388 lines
50 KiB
XML
388 lines
50 KiB
XML
<document id="7E6FFF5D673FB89515B51FDD87E02F7A" ID-DOI="10.5852/ejt.2024.935.2547" ID-ISSN="2118-9773" ID-Zenodo-Dep="11284619" ID-ZooBank="C6FAFB3A-1B4B-4245-A424-0E4EDDA8AF05" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1716568146815" checkinUser="plazi" docAuthor="Jaume-Schinkel, Santiago & Kvifte, Gunnar Mikalsen" docDate="2024" docId="BB4F87F6FFC2F530FD23FA69FED4FDB6" docLanguage="en" docName="EJT.2024.935.81-100.pdf" docOrigin="European Journal of Taxonomy 935" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2547/11497" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Eugenys upsilon Jaume-Schinkel & Kvifte 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="97" masterDocId="4776FF8EFFCEF520FFA0FFCBFF93FFB1" masterDocTitle="Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species" masterLastPageNumber="100" masterPageNumber="81" pageNumber="93" updateTime="1717032848221" updateUser="jonas" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="6FCFC137E1850A468C71814BE53C8864" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FF1D2E56DE0CE9F9185D022C4D8C9B1A">
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<mods:title id="BFB3516A79CDE7C61AFEB85F415F83B9">Revision of the genus Eugenys Quate, 1996 (Diptera: Psychodidae), with the description of three new species</mods:title>
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<mods:name id="A6001F36EC34B97FE0F7E2ADF4D18AEF" type="personal">
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<mods:roleTerm id="7EF50BEE65FF6FD1FE446DCBBDFB38C6">Author</mods:roleTerm>
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<mods:namePart id="6337711F1D0E315AE85ACD6B34BA8F9A">Jaume-Schinkel, Santiago</mods:namePart>
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<mods:affiliation id="EEDDE9815EE7851419D01B0ECE4815DF">Museum Koenig, Leibniz-Institut zur Analyse des Biodiversitätswandels, Adenauerallee 127, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="33026FC662CF847026C607B0D25E3428" type="email">santijaumes@hotmail.com & s.jaume@leibniz-lib.de</mods:nameIdentifier>
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<mods:name id="6F05A60537420AF056E1C07D3AAA12A0" type="personal">
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<mods:roleTerm id="4F4D6AE477145BB127B7F0742C8C8355">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FBE78FEDCBA9C85A633BEEE65C7BD857">Kvifte, Gunnar Mikalsen</mods:namePart>
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<mods:nameIdentifier id="BEB4715D15893342E9028F848B5203F7" type="LSID">A9E0868C-34C9-474C-BFB6-2D35CCBD8371</mods:nameIdentifier>
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<mods:affiliation id="5493BE72BC5046B430B2C3ABEF222F09">Department of Biosciences and Aquaculture, Nord University, P. O. Box 2501, 7729 Steinkjer, Norway.</mods:affiliation>
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<mods:nameIdentifier id="4327BD994EEC9E4F872EA93A46B25846" type="email">gunnar.mikalsen-kvifte@nord.no</mods:nameIdentifier>
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<mods:typeOfResource id="59329A38A514C95FF6B4FC9977AA7195">text</mods:typeOfResource>
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<mods:title id="21286AC8A6C7A3D24AB2851235DE5BEE">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="3F4E04E9E1E96909A0813CD23F6F68E1">
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<mods:date id="69A8053541B323BF5A405484A0707665">2024</mods:date>
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<mods:number id="8A6284E3665182588671F406291C281A">2024-05-23</mods:number>
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<mods:detail id="2A69D84A494E6411B3BEF03EDB149088" type="volume">
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<mods:number id="E63926B5905F6B31F90CB7B40CD4526C">935</mods:number>
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<mods:start id="0EE1800CCA4AA368559352FF74BA8E7C">81</mods:start>
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<mods:url id="C5F54BEF36F56E47619220546F186945">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2547/11497</mods:url>
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<mods:classification id="6F56E371F738592A4C84114056A20EFC">journal article</mods:classification>
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<mods:identifier id="C39F3D5060BFBAB72E248413FEA0EB0D" type="DOI">10.5852/ejt.2024.935.2547</mods:identifier>
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<mods:identifier id="3076FF5B170B54D65C3CA6A09D615AA6" type="ISSN">2118-9773</mods:identifier>
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<treatment id="BB4F87F6FFC2F530FD23FA69FED4FDB6" LSID="urn:lsid:plazi:treatment:BB4F87F6FFC2F530FD23FA69FED4FDB6" httpUri="http://treatment.plazi.org/id/BB4F87F6FFC2F530FD23FA69FED4FDB6" lastPageId="16" lastPageNumber="97" pageId="12" pageNumber="93">
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<subSubSection id="7BFC656BFFC2F52CFD23FA69FC22FA0C" box="[643,945,1442,1469]" pageId="12" pageNumber="93" type="nomenclature">
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<paragraph id="335936E0FFC2F52CFD23FA69FC22FA0C" blockId="12.[372,1215,1442,1539]" box="[643,945,1442,1469]" pageId="12" pageNumber="93">
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<heading id="6811818CFFC2F52CFD23FA69FC22FA0C" box="[643,945,1442,1469]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="93" reason="2">
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<taxonomicName id="F4E64D63FFC2F52CFD23FA69FCDCFA0C" authority="Jaume-Schinkel & Kvifte, 2024" authorityName="Jaume-Schinkel & Kvifte" authorityYear="2024" box="[643,847,1442,1469]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="upsilon" status="sp. nov.">
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<emphasis id="0192EAF2FFC2F52CFD23FA69FCDCFA0C" bold="true" box="[643,847,1442,1469]" italics="true" pageId="12" pageNumber="93">Eugenys upsilon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1AA15789FFC2F52CFCF6FA68FC22FA0C" box="[854,945,1443,1469]" pageId="12" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7BFC656BFFC2F52CFED4FA0EFCE6F9B2" pageId="12" pageNumber="93" type="description">
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<paragraph id="335936E0FFC2F52CFED4FA0EFB2CFA51" blockId="12.[372,1215,1442,1539]" box="[372,1215,1477,1504]" pageId="12" pageNumber="93">
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<uri id="47773AE2FFC2F52CFED4FA0EFB2CFA51" box="[372,1215,1477,1504]" pageId="12" pageNumber="93">
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urn:lsid:zoobank.org:act:
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<uuid id="47400C35FFC2F52CFD36FA0EFB2CFA51" box="[662,1215,1477,1504]" pageId="12" pageNumber="93">56AB3A46-1F40-471F-B126-5C667FA33B48</uuid>
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</uri>
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</paragraph>
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<paragraph id="335936E0FFC2F52CFD1FFA23FCE6F9B2" blockId="12.[372,1215,1442,1539]" box="[703,885,1512,1539]" pageId="12" pageNumber="93">
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<figureCitation id="ABDD2A65FFC2F52CFD1FFA23FC9BF9B2" box="[703,776,1512,1539]" captionStart="Fig" captionStartId="3.[187,230,1998,2024]" captionTargetBox="[189,1398,1352,1957]" captionTargetId="figure-251@3.[189,1398,1352,1957]" captionTargetPageId="3" captionText="Fig. 1. Distribution map of the species of Eugenys Quate, 1996." figureDoi="http://doi.org/10.5281/zenodo.11284621" httpUri="https://zenodo.org/record/11284621/files/figure.png" pageId="12" pageNumber="93">Figs 1</figureCitation>
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,
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<figureCitation id="ABDD2A65FFC2F52CFCB6FA22FCAFF9B2" box="[790,828,1513,1539]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="12" pageNumber="93">3D</figureCitation>
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,
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<figureCitation id="ABDD2A65FFC2F52CFCE9FA22FCE6F9B2" box="[841,885,1513,1539]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="13.[189,232,1822,1848]" captionStartId-1="14.[189,232,1963,1989]" captionTargetBox-0="[258,1333,330,1766]" captionTargetBox-1="[249,1354,380,1923]" captionTargetId-0="figure-12@13.[226,1345,302,1787]" captionTargetId-1="figure-15@14.[208,1380,265,1923]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="Fig. 7. Eugenys upsilon sp. nov. A. ♂, paratype (ZFMK-DIP-00097921), head. B. ♂, paratype (ZFMK- DIP-00097914), wing. C. ♂, paratype (ZFMK-DIP-00097921), mouth parts. D. ♂, paratype (ZFMK- DIP-00097921), first antennal segments. E–H. ♂, holotype (ZFMK-DIP-00097918). E. Aedeagus (lateral view). F. Aedeagus (ventral view). G. Epandrium and hypopods (ventral view). H. Aedeagus, gonocoxites, and gonostyli (ventral view). Scales in millimeters (mm). Abbreviations: see Material and methods." captionText-1="Fig. 8. Eugenys upsilon sp. nov., ♀, paratype (ZFMK-DIP-00097916). A. Head. B. Wing. C. Mouth parts. D. Genitalia (ventral view). Scales in millimeters (mm)." figureDoi-0="http://doi.org/10.5281/zenodo.11284647" figureDoi-1="http://doi.org/10.5281/zenodo.11284649" httpUri-0="https://zenodo.org/record/11284647/files/figure.png" httpUri-1="https://zenodo.org/record/11284649/files/figure.png" pageId="12" pageNumber="93">7–8</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7BFC656BFFC2F52CFF1DF9E1FAE7F94E" pageId="12" pageNumber="93" type="diagnosis">
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||
<paragraph id="335936E0FFC2F52CFF1DF9E1FE5FF9F5" blockId="12.[189,460,1578,1604]" box="[189,460,1578,1604]" pageId="12" pageNumber="93">
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||
<heading id="6811818CFFC2F52CFF1DF9E1FE5FF9F5" bold="true" box="[189,460,1578,1604]" fontSize="11" level="3" pageId="12" pageNumber="93" reason="3">
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<emphasis id="0192EAF2FFC2F52CFF1DF9E1FE5FF9F5" bold="true" box="[189,460,1578,1604]" pageId="12" pageNumber="93">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="335936E0FFC2F52CFF1DF993FAE7F94E" blockId="12.[187,1399,1624,1792]" pageId="12" pageNumber="93">
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This species can be easily differentiated from all other species of
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<taxonomicName id="F4E64D63FFC2F52CFC61F992FBB6F9C3" authorityName="Quate" authorityYear="1996" box="[961,1061,1625,1650]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="93" phylum="Arthropoda" rank="genus">
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<emphasis id="0192EAF2FFC2F52CFC61F992FBB6F9C3" box="[961,1061,1625,1650]" italics="true" pageId="12" pageNumber="93">Eugenys</emphasis>
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</taxonomicName>
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by the following characters: hypopods with singular apical tenaculum (10–14 present in
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<taxonomicName id="F4E64D63FFC2F52CFC28F9B7FBB1F927" authority="Jaume-Schinkel & Kvifte, 2024" authorityName="Jaume-Schinkel & Kvifte" authorityYear="2024" box="[904,1058,1660,1686]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
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<emphasis id="0192EAF2FFC2F52CFC28F9B7FBB1F927" box="[904,1058,1660,1686]" italics="true" pageId="12" pageNumber="93">E. singularis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1AA15789FFC2F52CFB8BF9B7FB14F927" box="[1067,1159,1660,1686]" pageId="12" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="F4E64D63FFC2F52CFB62F9B7FAAFF924" authorityName="Quate" authorityYear="1999" box="[1218,1340,1660,1685]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="cymosa">
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<emphasis id="0192EAF2FFC2F52CFB62F9B7FAAFF924" box="[1218,1340,1660,1685]" italics="true" pageId="12" pageNumber="93">E. cymosa</emphasis>
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</taxonomicName>
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, one apical tenaculum in
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<taxonomicName id="F4E64D63FFC2F52CFE0FF96BFD84F908" authority="Jaume-Schinkel & Kvifte, 2024" authorityName="Jaume-Schinkel & Kvifte" authorityYear="2024" box="[431,535,1696,1721]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="micra" status="sp. nov.">
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<emphasis id="0192EAF2FFC2F52CFE0FF96BFD84F908" box="[431,535,1696,1721]" italics="true" pageId="12" pageNumber="93">E. micra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1AA15789FFC2F52CFD80F954FDEDF908" box="[544,638,1695,1721]" pageId="12" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="F4E64D63FFC2F52CFD2BF96BFCAAF908" authorityName="Quate" authorityYear="1999" box="[651,825,1696,1721]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="panamensis">
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<emphasis id="0192EAF2FFC2F52CFD2BF96BFCAAF908" box="[651,825,1696,1721]" italics="true" pageId="12" pageNumber="93">E. panamensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="F4E64D63FFC2F52CFCD4F96BFC7EF908" authority="Jaume-Schinkel & Kvifte, 2024" authorityName="Jaume-Schinkel & Kvifte" authorityYear="2024" box="[884,1005,1695,1721]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="upsilon" status="sp. nov.">
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<emphasis id="0192EAF2FFC2F52CFCD4F96BFC7EF908" box="[884,1005,1695,1721]" italics="true" pageId="12" pageNumber="93">E. upsilon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1AA15789FFC2F52CFC58F954FBC5F908" box="[1016,1110,1695,1721]" pageId="12" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
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); antennal flagellomeres with patch of darkened sensilla (present in
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<taxonomicName id="F4E64D63FFC2F52CFD02F908FCDEF96D" authorityName="Quate" authorityYear="1999" box="[674,845,1731,1756]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="panamensis">
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<emphasis id="0192EAF2FFC2F52CFD02F908FCDEF96D" box="[674,845,1731,1756]" italics="true" pageId="12" pageNumber="93">E. panamensis</emphasis>
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</taxonomicName>
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, absent in remaining species); aedeagus curved, without distinguishable paramere (aedeagus straight, with distinguishable paramere in
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<taxonomicName id="F4E64D63FFC2F52CFB16F92DFAF0F94E" authorityName="Quate" authorityYear="1999" box="[1206,1379,1766,1791]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="panamensis">
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<emphasis id="0192EAF2FFC2F52CFB16F92DFAF0F94E" box="[1206,1379,1766,1791]" italics="true" pageId="12" pageNumber="93">E. panamensis</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="7BFC656BFFC2F52CFF1DF8ECFE05F823" pageId="12" pageNumber="93" type="etymology">
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<paragraph id="335936E0FFC2F52CFF1DF8ECFED6F8F0" blockId="12.[189,325,1831,1857]" box="[189,325,1831,1857]" pageId="12" pageNumber="93">
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<heading id="6811818CFFC2F52CFF1DF8ECFED6F8F0" bold="true" box="[189,325,1831,1857]" fontSize="11" level="3" pageId="12" pageNumber="93" reason="3">
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<emphasis id="0192EAF2FFC2F52CFF1DF8ECFED6F8F0" bold="true" box="[189,325,1831,1857]" pageId="12" pageNumber="93">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="335936E0FFC2F52CFF1DF89EFE05F823" blockId="12.[189,1398,1876,1938]" pageId="12" pageNumber="93">The species epithet is derived from the Greek letter ‘ύψιλον’ (‘ýpsilon’), referring to the Y- to U-shaped parameral sclerite.</paragraph>
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</subSubSection>
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<caption id="67996668FFC3F52DFF1DF8D5FEBBF859" ID-DOI="http://doi.org/10.5281/zenodo.11284647" ID-Zenodo-Dep="11284647" httpUri="https://zenodo.org/record/11284647/files/figure.png" pageId="13" pageNumber="94" startId="13.[189,232,1822,1848]" targetBox="[258,1333,330,1766]" targetPageId="13">
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<paragraph id="335936E0FFC3F52DFF1DF8D5FEBBF859" blockId="13.[189,1399,1822,2024]" pageId="13" pageNumber="94">
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<emphasis id="0192EAF2FFC3F52DFF1DF8D5FE98F889" bold="true" box="[189,267,1822,1848]" pageId="13" pageNumber="94">Fig. 7.</emphasis>
|
||
<taxonomicName id="F4E64D63FFC3F52DFEB1F8D4FE40F889" authority="Jaume-Schinkel & Kvifte, 2024" authorityName="Jaume-Schinkel & Kvifte" authorityYear="2024" box="[273,467,1822,1848]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="94" phylum="Arthropoda" rank="species" species="upsilon" status="sp. nov.">
|
||
<emphasis id="0192EAF2FFC3F52DFEB1F8D4FE40F889" box="[273,467,1822,1848]" italics="true" pageId="13" pageNumber="94">Eugenys upsilon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1AA15789FFC3F52DFE7BF8D4FDA6F888" box="[475,565,1823,1849]" pageId="13" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="0192EAF2FFC3F52DFD9BF8D5FDC3F889" bold="true" box="[571,592,1822,1848]" pageId="13" pageNumber="94">A</emphasis>
|
||
. ♂, paratype
|
||
<emphasis id="0192EAF2FFC3F52DFD4FF8D5FD6AF889" bold="true" box="[751,761,1822,1848]" pageId="13" pageNumber="94">(</emphasis>
|
||
ZFMK-DIP-00097921), head.
|
||
<emphasis id="0192EAF2FFC3F52DFBFFF8D5FBE0F889" bold="true" box="[1119,1139,1822,1848]" pageId="13" pageNumber="94">B</emphasis>
|
||
. ♂, paratype
|
||
<emphasis id="0192EAF2FFC3F52DFAB1F8D5FA88F889" bold="true" box="[1297,1307,1822,1848]" pageId="13" pageNumber="94">(</emphasis>
|
||
ZFMK-DIP-00097914), wing.
|
||
<emphasis id="0192EAF2FFC3F52DFE72F88AFE75F8EA" bold="true" box="[466,486,1857,1883]" pageId="13" pageNumber="94">C</emphasis>
|
||
. ♂, paratype
|
||
<emphasis id="0192EAF2FFC3F52DFD2BF88AFD06F8EA" bold="true" box="[651,661,1857,1883]" pageId="13" pageNumber="94">(</emphasis>
|
||
ZFMK-DIP-00097921), mouth parts.
|
||
<emphasis id="0192EAF2FFC3F52DFBF8F889FBFFF8ED" bold="true" box="[1112,1132,1858,1884]" pageId="13" pageNumber="94">D</emphasis>
|
||
. ♂, paratype
|
||
<emphasis id="0192EAF2FFC3F52DFAB1F88AFA88F8EA" bold="true" box="[1297,1307,1857,1883]" pageId="13" pageNumber="94">(</emphasis>
|
||
ZFMK-DIP-00097921), first antennal segments.
|
||
<emphasis id="0192EAF2FFC3F52DFD1AF8AEFD67F8CE" bold="true" box="[698,756,1893,1919]" pageId="13" pageNumber="94">E–H</emphasis>
|
||
. ♂, holotype (ZFMK-DIP-00097918).
|
||
<emphasis id="0192EAF2FFC3F52DFB7BF8AEFB7CF8CE" bold="true" box="[1243,1263,1893,1919]" pageId="13" pageNumber="94">E</emphasis>
|
||
. Aedeagus (lateral view).
|
||
<emphasis id="0192EAF2FFC3F52DFECCF843FEEEF813" bold="true" box="[364,381,1928,1954]" pageId="13" pageNumber="94">F</emphasis>
|
||
. Aedeagus (ventral view).
|
||
<emphasis id="0192EAF2FFC3F52DFD60F843FD45F813" bold="true" box="[704,726,1928,1954]" pageId="13" pageNumber="94">G</emphasis>
|
||
. Epandrium and hypopods (ventral view).
|
||
<emphasis id="0192EAF2FFC3F52DFB76F843FB7FF813" bold="true" box="[1238,1260,1928,1954]" pageId="13" pageNumber="94">H</emphasis>
|
||
. Aedeagus, gonocoxites, and gonostyli (ventral view). Scales in millimeters (mm). Abbreviations: see Material and methods.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="67996668FFC0F52EFF1DF860FC05F859" ID-DOI="http://doi.org/10.5281/zenodo.11284649" ID-Zenodo-Dep="11284649" httpUri="https://zenodo.org/record/11284649/files/figure.png" pageId="14" pageNumber="95" startId="14.[189,232,1963,1989]" targetBox="[249,1354,380,1923]" targetPageId="14">
|
||
<paragraph id="335936E0FFC0F52EFF1DF860FC05F859" blockId="14.[189,1398,1963,2025]" pageId="14" pageNumber="95">
|
||
<emphasis id="0192EAF2FFC0F52EFF1DF860FE9EF874" bold="true" box="[189,269,1963,1989]" pageId="14" pageNumber="95">Fig. 8.</emphasis>
|
||
<taxonomicName id="F4E64D63FFC0F52EFEBAF867FE4CF874" authority="Jaume-Schinkel & Kvifte, 2024" authorityName="Jaume-Schinkel & Kvifte" authorityYear="2024" box="[282,479,1963,1989]" class="Insecta" family="Psychodidae" genus="Eugenys" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="95" phylum="Arthropoda" rank="species" species="upsilon" status="sp. nov.">
|
||
<emphasis id="0192EAF2FFC0F52EFEBAF867FE4CF874" box="[282,479,1963,1989]" italics="true" pageId="14" pageNumber="95">Eugenys upsilon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1AA15789FFC0F52EFE49F867FDD5F877" box="[489,582,1964,1990]" pageId="14" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♀, paratype (ZFMK-DIP-00097916).
|
||
<emphasis id="0192EAF2FFC0F52EFBB7F860FBBFF874" bold="true" box="[1047,1068,1963,1989]" pageId="14" pageNumber="95">A</emphasis>
|
||
. Head.
|
||
<emphasis id="0192EAF2FFC0F52EFB2CF860FB33F874" bold="true" box="[1164,1184,1963,1989]" pageId="14" pageNumber="95">B</emphasis>
|
||
. Wing.
|
||
<emphasis id="0192EAF2FFC0F52EFAA2F860FA84F874" bold="true" box="[1282,1303,1963,1989]" pageId="14" pageNumber="95">C</emphasis>
|
||
. Mouth parts.
|
||
<emphasis id="0192EAF2FFC0F52EFEA5F805FE89F859" bold="true" box="[261,282,1998,2024]" pageId="14" pageNumber="95">D</emphasis>
|
||
. Genitalia (ventral view). Scales in millimeters (mm).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="7BFC656BFFC1F52FFF1DFEC3FB3AFD42" pageId="15" pageNumber="96" type="materials_examined">
|
||
<paragraph id="335936E0FFC1F52FFF1DFEC3FE3EFE93" blockId="15.[189,429,264,290]" box="[189,429,264,290]" pageId="15" pageNumber="96">
|
||
<heading id="6811818CFFC1F52FFF1DFEC3FE3EFE93" bold="true" box="[189,429,264,290]" fontSize="11" level="3" pageId="15" pageNumber="96" reason="3">
|
||
<emphasis id="0192EAF2FFC1F52FFF1DFEC3FE3EFE93" bold="true" box="[189,429,264,290]" pageId="15" pageNumber="96">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="335936E0FFC1F52FFF4AFEFCFDD6FE0A" blockId="15.[189,1399,311,443]" pageId="15" pageNumber="96">
|
||
<materialsCitation id="838E3CBDFFC1F52FFF4AFEFCFDD6FE0A" collectingDate="2022-02-01" collectingDateMax="2022-02-09" collectingDateMin="2022-02-01" collectionCode="ZFMK, INABIO" collectorName="I. Kilian" country="Ecuador" county="Pedro Vicente Maldonado" elevation="750" latitude="0.11882" location="Parroquia Pedro Vicente Maldonato" longLatPrecision="1" longitude="-78.95802" pageId="15" pageNumber="96" specimenCode="ZFMK-TIS-2637135, ZFMK-DIP-00097918" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="holotype">
|
||
<emphasis id="0192EAF2FFC1F52FFF4AFEFCFECEFEE0" bold="true" box="[234,349,311,337]" pageId="15" pageNumber="96">
|
||
<typeStatus id="EC5D8842FFC1F52FFF4AFEFCFECEFEE0" box="[234,349,311,337]" pageId="15" pageNumber="96" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="4BF17670FFC1F52FFF1DFE91FED8FEC5" box="[189,331,346,372]" name="Ecuador" pageId="15" pageNumber="96">ECUADOR</collectingCountry>
|
||
–
|
||
<emphasis id="0192EAF2FFC1F52FFEC4FE91FE72FEC5" bold="true" box="[356,481,346,372]" pageId="15" pageNumber="96">
|
||
<collectingRegion id="F122F802FFC1F52FFEC4FE91FE72FEC5" box="[356,481,346,372]" country="Ecuador" name="Pichincha" pageId="15" pageNumber="96">Pichincha</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="25E0FD69FFC1F52FFE59FE90FD81FEC4" box="[505,530,347,373]" count="1" pageId="15" pageNumber="96" type="male">♂</specimenCount>
|
||
;
|
||
<collectingCounty id="DA384E6CFFC1F52FFDBDFE91FCC3FEC5" box="[541,848,346,372]" pageId="15" pageNumber="96">Pedro Vicente Maldonado</collectingCounty>
|
||
,
|
||
<location id="3639603BFFC1F52FFCFAFE91FB6EFEC5" LSID="urn:lsid:plazi:treatment:BB4F87F6FFC2F530FD23FA69FED4FDB6:3639603BFFC1F52FFCFAFE91FB6EFEC5" box="[858,1277,346,372]" country="Ecuador" county="Pedro Vicente Maldonado" latitude="0.11882" longLatPrecision="1" longitude="-78.95802" municipality="Parroquia Pedro Vicente Maldonato" name="Parroquia Pedro Vicente Maldonato" pageId="15" pageNumber="96" stateProvince="Pichincha">Parroquia Pedro Vicente Maldonato</location>
|
||
,
|
||
<location id="3639603BFFC1F52FFAA8FE91FEADFE26" LSID="urn:lsid:plazi:treatment:BB4F87F6FFC2F530FD23FA69FED4FDB6:3639603BFFC1F52FFAA8FE91FEADFE26" country="Ecuador" county="Pedro Vicente Maldonado" latitude="0.11882" longLatPrecision="1" longitude="-78.95802" municipality="Parroquia Pedro Vicente Maldonato" name="Roadway to Pachijal" pageId="15" pageNumber="96" stateProvince="Pichincha">Roadway to Pachijal</location>
|
||
;
|
||
<geoCoordinate id="56D25027FFC1F52FFEEBFEB5FE46FE29" box="[331,469,382,408]" degrees="0.11882" direction="north" orientation="latitude" pageId="15" pageNumber="96" precision="1" value="0.11882">0.11882° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="56D25027FFC1F52FFE42FEB6FD12FE29" box="[482,641,381,408]" degrees="78.95802" direction="west" orientation="longitude" pageId="15" pageNumber="96" precision="1" value="-78.95802">78.95802° W</geoCoordinate>
|
||
; alt.
|
||
<quantity id="F41E9B05FFC1F52FFD1BFEB6FC9AFE29" box="[699,777,381,408]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="15" pageNumber="96" unit="m" value="750.0">
|
||
<elevation id="B8CBD1D3FFC1F52FFD1BFEB6FC9AFE29" box="[699,777,381,408]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="15" pageNumber="96" unit="m" value="750.0">750 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="47581020FFC1F52FFCB6FEB5FC50FE29" box="[790,963,381,408]" pageId="15" pageNumber="96" value="2022-02-01" valueMax="2022-02-09" valueMin="2022-02-01">
|
||
<collectingDate id="571CE9C8FFC1F52FFCB6FEB5FC50FE29" box="[790,963,381,408]" pageId="15" pageNumber="96" value="2022-02-01" valueMax="2022-02-09" valueMin="2022-02-01">1–9 Feb. 2022</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="9E135336FFC1F52FFC70FEB5FBA0FE26" box="[976,1075,381,408]" pageId="15" pageNumber="96">I. Kilian</collectorName>
|
||
leg.;
|
||
<specimenCode id="63409E9BFFC1F52FFBD7FEB6FAE1FE29" box="[1143,1394,381,408]" pageId="15" pageNumber="96">ZFMK-TIS-2637135</specimenCode>
|
||
;
|
||
<specimenCode id="63409E9BFFC1F52FFF1EFE6AFE5FFE0A" box="[190,460,417,443]" pageId="15" pageNumber="96">ZFMK-DIP-00097918</specimenCode>
|
||
;
|
||
<collectionCode id="55F7AE25FFC1F52FFE77FE6AFDD1FE0A" box="[471,578,417,443]" pageId="15" pageNumber="96">INABIO</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="335936E0FFC1F52FFF4AFE29FB3AFD42" blockId="15.[189,1399,482,755]" pageId="15" pageNumber="96">
|
||
<materialsCitation id="838E3CBDFFC1F52FFF4AFE29FEB7FDF2" collectingDate="2022-02-01" collectingDateMax="2022-02-09" collectingDateMin="2022-02-01" collectionCode="ZFMK, INABIO" collectorName="I. Kilian" country="Ecuador" county="Pedro Vicente Maldonado" elevation="750" latitude="0.11882" location="Parroquia Pedro Vicente Maldonato" longLatPrecision="1" longitude="-78.95802" pageId="15" pageNumber="96" specimenCode="ZFMK-TIS-2637099, ZFMK-DIP-00097913" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
|
||
<emphasis id="0192EAF2FFC1F52FFF4AFE29FEFAFE4D" bold="true" box="[234,361,482,508]" pageId="15" pageNumber="96">
|
||
<typeStatus id="EC5D8842FFC1F52FFF4AFE29FEFAFE4D" box="[234,361,482,508]" pageId="15" pageNumber="96" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="4BF17670FFC1F52FFF1DFDCEFED8FDAE" box="[189,331,517,543]" name="Ecuador" pageId="15" pageNumber="96">ECUADOR</collectingCountry>
|
||
•
|
||
<specimenCount id="25E0FD69FFC1F52FFEC5FDCEFE06FD91" box="[357,405,517,544]" count="1" pageId="15" pageNumber="96" type="male">1 ♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="63409E9BFFC1F52FFCFDFDCEFBCBFDAE" box="[861,1112,517,543]" pageId="15" pageNumber="96">ZFMK-TIS-2637099</specimenCode>
|
||
;
|
||
<specimenCode id="63409E9BFFC1F52FFBC5FDCEFAE1FDAE" box="[1125,1394,517,543]" pageId="15" pageNumber="96">ZFMK-DIP-00097913</specimenCode>
|
||
;
|
||
<collectionCode id="55F7AE25FFC1F52FFF1DFDE2FEB7FDF2" box="[189,292,553,579]" pageId="15" pageNumber="96">INABIO</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="838E3CBDFFC1F52FFEE5FDE2FE9DFDD7" collectingDate="2022-02-01" collectingDateMax="2022-02-09" collectingDateMin="2022-02-01" collectionCode="ZFMK" collectorName="I. Kilian" country="Ecuador" county="Pedro Vicente Maldonado" elevation="750" latitude="0.11882" location="Parroquia Pedro Vicente Maldonato" longLatPrecision="1" longitude="-78.95802" pageId="15" pageNumber="96" specimenCode="ZFMK-TIS-2637105, ZFMK-DIP-00097914" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
|
||
<specimenCount id="25E0FD69FFC1F52FFEE5FDE2FEEBFDF2" box="[325,376,553,579]" count="1" pageId="15" pageNumber="96" type="male">1 ♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="63409E9BFFC1F52FFCF9FDE3FBC6FDF2" box="[857,1109,552,579]" pageId="15" pageNumber="96">ZFMK-TIS-2637105</specimenCode>
|
||
;
|
||
<specimenCode id="63409E9BFFC1F52FFBC5FDE2FAE1FDF2" box="[1125,1394,553,579]" pageId="15" pageNumber="96">ZFMK-DIP-00097914</specimenCode>
|
||
;
|
||
<collectionCode id="55F7AE25FFC1F52FFF1DFD87FE9DFDD7" box="[189,270,588,614]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="15" pageNumber="96" type="Museum">ZFMK</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="838E3CBDFFC1F52FFE84FD87FAE5FDD7" box="[292,1398,587,614]" collectingDate="2022-02-01" collectingDateMax="2022-02-09" collectingDateMin="2022-02-01" collectionCode="ZFMK" collectorName="I. Kilian" country="Ecuador" county="Pedro Vicente Maldonado" elevation="750" latitude="0.11882" location="Parroquia Pedro Vicente Maldonato" longLatPrecision="1" longitude="-78.95802" pageId="15" pageNumber="96" specimenCode="ZFMK-TIS-2637111, ZFMK-DIP-00097915" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
|
||
<specimenCount id="25E0FD69FFC1F52FFE84FD87FEC1FDD7" box="[292,338,588,614]" count="1" pageId="15" pageNumber="96" type="male">1 ♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="63409E9BFFC1F52FFCAAFD87FB90FDD7" box="[778,1027,588,614]" pageId="15" pageNumber="96">ZFMK-TIS-2637111</specimenCode>
|
||
;
|
||
<specimenCode id="63409E9BFFC1F52FFBADFD80FA88FDD7" box="[1037,1307,587,614]" pageId="15" pageNumber="96">ZFMK-DIP-00097915</specimenCode>
|
||
;
|
||
<collectionCode id="55F7AE25FFC1F52FFA85FD87FAE5FDD7" box="[1317,1398,588,614]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="15" pageNumber="96" type="Museum">ZFMK</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="838E3CBDFFC1F52FFF6EFDA4FAB8FD38" box="[206,1323,623,649]" collectingDate="2022-02-01" collectingDateMax="2022-02-09" collectingDateMin="2022-02-01" collectionCode="ZFMK" collectorName="I. Kilian" country="Ecuador" county="Pedro Vicente Maldonado" elevation="750" latitude="0.11882" location="Parroquia Pedro Vicente Maldonato" longLatPrecision="1" longitude="-78.95802" pageId="15" pageNumber="96" specimenCode="ZFMK-TIS-2637140, ZFMK-DIP-00097919" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
|
||
<specimenCount id="25E0FD69FFC1F52FFF6EFDA4FF6FFD38" box="[206,252,623,649]" count="1" pageId="15" pageNumber="96" type="male">1 ♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="63409E9BFFC1F52FFD1BFDA4FC24FD38" box="[699,951,623,649]" pageId="15" pageNumber="96">ZFMK-TIS-2637140</specimenCode>
|
||
;
|
||
<specimenCode id="63409E9BFFC1F52FFC61FDA4FB5CFD38" box="[961,1231,623,649]" pageId="15" pageNumber="96">ZFMK-DIP-00097919</specimenCode>
|
||
;
|
||
<collectionCode id="55F7AE25FFC1F52FFB7AFDA4FAB8FD38" box="[1242,1323,623,649]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="15" pageNumber="96" type="Museum">ZFMK</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="838E3CBDFFC1F52FFAE3FDA4FB76FD1C" collectingDate="2022-02-01" collectingDateMax="2022-02-09" collectingDateMin="2022-02-01" collectionCode="ZFMK" collectorName="I. Kilian" country="Ecuador" county="Pedro Vicente Maldonado" elevation="750" latitude="0.11882" location="Parroquia Pedro Vicente Maldonato" longLatPrecision="1" longitude="-78.95802" municipality="Parroquia Pedro Vicente Maldonato" pageId="15" pageNumber="96" specimenCode="ZFMK-TIS-2637141, ZFMK-DIP-00097920" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
|
||
<specimenCount id="25E0FD69FFC1F52FFAE3FDA4FAE2FD38" box="[1347,1393,623,649]" count="1" pageId="15" pageNumber="96" type="male">1 ♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="63409E9BFFC1F52FFDD4FD59FCFCFD1D" box="[628,879,658,684]" pageId="15" pageNumber="96">ZFMK-TIS-2637141</specimenCode>
|
||
;
|
||
<specimenCode id="63409E9BFFC1F52FFCDBFD59FB1BFD1D" box="[891,1160,658,684]" pageId="15" pageNumber="96">ZFMK-DIP-00097920</specimenCode>
|
||
;
|
||
<collectionCode id="55F7AE25FFC1F52FFB34FD58FB76FD1C" box="[1172,1253,659,685]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="15" pageNumber="96" type="Museum">ZFMK</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="838E3CBDFFC1F52FFB5FFD59FB5FFD61" collectingDate="2022-02-01" collectingDateMax="2022-02-09" collectingDateMin="2022-02-01" collectionCode="ZFMK" collectorName="I. Kilian" country="Ecuador" county="Pedro Vicente Maldonado" elevation="750" latitude="0.11882" location="Parroquia Pedro Vicente Maldonato" longLatPrecision="1" longitude="-78.95802" pageId="15" pageNumber="96" specimenCode="ZFMK-TIS-2637144, ZFMK-DIP-00097921" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
|
||
<specimenCount id="25E0FD69FFC1F52FFB5FFD59FABDFD1C" box="[1279,1326,658,685]" count="1" pageId="15" pageNumber="96" type="male">1 ♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="63409E9BFFC1F52FFDEFFD7EFCD8FD7E" box="[591,843,693,719]" pageId="15" pageNumber="96">ZFMK-TIS-2637144</specimenCode>
|
||
;
|
||
<specimenCode id="63409E9BFFC1F52FFCFCFD7EFBF9FD7E" box="[860,1130,693,719]" pageId="15" pageNumber="96">ZFMK-DIP-00097921</specimenCode>
|
||
;
|
||
<collectionCode id="55F7AE25FFC1F52FFBDBFD7DFB5FFD61" box="[1147,1228,694,720]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="15" pageNumber="96" type="Museum">ZFMK</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="838E3CBDFFC1F52FFB52FD7EFB36FD42" collectingDate="2022-02-01" collectingDateMax="2022-02-09" collectingDateMin="2022-02-01" collectionCode="ZFMK" collectorName="I. Kilian" country="Ecuador" county="Pedro Vicente Maldonado" elevation="750" latitude="0.11882" location="Parroquia Pedro Vicente Maldonato" longLatPrecision="1" longitude="-78.95802" pageId="15" pageNumber="96" specimenCode="ZFMK-TIS-2637123, ZFMK-DIP-00097916" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha" typeStatus="paratype">
|
||
<specimenCount id="25E0FD69FFC1F52FFB52FD7EFAB6FD61" box="[1266,1317,693,720]" count="1" pageId="15" pageNumber="96" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="63409E9BFFC1F52FFD90FD13FCB8FD43" box="[560,811,728,754]" pageId="15" pageNumber="96">ZFMK-TIS-2637123</specimenCode>
|
||
;
|
||
<specimenCode id="63409E9BFFC1F52FFC97FD13FBD6FD43" box="[823,1093,728,754]" pageId="15" pageNumber="96">ZFMK-DIP-00097916</specimenCode>
|
||
;
|
||
<collectionCode id="55F7AE25FFC1F52FFBF0FD12FB36FD42" box="[1104,1189,729,755]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="15" pageNumber="96" type="Museum">ZFMK</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BFC656BFFC1F52FFF1DFCD1FC27F869" pageId="15" pageNumber="96" type="description">
|
||
<paragraph id="335936E0FFC1F52FFF1DFCD1FEDCFC85" blockId="15.[189,335,794,820]" box="[189,335,794,820]" pageId="15" pageNumber="96">
|
||
<heading id="6811818CFFC1F52FFF1DFCD1FEDCFC85" bold="true" box="[189,335,794,820]" fontSize="11" level="3" pageId="15" pageNumber="96" reason="3">
|
||
<emphasis id="0192EAF2FFC1F52FFF1DFCD1FEDCFC85" bold="true" box="[189,335,794,820]" pageId="15" pageNumber="96">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="335936E0FFC1F52FFF4AFC82FEB9FCD2" blockId="15.[189,1399,841,973]" box="[234,298,841,867]" pageId="15" pageNumber="96">
|
||
<emphasis id="0192EAF2FFC1F52FFF4AFC82FEB9FCD2" bold="true" box="[234,298,841,867]" pageId="15" pageNumber="96">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="335936E0FFC1F52FFF1DFCA7FBCFFC7C" blockId="15.[189,1399,841,973]" pageId="15" pageNumber="96">
|
||
<smallCapsWord id="35BFA03CFFC1F52FFF1DFCA7FEFDFC35" baselines="895,895" box="[189,366,876,902]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="15" pageNumber="96">MEASUREMENTS</smallCapsWord>
|
||
(in mm; n = 7). Wing length 3.04 (2.8–3.4), width 1.53 (1.3–1.63); head length 0.53 (0.56–0.50), width 0.63 (0.66–0.60); antennal segments, scape 0.13, pedicel 0.08, flagellomeres 1: 0.11, 2–12: 0.10, 13:0.11, 14: 0.36; palpal segments 1: 0.05, 2: 0.01, 3: 0.08, 4: 0.05.
|
||
</paragraph>
|
||
<paragraph id="335936E0FFC1F52FFF1DFC3FFE6DFA96" blockId="15.[188,1399,1011,1319]" pageId="15" pageNumber="96">
|
||
<smallCapsWord id="35BFA03CFFC1F52FFF1DFC3FFF6FFBBD" baselines="1031,1031" box="[189,252,1012,1038]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="15" pageNumber="96">HEAD</smallCapsWord>
|
||
(
|
||
<typeStatus id="EC5D8842FFC1F52FFEAEFC38FEE5FBBC" box="[270,374,1011,1037]" pageId="15" pageNumber="96" type="holotype">holotype</typeStatus>
|
||
). Wider than long; eye bridge contiguous, eye bridge with two facet rows on narrowest part and three facet rows on remaining part of bridge; no enlarged alveoli on postocular area; interocular suture appears absent (
|
||
<figureCitation id="ABDD2A65FFC1F52FFE6AFBF1FDB9FBE5" box="[458,554,1082,1108]" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[258,1333,330,1766]" captionTargetId="figure-12@13.[226,1345,302,1787]" captionTargetPageId="13" captionText="Fig. 7. Eugenys upsilon sp. nov. A. ♂, paratype (ZFMK-DIP-00097921), head. B. ♂, paratype (ZFMK- DIP-00097914), wing. C. ♂, paratype (ZFMK-DIP-00097921), mouth parts. D. ♂, paratype (ZFMK- DIP-00097921), first antennal segments. E–H. ♂, holotype (ZFMK-DIP-00097918). E. Aedeagus (lateral view). F. Aedeagus (ventral view). G. Epandrium and hypopods (ventral view). H. Aedeagus, gonocoxites, and gonostyli (ventral view). Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284647" httpUri="https://zenodo.org/record/11284647/files/figure.png" pageId="15" pageNumber="96">Fig. 7A</figureCitation>
|
||
). Antenna with scape wider than long, asymmetrical; pedicel spherical, about 2 × as wide as long; 14 flagellomeres, 1–13 fusiform, flagellomere 14 elongated, 3 × as long as previous flagellomere, ending in digitiform apiculus, flagellomeres with patch of darkened sensilla; ascoids very long, at least length of three flagellomeres, rod-like and with zigzag shape; frontal alveoli patch egg-shaped; mouthparts reduced, not extending beyond margin of head. Palpal segments short and sclerotized, apical segment not corrugated; not extending more than half of flagellomere 4, palpal proportions 1.0:2.0:1.6:1.0.
|
||
</paragraph>
|
||
<paragraph id="335936E0FFC1F52FFF1DFA84FCD5FA1E" blockId="15.[189,1399,1358,1456]" pageId="15" pageNumber="96">
|
||
<smallCapsWord id="35BFA03CFFC1F52FFF1DFA84FE8AFAD6" baselines="1378,1378" box="[189,281,1359,1385]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="15" pageNumber="96">THORAX</smallCapsWord>
|
||
. Without allurement organs; wing length about 2 × its width, hyaline, with alveoli in whole surface (
|
||
<figureCitation id="ABDD2A65FFC1F52FFE82FAB9FEEDFA3D" box="[290,382,1394,1420]" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[258,1333,330,1766]" captionTargetId="figure-12@13.[226,1345,302,1787]" captionTargetPageId="13" captionText="Fig. 7. Eugenys upsilon sp. nov. A. ♂, paratype (ZFMK-DIP-00097921), head. B. ♂, paratype (ZFMK- DIP-00097914), wing. C. ♂, paratype (ZFMK-DIP-00097921), mouth parts. D. ♂, paratype (ZFMK- DIP-00097921), first antennal segments. E–H. ♂, holotype (ZFMK-DIP-00097918). E. Aedeagus (lateral view). F. Aedeagus (ventral view). G. Epandrium and hypopods (ventral view). H. Aedeagus, gonocoxites, and gonostyli (ventral view). Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284647" httpUri="https://zenodo.org/record/11284647/files/figure.png" pageId="15" pageNumber="96">Fig. 7B</figureCitation>
|
||
); Sc vein almost straight, ending at origin of R
|
||
<subScript id="AF6234A5FFC1F52FFC03FA4FFC3FFA23" attach="left" box="[931,940,1412,1426]" fontSize="6" pageId="15" pageNumber="96">5</subScript>
|
||
, fork of R
|
||
<subScript id="AF6234A5FFC1F52FFB86FA4EFBD2FA22" attach="left" box="[1062,1089,1413,1427]" fontSize="6" pageId="15" pageNumber="96">2+3</subScript>
|
||
basal to M
|
||
<subScript id="AF6234A5FFC1F52FFB64FA4EFB4CFA22" attach="left" box="[1220,1247,1413,1427]" fontSize="6" pageId="15" pageNumber="96">1+2</subScript>
|
||
, R
|
||
<subScript id="AF6234A5FFC1F52FFAA1FA4EFA8FFA22" attach="left" box="[1281,1308,1413,1427]" fontSize="6" pageId="15" pageNumber="96">2+3</subScript>
|
||
joining R
|
||
<subScript id="AF6234A5FFC1F52FFF71FA63FF49FA07" attach="left" box="[209,218,1448,1462]" fontSize="6" pageId="15" pageNumber="96">4</subScript>
|
||
, wing apex rounded, R5 ending slightly below apex.
|
||
</paragraph>
|
||
<paragraph id="335936E0FFC1F52FFF1DFA1CFD77F89C" blockId="15.[188,1401,1494,1837]" pageId="15" pageNumber="96">
|
||
<smallCapsWord id="35BFA03CFFC1F52FFF1DFA1CFED0FA5E" baselines="1514,1514" box="[189,323,1495,1521]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Terminalia" pageId="15" pageNumber="96">TERMINALIA</smallCapsWord>
|
||
. Hypandrium narrow, plate-like, and rounded, Gonocoxites about half length of gonostyli, about as wide as long, gonocoxites joined in middle, and together giving origin to semi-sclerotized parameral sheath protruding beyond aedeagus, covering aedeagus only on sternal surface (
|
||
<figureCitation id="ABDD2A65FFC1F52FFB44F9D6FAD5F986" box="[1252,1350,1565,1591]" captionStart="Fig" captionStartId="6.[189,232,1822,1848]" captionTargetBox="[333,1275,301,1755]" captionTargetId="figure-14@6.[256,1331,265,1784]" captionTargetPageId="6" captionText="Fig. 3. Male genitalia, ventral view. A. Eugenys cymosa Quate, 1999, paratype (BMNH(E)2001-8). B. Eugenys panamensis Quate, 1999, paratype (BMNH(E)-2001-8). C. Eugenys clavellata Quate, 1996, paratype (USNMENT00857092). D. Eugenys upsilon sp. nov., holotype (ZFMK-DIP-00097918). E. Eugenys micra sp. nov., holotype (ZFMK-DIP-00097917). F. Eugenis singularis sp. nov., holotype (INBIO-CRI001470242). Red arrows point to the parameral sclerites. Photos: A–B, D–F by Santiago Jaume-Schinkel, C by David Pecor, Smithsonian Institute Museum. Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284629" httpUri="https://zenodo.org/record/11284629/files/figure.png" pageId="15" pageNumber="96">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation id="ABDD2A65FFC1F52FFAF2F9D6FF45F9EB" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[258,1333,330,1766]" captionTargetId="figure-12@13.[226,1345,302,1787]" captionTargetPageId="13" captionText="Fig. 7. Eugenys upsilon sp. nov. A. ♂, paratype (ZFMK-DIP-00097921), head. B. ♂, paratype (ZFMK- DIP-00097914), wing. C. ♂, paratype (ZFMK-DIP-00097921), mouth parts. D. ♂, paratype (ZFMK- DIP-00097921), first antennal segments. E–H. ♂, holotype (ZFMK-DIP-00097918). E. Aedeagus (lateral view). F. Aedeagus (ventral view). G. Epandrium and hypopods (ventral view). H. Aedeagus, gonocoxites, and gonostyli (ventral view). Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284647" httpUri="https://zenodo.org/record/11284647/files/figure.png" pageId="15" pageNumber="96">7F, H</figureCitation>
|
||
); gonostyli narrow, outwardly curved; ejaculatory apodeme slightly longer than aedeagus, narrow but broadening towards basal margin, aedeagus curved in ventral view, and claw-shaped in lateral view, aedeagus with heavily sclerotized lump at base, which in lateral view extends towards dorsal surface (
|
||
<figureCitation id="ABDD2A65FFC1F52FFF68F962FEB6F975" box="[200,293,1705,1732]" captionStart="Fig" captionStartId="13.[189,232,1822,1848]" captionTargetBox="[258,1333,330,1766]" captionTargetId="figure-12@13.[226,1345,302,1787]" captionTargetPageId="13" captionText="Fig. 7. Eugenys upsilon sp. nov. A. ♂, paratype (ZFMK-DIP-00097921), head. B. ♂, paratype (ZFMK- DIP-00097914), wing. C. ♂, paratype (ZFMK-DIP-00097921), mouth parts. D. ♂, paratype (ZFMK- DIP-00097921), first antennal segments. E–H. ♂, holotype (ZFMK-DIP-00097918). E. Aedeagus (lateral view). F. Aedeagus (ventral view). G. Epandrium and hypopods (ventral view). H. Aedeagus, gonocoxites, and gonostyli (ventral view). Scales in millimeters (mm). Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.11284647" httpUri="https://zenodo.org/record/11284647/files/figure.png" pageId="15" pageNumber="96">Fig. 7E</figureCitation>
|
||
); parameral sclerite forming single Y-shaped sclerotized structure, linked to aedeagal sheath and aedeagus; epandrium about 4 × as wide as long; hypopods elongate, longer than epandrium, tapering towards apex, apex of hypopods with single tenaculum with crowned apex. Epiproct tongue-shaped with rounded margin, covered in small setulae.
|
||
</paragraph>
|
||
<paragraph id="335936E0FFC1F52FFF4AF89EFED5F8DE" blockId="15.[189,1399,1877,2009]" box="[234,326,1877,1903]" pageId="15" pageNumber="96">
|
||
<emphasis id="0192EAF2FFC1F52FFF4AF89EFED5F8DE" bold="true" box="[234,326,1877,1903]" pageId="15" pageNumber="96">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="335936E0FFC1F52FFF1DF8B3FC27F869" blockId="15.[189,1399,1877,2009]" pageId="15" pageNumber="96">
|
||
Same as male except for palpi consisting of three segments; terminal flagellomeres absent in material examined; maximum number of flagellomeres present: 6. Mouthparts slightly more developed than in male, mouthparts with teeth-like sclerotized structures (
|
||
<figureCitation id="ABDD2A65FFC1F52FFCEAF875FC34F869" box="[842,935,1982,2008]" captionStart="Fig" captionStartId="14.[189,232,1963,1989]" captionTargetBox="[249,1354,380,1923]" captionTargetId="figure-15@14.[208,1380,265,1923]" captionTargetPageId="14" captionText="Fig. 8. Eugenys upsilon sp. nov., ♀, paratype (ZFMK-DIP-00097916). A. Head. B. Wing. C. Mouth parts. D. Genitalia (ventral view). Scales in millimeters (mm)." figureDoi="http://doi.org/10.5281/zenodo.11284649" httpUri="https://zenodo.org/record/11284649/files/figure.png" pageId="15" pageNumber="96">Fig. 8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BFC656BFFDEF530FF1DFEC3FD72FEE0" box="[189,737,264,338]" pageId="16" pageNumber="97" type="distribution">
|
||
<paragraph id="335936E0FFDEF530FF1DFEC3FECBFE93" blockId="16.[189,344,264,290]" box="[189,344,264,290]" pageId="16" pageNumber="97">
|
||
<heading id="6811818CFFDEF530FF1DFEC3FECBFE93" bold="true" box="[189,344,264,290]" fontSize="11" level="3" pageId="16" pageNumber="97" reason="3">
|
||
<emphasis id="0192EAF2FFDEF530FF1DFEC3FECBFE93" bold="true" box="[189,344,264,290]" pageId="16" pageNumber="97">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="335936E0FFDEF530FF1DFEFCFD72FEE0" blockId="16.[189,737,311,338]" box="[189,737,311,338]" pageId="16" pageNumber="97">
|
||
Only known from the
|
||
<typeStatus id="EC5D8842FFDEF530FE62FEF3FE66FEE3" box="[450,501,312,338]" pageId="16" pageNumber="97">type</typeStatus>
|
||
locality in
|
||
<collectingCountry id="4BF17670FFDEF530FDD9FEFCFD4EFEE0" box="[633,733,311,337]" name="Ecuador" pageId="16" pageNumber="97">Ecuador</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BFC656BFFDEF530FF1DFEB3FED4FDB6" pageId="16" pageNumber="97" type="biology_ecology">
|
||
<paragraph id="335936E0FFDEF530FF1DFEB3FEB9FE23" blockId="16.[189,298,376,402]" box="[189,298,376,402]" pageId="16" pageNumber="97">
|
||
<heading id="6811818CFFDEF530FF1DFEB3FEB9FE23" bold="true" box="[189,298,376,402]" fontSize="11" level="3" pageId="16" pageNumber="97" reason="3">
|
||
<emphasis id="0192EAF2FFDEF530FF1DFEB3FEB9FE23" bold="true" box="[189,298,376,402]" pageId="16" pageNumber="97">Genetics</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="335936E0FFDEF530FF1DFE6CFED4FDB6" blockId="16.[187,1398,423,519]" pageId="16" pageNumber="97">All specimens examined were successfully sequenced.All sequences are identical. Genebank’s accession numbers are OQ706367, OQ706373, OQ706374, OQ706368, OQ706372, OQ706366, OQ706371, OQ706369.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |