sternal surface (see
<figureCitation id="ABDD2A65FFC2F52CFBC6FCC4FB7DFC98" box="[1126,1262,783,809]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[300,1338,296,1820]" captionTargetId="figure-12@11.[220,1367,260,1850]" captionTargetPageId="11" captionText="Fig. 6. Eugenys singularis sp. nov., ♂, holotype (INBIO-CRI001470242). A. Head. B. Aedeagus, gonocoxites and gonostyli (ventral view). C. Hypopods (ventral view). D. Aedeagus, gonocoxites and gonostyli (ventral view). E. Hypopods. Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284645" httpUri="https://zenodo.org/record/11284645/files/figure.png" pageId="12" pageNumber="93">Fig. 6B, D</figureCitation>
); gonostyli digitiform, tapering towards apex, with rounded apex, outwardly curved; ejaculatory apodeme about 2 × as long as aedeagus, narrow but broadening towards basal margin; aedeagus digitiform and tapering towards the apex, apex pointed; parameral sclerites fused, forming single Y-shaped sclerotized segment, linked to aedeagal sheath and aedeagus; epandrium not discernible in examined material; hypopods cylindrical, with outer laterally curved projection (
<figureCitation id="ABDD2A65FFC2F52CFCBFFC74FC32FC68" box="[799,929,959,985]" captionStart="Fig" captionStartId="11.[189,232,1891,1917]" captionTargetBox="[300,1338,296,1820]" captionTargetId="figure-12@11.[220,1367,260,1850]" captionTargetPageId="11" captionText="Fig. 6. Eugenys singularis sp. nov., ♂, holotype (INBIO-CRI001470242). A. Head. B. Aedeagus, gonocoxites and gonostyli (ventral view). C. Hypopods (ventral view). D. Aedeagus, gonocoxites and gonostyli (ventral view). E. Hypopods. Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284645" httpUri="https://zenodo.org/record/11284645/files/figure.png" pageId="12" pageNumber="93">Fig. 6C, E</figureCitation>
), lateral projection carrying two apical tenacula, while margin prior to projection in apex of hypopods carries 1214 tenacula, all tenacula with rounded apex; epiproct v-shaped, covered in small setulae.
</paragraph>
<paragraph id="335936E0FFC2F52CFF4AFB8CFED5FBD0" blockId="12.[189,326,1095,1156]" box="[234,326,1095,1121]" pageId="12" pageNumber="93">
<emphasis id="0192EAF2FFC2F52CFF4AFB8CFED5FBD0" bold="true" box="[234,326,1095,1121]" pageId="12" pageNumber="93">Female</emphasis>
</paragraph>
<paragraph id="335936E0FFC2F52CFF1DFBA1FEAAFB35" blockId="12.[189,326,1095,1156]" box="[189,313,1130,1156]" pageId="12" pageNumber="93">Unknown.</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFC2F52CFF1DFB60FC92FB45" box="[189,769,1195,1269]" pageId="12" pageNumber="93" type="distribution">
<paragraph id="335936E0FFC2F52CFF1DFB60FECBFB74" blockId="12.[189,344,1195,1221]" box="[189,344,1195,1221]" pageId="12" pageNumber="93">
<heading id="6811818CFFC2F52CFF1DFB60FECBFB74" bold="true" box="[189,344,1195,1221]" fontSize="11" level="3" pageId="12" pageNumber="93" reason="3">
<emphasis id="0192EAF2FFC2F52CFF1DFB60FECBFB74" bold="true" box="[189,344,1195,1221]" pageId="12" pageNumber="93">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="335936E0FFC2F52CFF1DFB11FC92FB45" blockId="12.[189,769,1242,1269]" box="[189,769,1242,1269]" pageId="12" pageNumber="93">
Only known from the
<typeStatus id="EC5D8842FFC2F52CFE62FB10FE66FB44" box="[450,501,1243,1269]" pageId="12" pageNumber="93">type</typeStatus>
locality in
<collectingCountry id="4BF17670FFC2F52CFDD9FB11FD6EFB45" box="[633,765,1242,1268]" name="Costa Rica" pageId="12" pageNumber="93">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7BFC656BFFC2F52CFF1DFAD0FD42FAD5" box="[189,721,1307,1381]" pageId="12" pageNumber="93" type="biology_ecology">
<paragraph id="335936E0FFC2F52CFF1DFAD0FEB9FA84" blockId="12.[189,298,1307,1333]" box="[189,298,1307,1333]" pageId="12" pageNumber="93">
<heading id="6811818CFFC2F52CFF1DFAD0FEB9FA84" bold="true" box="[189,298,1307,1333]" fontSize="11" level="3" pageId="12" pageNumber="93" reason="3">
<emphasis id="0192EAF2FFC2F52CFF1DFAD0FEB9FA84" bold="true" box="[189,298,1307,1333]" pageId="12" pageNumber="93">Genetics</emphasis>
</heading>
</paragraph>
<paragraph id="335936E0FFC2F52CFF1DFA80FD42FAD5" blockId="12.[189,721,1354,1381]" box="[189,721,1354,1381]" pageId="12" pageNumber="93">No specimens were available for sequencing.</paragraph>
</subSubSection>
</treatment>
</document>