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247 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.745.23685" ID-PMC="PMC5904534" ID-Pensoft-Pub="1313-2970-745-101" ID-Pensoft-UUID="047DFFCAFFA5F32EA97C873F4708943F" ID-PubMed="29670449" ID-Zenodo-Dep="1222435" ID-ZooBank="8785DC6BC2A244FD94B6243EB07C717F" ModsDocID="1313-2970-745-101" checkinTime="1521799180220" checkinUser="pensoft" docAuthor="Moore, Matthew R., Cave, Ronald D. & Branham, Marc A." docDate="2018" docId="B15F91733361DB3B2A1AC7662A33ECB8" docLanguage="en" docName="ZooKeys 745: 101-378" docOrigin="ZooKeys 745" docPubDate="2018-03-22" docSource="http://dx.doi.org/10.3897/zookeys.745.23685" docTitle="Ancognatha ustulata" docType="treatment" docVersion="6" id="047DFFCAFFA5F32EA97C873F4708943F" lastPageNumber="117" masterDocId="047DFFCAFFA5F32EA97C873F4708943F" masterDocTitle="Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)" masterLastPageNumber="378" masterPageNumber="101" pageNumber="116" updateTime="1668165599527" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moore, Matthew R.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
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<mods:nameIdentifier type="email">cyclocephala@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cave, Ronald D.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of Florida, Indian River Research and Education Center, 2199 South Rock Road, Fort Pierce, FL 34945, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Branham, Marc A.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-03-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>745</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>101</mods:start>
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<mods:end>378</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.745.23685</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.745.23685</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-745-101</mods:identifier>
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<mods:identifier type="ZooBank">8785DC6BC2A244FD94B6243EB07C717F</mods:identifier>
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<mods:identifier type="Pensoft-UUID">047DFFCAFFA5F32EA97C873F4708943F</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1222435</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="142480108" LSID="urn:lsid:plazi:treatment:B15F91733361DB3B2A1AC7662A33ECB8" httpUri="http://treatment.plazi.org/id/B15F91733361DB3B2A1AC7662A33ECB8" lastPageId="16" lastPageNumber="117" pageId="15" pageNumber="116">
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<subSubSection pageId="15" pageNumber="116" type="nomenclature">
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<paragraph pageId="15" pageNumber="116">
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<taxonomicName LSID="B15F9173-3361-DB3B-2A1A-C7662A33ECB8" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Ancognatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancognatha ustulata" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="ustulata">Ancognatha ustulata (Burmeister, 1847)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="116" type="reference_group">
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<paragraph pageId="15" pageNumber="116">
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<taxonomicName authorityName="Burmeister" authorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclocephala ustulata" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="ustulata">Cyclocephala ustulata</taxonomicName>
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Burmeister, 1847: 39 [original combination].
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</paragraph>
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<paragraph pageId="15" pageNumber="116">
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<taxonomicName baseAuthorityName="Burmeister" baseAuthorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Ancognatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancognatha ustulata" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="ustulata">Ancognatha ustulata</taxonomicName>
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(Burmeister) [new combination by
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<bibRefCitation author="Bates, HW" pageId="164" pageNumber="265" refId="B65" refString="Bates, HW, 1888. Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296-342." title="Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296 - 342." year="1888">Bates 1888</bibRefCitation>
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: 297].
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</paragraph>
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<paragraph pageId="15" pageNumber="116">
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<emphasis bold="true" pageId="15" pageNumber="116">syn.</emphasis>
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<taxonomicName class="Insecta" family="Scarabaeidae" genus="Ancognatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancognatha ustulata subsp. ustulatoides" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="subSpecies" species="ustulata" subSpecies="ustulatoides">Ancognatha ustulata ustulatoides</taxonomicName>
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<normalizedToken originalValue="Höhne">Hoehne</normalizedToken>
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, 1922d: 373-374 [original combination].
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</paragraph>
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<paragraph pageId="15" pageNumber="116">
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<taxonomicName class="Insecta" family="Scarabaeidae" genus="Ancognatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancognatha ustulata subsp. var. var. ustulatoides" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="variety" species="ustulata" subSpecies="var." variety="ustulatoides">Ancognatha ustulata var. ustulatoides</taxonomicName>
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<normalizedToken originalValue="Höhne">Hoehne</normalizedToken>
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[new infrasubspecific status by
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<bibRefCitation author="Arrow, GJ" pageId="164" pageNumber="265" refId="B51" refString="Arrow, GJ, 1937b. Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1-124." title="Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1 - 124." year="1937 b">Arrow 1937b</bibRefCitation>
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: 6].
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</paragraph>
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<paragraph pageId="15" pageNumber="116">
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<taxonomicName authority="ustulatoides" class="Insecta" family="Scarabaeidae" genus="Ancognatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancognatha ustulata subsp. ab." order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="subSpecies" species="ustulata" subSpecies="ab.">Ancognatha ustulata ab. ustulatoides</taxonomicName>
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<normalizedToken originalValue="Höhne">Hoehne</normalizedToken>
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[revised infrasubspecific status by
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<bibRefCitation author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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1985a
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</bibRefCitation>
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: 162].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="116" type="types">
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<paragraph pageId="15" pageNumber="116">Types.</paragraph>
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<paragraph pageId="15" pageNumber="116">
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Lectotype ♂ of
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<taxonomicName family="Scarabaeidae" kingdom="Animalia" lsidName="C. ustulata" order="Coleoptera" pageId="15" pageNumber="116" rank="species" species="ustulata">
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<emphasis italics="true" pageId="15" pageNumber="116">C. ustulata</emphasis>
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</taxonomicName>
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at MLUH (
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<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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1966
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</bibRefCitation>
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). Lectotype ♂ of
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<taxonomicName lsidName="A. ustulata subsp. ustulatoides" pageId="15" pageNumber="116" rank="subSpecies" species="ustulata" subSpecies="ustulatoides">
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<emphasis italics="true" pageId="15" pageNumber="116">A. ustulata ustulatoides</emphasis>
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</taxonomicName>
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at ZMHB (
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<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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1966
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="116" type="distribution">
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<paragraph pageId="15" pageNumber="116">Distribution.</paragraph>
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<paragraph pageId="15" pageNumber="116">
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COLOMBIA: Antioquia,
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<normalizedToken originalValue="Bogotá">Bogota</normalizedToken>
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D. C.,
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<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
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, Caldas, Cauca,
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<normalizedToken originalValue="Chocó">Choco</normalizedToken>
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, Cundinamarca, Tolima, Valle del Cauca. ECUADOR. PERU: Pasco. VENEZUELA:
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<normalizedToken originalValue="Mérida">Merida</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="116" type="references">
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<paragraph pageId="15" pageNumber="116">References.</paragraph>
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<paragraph pageId="15" pageNumber="116">
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<bibRefCitation author="Dejean, PFMA" pageId="174" pageNumber="275" refId="B252" refString="Dejean, PFMA, 1833. Catalogue des Coleopteres de la collection de M. le Comte Dejean. Mequignon-Marvis,Paris, 443 pp. [Livraisons 1-2 (1833) : 1-176; Livraison 3 (1834): 177-256; Livraison 4 (1835): 257-360; Livraison 5 (1836a): 361-443]." title="Catalogue des Coleopteres de la collection de M. le Comte Dejean. Mequignon-Marvis, Paris, 443 pp. [Livraisons 1 - 2 (1833): 1 - 176; Livraison 3 (1834): 177 - 256; Livraison 4 (1835): 257 - 360; Livraison 5 (1836 a): 361 - 443]." year="1833">Dejean 1833</bibRefCitation>
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,
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<bibRefCitation author="Dejean, PFMA" pageId="174" pageNumber="275" refId="B253" refString="Dejean, PFMA, 1836b. Catalogue des coleopteres de la collection de M. le comte Dejean. Troisieme edition, revue, corrigee et augmentee. Mequignon-Marvis Pere et Fils, Paris, xiv, 503 pp. [Livraisons 1-4 (1836b) : 1-384; Livraison 5 (1837): i-xiv, 385-503]." title="Catalogue des coleopteres de la collection de M. le comte Dejean. Troisieme edition, revue, corrigee et augmentee. Mequignon-Marvis Pere et Fils, Paris, xiv, 503 pp. [Livraisons 1 - 4 (1836 b): 1 - 384; Livraison 5 (1837): i-xiv, 385 - 503]." year="1836 b">1836b</bibRefCitation>
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,
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.37837" author="Sturm, J" pageId="227" pageNumber="328" refId="B1135" refString="Sturm, J, 1843. Catalog der Kaefer-Sammlung von Jacob Sturm. Sebald's Buchdruckerei, Nuremberg, 386 pp. [+ 6 pls] https://doi.org/10.5962/bhl.title.37837" title="Catalog der Kaefer-Sammlung von Jacob Sturm. Sebald's Buchdruckerei, Nuremberg, 386 pp. [+ 6 pls]" url="https://doi.org/10.5962/bhl.title.37837" year="1843">Sturm 1843</bibRefCitation>
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,
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<bibRefCitation author="Burmeister, H" pageId="168" pageNumber="269" refId="B140" refString="Burmeister, H, 1847. Handbuch der Entomologie, volume 5. TCF Enslin, Berlin, 584 pp." title="Handbuch der Entomologie, volume 5. TCF Enslin, Berlin, 584 pp." year="1847">Burmeister 1847</bibRefCitation>
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,
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<bibRefCitation author="Hoehne, W" pageId="194" pageNumber="295" refId="B566" refString="Hoehne, W, 1922d. Ancognatha ustulata Burm. n. subsp. ustulatoides (Col. Dyn.). Deutsche Entomologische Zeitchrift 1922: 373-374." title="Ancognatha ustulata Burm. n. subsp. ustulatoides (Col. Dyn.). Deutsche Entomologische Zeitchrift 1922: 373 - 374." year="1922 d">
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<normalizedToken originalValue="Höhne">Hoehne</normalizedToken>
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1922d
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</bibRefCitation>
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,
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<bibRefCitation author="Arrow, GJ" pageId="164" pageNumber="265" refId="B51" refString="Arrow, GJ, 1937b. Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1-124." title="Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1 - 124." year="1937 b">Arrow 1937b</bibRefCitation>
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,
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<bibRefCitation author="Blackwelder, RE" pageId="167" pageNumber="268" refId="B101" refString="Blackwelder, RE, 1944. Checklist of coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2. Bulletin of the United States National Museum 185: 189-341." title="Checklist of coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2. Bulletin of the United States National Museum 185: 189 - 341." year="1944">Blackwelder 1944</bibRefCitation>
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,
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<bibRefCitation author="Apolinar Maria, H" pageId="163" pageNumber="264" refId="B33" refString="Apolinar Maria, H, 1946. Miscelanea entomologica. Revista de la Academia Colombiana de Ciencias Exactas, Fiscas y Naturales 5: 552-553." title="Miscelanea entomologica. Revista de la Academia Colombiana de Ciencias Exactas, Fiscas y Naturales 5: 552 - 553." year="1946">Apolinar Maria 1946</bibRefCitation>
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,
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<bibRefCitation author="Pike, KS" pageId="213" pageNumber="314" refId="B899" refString="Pike, KS, River, RL, Ratcliffe, BC, Oseto, CY, Mayo, ZB, 1976. A world bibliography of the genus Cyclocephala (Coleoptera: Scarabaeidae). Miscellaneous Publication of the University of Nebraska Agricultural Experiment Station 32: 1-32." title="A world bibliography of the genus Cyclocephala (Coleoptera: Scarabaeidae). Miscellaneous Publication of the University of Nebraska Agricultural Experiment Station 32: 1 - 32." year="1976">Pike et al. 1976</bibRefCitation>
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,
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<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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1966
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</bibRefCitation>
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,
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<bibRefCitation author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
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, Restrepo et al. 2003,
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Neita-Moreno and
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<normalizedToken originalValue="Morón">Moron</normalizedToken>
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2008
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</bibRefCitation>
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.34.309" author="Gasca-Alvarez, HJ" pageId="182" pageNumber="283" refId="B381" refString="Gasca-Alvarez, HJ, Amat-Garcia, G, 2010. Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia. ZooKeys 34: 153-192. https://doi.org/10.3897/zookeys.34.309" title="Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia. ZooKeys 34: 153 - 192." url="https://doi.org/10.3897/zookeys.34.309" year="2010">
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<normalizedToken originalValue="Gasca-Álvarez">Gasca-Alvarez</normalizedToken>
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et al. 2015
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<bibRefCitation DOI="https://doi.org/10.1649/072.070.0107" author="Figueroa, L" pageId="180" pageNumber="281" refId="B348" refString="Figueroa, L, Ratcliffe, BC, 2016. A new species of Ancognatha Erichson (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) from Peru, with distributions of Peruvian Ancognatha species. The Coleopterists Bulletin 70: 65-72. https://doi.org/10.1649/072.070.0107" title="A new species of Ancognatha Erichson (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) from Peru, with distributions of Peruvian Ancognatha species. The Coleopterists Bulletin 70: 65 - 72." url="https://doi.org/10.1649/072.070.0107" year="2016">Figueroa and Ratcliffe 2016</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="117" type="remarks">
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<paragraph pageId="16" pageNumber="117">
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<pageBreakToken pageId="16" pageNumber="117" start="start">Remarks</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="16" pageNumber="117">
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<taxonomicName class="Insecta" family="Scarabaeidae" genus="Ancognatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ancognatha ustulata subsp. ustulatoides" order="Coleoptera" pageId="16" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="ustulata" subSpecies="ustulatoides">
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<emphasis italics="true" pageId="16" pageNumber="117">Ancognatha ustulata ustulatoides</emphasis>
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</taxonomicName>
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was a validly described subspecies (
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<bibRefCitation author="Hoehne, W" pageId="194" pageNumber="295" refId="B566" refString="Hoehne, W, 1922d. Ancognatha ustulata Burm. n. subsp. ustulatoides (Col. Dyn.). Deutsche Entomologische Zeitchrift 1922: 373-374." title="Ancognatha ustulata Burm. n. subsp. ustulatoides (Col. Dyn.). Deutsche Entomologische Zeitchrift 1922: 373 - 374." year="1922 d">
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<normalizedToken originalValue="Höhne">Hoehne</normalizedToken>
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1922d
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</bibRefCitation>
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). The subspecies was considered an infrasubspecific entity after
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<bibRefCitation author="Arrow, GJ" pageId="164" pageNumber="265" refId="B51" refString="Arrow, GJ, 1937b. Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1-124." title="Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1 - 124." year="1937 b">Arrow (1937b)</bibRefCitation>
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and was referred to as an
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<normalizedToken originalValue="“ab.”">"ab."</normalizedToken>
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or
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<normalizedToken originalValue="“var.”">"var."</normalizedToken>
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(e.g., see
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<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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1966
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</bibRefCitation>
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,
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<bibRefCitation author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
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). The name has not been clearly synonymized with
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<taxonomicName lsidName="A. ustulata" pageId="16" pageNumber="117" rank="species" species="ustulata">
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<emphasis italics="true" pageId="16" pageNumber="117">A. ustulata</emphasis>
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</taxonomicName>
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(Burmeister), but was listed as a synonym by
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<bibRefCitation author="Krajcik, M" pageId="200" pageNumber="301" refId="B658" refString="Krajcik, M, 2005. Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma.X, Supplement No. 2, Pilsen, 122 pp." title="Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma. X, Supplement No. 2, Pilsen, 122 pp." year="2005">Krajcik (2005)</bibRefCitation>
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.
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<bibRefCitation author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
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<normalizedToken originalValue="Endrődi">Endrodi</normalizedToken>
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(1966
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</bibRefCitation>
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,
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<bibRefCitation author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
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) reported
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<taxonomicName lsidName="A. ustulata" pageId="16" pageNumber="117" rank="species" species="ustulata">
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<emphasis italics="true" pageId="16" pageNumber="117">A. ustulata</emphasis>
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|
</taxonomicName>
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|
from Mexico and Panama, but major faunistic studies have not found additional specimens from these countries (
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<bibRefCitation author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B974" refString="Ratcliffe, BC, 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1-506." title="The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." year="2003">Ratcliffe 2003</bibRefCitation>
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,
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<bibRefCitation author="Ratcliffe, BC" pageId="219" pageNumber="320" refId="B985" refString="Ratcliffe, BC, Cave, RD, Cano, EB, 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1-666." title="The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 666." year="2013">Ratcliffe et al. 2013</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |