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<document id="487403EBEC75A70E5129721CCAA60EFF" ID-DOI="10.5281/zenodo.171270" ID-GBIF-Dataset="0205a7f4-1fa9-460d-95f0-31e5b931aa84" ID-ISSN="1175­5326" ID-Zenodo-Dep="171270" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459719919277" checkinUser="plazi" docAuthor="Macgowan, Iain" docDate="2005" docId="BB2287DBFFC6FFAB1B54FB6532D4CD0C" docLanguage="en" docName="zt00967.pdf" docOrigin="Zootaxa 967" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Lonchaea" docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="471BFFA3FFC5FFAD1A5CFFD13376CA62" masterDocTitle="New species of Lonchaeidae (Diptera: Schizophora) from central and southern Africa" masterLastPageNumber="23" masterPageNumber="1" pageNumber="4" updateTime="1698208965858" updateUser="plazi">
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<mods:title id="2752965A11064243C65BBB34F8EA24FB">New species of Lonchaeidae (Diptera: Schizophora) from central and southern Africa</mods:title>
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<mods:namePart id="B31CDF7D28B659733626B0D941F43230">Macgowan, Iain</mods:namePart>
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<mods:date id="392ABEA47EA0EFA8392D1BD8EA58DF9B">2005</mods:date>
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<treatment id="BB2287DBFFC6FFAB1B54FB6532D4CD0C" ID-DOI="http://doi.org/10.5281/zenodo.6267026" ID-GBIF-Taxon="119335443" ID-Zenodo-Dep="6267026" LSID="urn:lsid:plazi:treatment:BB2287DBFFC6FFAB1B54FB6532D4CD0C" httpUri="http://treatment.plazi.org/id/BB2287DBFFC6FFAB1B54FB6532D4CD0C" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
<subSubSection id="7B916546FFC6FFAE1B54FB65306CCEAC" box="[264,794,1204,1230]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="333436CDFFC6FFAE1B54FB65306CCEAC" blockId="3.[264,794,1204,1230]" box="[264,794,1204,1230]" pageId="3" pageNumber="4">
<heading id="687C81A1FFC6FFAE1B54FB65306CCEAC" bold="true" box="[264,794,1204,1230]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="01FFEADFFFC6FFAE1B54FB65306CCEAC" bold="true" box="[264,794,1204,1230]" pageId="3" pageNumber="4">
GENUS:
<taxonomicName id="F48B4D4EFFC6FFAE1B22FB653169CEAC" ID-CoL="9CJN5" box="[382,543,1204,1230]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="01FFEADFFFC6FFAE1B22FB653169CEAC" bold="true" box="[382,543,1204,1230]" italics="true" pageId="3" pageNumber="4">LONCHAEA</emphasis>
</taxonomicName>
FALLÉN, 1820: 25.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC6FFA91B54FB2F3027CE34" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="333436CDFFC6FFAE1B54FB2F31F5CF7B" blockId="3.[264,643,1278,1305]" box="[264,643,1278,1305]" pageId="3" pageNumber="4">
<taxonomicName id="F48B4D4EFFC6FFAE1B54FB2F3157CF7A" box="[264,545,1278,1304]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="asymmetrica" status="sp. nov.">
<emphasis id="01FFEADFFFC6FFAE1B54FB2F3157CF7A" bold="true" box="[264,545,1278,1304]" italics="true" pageId="3" pageNumber="4">Lonchaea asymmetrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ACC57A4FFC6FFAE1874FB2E31F5CF7B" box="[552,643,1279,1305]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="333436CDFFC6FFAE1B54FA9830E3CC49" blockId="3.[264,1324,1353,1899]" pageId="3" pageNumber="4">Description.Male: Head: Eyes bare. Frons rather wide, narrowing only slightly from the level of the ocellar triangle to the level of the antennal bases where it is approximately 60% the width of an eye, subshining with short hairs on mid and upper parts with some longer hairs present on the anterior margin above the lunule, all these hairs less than half the length of the orbital bristle. Orbital plate brightly shining and bare. When viewed from above, lunule, face and parafacials with a silvery sheen.</paragraph>
<paragraph id="333436CDFFC6FFAE1B64F9E83030CCC1" blockId="3.[264,1324,1353,1899]" pageId="3" pageNumber="4">Antenna all black. First flagellomere large and long, three times as long as it is deep reaching over the mouth edge. Arista yellow basally with very short pubescence, hairs on each side about as long as the width of the arista.</paragraph>
<paragraph id="333436CDFFC6FFA91B64F9603136CBB4" blockId="3.[264,1324,1353,1899]" lastBlockId="4.[264,1324,284,1310]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">Thorax: Thoracic dorsum subshining blue black with very short hairs these not more than one third the length of the orbital bristle. One propleural and one stigmatical bristle each about as strong as the other. Anepisternum with three strong anterior bristles, the lower one being rather weak, and two strong posterior bristles. Katepisternum with one strong bristle, no hairs posterior to or below this bristle. Scutellum with disc bare, rather brassy black, on the margin between the strong lateral and apical bristles with one short hair, no hairs between apical bristles. Squamae and fringes whitish, wings clear with light yellow veins. Wing length 3.0 mm. Legs black but with basal and second tarsomere of all legs clear yellow with a ventral fringe of yellow setulae, remaining tarsomeres darkened. Abdomen rather spherical.</paragraph>
<paragraph id="333436CDFFC1FFA91B64FE353799C974" blockId="4.[264,1324,284,1310]" pageId="4" pageNumber="5">
Male genitalia;
<figureCitation id="ABB02A48FFC1FFA91BACFE353143CB9C" box="[496,565,484,510]" captionStart="FIGURE 1" captionStartId="2.[264,367,1748,1772]" captionTargetBox="[305,1280,334,1709]" captionTargetId="figure@2.[295,1292,329,1721]" captionTargetPageId="2" captionText="FIGURE 1. Male genitalia of: Lamprolonchaea longicercus sp. nov. 1. epandrium and associated structures, lateral view. 2. posterior view. 3. aedeagus lateral view. Lonchaea asymmetrica sp. nov. 4. epandrium and associated structures, lateral view. 5. ventral view. Lonchaea grandiseta sp. nov. 6. epandrium and associated structures, lateral view. 7. ventral view. 8. aedeagus lateral view." httpUri="https://zenodo.org/record/171271/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
;
<emphasis id="01FFEADFFFC1FFA91818FE35311ECB9F" box="[580,616,484,509]" italics="true" pageId="4" pageNumber="5">4,5</emphasis>
. Cerci fused, with a sclerotized swelling near the base similar to that found in
<taxonomicName id="F48B4D4EFFC1FFA91BA4FDDD3040C844" authority="McAlpine, 1960" authorityName="McAlpine" authorityYear="1960" box="[504,822,524,550]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="viriosa">
<emphasis id="01FFEADFFFC1FFA91BA4FDDD311FC847" box="[504,617,524,549]" italics="true" pageId="4" pageNumber="5">L. viriosa</emphasis>
<bibRefCitation id="571A4B3CFFC1FFA9182EFDDD3040C844" author="McAlpine" box="[626,822,524,550]" lastPageId="4" lastPageNumber="23" pageId="4" pageNumber="22" refString="McAlpine, J. F. (1960) Chapter XVI. Diptera (Brachycera): Lonchaeidae. In Hanstrom, B., Brink, P. &amp; Rudebeck, G. (Eds.). South African animal life. Results of the Lund University Expedition 1950 - 1951. Volume 7. Almqvist and Wisell, Goteborg, Stockholm, Upsalla, pp. 327 - 376." type="book chapter" year="1960">McAlpine, 1960</bibRefCitation>
</taxonomicName>
, this is best seen in lateral view and gives the cerci an extended, sinuous appearance. Epandrium in lateral view slightly higher than it is wide, rather rectangular. In lateral view surstyli only extending slightly beyond shell of epandrium along its ventral edge, no posteroventral process. Surstyli differing in shape, right one broader than the left and with a deep circular excavation posteriorly taking up more than a third of the depth of the surstylus, the left one with a smaller excavation taking up about a quarter of the total depth. Aedeagus long and sinuate with a bulbous apex.
</paragraph>
<paragraph id="333436CDFFC1FFA91B64FCF5316FC95C" blockId="4.[264,1324,284,1310]" box="[312,537,804,830]" pageId="4" pageNumber="5">Female: Unknown.</paragraph>
<paragraph id="333436CDFFC1FFA91B64FC9D3027CE34" blockId="4.[264,1324,284,1310]" pageId="4" pageNumber="5">
Differential diagnosis: This species belongs to the
<taxonomicName id="F48B4D4EFFC1FFA919C2FC9D37CFC907" box="[926,1209,844,869]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="impressifrons">
<emphasis id="01FFEADFFFC1FFA919C2FC9D37CFC907" box="[926,1209,844,869]" italics="true" pageId="4" pageNumber="5">Lonchaea impressifrons</emphasis>
</taxonomicName>
­group of species recognised by
<bibRefCitation id="571A4B3CFFC1FFA9184BFCA5319FC9EC" author="McAlpine" box="[535,745,884,910]" lastPageId="4" lastPageNumber="23" pageId="4" pageNumber="22" refString="McAlpine, J. F. (1960) Chapter XVI. Diptera (Brachycera): Lonchaeidae. In Hanstrom, B., Brink, P. &amp; Rudebeck, G. (Eds.). South African animal life. Results of the Lund University Expedition 1950 - 1951. Volume 7. Almqvist and Wisell, Goteborg, Stockholm, Upsalla, pp. 327 - 376." type="book chapter" year="1960">McAlpine (1960)</bibRefCitation>
. The long sinuous aedeagus with a bulbous tip shows it to be close to species such as
<taxonomicName id="F48B4D4EFFC1FFA9188AFC4D375CC9D4" authority="Bezzi, 1920" authorityName="Bezzi" authorityYear="1920" box="[726,1066,924,950]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="impressifrons">
<emphasis id="01FFEADFFFC1FFA9188AFC4D30E2C9D7" box="[726,916,924,949]" italics="true" pageId="4" pageNumber="5">L. impressifrons</emphasis>
Bezzi, 1920
</taxonomicName>
and
<emphasis id="01FFEADFFFC1FFA91E38FC4D37A8C9D7" box="[1124,1246,924,949]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="F48B4D4EFFC1FFA91E38FC4D37ACC9D7" box="[1124,1242,924,949]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="viriosa">L. viriosa</taxonomicName>
.
</emphasis>
It differs from both these species however in that the surstyli do not extend posterioventraly beyond the shell of the epandrium and in the shape of the surstyli. McAlpine noted that in species of this group the pregonites are unequally developed, in
<taxonomicName id="F48B4D4EFFC1FFA91E64FBC53785CE4F" box="[1080,1267,1044,1069]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="asymmetrica">
<emphasis id="01FFEADFFFC1FFA91E64FBC53785CE4F" box="[1080,1267,1044,1069]" italics="true" pageId="4" pageNumber="5">L. asymmetrica</emphasis>
</taxonomicName>
this unequal development has extended to the surstyli.
</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC1FFA91B64FBB532C3CF7C" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="333436CDFFC1FFA91B64FBB5305FCEAC" blockId="4.[264,1324,284,1310]" pageId="4" pageNumber="5">
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<collectionCode id="559AAE08FFC1FFA91B64FBB532F7CE1C" box="[312,385,1124,1150]" pageId="4" pageNumber="5">TYPE</collectionCode>
</typeStatus>
MATERIAL:
<typeStatus id="EC30886FFFC1FFA91869FBB531D4CE1C" box="[565,674,1124,1150]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
male.
<emphasis id="01FFEADFFFC1FFA918A5FBB530E5CE1C" bold="true" box="[761,915,1124,1150]" pageId="4" pageNumber="5">
<collectingCountry id="4B9C765DFFC1FFA918A5FBB530E5CE1C" box="[761,915,1124,1150]" name="Tanzania" pageId="4" pageNumber="5">TANZANIA</collectingCountry>
</emphasis>
: East Usambara, Amani,
<quantity id="F4739B28FFC1FFA91E8EFBB53650CE1C" box="[1234,1318,1124,1150]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1000.0">1000m</quantity>
,
<date id="4735100DFFC1FFA91B54FB5D320CCEC4" box="[264,378,1164,1190]" pageId="4" pageNumber="5" value="1977-01-21">21.i.1977</date>
. H.Enghoff, O. Lomholdt, O. Martin leg. Only known from the
<typeStatus id="EC30886FFFC1FFA91E2EFB5D37ABCEC4" box="[1138,1245,1164,1190]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
. Specimen in the Zoological Museum, Copenhagen.
</paragraph>
<paragraph id="333436CDFFC1FFA91B64FB0D32C3CF7C" blockId="4.[264,1324,284,1310]" pageId="4" pageNumber="5">Etymology. The specific epithet refers to the asymetrical shape of the surstyli of the male genitalia.</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC1FFA91B54FAAD31BCCFF4" box="[264,714,1404,1430]" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="333436CDFFC1FFA91B54FAAD31BCCFF4" blockId="4.[264,714,1404,1430]" box="[264,714,1404,1430]" pageId="4" pageNumber="5">
<taxonomicName id="F48B4D4EFFC1FFA91B54FAAD31BCCFF4" ID-CoL="3VXH7" authority="McAlpine, 1960" authorityName="McAlpine" authorityYear="1960" box="[264,714,1404,1430]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="conicoura">
<emphasis id="01FFEADFFFC1FFA91B54FAAD3174CFF4" bold="true" box="[264,514,1404,1430]" italics="true" pageId="4" pageNumber="5">Lonchaea conicoura</emphasis>
<bibRefCitation id="571A4B3CFFC1FFA91855FAAD31BCCFF4" author="McAlpine" box="[521,714,1404,1430]" lastPageId="4" lastPageNumber="23" pageId="4" pageNumber="22" refString="McAlpine, J. F. (1960) Chapter XVI. Diptera (Brachycera): Lonchaeidae. In Hanstrom, B., Brink, P. &amp; Rudebeck, G. (Eds.). South African animal life. Results of the Lund University Expedition 1950 - 1951. Volume 7. Almqvist and Wisell, Goteborg, Stockholm, Upsalla, pp. 327 - 376." type="book chapter" year="1960">McAlpine, 1960</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC1FFA81B54FA1631A9CF0C" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="distribution">
<paragraph id="333436CDFFC1FFA91B54FA1630B6CC6B" blockId="4.[264,1323,1479,1625]" pageId="4" pageNumber="5">
Distribution: Recorded by
<bibRefCitation id="571A4B3CFFC1FFA9181BFA16306ECF83" author="McAlpine" box="[583,792,1479,1505]" lastPageId="4" lastPageNumber="23" pageId="4" pageNumber="22" refString="McAlpine, J. F. (1960) Chapter XVI. Diptera (Brachycera): Lonchaeidae. In Hanstrom, B., Brink, P. &amp; Rudebeck, G. (Eds.). South African animal life. Results of the Lund University Expedition 1950 - 1951. Volume 7. Almqvist and Wisell, Goteborg, Stockholm, Upsalla, pp. 327 - 376." type="book chapter" year="1960">McAlpine (1960)</bibRefCitation>
, who figured the male and female genitalia, from
<collectingCountry id="4B9C765DFFC1FFA91B14FA3E32B3CC6B" box="[328,453,1519,1545]" name="Zimbabwe" pageId="4" pageNumber="5">Zimbabwe</collectingCountry>
and
<collectingCountry id="4B9C765DFFC1FFA91BA2FA3E3136CC6B" box="[510,576,1519,1545]" name="South Africa" pageId="4" pageNumber="5">Natal</collectingCountry>
,
<collectingCountry id="4B9C765DFFC1FFA91811FA3E3190CC6B" box="[589,742,1519,1545]" name="South Africa" pageId="4" pageNumber="5">South Africa</collectingCountry>
. New to
<collectingCountry id="4B9C765DFFC1FFA91912FA3E30CDCC6B" box="[846,955,1519,1545]" name="Tanzania" pageId="4" pageNumber="5">Tanzania</collectingCountry>
.
</paragraph>
<paragraph id="333436CDFFC1FFA91B64F9C63656CC3B" blockId="4.[264,1323,1479,1625]" pageId="4" pageNumber="5">
Material examined:
<emphasis id="01FFEADFFFC1FFA91879F9C731C9CC52" bold="true" box="[549,703,1558,1584]" pageId="4" pageNumber="5">
<collectingCountry id="4B9C765DFFC1FFA91879F9C731C9CC52" box="[549,703,1558,1584]" name="Tanzania" pageId="4" pageNumber="5">TANZANIA</collectingCountry>
</emphasis>
: Uluguru Mts. Lupanga, West,
<quantity id="F4739B28FFC1FFA91E61F9C637E5CC53" box="[1085,1171,1559,1585]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="5" unit="m" value="1600.0">1600m</quantity>
.
<date id="4735100DFFC1FFA91EC2F9C63650CC53" box="[1182,1318,1559,1585]" pageId="4" pageNumber="5" value="1981-07-01">01.vii.1981</date>
, M. Stoltze &amp; N. Scharff leg.
<specimenCount id="258DFD44FFC1FFA9183EF9EE31C3CC3B" box="[610,693,1599,1625]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
. Specimen in the Zoological Museum, Copenhagen.
</paragraph>
<paragraph id="333436CDFFC1FFA91B54F967311FCCB3" blockId="4.[264,617,1718,1745]" box="[264,617,1718,1745]" pageId="4" pageNumber="5">
<taxonomicName id="F48B4D4EFFC1FFA91B54F9673171CCB2" box="[264,519,1718,1744]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="grandiseta" status="sp. nov.">
<emphasis id="01FFEADFFFC1FFA91B54F9673171CCB2" bold="true" box="[264,519,1718,1744]" italics="true" pageId="4" pageNumber="5">Lonchaea grandiseta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ACC57A4FFC1FFA91852F966311FCCB3" box="[526,617,1719,1745]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="333436CDFFC1FFA81B54F8D0313ECBCC" blockId="4.[264,1321,1793,1899]" lastBlockId="5.[264,1324,284,1870]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Description. Male: Head: Eyes bare, face and parafacials with a silvery white tomentum. Frons wide, above antennal bases about 25% of the head width, with a slight depression centrally. Hairs on upper part of frons very short, not more than one quarter the length of the orbital bristle. Antennae with first flagellomere very long and narrow, approximately 3.
<date id="4735100DFFC0FFA81B40FE953233CB3C" box="[284,325,324,350]" pageId="5" pageNumber="6">5 x</date>
as long as wide, reaching well below mouth edge, all black apart from basal half being orange ventrally and basally on its inner surface. Arista yellow orange at base, microscopically pubescent.
</paragraph>
<paragraph id="333436CDFFC0FFA81B64FE6D30D0C814" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
Thorax: One propleural and one stigmatical bristle. Katepisternum with one strong bristle with no hairs posterior to it. Anepisternum with three strong anterior bristles and four strong posterior bristles. Disc of scutellum bare. Scutellar margin with two hairs on the right side and one on the left side between the lateral and apical bristles. One hair between apical bristles (this only occurs in the
<typeStatus id="EC30886FFFC0FFA81972FD8D30ECC814" box="[814,922,604,630]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="333436CDFFC0FFA81B64FD553157C88C" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">Legs: Tarsomeres all yellow with slight darkening on apical segments. Squamae and fringes whitish, wings clear with veins light yellow. Pterostigmal section 1.5 times the length of the cross vein.</paragraph>
<paragraph id="333436CDFFC0FFA81B64FD2D3770C9BC" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
Male genitalia.
<figureCitation id="ABB02A48FFC0FFA81BA2FD2D3130C974" box="[510,582,764,790]" captionStart="FIGURE 1" captionStartId="2.[264,367,1748,1772]" captionTargetBox="[305,1280,334,1709]" captionTargetId="figure@2.[295,1292,329,1721]" captionTargetPageId="2" captionText="FIGURE 1. Male genitalia of: Lamprolonchaea longicercus sp. nov. 1. epandrium and associated structures, lateral view. 2. posterior view. 3. aedeagus lateral view. Lonchaea asymmetrica sp. nov. 4. epandrium and associated structures, lateral view. 5. ventral view. Lonchaea grandiseta sp. nov. 6. epandrium and associated structures, lateral view. 7. ventral view. 8. aedeagus lateral view." httpUri="https://zenodo.org/record/171271/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
;
<emphasis id="01FFEADFFFC0FFA8180BFD2D31E4C977" box="[599,658,764,789]" italics="true" pageId="5" pageNumber="6">6,7,8</emphasis>
. Cerci large and broad, forming a hood­like structure. Epandrium wider than high with three exceptionally strong bristles at posteroventral corner which are almost as long as epandrium is high, a small group of hairs anteriorly below these. Surstyli extending posteriorly from the shell of the epandrium as a rounded process produced into elongate spinules of various shape. Aedeagus U­shaped, without projections, basal portion relatively broad, apical part short and tubular.
</paragraph>
<paragraph id="333436CDFFC0FFA81B64FC3D311BCE34" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">Female similar to the male. Frons width approximately 30% of head width. Ovipositor rather simple, apical segment ventrally with two strong hairs subapically, these just over half the length of the segment.</paragraph>
<paragraph id="333436CDFFC0FFA81B64FBB531A9CF0C" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
Differential diagnosis: This species belongs to the
<taxonomicName id="F48B4D4EFFC0FFA819C2FBB537CFCE1F" box="[926,1209,1124,1149]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="impressifrons">
<emphasis id="01FFEADFFFC0FFA819C2FBB537CFCE1F" box="[926,1209,1124,1149]" italics="true" pageId="5" pageNumber="6">Lonchaea impressifrons</emphasis>
</taxonomicName>
group of species identified by
<bibRefCitation id="571A4B3CFFC0FFA8185DFB5D31A7CEC4" author="McAlpine" box="[513,721,1164,1190]" pageId="5" pageNumber="23" refString="McAlpine, J. F. (1964) Descriptions of new Lonchaeidae (Diptera). Canadian Entomologist, 96, 661 - 757." type="journal article" year="1964">McAlpine (1964)</bibRefCitation>
. This group is composed of a number of very similar small species which have a combination of yellow tarsomeres, clear wings and pale squamae, antennae with first flagellomere quite long, reaching to or beyond mouth margin and the frons marked by a more or less distinct, oval depression just below anterior ocellus. The scutellum has one or more lateral setulae on each side in addition to four scutellar bristles; without apical or discal setulae.
</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC0FFA81B64FAAD320DCC6C" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="333436CDFFC0FFA81B64FAAD320DCC6C" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
<taxonomicName id="F48B4D4EFFC0FFA81B64FAAD32AFCFF7" ID-CoL="3VXJN" box="[312,473,1404,1429]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="grandiseta">
<emphasis id="01FFEADFFFC0FFA81B64FAAD32AFCFF7" box="[312,473,1404,1429]" italics="true" pageId="5" pageNumber="6">L. grandiseta</emphasis>
</taxonomicName>
is perhaps most similar to
<taxonomicName id="F48B4D4EFFC0FFA8196EFAAD37FECFF4" ID-CoL="72N9M" authority="McAlpine, 1964" authorityName="McAlpine" authorityYear="1964" box="[818,1160,1404,1430]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="teratosa">
<emphasis id="01FFEADFFFC0FFA8196EFAAD30C0CFF7" box="[818,950,1404,1429]" italics="true" pageId="5" pageNumber="6">L. teratosa</emphasis>
<bibRefCitation id="571A4B3CFFC0FFA8199EFAAD37FECFF4" author="McAlpine" box="[962,1160,1404,1430]" pageId="5" pageNumber="23" refString="McAlpine, J. F. (1964) Descriptions of new Lonchaeidae (Diptera). Canadian Entomologist, 96, 661 - 757." type="journal article" year="1964">McAlpine, 1964</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="F48B4D4EFFC0FFA81E96FAAD315ECFDC" ID-CoL="3VXMG" authority="McAlpine, 1964" authorityName="McAlpine" authorityYear="1964" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="polyhamata">
<emphasis id="01FFEADFFFC0FFA81E96FAAD3229CFDF" italics="true" pageId="5" pageNumber="6">L. polyhamata</emphasis>
<bibRefCitation id="571A4B3CFFC0FFA81B3BFA75315ECFDC" author="McAlpine" box="[359,552,1444,1470]" pageId="5" pageNumber="23" refString="McAlpine, J. F. (1964) Descriptions of new Lonchaeidae (Diptera). Canadian Entomologist, 96, 661 - 757." type="journal article" year="1964">McAlpine, 1964</bibRefCitation>
</taxonomicName>
which both have spinules on the surstyli, both these species however lack the distinctive large bristles on the epandrium and both have a more complex aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC0FFAB1B64F9CD32C3CB3C" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="333436CDFFC0FFA81B64F9CD30D5CCE4" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
<typeStatus id="EC30886FFFC0FFA81B64F9CD32F7CC54" box="[312,385,1564,1590]" pageId="5" pageNumber="6">
<collectionCode id="559AAE08FFC0FFA81B64F9CD32F7CC54" box="[312,385,1564,1590]" pageId="5" pageNumber="6">TYPE</collectionCode>
</typeStatus>
MATERIAL:
<typeStatus id="EC30886FFFC0FFA8186CF9CD31EBCC54" box="[560,669,1564,1590]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
male,
<taxonomicName id="F48B4D4EFFC0FFA818ACF9CD3003CC54" box="[752,885,1564,1590]" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="17" phylum="Arthropoda" rank="species" species="namibia">
<emphasis id="01FFEADFFFC0FFA818ACF9CD3003CC54" bold="true" box="[752,885,1564,1590]" pageId="5" pageNumber="6">
<collectingCountry id="4B9C765DFFC0FFA818ACF9CD3003CC54" box="[752,885,1564,1590]" name="Namibia" pageId="5" pageNumber="6">NAMIBIA</collectingCountry>
</emphasis>
</taxonomicName>
: Khorixas District, Leekop 664, 19 53 15” S 14 21 44 E,
<date id="4735100DFFC0FFA8187AF99531BCCC3C" box="[550,714,1604,1630]" pageId="5" pageNumber="6" value="2001-10-26" valueMax="2001-10-30" valueMin="2001-10-26">2630.x.2001</date>
, A.H. Kirk­Spriggs &amp; E. Marais, Malaise trap sample riverbed. Specimen in National Museum of
<taxonomicName id="F48B4D4EFFC0FFA8196AF9BD30E9CCE4" box="[822,927,1644,1670]" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="17" phylum="Arthropoda" rank="species" species="namibia">
<collectingCountry id="4B9C765DFFC0FFA8196AF9BD30E9CCE4" box="[822,927,1644,1670]" name="Namibia" pageId="5" pageNumber="6">Namibia</collectingCountry>
</taxonomicName>
.
</paragraph>
<paragraph id="333436CDFFC0FFA81B64F9453203CC9C" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
<typeStatus id="EC30886FFFC0FFA81B64F94532D8CCCC" box="[312,430,1684,1710]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
.
<taxonomicName id="F48B4D4EFFC0FFA81BE2F9453135CCCC" box="[446,579,1684,1710]" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="17" phylum="Arthropoda" rank="species" species="namibia">
<collectingCountry id="4B9C765DFFC0FFA81BE2F9453135CCCC" box="[446,579,1684,1710]" name="Namibia" pageId="5" pageNumber="6">
<emphasis id="01FFEADFFFC0FFA81BE2F9453135CCCC" bold="true" box="[446,579,1684,1710]" pageId="5" pageNumber="6">NAMIBIA</emphasis>
</collectingCountry>
</taxonomicName>
: Windhoek, SE 2217 Ca,
<date id="4735100DFFC0FFA819CCF9453725CCCC" box="[912,1107,1684,1710]" pageId="5" pageNumber="6" value="1973-11-28" valueMax="1973-12-05" valueMin="1973-11-28">28.xi5.xii.1973</date>
,
<specimenCount id="258DFD44FFC0FFA81E39F94537CBCCCC" box="[1125,1213,1684,1710]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
, Malaise trap.
<specimenCount id="258DFD44FFC0FFA81B11F96D32C9CCB4" box="[333,447,1724,1750]" pageId="5" pageNumber="6" type="female">1 female</specimenCount>
, same locality,
<date id="4735100DFFC0FFA818C3F96D302DCCB4" box="[671,859,1724,1750]" pageId="5" pageNumber="6" value="1973-11-05" valueMax="1973-11-12" valueMin="1973-11-05">5.xi12.xi.1973</date>
. Specimens in National Museum of
<taxonomicName id="F48B4D4EFFC0FFA81B54F9353207CC9C" box="[264,369,1764,1790]" class="Insecta" family="Lonchaeidae" genus="Silba" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="17" phylum="Arthropoda" rank="species" species="namibia">
<collectingCountry id="4B9C765DFFC0FFA81B54F9353207CC9C" box="[264,369,1764,1790]" name="Namibia" pageId="5" pageNumber="6">Namibia</collectingCountry>
</taxonomicName>
.
</paragraph>
<paragraph id="333436CDFFC0FFA81B64F8DD37E4CD2C" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
<emphasis id="01FFEADFFFC0FFA81B64F8DD3296CD44" bold="true" box="[312,480,1804,1830]" pageId="5" pageNumber="6">
<collectingCountry id="4B9C765DFFC0FFA81B64F8DD3296CD44" box="[312,480,1804,1830]" name="Botswana" pageId="5" pageNumber="6">BOTSWANA</collectingCountry>
</emphasis>
:
<specimenCount id="258DFD44FFC0FFA81BA9F8DD3138CD44" box="[501,590,1804,1830]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
, Serowe, Gabarone Village, Museum garden, Malaise trap
<date id="4735100DFFC0FFA81B54F8E532F6CD2C" box="[264,384,1844,1870]" pageId="5" pageNumber="6" value="1993-09-18">18.9.1993</date>
, leg B.Viklund. Specimen in Natural History Museum, Stockholm.
</paragraph>
<paragraph id="333436CDFFC3FFAB1B64FECD32C3CB3C" blockId="6.[264,1322,284,350]" pageId="6" pageNumber="7">Etymology. The specific epithet refers to the large setae present on the epndrium of the male genitalia.</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC3FFAB1B54FE6037D1C8C9" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="333436CDFFC3FFAB1B54FE603198CBA9" blockId="6.[264,750,433,459]" box="[264,750,433,459]" pageId="6" pageNumber="7">
<taxonomicName id="F48B4D4EFFC3FFAB1B54FE603198CBA9" ID-CoL="3VXJR" authority="McAlpine, 1960" authorityName="McAlpine" authorityYear="1960" box="[264,750,433,459]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="haplosetifera">
<emphasis id="01FFEADFFFC3FFAB1B54FE603150CBA9" bold="true" box="[264,550,433,459]" italics="true" pageId="6" pageNumber="7">Lonchaea haplosetifera</emphasis>
<bibRefCitation id="571A4B3CFFC3FFAB1871FE603198CBA9" author="McAlpine" box="[557,750,433,459]" lastPageId="6" lastPageNumber="23" pageId="6" pageNumber="22" refString="McAlpine, J. F. (1960) Chapter XVI. Diptera (Brachycera): Lonchaeidae. In Hanstrom, B., Brink, P. &amp; Rudebeck, G. (Eds.). South African animal life. Results of the Lund University Expedition 1950 - 1951. Volume 7. Almqvist and Wisell, Goteborg, Stockholm, Upsalla, pp. 327 - 376." type="book chapter" year="1960">McAlpine, 1960</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="333436CDFFC3FFAB1B54FE2037D1C8C9" blockId="6.[264,1324,497,1603]" pageId="6" pageNumber="7">
<bibRefCitation id="571A4B3CFFC3FFAB1B54FE2032A2C869" author="McAlpine" box="[264,468,497,523]" lastPageId="6" lastPageNumber="23" pageId="6" pageNumber="22" refString="McAlpine, J. F. (1960) Chapter XVI. Diptera (Brachycera): Lonchaeidae. In Hanstrom, B., Brink, P. &amp; Rudebeck, G. (Eds.). South African animal life. Results of the Lund University Expedition 1950 - 1951. Volume 7. Almqvist and Wisell, Goteborg, Stockholm, Upsalla, pp. 327 - 376." type="book chapter" year="1960">McAlpine (1960)</bibRefCitation>
described this species from a series of females. They were characterised in his key by having a combination of a single katepisternal bristle, basal tarsomeres yellow and dark squamal fringes. In addition the lateral margin of scutellum has eight or ten fine setulae in multiserial rows between the usual scutellar bristles. The male described here also exhibits this combination of characters and is ascribed to this species.
</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC3FFAB1B64FD68323ACC21" pageId="6" pageNumber="7" type="description">
<paragraph id="333436CDFFC3FFAB1B64FD6832BAC9F9" blockId="6.[264,1324,497,1603]" pageId="6" pageNumber="7">Description. Male: Head: Eyes with very short and sparse hairs. Frons with very short hairs, no hairs present on the orbital plate above the orbital bristle. Lunule orange Frons and face with slight silver pollinosity. Antennae with first flagellomere long, extending down past the mouth margin, black dorsally and apically with orange colouration ventrally, becoming almost entirely orange basally. Pedicel also orange. Arista microscopically pubescent..</paragraph>
<paragraph id="333436CDFFC3FFAB1B64FC7832BDCE01" blockId="6.[264,1324,497,1603]" pageId="6" pageNumber="7">Thorax: Katepisternum with one single strong bristle with a single hair posterior to it. Scutellum on margin with multiserial rows of about 10 hairs on each side between apical and lateral bristles, no hairs between apical bristles. Squamae with fringe dark and rather densely haired. Legs black, tarsomeres almost entirely yellow, only slightly darkened on apical segments.</paragraph>
<paragraph id="333436CDFFC3FFAB1B64FBA032C4CF31" blockId="6.[264,1324,497,1603]" pageId="6" pageNumber="7">
Male genitalia:
<figureCitation id="ABB02A48FFC3FFAB185CFBA03137CEE9" box="[512,577,1137,1163]" captionStart="FIGURE 2" captionStartId="7.[264,367,1538,1562]" captionTargetBox="[357,1234,290,1522]" captionTargetId="figure@7.[346,1241,288,1526]" captionTargetPageId="7" captionText="FIGURE 2. Male genitalia of: Lonchaea haplosetifera McAlpine. 1. epandrium and associated structures, lateral view. 2. aedeagus lateral view. Silba budongo sp. nov. 3. epandrium and associated structures, lateral view. 4. ventral view. 5. aedeagus lateral view." httpUri="https://zenodo.org/record/171272/files/figure.png" pageId="6" pageNumber="7">Fig.2</figureCitation>
;
<emphasis id="01FFEADFFFC3FFAB1809FBA0310FCEE8" box="[597,633,1137,1162]" italics="true" pageId="6" pageNumber="7">1,2</emphasis>
. Epandrium slightly wider than high with a marked anteroventral process. Cerci as high as the epandrium is deep with a long basal stalk and with club shaped apex,, with short hairs on the apical club. Surstyli extending posteriorly from the epandrium as a broad column with a concave apex, bearing hairs on the dorsal margin at the apex. Aedeagus relatively simple and U­shaped but with more ornate processes basally.
</paragraph>
<paragraph id="333436CDFFC3FFAB1B64FAB03184CFC1" blockId="6.[264,1324,497,1603]" pageId="6" pageNumber="7">
Distribution: Recorded by
<bibRefCitation id="571A4B3CFFC3FFAB18D1FAB03029CF19" author="McAlpine" box="[653,863,1377,1403]" lastPageId="6" lastPageNumber="23" pageId="6" pageNumber="22" refString="McAlpine, J. F. (1960) Chapter XVI. Diptera (Brachycera): Lonchaeidae. In Hanstrom, B., Brink, P. &amp; Rudebeck, G. (Eds.). South African animal life. Results of the Lund University Expedition 1950 - 1951. Volume 7. Almqvist and Wisell, Goteborg, Stockholm, Upsalla, pp. 327 - 376." type="book chapter" year="1960">McAlpine (1960)</bibRefCitation>
from
<collectingCountry id="4B9C765DFFC3FFAB19EDFAB03759CF19" box="[945,1071,1377,1403]" name="Zimbabwe" pageId="6" pageNumber="7">Zimbabwe</collectingCountry>
and the
<collectingCountry id="4B9C765DFFC3FFAB1EFFFAB0329DCFC1" name="Democratic Republic of the Congo" pageId="6" pageNumber="7">Democratic Republic of Congo</collectingCountry>
. New to
<collectingCountry id="4B9C765DFFC3FFAB180FFA583198CFC1" box="[595,750,1417,1443]" name="South Africa" pageId="6" pageNumber="7">South Africa</collectingCountry>
.
</paragraph>
<paragraph id="333436CDFFC3FFAB1B64FA60323ACC21" blockId="6.[264,1324,497,1603]" pageId="6" pageNumber="7">
Material examined:
<emphasis id="01FFEADFFFC3FFAB1874FA60307FCFA9" bold="true" box="[552,777,1457,1483]" pageId="6" pageNumber="7">
<collectingCountry id="4B9C765DFFC3FFAB1874FA60307FCFA9" box="[552,777,1457,1483]" name="South Africa" pageId="6" pageNumber="7">SOUTH AFRICA</collectingCountry>
</emphasis>
: Cape Province; Bloukrans Pas, at Varkrivier (S
<geoCoordinate id="56BF500AFFC3FFAB1B7CFA0832E5CF91" box="[288,403,1497,1523]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="33.95">33 57 E</geoCoordinate>
23 38), Malaise trap, coastal rain forest in ravine.
<date id="4735100DFFC3FFAB1E42FA0837C8CF91" box="[1054,1214,1497,1523]" pageId="6" pageNumber="7" value="1994-10-14" valueMax="1994-10-19" valueMin="1994-10-14">14­19.x.1994</date>
. leg. M. Soderlund.
<specimenCount id="258DFD44FFC3FFAB1BCFF9D032A8CC79" box="[403,478,1537,1563]" pageId="6" pageNumber="7" type="male">1male</specimenCount>
. Specimen in the collections of the Natural History Museum, Stockholm.
</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC3FFAB1B54F94031ABCCC9" box="[264,733,1681,1707]" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="333436CDFFC3FFAB1B54F94031ABCCC9" blockId="6.[264,733,1681,1707]" box="[264,733,1681,1707]" pageId="6" pageNumber="7">
<taxonomicName id="F48B4D4EFFC3FFAB1B54F94031ABCCC9" ID-CoL="3VXMG" authority="McAlpine, 1964" authorityName="McAlpine" authorityYear="1964" box="[264,733,1681,1707]" class="Insecta" family="Lonchaeidae" genus="Lonchaea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="polyhamata">
<emphasis id="01FFEADFFFC3FFAB1B54F9403163CCC9" bold="true" box="[264,533,1681,1707]" italics="true" pageId="6" pageNumber="7">Lonchaea polyhamata</emphasis>
<bibRefCitation id="571A4B3CFFC3FFAB1840F94031ABCCC9" author="McAlpine" box="[540,733,1681,1707]" pageId="6" pageNumber="23" refString="McAlpine, J. F. (1964) Descriptions of new Lonchaeidae (Diptera). Canadian Entomologist, 96, 661 - 757." type="journal article" year="1964">McAlpine, 1964</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7B916546FFC3FFAB1B54F90D32D4CD0C" pageId="6" pageNumber="7" type="distribution">
<paragraph id="333436CDFFC3FFAB1B54F90D3668CC94" blockId="6.[264,1323,1756,1902]" box="[264,1310,1756,1782]" pageId="6" pageNumber="7">
Distribution: Recorded by
<bibRefCitation id="571A4B3CFFC3FFAB181DF90D307BCC94" author="McAlpine" box="[577,781,1756,1782]" pageId="6" pageNumber="23" refString="McAlpine, J. F. (1964) Descriptions of new Lonchaeidae (Diptera). Canadian Entomologist, 96, 661 - 757." type="journal article" year="1964">McAlpine (1964)</bibRefCitation>
from
<collectingCountry id="4B9C765DFFC3FFAB1900F90D30E8CC94" box="[860,926,1756,1782]" name="South Africa" pageId="6" pageNumber="7">Natal</collectingCountry>
,
<collectingCountry id="4B9C765DFFC3FFAB19F7F90D3733CC94" box="[939,1093,1756,1782]" name="South Africa" pageId="6" pageNumber="7">South Africa</collectingCountry>
. New to
<collectingCountry id="4B9C765DFFC3FFAB1EF1F90D366CCC94" box="[1197,1306,1756,1782]" name="Tanzania" pageId="6" pageNumber="7">Tanzania</collectingCountry>
.
</paragraph>
<paragraph id="333436CDFFC3FFAB1B64F8D532D4CD0C" blockId="6.[264,1323,1756,1902]" pageId="6" pageNumber="7">
Material examined:
<collectingCountry id="4B9C765DFFC3FFAB1819F8D531A9CD7C" box="[581,735,1796,1822]" name="Tanzania" pageId="6" pageNumber="7">
<emphasis id="01FFEADFFFC3FFAB1819F8D531A9CD7C" bold="true" box="[581,735,1796,1822]" pageId="6" pageNumber="7">TANZANIA</emphasis>
</collectingCountry>
: West Usambara Mts., Mazumbai,
<quantity id="F4739B28FFC3FFAB1E8EF8D53650CD7C" box="[1234,1318,1796,1822]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="7" unit="m" value="1600.0">1600m</quantity>
,
<date id="4735100DFFC3FFAB1B54F8FD32EFCD24" box="[264,409,1836,1862]" pageId="6" pageNumber="7" value="1980-08-01">01.viii.1980</date>
, leg. M.Stoltze &amp; N. Scharff.
<specimenCount id="258DFD44FFC3FFAB195AF8FD302DCD24" box="[774,859,1836,1862]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
. Specimen in the Zoological Museum, Copenhagen.
</paragraph>
</subSubSection>
</treatment>
</document>