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<mods:title id="E45E9F8FCA5FBE52C77C7E5AA7378CC6">Two new species of the Hypostomus cochliodon group (Siluriformes: Loricariidae) from the rio Paraguay basin, with a redescription of Hypostomus cochliodon Kner, 1854</mods:title>
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<mods:namePart id="21AEC82845D14168018614C052D4B9F3">Tencatt, Luiz F. C.</mods:namePart>
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<emphasis id="17ECEAD1055F1B13BDD362E7FB3E0D41" bold="true" box="[942,1191,1005,1029]" italics="true" pageId="12" pageNumber="597">Hypostomus khimaera</emphasis>
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<figureCitation id="BDA32A46055F1B13BD906504FBD90D61" box="[1005,1088,1037,1061]" captionStart="Fig" captionStartId="4.[136,175,1321,1345]" captionTargetBox="[136,779,174,1299]" captionTargetId="figure-618@4.[136,779,174,1299]" captionTargetPageId="4" captionText="Fig. 3. Opercle morphology of (a) Hypostomus cochliodon, NUP 10809, 215.8 mm SL, showing indistinct opercle without odontodes; (b) Hypostomus basilisko, holotype, MZUSP 111110, 182.5 mm SL and (c) Hypostomus khimaera, holotype, MZUSP 111129, 139.1 mm SL, showing layer of skin surrounding opercle that supports odontodes." figureDoi="http://doi.org/10.5281/zenodo.12718833" httpUri="https://zenodo.org/record/12718833/files/figure.png" pageId="12" pageNumber="597">Figs. 3c</figureCitation>
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<figureCitation id="BDA32A46055F1B13BA376504FBFD0D61" box="[1098,1124,1037,1061]" captionStart="Fig" captionStartId="5.[129,168,1840,1864]" captionTargetBox="[129,772,1051,1805]" captionTargetId="figure-947@5.[129,772,1051,1805]" captionTargetPageId="5" captionText="Fig. 4. Lateral view of juvenile in (a) Hypostomus cochliodon, NUP 11956, 92.8 mm SL, (b) Hypostomus basilisko, paratype, ZUFMS-PIS 3083, 64.2 mm SL and (c) Hypostomus khimaera, NUP13494,68.8 mmSL." figureDoi="http://doi.org/10.5281/zenodo.12718835" httpUri="https://zenodo.org/record/12718835/files/figure.png" pageId="12" pageNumber="597">4c</figureCitation>
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<figureCitation id="BDA32A46055F1B13BA136504FB100D61" box="[1134,1161,1037,1061]" captionStart="Fig" captionStartId="7.[129,168,1567,1591]" captionTargetBox="[129,772,176,1543]" captionTargetId="figure-69@7.[129,772,176,1543]" captionTargetPageId="7" captionText="Fig. 5. Mouth and teeth of (a) Hypostomus cochliodon, NUP 10807, 208.0 mm SL, showing one series of papillae in inner face of upper and lower jaws; (b) Hypostomus basilisko, holotype, MZUSP 111110, 182.5 mm SL, showing the agglomerated papillae on inner face of upper and lower jaws; and (c) Hypostomus khimaera, MZUSP 111129, holotype, 139.1 mm SL, showing weakly developed papillae not forming series." figureDoi="http://doi.org/10.5281/zenodo.12718839" httpUri="https://zenodo.org/record/12718839/files/figure.png" pageId="12" pageNumber="597">5c</figureCitation>
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<figureCitation id="BDA32A46055F1B13BAEF6504FB340D61" box="[1170,1197,1037,1061]" captionStart="Fig" captionStartId="7.[809,850,1441,1465]" captionTargetBox="[809,1452,174,1419]" captionTargetId="figure-141@7.[809,1452,174,1419]" captionTargetPageId="7" captionText="Fig. 6. Keels, ridges and odontodes of (a) Hypostomus cochliodon, NUP 10807, 208.0 mm SL, showing two weakly developed ridges in compound pterotic, weakly developed keels, few and weakly developed odontodes and predorsal keels weakly developed; (b) Hypostomus basilisko, holotype, MZUSP 111110, 182.5 mm SL, showing the three ridges on top of head, strongly developed keels, numerous welldeveloped odontodes on plates covering body and head and strong V-shaped predorsal keels; (c) Hypostomus khimaera, holotype, MZUSP 111129, 139.1 mm SL, showing moderately developed keels and dermal plates supporting moderately developed odontodes." figureDoi="http://doi.org/10.5281/zenodo.12718837" httpUri="https://zenodo.org/record/12718837/files/figure.png" pageId="12" pageNumber="597">6c</figureCitation>
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<figureCitation id="BDA32A46055F1B13BACB6504FB4B0D61" box="[1206,1234,1037,1061]" captionStart="Fig" captionStartId="13.[129,168,1904,1928]" captionTargetBox="[129,1452,401,1873]" captionTargetId="figure-170@13.[129,1452,401,1873]" captionTargetPageId="13" captionText="Fig. 13. Hypostomus khimaera, holotype, MZUSP 111129, 139.1 mm SL, from córrego Salobo, rio Paraguay basin, Mato Grosso State, Brazil; dorsal (top), lateral (middle), and ventral (bottom) views." figureDoi="http://doi.org/10.5281/zenodo.12718853" httpUri="https://zenodo.org/record/12718853/files/figure.png" pageId="12" pageNumber="597">13</figureCitation>
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<figureCitation id="BDA32A46055F1B13BAA16504FB6E0D61" box="[1244,1271,1037,1061]" captionStart="Fig" captionStartId="15.[129,168,922,946]" captionTargetBox="[129,1452,176,902]" captionTargetId="figure-474@15.[129,1452,176,902]" captionTargetPageId="15" captionText="Fig. 14. Color pattern of live specimen of Hypostomus khimaera, ZUFMS-PIS 3738, 10.6 mm SL, rioAquidauana, Mato Grosso do Sul, Brazil, showing the dark tan stripe along midline of flank and diffuse spots on the body." figureDoi="http://doi.org/10.5281/zenodo.12718855" httpUri="https://zenodo.org/record/12718855/files/figure.png" pageId="12" pageNumber="597">14</figureCitation>
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SL; 
<collectionCode id="4389AE06055F1B13BDA2643EFBB90C08" box="[991,1056,1335,1356]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="12" pageNumber="597">MNRJ</collectionCode>
<materialsCitation id="95F03C9E055F1B13BA5A643EFA2B0C08" box="[1063,1458,1335,1357]" collectionCode="MZUEL" pageId="12" pageNumber="597" specimenCode="2, 105.5" specimenCount="1" typeStatus="paratype">
40185, 
<specimenCode id="753E9EB8055F1B13BA0C643EFBE60C08" box="[1137,1151,1335,1356]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="12" pageNumber="597">2</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BAF6643EFB5A0C09" box="[1163,1219,1335,1357]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="12" pageNumber="597">105.5</specimenCode>
- 
<quantity id="E2609B26055F1B13BAB7643EFAB70C08" box="[1226,1326,1335,1357]" metricMagnitude="-1" metricUnit="m" metricValue="1.281" pageId="12" pageNumber="597" unit="mm" value="128.1">128.1 mm</quantity>
SL; 
<collectionCode id="4389AE06055F1B13BB22643EFA2B0C08" box="[1375,1458,1335,1356]" pageId="12" pageNumber="597">MZUEL</collectionCode>
</materialsCitation>
<materialsCitation id="95F03C9E055F1B13BD4D645AFB350C2D" box="[816,1196,1363,1385]" collectionCode="NUP" pageId="12" pageNumber="597" specimenCode="16" specimenCount="1" typeStatus="paratype">
5821, 
<specimenCode id="753E9EB8055F1B13BD10645AFC1E0C2D" box="[877,903,1363,1385]" collectionCode="MZUEL" pageId="12" pageNumber="597">16</specimenCode>
, 
<quantity id="E2609B26055F1B13BDED645AFBBF0C2D" box="[912,1062,1363,1385]" metricMagnitude="-2" metricUnit="m" metricValue="7.534999999999999" metricValueMax="11.32" metricValueMin="3.75" pageId="12" pageNumber="597" unit="mm" value="75.35" valueMax="113.2" valueMin="37.5">37.5-113.2 mm</quantity>
SL; and 
<collectionCode id="4389AE06055F1B13BA01645DFB350C2D" box="[1148,1196,1364,1385]" pageId="12" pageNumber="597">NUP</collectionCode>
</materialsCitation>
<materialsCitation id="95F03C9E055F1B13BACF645AFBDD0CC2" collectingDate="1999-08-02" collectingDateMax="1999-08-03" collectingDateMin="1999-08-02" collectionCode="NUP" collectorName="Serra das Araras &amp; O. A. Shibatta." country="Brazil" latitude="-15.650833" location="Salobo" longLatPrecision="21" longitude="-57.215" municipality="Porto Estrela" pageId="12" pageNumber="597" specimenCode="4, 125.7" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
14506, 
<specimenCode id="753E9EB8055F1B13BA87645DFA910C2D" box="[1274,1288,1364,1385]" collectionCode="NUP" pageId="12" pageNumber="597">4</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BB6F645AFAD30C2D" box="[1298,1354,1363,1385]" collectionCode="NUP" pageId="12" pageNumber="597">125.7</specimenCode>
- 
<quantity id="E2609B26055F1B13BB2D645AFA2A0C2D" box="[1360,1459,1363,1385]" metricMagnitude="-1" metricUnit="m" metricValue="1.415" pageId="12" pageNumber="597" unit="mm" value="141.5">141.5 mm</quantity>
SL, collected with holotype
</materialsCitation>
.
</paragraph>
<paragraph id="252736C3055F1B11BD4D64A3FDF608D2" blockId="12.[816,1460,1450,1961]" lastBlockId="14.[136,779,183,406]" lastPageId="14" lastPageNumber="599" pageId="12" pageNumber="597">
<emphasis id="17ECEAD1055F1B13BD4D64A3FC6E0C84" bold="true" box="[816,1015,1450,1472]" pageId="12" pageNumber="597">Non-type material.</emphasis>
<materialsCitation id="95F03C9E055F1B11BA7C64A3FDF508D2" collectionCode="LBP, MCP, MZUSP, NUP" collectorName="Chapada dos Guimaraes &amp; de Melgaco &amp; de Chacorore. &amp; da Serra &amp; da Onca. &amp; das Antas. &amp; Terenos" country="Brazil" county="rio Paraguay" lastPageId="14" lastPageNumber="599" location="rio Salobra." municipality="rio Coxipo." pageId="12" pageNumber="597" specimenCode="LBP 5825, 1, LBP 8496, 10, MCP 15621, MCP 15785, 3, 85.4, MZUSP 44313, 4, MZUSP 78741, 5, MZUSP 78783, 7, MZUSP 78785, MZUSP 78853, MZUSP 79135, 2, MZUSP 82036, MZUSP 90458, NUP 915, 12, NUP 12393, NUP 2843, NUP 3000, 114.9, NUP 12014, NUP 3037, NUP 9723, NUP 11825, NUP 12005, NUP 12495, NUP 12006, NUP 12013, NUP 12015, NUP 12016, NUP 12121, NUP 13399, 136.2, NUP 13400, NUP 13401, NUP 13423, NUP 13480, 11, NUP 10982, NUP 11819, NUP 13269, NUP 13270, 9, 43.4, NUP 13494, ZUFMS-PIS 1316, 27.5, ZUFMS-PIS 1978, ZUFMS-PIS 3539, 20, ZUFMS-PIS 3557, 32, ZUFMS-PIS 3738" specimenCount="169" stateProvince="Mato Grosso">
<specimenCount id="339EFD4A055F1B13BA7C64A3FB0F0C84" box="[1025,1174,1450,1472]" count="169" pageId="12" pageNumber="597" type="generic">169 specimens</specimenCount>
. 
<emphasis id="17ECEAD1055F1B13BADE64A3FB7B0C84" bold="true" box="[1187,1250,1450,1472]" pageId="12" pageNumber="597">
<collectingCountry id="5D8F7653055F1B13BADE64A3FB7B0C84" box="[1187,1250,1450,1472]" name="Brazil" pageId="12" pageNumber="597">Brazil</collectingCountry>
</emphasis>
. 
<collectingCounty id="CC464E4F055F1B13BA8C64A3FAE90CFB" box="[1265,1392,1450,1472]" pageId="12" pageNumber="597">rio Paraguay</collectingCounty>
basin. 
<collectingRegion id="E75CF821055F1B13BD4D64CEFC290C98" box="[816,944,1479,1500]" country="Brazil" name="Mato Grosso" pageId="12" pageNumber="597">Mato Grosso</collectingRegion>
. 
<specimenCode id="753E9EB8055F1B13BDC464CEFB840C99" box="[953,1053,1479,1501]" collectionCode="LBP" pageId="12" pageNumber="597">LBP 5825</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BA5A64CEFBAA0C98" box="[1063,1075,1479,1500]" collectionCode="LBP" pageId="12" pageNumber="597">1</specimenCode>
, 
<quantity id="E2609B26055F1B13BA4264CEFB0D0C98" box="[1087,1172,1479,1501]" metricMagnitude="-2" metricUnit="m" metricValue="9.58" pageId="12" pageNumber="597" unit="mm" value="95.8">95.8 mm</quantity>
SL, Cuiabá, 
<collectingMunicipality id="C543ACB9055F1B13BB6E64CEFA1B0C99" box="[1299,1410,1479,1501]" pageId="12" pageNumber="597">rio Coxipó.</collectingMunicipality>
<specimenCode id="753E9EB8055F1B13BBFA64CEFCFB0CBD" collectionCode="LBP" pageId="12" pageNumber="597">LBP 8496</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BD1764EDFC1D0CBD" box="[874,900,1508,1529]" collectionCode="LBP" pageId="12" pageNumber="597">10</specimenCode>
, 
<quantity id="E2609B26055F1B13BDF164EAFB850CBD" box="[908,1052,1507,1529]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" metricValueMax="10.45" metricValueMin="4.75" pageId="12" pageNumber="597" unit="mm" value="76.0" valueMax="104.5" valueMin="47.5">47.5-104.5 mm</quantity>
SL, Porto Estrela, 
<location id="20476018055F1B13BAB764EAFAAE0CBD" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BAB764EAFAAE0CBD" box="[1226,1335,1507,1529]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Salobra." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Salobra.</location>
<specimenCode id="753E9EB8055F1B13BB4664EDFA290CBD" box="[1339,1456,1507,1529]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="12" pageNumber="597">MCP 15621</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BD4D6709FCA50F51" box="[816,828,1536,1557]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="12" pageNumber="597">1</specimenCode>
, 
<quantity id="E2609B26055F1B13BD376709FC390F52" box="[842,928,1536,1558]" metricMagnitude="-2" metricUnit="m" metricValue="8.65" pageId="12" pageNumber="597" unit="mm" value="86.5">86.5 mm</quantity>
SL; and 
<specimenCode id="753E9EB8055F1B13BD866709FBEA0F52" box="[1019,1139,1536,1558]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="12" pageNumber="597">MCP 15785</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BA036709FB120F51" box="[1150,1163,1536,1557]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="12" pageNumber="597">3</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BAEA6709FB5B0F52" box="[1175,1218,1536,1558]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="12" pageNumber="597">85.4</specimenCode>
-113.0 mm SL, unnamed stream. 
<specimenCode id="753E9EB8055F1B13BDFE6714FBB90F76" box="[899,1056,1565,1586]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">MZUSP 44313</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BA506714FBA20F76" box="[1069,1083,1565,1586]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">4</specimenCode>
, 
<quantity id="E2609B26055F1B13BA346714FB7C0F77" box="[1097,1253,1565,1587]" metricMagnitude="-2" metricUnit="m" metricValue="7.21" metricValueMax="11.39" metricValueMin="3.03" pageId="12" pageNumber="597" unit="mm" value="72.1" valueMax="113.9" valueMin="30.3">30.3-113.9 mm</quantity>
SL, Nova Olímpia, unnamed stream. 
<specimenCode id="753E9EB8055F1B13BDA56733FBF20F0B" box="[984,1131,1594,1615]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">MZUSP 78741</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BA096733FB180F14" box="[1140,1153,1594,1616]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">5</specimenCode>
, 
<quantity id="E2609B26055F1B13BAF66733FA850F0B" box="[1163,1308,1594,1616]" metricMagnitude="-2" metricUnit="m" metricValue="7.65" metricValueMax="10.129999999999999" metricValueMin="5.17" pageId="12" pageNumber="597" unit="mm" value="76.5" valueMax="101.3" valueMin="51.7">51.7-101.3 mm</quantity>
SL, 
<location id="20476018055F1B13BB346733FCE60F29" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BB346733FCE60F29" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="Reserva do Cabacal" pageId="12" pageNumber="597" stateProvince="Mato Grosso">Reserva do Cabaçal</location>
, ribeirão 
<location id="20476018055F1B13BDA9675EFBE70F29" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BDA9675EFBE70F29" box="[980,1150,1623,1645]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="Sete de Setembro." pageId="12" pageNumber="597" stateProvince="Mato Grosso">Sete de Setembro.</location>
<specimenCode id="753E9EB8055F1B13BAFC675EFA8A0F28" box="[1153,1299,1623,1644]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">MZUSP 78783</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BB67675EFABF0F28" box="[1306,1318,1623,1644]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">7</specimenCode>
of 8, 13.2-96.0 mm SL (13.2-96.0 mm SL), Indiavaí, 
<location id="20476018055F1B13BADD677AFB650FCD" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BADD677AFB650FCD" box="[1184,1276,1651,1673]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Jauru." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Jaurú.</location>
<specimenCode id="753E9EB8055F1B13BB7F677DFA0E0FCD" box="[1282,1431,1651,1673]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">MZUSP 78785</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BBDC677DFA360FCD" box="[1441,1455,1652,1673]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">4</specimenCode>
, 
<quantity id="E2609B26055F1B13BD4D6799FC2C0FE2" box="[816,949,1680,1702]" metricMagnitude="-2" metricUnit="m" metricValue="6.14" metricValueMax="6.63" metricValueMin="5.65" pageId="12" pageNumber="597" unit="mm" value="61.4" valueMax="66.3" valueMin="56.5">56.5-66.3 mm</quantity>
SL, 
<location id="20476018055F1B13BD9C6798FB070FE2" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BD9C6798FB070FE2" box="[993,1182,1680,1702]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="Reserva do Cabacal" pageId="12" pageNumber="597" stateProvince="Mato Grosso">Reserva do Cabaçal</location>
, 
<location id="20476018055F1B13BADB6799FA800FE2" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BADB6799FA800FE2" box="[1190,1305,1680,1702]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Cabacal." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Cabaçal.</location>
<specimenCode id="753E9EB8055F1B13BB606799FA360FE2" box="[1309,1455,1680,1702]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">MZUSP 78853</specimenCode>
, 
<quantity id="E2609B26055F1B13BD4D67A4FC3E0F87" box="[816,935,1709,1731]" metricMagnitude="0" metricUnit="m" metricValue="1.1567" pageId="12" pageNumber="597" unit="mm" value="1156.7">1, 156.7 mm</quantity>
SL, Diamantino, 
<location id="20476018055F1B13BA3367A4FB550F87" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BA3367A4FB550F87" box="[1102,1228,1709,1731]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Paraguay." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Paraguay.</location>
<specimenCode id="753E9EB8055F1B13BAAD67A4FAF80F87" box="[1232,1377,1709,1731]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">MZUSP 79135</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BB1767A4FAE10F86" box="[1386,1400,1709,1730]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">2</specimenCode>
, 
<quantity id="E2609B26055F1B13BBFC67A4FC1A0F9B" metricMagnitude="-2" metricUnit="m" metricValue="7.919999999999999" metricValueMax="8.879999999999999" metricValueMin="6.959999999999999" pageId="12" pageNumber="597" unit="mm" value="79.19999999999999" valueMax="88.8" valueMin="69.6">69.6- 88.8 mm</quantity>
SL, Cuiabá, 
<location id="20476018055F1B13BD8467C3FBF80FA4" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BD8467C3FBF80FA4" box="[1017,1121,1738,1760]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Cuiaba." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Cuiabá.</location>
<specimenCode id="753E9EB8055F1B13BA1E67C3FB6D0FA4" box="[1123,1268,1738,1760]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">MZUSP 82036</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BA8667C3FA9E0F9B" box="[1275,1287,1738,1759]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">1</specimenCode>
, 109.0 mm SL, 
<location id="20476018055F1B13BBE467C3FC090FB9" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BBE467C3FC090FB9" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Sepotuba." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Sepotuba.</location>
<specimenCode id="753E9EB8055F1B13BDE867EEFBB00FB9" box="[917,1065,1767,1789]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">MZUSP 90458</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BA4E67EEFBA60FB8" box="[1075,1087,1767,1788]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="597">1</specimenCode>
, 
<quantity id="E2609B26055F1B13BA3667EEFB060FB8" box="[1099,1183,1767,1789]" metricMagnitude="-2" metricUnit="m" metricValue="9.45" pageId="12" pageNumber="597" unit="mm" value="94.5">94.5 mm</quantity>
SL, 
<location id="20476018055F1B13BAB367EEFAD70FB9" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BAB367EEFAD70FB9" box="[1230,1358,1767,1789]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Sepotuba." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Sepotuba.</location>
<specimenCode id="753E9EB8055F1B13BB2E67EEFA360FB9" box="[1363,1455,1767,1789]" collectionCode="NUP" pageId="12" pageNumber="597">NUP 915</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BD4D660DFCD30E5D" box="[816,842,1796,1817]" collectionCode="NUP" pageId="12" pageNumber="597">12</specimenCode>
, 
<quantity id="E2609B26055F1B13BD29660AFC7F0E5D" box="[852,998,1795,1817]" metricMagnitude="-2" metricUnit="m" metricValue="9.94" metricValueMax="11.61" metricValueMin="8.27" pageId="12" pageNumber="597" unit="mm" value="99.4" valueMax="116.1" valueMin="82.7">82.7-116.1 mm</quantity>
SL; and 
<specimenCode id="753E9EB8055F1B13BA41660DFB290E5D" box="[1084,1200,1796,1817]" collectionCode="NUP" pageId="12" pageNumber="597">NUP 12393</specimenCode>
, 
<quantity id="E2609B26055F1B13BAC4660DFAA80E5D" box="[1209,1329,1795,1817]" metricMagnitude="0" metricUnit="m" metricValue="1.1465999999999998" pageId="12" pageNumber="597" unit="mm" value="1146.6">1, 146.6 mm</quantity>
SL, 
<collectorName id="886D5315055F1B13BB22660AFC5B0E72" pageId="12" pageNumber="597">Chapada dos Guimarães</collectorName>
, Manso Reservoir. 
<specimenCode id="753E9EB8055F1B13BA066628FB7B0E71" box="[1147,1250,1824,1846]" collectionCode="NUP" pageId="12" pageNumber="597">NUP 2843</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BA916629FB630E71" box="[1260,1274,1824,1845]" collectionCode="NUP" pageId="12" pageNumber="597">2</specimenCode>
, 
<quantity id="E2609B26055F1B13BB796629FA130E72" box="[1284,1418,1824,1846]" metricMagnitude="-2" metricUnit="m" metricValue="8.265" metricValueMax="8.95" metricValueMin="7.58" pageId="12" pageNumber="597" unit="mm" value="82.65" valueMax="89.5" valueMin="75.8">75.8-89.5 mm</quantity>
SL, 
<collectorName id="886D5315055F1B13BD4D6634FB850E17" box="[816,1052,1853,1875]" pageId="12" pageNumber="597">Chapada dos Guimarães</collectorName>
, 
<location id="20476018055F1B13BA5B6634FB360E17" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BA5B6634FB360E17" box="[1062,1199,1853,1875]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Quilombo." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Quilombo.</location>
<specimenCode id="753E9EB8055F1B13BAC96637FA840E16" box="[1204,1309,1853,1875]" collectionCode="NUP" pageId="12" pageNumber="597">NUP 3000</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BB5A6634FAAC0E16" box="[1319,1333,1853,1874]" collectionCode="NUP" pageId="12" pageNumber="597">2</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BB426634FAEF0E17" box="[1343,1398,1853,1875]" collectionCode="NUP" pageId="12" pageNumber="597">114.9</specimenCode>
- 
<quantity id="E2609B26055F1B13BB006634FCCC0E2B" metricMagnitude="-1" metricUnit="m" metricValue="1.225" pageId="12" pageNumber="597" unit="mm" value="122.5">122.5 mm</quantity>
SL; and 
<specimenCode id="753E9EB8055F1B13BDDA6653FB810E2B" box="[935,1048,1882,1903]" collectionCode="NUP" pageId="12" pageNumber="597">NUP 12014</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BA626653FBB20E2B" box="[1055,1067,1882,1903]" collectionCode="NUP" pageId="12" pageNumber="597">1</specimenCode>
, 
<quantity id="E2609B26055F1B13BA486653FB110E2B" box="[1077,1160,1882,1903]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="12" pageNumber="597" unit="mm" value="27.2">27.2 mm</quantity>
SL, 
<location id="20476018055F1B13BACF6653FA290E2B" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BACF6653FA290E2B" box="[1202,1456,1882,1904]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="Santo Antonio do Leverger" pageId="12" pageNumber="597" stateProvince="Mato Grosso">Santo Antônio do Leverger</location>
, 
<location id="20476018055F1B13BD4D667EFC020EC9" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BD4D667EFC020EC9" box="[816,923,1911,1933]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Cuiaba." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Cuiabá.</location>
<specimenCode id="753E9EB8055F1B13BDE2667EFB9F0EC8" box="[927,1030,1911,1932]" collectionCode="NUP" pageId="12" pageNumber="597">NUP 3037</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BA72667EFB820EC8" box="[1039,1051,1911,1932]" collectionCode="NUP" pageId="12" pageNumber="597">1</specimenCode>
, 
<quantity id="E2609B26055F1B13BA5B667EFBE30EC8" box="[1062,1146,1911,1932]" metricMagnitude="-2" metricUnit="m" metricValue="8.92" pageId="12" pageNumber="597" unit="mm" value="89.2">89.2 mm</quantity>
SL, 
<collectorName id="886D5315055F1B13BADA667EFA090EC9" box="[1191,1424,1911,1933]" pageId="12" pageNumber="597">Chapada dos Guimarães</collectorName>
, 
<location id="20476018055F1B13BBE5667EFCE90EED" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BBE5667EFCE90EED" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Casca." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Casca.</location>
<specimenCode id="753E9EB8055F1B13BD0A669DFC780EED" box="[887,993,1940,1961]" collectionCode="NUP" pageId="12" pageNumber="597">NUP 9723</specimenCode>
, 
<specimenCode id="753E9EB8055F1B13BD91669DFC630EED" box="[1004,1018,1940,1961]" collectionCode="NUP" pageId="12" pageNumber="597">2</specimenCode>
, 
<quantity id="E2609B26055F1B13BA7B669AFB170EED" box="[1030,1166,1939,1961]" metricMagnitude="-2" metricUnit="m" metricValue="6.709999999999999" metricValueMax="7.24" metricValueMin="6.18" pageId="12" pageNumber="597" unit="mm" value="67.1" valueMax="72.4" valueMin="61.8">61.8-72.4 mm</quantity>
SL, Cáceres, 
<location id="20476018055F1B13BB64669AFA2D0EED" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055F1B13BB64669AFA2D0EED" box="[1305,1460,1939,1961]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Piraputanga." pageId="12" pageNumber="597" stateProvince="Mato Grosso">rio Piraputanga.</location>
<specimenCode id="753E9EB8055E1B12BEFC61B8FF680983" box="[129,241,177,199]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 11825</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BE8761B8FE9F0982" box="[250,262,177,198]" collectionCode="NUP" pageId="13" pageNumber="598">1</specimenCode>
c&amp;s, 
<quantity id="E2609B26055E1B12BF4761B8FE140982" box="[314,397,177,199]" metricMagnitude="-2" metricUnit="m" metricValue="6.95" pageId="13" pageNumber="598" unit="mm" value="69.5">69.5 mm</quantity>
SL, Cáceres, córrego Piraputanga. 
<specimenCode id="753E9EB8055E1B12BCAE61B8FF2609A7" collectionCode="NUP" pageId="13" pageNumber="598">NUP 12005</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BEB561C7FF4D09A7" box="[200,212,206,227]" collectionCode="NUP" pageId="13" pageNumber="598">1</specimenCode>
, 139.0 mm SL; and 
<specimenCode id="753E9EB8055E1B12BFEB61C7FD9309A7" box="[406,522,205,227]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 12495</specimenCode>
, 
<quantity id="E2609B26055E1B12BC6E61C7FD1209A7" box="[531,651,205,227]" metricMagnitude="0" metricUnit="m" metricValue="1.1084999999999998" pageId="13" pageNumber="598" unit="mm" value="1108.5">1, 108.5 mm</quantity>
SL, Rosário Oeste, 
<location id="20476018055E1B12BEBA61E3FEAD0844" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BEBA61E3FEAD0844" box="[199,308,234,256]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Manso." pageId="13" pageNumber="598" stateProvince="Mato Grosso">rio Manso.</location>
<specimenCode id="753E9EB8055E1B12BF4161E2FE2C0844" box="[316,437,234,256]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 12006</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BFBF61E2FE490844" box="[450,464,235,256]" collectionCode="NUP" pageId="13" pageNumber="598">2</specimenCode>
, 
<quantity id="E2609B26055E1B12BFA061E2FDED0844" box="[477,628,235,256]" metricMagnitude="-2" metricUnit="m" metricValue="9.944999999999999" metricValueMax="10.709999999999999" metricValueMin="9.18" pageId="13" pageNumber="598" unit="mm" value="99.44999999999999" valueMax="107.1" valueMin="91.8">91.8-107.1 mm</quantity>
SL, 
<location id="20476018055E1B12BCD461E2FF4E0859" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BCD461E2FF4E0859" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="Barao de Melgaco" pageId="13" pageNumber="598" stateProvince="Mato Grosso">
Barão 
<collectorName id="886D5315055E1B12BC9161E3FF4E0859" pageId="13" pageNumber="598">de Melgaço</collectorName>
</location>
, 
<location id="20476018055E1B12BE9F600EFED60859" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BE9F600EFED60859" box="[226,335,263,285]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Cuiaba." pageId="13" pageNumber="598" stateProvince="Mato Grosso">rio Cuiabá.</location>
<specimenCode id="753E9EB8055E1B12BF2B6001FE550858" box="[342,460,263,285]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 12013</specimenCode>
, 
<quantity id="E2609B26055E1B12BFA5600EFDCD0859" box="[472,596,263,285]" metricMagnitude="0" metricUnit="m" metricValue="1.1127" pageId="13" pageNumber="598" unit="mm" value="1112.7">1, 112.7 mm</quantity>
SL, 
<collectorName id="886D5315055E1B12BCFB600EFF7E087E" pageId="13" pageNumber="598">Chapada dos Guimarães</collectorName>
, 
<location id="20476018055E1B12BE93602DFECA087E" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BE93602DFECA087E" box="[238,339,292,314]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Cuiaba." pageId="13" pageNumber="598" stateProvince="Mato Grosso">rio Cuiabá.</location>
<specimenCode id="753E9EB8055E1B12BF28602DFE5D087E" box="[341,452,292,314]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 12015</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BFB6602DFE4E087D" box="[459,471,292,313]" collectionCode="NUP" pageId="13" pageNumber="598">1</specimenCode>
, 
<quantity id="E2609B26055E1B12BF9D602DFDA8087D" box="[480,561,292,314]" metricMagnitude="-2" metricUnit="m" metricValue="9.59" pageId="13" pageNumber="598" unit="mm" value="95.9">95.9 mm</quantity>
SL, 
<location id="20476018055E1B12BC25602DFC99087E" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BC25602DFC99087E" box="[600,768,292,314]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="Barao de Melgaco" pageId="13" pageNumber="598" stateProvince="Mato Grosso">
Barão 
<collectorName id="886D5315055E1B12BCEE602DFC99087E" box="[659,768,292,314]" pageId="13" pageNumber="598">de Melgaço</collectorName>
</location>
, baia 
<collectorName id="886D5315055E1B12BED26048FEAE0813" box="[175,311,321,343]" pageId="13" pageNumber="598">de Chacororé.</collectorName>
<specimenCode id="753E9EB8055E1B12BF406048FE280813" box="[317,433,321,343]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 12016</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BFC66048FE5E0812" box="[443,455,321,342]" collectionCode="NUP" pageId="13" pageNumber="598">1</specimenCode>
, 
<quantity id="E2609B26055E1B12BFA96048FDB30812" box="[468,554,321,343]" metricMagnitude="-2" metricUnit="m" metricValue="9.36" pageId="13" pageNumber="598" unit="mm" value="93.6">93.6 mm</quantity>
SL, 
<location id="20476018055E1B12BC276048FF450837" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BC276048FF450837" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="Santo Antonio do Leverger." pageId="13" pageNumber="598" stateProvince="Mato Grosso">Santo Antônio do Leverger.</location>
<specimenCode id="753E9EB8055E1B12BEA26057FEC80837" box="[223,337,350,371]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 12121</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BF246057FEFC0837" box="[345,357,350,371]" collectionCode="NUP" pageId="13" pageNumber="598">1</specimenCode>
, 
<quantity id="E2609B26055E1B12BF126057FE5B0837" box="[367,450,350,371]" metricMagnitude="-2" metricUnit="m" metricValue="9.14" pageId="13" pageNumber="598" unit="mm" value="91.4">91.4 mm</quantity>
SL, Nova Brasilândia, 
<location id="20476018055E1B12BCE06057FC9D0837" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BCE06057FC9D0837" box="[669,772,350,372]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Manso." pageId="13" pageNumber="598" stateProvince="Mato Grosso">rio Manso.</location>
<specimenCode id="753E9EB8055E1B12BD5461B8FC020982" box="[809,923,177,198]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 13399</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BDDE61B8FC280982" box="[931,945,177,198]" collectionCode="NUP" pageId="13" pageNumber="598">4</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BDC461B8FC690983" box="[953,1008,177,199]" collectionCode="NUP" pageId="13" pageNumber="598">136.2</specimenCode>
- 
<quantity id="E2609B26055E1B12BD8A61B8FBCF0982" box="[1015,1110,177,199]" metricMagnitude="-1" metricUnit="m" metricValue="1.638" pageId="13" pageNumber="598" unit="mm" value="163.8">163.8 mm</quantity>
SL, 
<location id="20476018055E1B12BAFC61B8FABA0982" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BAFC61B8FABA0982" box="[1153,1315,177,199]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="Tangara da Serra" pageId="13" pageNumber="598" stateProvince="Mato Grosso">
Tangará 
<collectorName id="886D5315055E1B12BAAF61B8FABA0982" box="[1234,1315,177,199]" pageId="13" pageNumber="598">da Serra</collectorName>
</location>
, córrego Rico. 
<specimenCode id="753E9EB8055E1B12BD5461C7FC0409A7" box="[809,925,206,227]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 13400</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BDDB61C7FC2B09A7" box="[934,946,206,227]" collectionCode="NUP" pageId="13" pageNumber="598">1</specimenCode>
, 
<quantity id="E2609B26055E1B12BDC361C4FB8B09A7" box="[958,1042,205,227]" metricMagnitude="-2" metricUnit="m" metricValue="5.91" pageId="13" pageNumber="598" unit="mm" value="59.1">59.1 mm</quantity>
SL, Rondonópolis, córrego do Esparramo. 
<specimenCode id="753E9EB8055E1B12BD5461E2FC040844" box="[809,925,235,256]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 13401</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BDDB61E2FC2B0844" box="[934,946,235,256]" collectionCode="NUP" pageId="13" pageNumber="598">1</specimenCode>
, 
<quantity id="E2609B26055E1B12BDC361E3FB8A0844" box="[958,1043,234,256]" metricMagnitude="-2" metricUnit="m" metricValue="6.18" pageId="13" pageNumber="598" unit="mm" value="61.8">61.8 mm</quantity>
SL, Rondonópolis, córrego Pitaluga. 
<specimenCode id="753E9EB8055E1B12BB0161E2FCFE0858" collectionCode="NUP" pageId="13" pageNumber="598">NUP 13423</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BD12600EFCE50858" box="[879,892,263,284]" collectionCode="NUP" pageId="13" pageNumber="598">3</specimenCode>
, 
<quantity id="E2609B26055E1B12BDFB600EFB920859" box="[902,1035,263,285]" metricMagnitude="-2" metricUnit="m" metricValue="9.115" metricValueMax="9.61" metricValueMin="8.62" pageId="13" pageNumber="598" unit="mm" value="91.15" valueMax="96.1" valueMin="86.2">86.2-96.1 mm</quantity>
SL, 
<collectorName id="886D5315055E1B12BA4B600EFA870859" box="[1078,1310,263,285]" pageId="13" pageNumber="598">Chapada dos Guimarães</collectorName>
, 
<location id="20476018055E1B12BB5B600EFA340859" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BB5B600EFA340859" box="[1318,1453,263,285]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Quilombo." pageId="13" pageNumber="598" stateProvince="Mato Grosso">rio Quilombo.</location>
<specimenCode id="753E9EB8055E1B12BD54602DFC39087D" box="[809,928,292,313]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 13480</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BDD1602DFC5C087D" box="[940,965,292,313]" collectionCode="NUP" pageId="13" pageNumber="598">11</specimenCode>
, 50.0- 
<quantity id="E2609B26055E1B12BA78602DFBF0087D" box="[1029,1129,292,314]" metricMagnitude="-1" metricUnit="m" metricValue="1.302" pageId="13" pageNumber="598" unit="mm" value="130.2">130.2 mm</quantity>
SL, Jangada, 
<location id="20476018055E1B12BA84602DFAE8087E" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055E1B12BA84602DFAE8087E" box="[1273,1393,292,314]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Jangada." pageId="13" pageNumber="598" stateProvince="Mato Grosso">rio Jangada.</location>
<collectingRegion id="E75CF821055E1B12BB04602DFC7D0812" country="Brazil" name="Mato Grosso do Sul" pageId="13" pageNumber="598">Mato Grosso do Sul State</collectingRegion>
. 
<specimenCode id="753E9EB8055E1B12BD966048FBC40812" box="[1003,1117,321,342]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 10982</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BA186048FBE80812" box="[1125,1137,321,342]" collectionCode="NUP" pageId="13" pageNumber="598">1</specimenCode>
, 
<quantity id="E2609B26055E1B12BA066048FB570812" box="[1147,1230,321,342]" metricMagnitude="-2" metricUnit="m" metricValue="9.91" pageId="13" pageNumber="598" unit="mm" value="99.1">99.1 mm</quantity>
SL; and 
<specimenCode id="753E9EB8055E1B12BB5D6048FA0B0812" box="[1312,1426,321,342]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 11819</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BBE76048FA3E0812" box="[1434,1447,321,342]" collectionCode="NUP" pageId="13" pageNumber="598">3</specimenCode>
, 
<quantity id="E2609B26055E1B12BD546057FC250837" box="[809,956,350,372]" metricMagnitude="-2" metricUnit="m" metricValue="9.005" metricValueMax="10.37" metricValueMin="7.64" pageId="13" pageNumber="598" unit="mm" value="90.05" valueMax="103.7" valueMin="76.4">76.4-103.7 mm</quantity>
SL, Coxim, córrego 
<collectorName id="886D5315055E1B12BAFA6057FB470837" box="[1159,1246,350,372]" pageId="13" pageNumber="598">da Onça.</collectorName>
<specimenCode id="753E9EB8055E1B12BA9E6057FACE0830" box="[1251,1367,350,372]" collectionCode="NUP" pageId="13" pageNumber="598">NUP 13269</specimenCode>
, 
<specimenCode id="753E9EB8055E1B12BB1C6057FAF70837" box="[1377,1390,350,371]" collectionCode="NUP" pageId="13" pageNumber="598">3</specimenCode>
, 
<quantity id="E2609B26055E1B11BB076057FF440988" lastPageId="14" lastPageNumber="599" metricMagnitude="-2" metricUnit="m" metricValue="7.279999999999999" metricValueMax="8.15" metricValueMin="6.409999999999999" pageId="13" pageNumber="598" unit="mm" value="72.8" valueMax="81.5" valueMin="64.1">64.1- 81.5 mm</quantity>
SL, Rochedo, córrego Formiga. 
<specimenCode id="753E9EB8055D1B11BC6461BEFD140988" box="[537,653,183,204]" collectionCode="NUP" pageId="14" pageNumber="599">NUP 13270</specimenCode>
, 
<specimenCode id="753E9EB8055D1B11BCEA61BEFD3C0988" box="[663,677,183,204]" collectionCode="NUP" pageId="14" pageNumber="599">9</specimenCode>
, 
<specimenCode id="753E9EB8055D1B11BCD261BEFD430988" box="[687,730,183,204]" collectionCode="NUP" pageId="14" pageNumber="599">43.4</specimenCode>
-93.0 mm SL, Rochedo, córrego Tamanduá. 
<specimenCode id="753E9EB8055D1B11BF8961DDFDFF09AD" box="[500,614,212,233]" collectionCode="NUP" pageId="14" pageNumber="599">NUP 13494</specimenCode>
, 
<specimenCode id="753E9EB8055D1B11BC1361DDFDE509AD" box="[622,636,212,233]" collectionCode="NUP" pageId="14" pageNumber="599">4</specimenCode>
, 
<quantity id="E2609B26055D1B11BCF861DAFC9309AD" box="[645,778,211,233]" metricMagnitude="-2" metricUnit="m" metricValue="6.715" metricValueMax="7.31" metricValueMin="6.12" pageId="14" pageNumber="599" unit="mm" value="67.15" valueMax="73.1" valueMin="61.2">61.2-73.1 mm</quantity>
SL, Terenos. 
<specimenCode id="753E9EB8055D1B11BF7861F9FE2F0842" box="[261,438,240,262]" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">ZUFMS-PIS 1316</specimenCode>
, 
<specimenCode id="753E9EB8055D1B11BFC261F9FE410841" box="[447,472,240,261]" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">11</specimenCode>
, 
<specimenCode id="753E9EB8055D1B11BF9C61F9FD920842" box="[481,523,240,262]" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">27.5</specimenCode>
-71.0 mm SL, Aquidauana, córrego 
<collectorName id="886D5315055D1B11BEAB6004FEA00866" box="[214,313,269,291]" pageId="14" pageNumber="599">das Antas.</collectorName>
<specimenCode id="753E9EB8055D1B11BF406004FE770866" box="[317,494,269,290]" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">ZUFMS-PIS 1978</specimenCode>
, 
<quantity id="E2609B26055D1B11BF8A6004FDF40867" box="[503,621,269,291]" metricMagnitude="0" metricUnit="m" metricValue="1.1157" pageId="14" pageNumber="599" unit="mm" value="1115.7">1, 115.7 mm</quantity>
SL, 
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, 
<location id="20476018055D1B11BE8D6023FEC40804" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055D1B11BE8D6023FEC40804" box="[240,349,298,320]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Coxim." pageId="14" pageNumber="599" stateProvince="Mato Grosso">rio Coxim.</location>
<specimenCode id="753E9EB8055D1B11BF1B6023FD870804" box="[358,542,298,320]" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">ZUFMS-PIS 3539</specimenCode>
, 
<specimenCode id="753E9EB8055D1B11BC566023FDDC087B" box="[555,581,298,319]" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">20</specimenCode>
, 39.0-73.0 mm SL, Jaraguari, córrego 
<location id="20476018055D1B11BF44604EFE6F0819" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055D1B11BF44604EFE6F0819" box="[313,502,327,349]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="Furnas do Dionisio." pageId="14" pageNumber="599" stateProvince="Mato Grosso">Furnas do Dionísio.</location>
<specimenCode id="753E9EB8055D1B11BF86604EFD350819" box="[507,684,327,349]" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">ZUFMS-PIS 3557</specimenCode>
, 
<specimenCode id="753E9EB8055D1B11BCC8604EFD560818" box="[693,719,327,348]" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">32</specimenCode>
, 26.0- 77.0 mm SL, Corguinho, tributary to 
<location id="20476018055D1B11BF8D606AFD12083D" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055D1B11BF8D606AFD12083D" box="[496,651,355,377]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Aquidauana." pageId="14" pageNumber="599" stateProvince="Mato Grosso">rio Aquidauana.</location>
<specimenCode id="753E9EB8055D1B11BCED606DFF2308D1" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">ZUFMS-PIS 3738</specimenCode>
, 
<specimenCode id="753E9EB8055D1B11BEBE6089FF4908D2" box="[195,208,384,406]" collectionCode="ZUFMS-PIS" pageId="14" pageNumber="599">5</specimenCode>
, 
<quantity id="E2609B26055D1B11BEA76089FECD08D2" box="[218,340,384,406]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" metricValueMax="10.6" metricValueMin="8.8" pageId="14" pageNumber="599" unit="mm" value="9.7" valueMax="10.6" valueMin="8.8">8.8-10.6 mm</quantity>
SL, 
<collectorName id="886D5315055D1B11BFFD6088FE5708D2" box="[384,462,385,406]" pageId="14" pageNumber="599">Terenos</collectorName>
, 
<location id="20476018055D1B11BFAB6089FDF508D2" LSID="urn:lsid:plazi:treatment:AD3187D5055F1B10BDD362E7FB690EEC:20476018055D1B11BFAB6089FDF508D2" box="[470,620,384,406]" country="Brazil" county="rio Paraguay" municipality="rio Coxipo." name="rio Aquidauana" pageId="14" pageNumber="599" stateProvince="Mato Grosso">rio Aquidauana</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="71E7664B055E1B12BEFC6679FC220EED" ID-DOI="http://doi.org/10.5281/zenodo.12718853" ID-Zenodo-Dep="12718853" httpUri="https://zenodo.org/record/12718853/files/figure.png" pageId="13" pageNumber="598" startId="13.[129,168,1904,1928]" targetBox="[129,1452,401,1873]" targetPageId="13" targetType="figure">
<paragraph id="252736C3055E1B12BEFC6679FC220EED" blockId="13.[129,1452,1904,1961]" pageId="13" pageNumber="598">
<emphasis id="17ECEAD1055E1B12BEFC6679FF4C0ECC" bold="true" box="[129,213,1904,1928]" pageId="13" pageNumber="598">Fig. 13.</emphasis>
<taxonomicName id="E2984D40055E1B12BEA06678FE490ECD" authorityName="Tencatt &amp; Zawadzki &amp; Froehlich" authorityYear="2014" box="[221,464,1905,1929]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="598" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055E1B12BEA06678FE490ECD" box="[221,464,1905,1929]" italics="true" pageId="13" pageNumber="598">Hypostomus khimaera</emphasis>
</taxonomicName>
, holotype, MZUSP 111129, 139.1 mm SL, from córrego Salobo, rio Paraguay basin, Mato Grosso State, Brazil; dorsal (top), lateral (middle), and ventral (bottom) views.
</paragraph>
</caption>
<subSubSection id="6D826548055D1B11BEF560B4FDBF0D71" pageId="14" pageNumber="599" type="diagnosis">
<paragraph id="252736C3055D1B11BEF560B4FDBF0D71" blockId="14.[136,779,445,1077]" pageId="14" pageNumber="599">
<emphasis id="17ECEAD1055D1B11BEF560B4FF650891" bold="true" box="[136,252,445,469]" pageId="14" pageNumber="599">Diagnosis.</emphasis>
<taxonomicName id="E2984D40055D1B11BF7860B7FE6F0891" authorityName="Tencatt &amp; Zawadzki &amp; Froehlich" authorityYear="2014" box="[261,502,445,469]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055D1B11BF7860B7FE6F0891" box="[261,502,445,469]" italics="true" pageId="14" pageNumber="599">Hypostomus khimaera</emphasis>
</taxonomicName>
is distinguished from all other 
<taxonomicName id="E2984D40055D1B11BEBA60D7FE3908B1" box="[199,416,477,501]" pageId="14" pageNumber="599">
<emphasis id="17ECEAD1055D1B11BEBA60D7FED308B1" box="[199,330,478,501]" italics="true" pageId="14" pageNumber="599">Hypostomus</emphasis>
species
</taxonomicName>
, except those belonging to the 
<taxonomicName id="E2984D40055D1B11BC8C60D7FF650B51" authorityName="Armbruster" authorityYear="2003" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="cochliodon">
<emphasis id="17ECEAD1055D1B11BC8C60D7FF650B51" italics="true" pageId="14" pageNumber="599">H. cochliodon</emphasis>
</taxonomicName>
group, by having the following unique combination of features: notch between metapterygoid and hyomandibula absent (
<emphasis id="17ECEAD1055D1B11BEA66337FF610B11" box="[219,248,574,597]" italics="true" pageId="14" pageNumber="599">vs.</emphasis>
notch present) and strongly angled dentaries, less than 80° (
<emphasis id="17ECEAD1055D1B11BE906357FE9A0B31" box="[237,259,606,629]" italics="true" pageId="14" pageNumber="599">vs</emphasis>
. shallow angle between dentaries, generally more than 80°). 
<taxonomicName id="E2984D40055D1B11BE8E6377FE450BD1" authorityName="Tencatt &amp; Zawadzki &amp; Froehlich" authorityYear="2014" box="[243,476,637,661]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055D1B11BE8E6377FE450BD1" box="[243,476,637,661]" italics="true" pageId="14" pageNumber="599">Hypostomus khimaera</emphasis>
</taxonomicName>
can be distinguished from all other species of the 
<taxonomicName id="E2984D40055D1B11BF106397FD910BF1" authorityName="Armbruster" authorityYear="2003" box="[365,520,669,693]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="cochliodon">
<emphasis id="17ECEAD1055D1B11BF106397FD910BF1" box="[365,520,669,693]" italics="true" pageId="14" pageNumber="599">H. cochliodon</emphasis>
</taxonomicName>
group, except from 
<taxonomicName id="E2984D40055D1B11BC8D6397FF7F0B91" authorityName="Tencatt &amp; Zawadzki &amp; Froehlich" authorityYear="2014" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="basilisko">
<emphasis id="17ECEAD1055D1B11BC8D6397FF7F0B91" italics="true" pageId="14" pageNumber="599">H. basilisko</emphasis>
</taxonomicName>
and 
<taxonomicName id="E2984D40055D1B11BF6163B7FE1B0B91" authorityName="Hollanda Carvalho &amp; Weber" authorityYear="2005" box="[284,386,702,725]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="soniae">
<emphasis id="17ECEAD1055D1B11BF6163B7FE1B0B91" box="[284,386,702,725]" italics="true" pageId="14" pageNumber="599">H. soniae</emphasis>
</taxonomicName>
, by the presence of a dark tan stripe along the flank (
<emphasis id="17ECEAD1055D1B11BF2D63D7FEF70BB1" box="[336,366,734,757]" italics="true" pageId="14" pageNumber="599">vs.</emphasis>
absence). The new species can be distinguished from 
<taxonomicName id="E2984D40055D1B11BF2F63F7FE560A51" authorityName="Tencatt &amp; Zawadzki &amp; Froehlich" authorityYear="2014" box="[338,463,765,789]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="basilisko">
<emphasis id="17ECEAD1055D1B11BF2F63F7FE560A51" box="[338,463,765,789]" italics="true" pageId="14" pageNumber="599">H. basilisko</emphasis>
</taxonomicName>
and 
<taxonomicName id="E2984D40055D1B11BC7D63F7FDFD0A51" authorityName="Hollanda Carvalho &amp; Weber" authorityYear="2005" box="[512,612,766,789]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="soniae">
<emphasis id="17ECEAD1055D1B11BC7D63F7FDFD0A51" box="[512,612,766,789]" italics="true" pageId="14" pageNumber="599">H. soniae</emphasis>
</taxonomicName>
by the presence of black spots on the body and fins or at least in one of these (
<emphasis id="17ECEAD1055D1B11BEEC6237FF370A11" box="[145,174,830,853]" italics="true" pageId="14" pageNumber="599">vs.</emphasis>
absence of spots). It can be additionally distinguished from 
<taxonomicName id="E2984D40055D1B11BEBE6257FEDB0A31" authorityName="Tencatt &amp; Zawadzki &amp; Froehlich" authorityYear="2014" box="[195,322,861,885]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="basilisko">
<emphasis id="17ECEAD1055D1B11BEBE6257FEDB0A31" box="[195,322,861,885]" italics="true" pageId="14" pageNumber="599">H. basilisko</emphasis>
</taxonomicName>
in having moderately developed keels (
<emphasis id="17ECEAD1055D1B11BC936257FC920A31" box="[750,779,862,885]" italics="true" pageId="14" pageNumber="599">vs.</emphasis>
highly developed keels) and more vertebrae (28 
<emphasis id="17ECEAD1055D1B11BCF16277FD300AD1" box="[652,681,894,917]" italics="true" pageId="14" pageNumber="599">vs.</emphasis>
27). The new species is additionally distinguished from 
<taxonomicName id="E2984D40055D1B11BC086297FC930AF1" authorityName="Armbruster" authorityYear="2003" box="[629,778,925,949]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="cochliodon">
<emphasis id="17ECEAD1055D1B11BC086297FC930AF1" box="[629,778,925,949]" italics="true" pageId="14" pageNumber="599">H. cochliodon</emphasis>
</taxonomicName>
by having comparatively more teeth (12-27 
<emphasis id="17ECEAD1055D1B11BC4662B7FDCE0A91" box="[571,599,958,981]" italics="true" pageId="14" pageNumber="599">vs.</emphasis>
7-9), externalized opercle, exposed region easily visible (
<figureCitation id="BDA32A46055D1B11BC5A62D4FDFC0AB1" box="[551,613,989,1013]" captionStart="Fig" captionStartId="4.[136,175,1321,1345]" captionTargetBox="[136,779,174,1299]" captionTargetId="figure-618@4.[136,779,174,1299]" captionTargetPageId="4" captionText="Fig. 3. Opercle morphology of (a) Hypostomus cochliodon, NUP 10809, 215.8 mm SL, showing indistinct opercle without odontodes; (b) Hypostomus basilisko, holotype, MZUSP 111110, 182.5 mm SL and (c) Hypostomus khimaera, holotype, MZUSP 111129, 139.1 mm SL, showing layer of skin surrounding opercle that supports odontodes." figureDoi="http://doi.org/10.5281/zenodo.12718833" httpUri="https://zenodo.org/record/12718833/files/figure.png" pageId="14" pageNumber="599">Fig. 3</figureCitation>
b-c) (
<emphasis id="17ECEAD1055D1B11BCDD62D7FD240AB1" box="[672,701,990,1013]" italics="true" pageId="14" pageNumber="599">vs.</emphasis>
almost entire internalized opercle, exposed region not easily visible (
<figureCitation id="BDA32A46055D1B11BEED6514FF410D71" box="[144,216,1053,1077]" captionStart="Fig" captionStartId="4.[136,175,1321,1345]" captionTargetBox="[136,779,174,1299]" captionTargetId="figure-618@4.[136,779,174,1299]" captionTargetPageId="4" captionText="Fig. 3. Opercle morphology of (a) Hypostomus cochliodon, NUP 10809, 215.8 mm SL, showing indistinct opercle without odontodes; (b) Hypostomus basilisko, holotype, MZUSP 111110, 182.5 mm SL and (c) Hypostomus khimaera, holotype, MZUSP 111129, 139.1 mm SL, showing layer of skin surrounding opercle that supports odontodes." figureDoi="http://doi.org/10.5281/zenodo.12718833" httpUri="https://zenodo.org/record/12718833/files/figure.png" pageId="14" pageNumber="599">Fig. 3a</figureCitation>
) and fewer vertebrae (28 
<emphasis id="17ECEAD1055D1B11BFA16517FE6B0D71" box="[476,498,1054,1077]" italics="true" pageId="14" pageNumber="599">vs</emphasis>
. 29).
</paragraph>
</subSubSection>
<subSubSection id="6D826548055D1B10BEF56559FD7A0C8D" lastPageId="15" lastPageNumber="600" pageId="14" pageNumber="599" type="description">
<paragraph id="252736C3055D1B11BEF56559FA3E0BCD" blockId="14.[136,780,1104,1961]" lastBlockId="14.[816,1460,177,1705]" pageId="14" pageNumber="599">
<emphasis id="17ECEAD1055D1B11BEF56559FE8B0D2C" bold="true" box="[136,274,1104,1128]" pageId="14" pageNumber="599">Description.</emphasis>
Morphometric data in Table 1. Overall view of body in 
<figureCitation id="BDA32A46055D1B11BE946578FEA50DCD" box="[233,316,1137,1161]" captionStart="Fig" captionStartId="13.[129,168,1904,1928]" captionTargetBox="[129,1452,401,1873]" captionTargetId="figure-170@13.[129,1452,401,1873]" captionTargetPageId="13" captionText="Fig. 13. Hypostomus khimaera, holotype, MZUSP 111129, 139.1 mm SL, from córrego Salobo, rio Paraguay basin, Mato Grosso State, Brazil; dorsal (top), lateral (middle), and ventral (bottom) views." figureDoi="http://doi.org/10.5281/zenodo.12718853" httpUri="https://zenodo.org/record/12718853/files/figure.png" pageId="14" pageNumber="599">Fig. 13</figureCitation>
, juvenile in 
<figureCitation id="BDA32A46055D1B11BFB26578FDB80DCD" box="[463,545,1137,1161]" captionStart="Fig" captionStartId="5.[129,168,1840,1864]" captionTargetBox="[129,772,1051,1805]" captionTargetId="figure-947@5.[129,772,1051,1805]" captionTargetPageId="5" captionText="Fig. 4. Lateral view of juvenile in (a) Hypostomus cochliodon, NUP 11956, 92.8 mm SL, (b) Hypostomus basilisko, paratype, ZUFMS-PIS 3083, 64.2 mm SL and (c) Hypostomus khimaera, NUP13494,68.8 mmSL." figureDoi="http://doi.org/10.5281/zenodo.12718835" httpUri="https://zenodo.org/record/12718835/files/figure.png" pageId="14" pageNumber="599">Fig. 4c</figureCitation>
. Head broad, rough, moderately deep and slightly compressed. Snout and anterior profile of head pointed, sometimes slightly round, in dorsal view. Eyes moderate in size, laterally positioned. Dorsal margin of orbit slightly elevated. Compound pterotic covered by well-developed odontodes in some specimens. Greatest body width at cleithrum, then decreasing to caudal peduncle. Dorsal profile of head straight from snout tip to supraocciptal region, and forming angle of 45° with ventral region of head; convex from that point to dorsal-fin origin; sloped downward to first dorsal caudal-fin procurrent rays, then elevating again to caudal-fin insertion. Ventral profile almost straight from snout tip to insertion of pelvic-fin unbranched ray; slightly straight from pelvic-fin insertion to first ventral caudal-fin procurrent rays, then descending to caudal-fin insertion. Caudal peduncle laterally compressed, broad on anterior region to very compressed on posterior region. Body dorsally covered with spinulose dermal plates, except of dorsal fin and small naked area on snout tip. Mesethmoid region covered by odontodes, more conspicuous anteriorly to nares. Moderate ridge originating laterally to nares, passing through supraorbital and extending through superior portion of compound pterotic. Exposed region of opercle ellipsoid and moderate in size; odontodes more developed distally. Opercle completely surrounded by thin layer of skin (
<figureCitation id="BDA32A46055D1B11BFA86678FD840ECD" box="[469,541,1905,1929]" captionStart="Fig" captionStartId="4.[136,175,1321,1345]" captionTargetBox="[136,779,174,1299]" captionTargetId="figure-618@4.[136,779,174,1299]" captionTargetPageId="4" captionText="Fig. 3. Opercle morphology of (a) Hypostomus cochliodon, NUP 10809, 215.8 mm SL, showing indistinct opercle without odontodes; (b) Hypostomus basilisko, holotype, MZUSP 111110, 182.5 mm SL and (c) Hypostomus khimaera, holotype, MZUSP 111129, 139.1 mm SL, showing layer of skin surrounding opercle that supports odontodes." figureDoi="http://doi.org/10.5281/zenodo.12718833" httpUri="https://zenodo.org/record/12718833/files/figure.png" pageId="14" pageNumber="599">Fig. 3c</figureCitation>
); covered by numerous odontodes. Oral disk round, relatively small in size, lower lip not reaching transversal through gill openings; ventral surface covered with numerous small papillae decreasing in size posteriorly (
<figureCitation id="BDA32A46055D1B11BDD161F8FC6C084D" box="[940,1013,241,265]" captionStart="Fig" captionStartId="7.[129,168,1567,1591]" captionTargetBox="[129,772,176,1543]" captionTargetId="figure-69@7.[129,772,176,1543]" captionTargetPageId="7" captionText="Fig. 5. Mouth and teeth of (a) Hypostomus cochliodon, NUP 10807, 208.0 mm SL, showing one series of papillae in inner face of upper and lower jaws; (b) Hypostomus basilisko, holotype, MZUSP 111110, 182.5 mm SL, showing the agglomerated papillae on inner face of upper and lower jaws; and (c) Hypostomus khimaera, MZUSP 111129, holotype, 139.1 mm SL, showing weakly developed papillae not forming series." figureDoi="http://doi.org/10.5281/zenodo.12718839" httpUri="https://zenodo.org/record/12718839/files/figure.png" pageId="14" pageNumber="599">Fig. 5c</figureCitation>
). Maxillary barbel equal to or slightly larger than eye pupil. Odontodes present on anterior surface of upper lip, just anterior snout. Buccal papilla (
<emphasis id="17ECEAD1055D1B11BAC5603BFB6B080D" box="[1208,1266,306,329]" italics="true" pageId="14" pageNumber="599">sensu</emphasis>
<bibRefCitation id="41094B32055D1B11BA886038FA35080D" author="Armbruster, J. W." box="[1269,1452,305,329]" pageId="14" pageNumber="599" pagination="1 - 60" refId="ref13139" refString="Armbruster, J. W. 2003. The species of the Hypostomus cochliodon group (Siluriformes: Loricariidae). Zootaxa, 249: 1 - 60." type="journal article" year="2003">Armbruster, 2003</bibRefCitation>
) absent. Dentaries acutely angled, averaging less than 80° between left and right dentary rami. Twelve to 27 teeth (mode 19, 
<typeStatus id="FA238861055D1B11BD256098FC2D08ED" box="[856,948,401,425]" pageId="14" pageNumber="599" type="holotype">holotype</typeStatus>
21) in maxilla, twelve to 27 teeth (mode 22*) in dentary. Teeth shovel-shaped, bicuspid; mesial cusp round to oblong in ventral view and conspicuously more developed than lateral cusp; lateral cusp distinct and generally well-developed, sometimes fused to mesial cusp (similar to condition found in 
<taxonomicName id="E2984D40055D1B11BDF0633BFB270B0D" authorityName="Armbruster" authorityYear="2003" box="[909,1214,561,585]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="hemicochliodon">
<emphasis id="17ECEAD1055D1B11BDF0633BFB270B0D" box="[909,1214,561,585]" italics="true" pageId="14" pageNumber="599">Hypostomus hemicochliodon</emphasis>
</taxonomicName>
, see 
<bibRefCitation id="41094B32055D1B11BA8D6338FA360B0D" author="Armbruster, J. W." box="[1264,1455,561,585]" pageId="14" pageNumber="599" pagination="1 - 60" refId="ref13139" refString="Armbruster, J. W. 2003. The species of the Hypostomus cochliodon group (Siluriformes: Loricariidae). Zootaxa, 249: 1 - 60." type="journal article" year="2003">Armbruster (2003</bibRefCitation>
, 
<bibRefCitation id="41094B32055D1B11BD4D6358FCFF0B2D" author="Armbruster, J. W." box="[816,870,593,617]" pageId="14" pageNumber="599" pagination="1 - 80" refId="ref13168" refString="Armbruster, J. W. 2004. Phylogenetic relationships of the suckermouth armored catfishes (Loricariidae) with emphasis on the Hypostominae and the Ancistrinae. Zoological Journal of the Linnean Society, 141: 1 - 80." type="journal article" year="2004">2004</bibRefCitation>
) and 
<taxonomicName id="E2984D40055D1B11BDDD635BFB9D0B2D" authorityName="Hollanda Carvalho &amp; Weber" authorityYear="2005" box="[928,1028,594,617]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="599" phylum="Chordata" rank="species" species="soniae">
<emphasis id="17ECEAD1055D1B11BDDD635BFB9D0B2D" box="[928,1028,594,617]" italics="true" pageId="14" pageNumber="599">H. soniae</emphasis>
</taxonomicName>
, see 
<bibRefCitation id="41094B32055D1B11BA456358FA310B2D" author="Hollanda Carvalho, P. &amp; C. Weber" box="[1080,1448,593,617]" pageId="14" pageNumber="599" pagination="953 - 978" refId="ref13313" refString="Hollanda Carvalho, P. &amp; C. Weber. 2004. Five new species of the Hypostomus cochliodon group (Siluriformes: Loricariidae) from middle and lower Amazon System. Revue suisse de Zoologie, 111: 953 - 978." type="journal article" year="2004">Hollanda Carvalho &amp; Weber, 2004</bibRefCitation>
). Juveniles with tooth morphology similar to adult specimens.
</paragraph>
<paragraph id="252736C3055D1B11BD2B6398FB850D8D" blockId="14.[816,1460,177,1705]" pageId="14" pageNumber="599">
Body covered by five rows of rough and spiny dermal plates. Odontodes more concentrated on dorsal surface of head and medially on dermal plates. Dorsal-fin base naked. Predorsal region medially keeled; these keels moderate developed, slightly diverging posteriorly (
<figureCitation id="BDA32A46055D1B11BA6A6218FBF80A6D" box="[1047,1121,785,809]" captionStart="Fig" captionStartId="7.[809,850,1441,1465]" captionTargetBox="[809,1452,174,1419]" captionTargetId="figure-141@7.[809,1452,174,1419]" captionTargetPageId="7" captionText="Fig. 6. Keels, ridges and odontodes of (a) Hypostomus cochliodon, NUP 10807, 208.0 mm SL, showing two weakly developed ridges in compound pterotic, weakly developed keels, few and weakly developed odontodes and predorsal keels weakly developed; (b) Hypostomus basilisko, holotype, MZUSP 111110, 182.5 mm SL, showing the three ridges on top of head, strongly developed keels, numerous welldeveloped odontodes on plates covering body and head and strong V-shaped predorsal keels; (c) Hypostomus khimaera, holotype, MZUSP 111129, 139.1 mm SL, showing moderately developed keels and dermal plates supporting moderately developed odontodes." figureDoi="http://doi.org/10.5281/zenodo.12718837" httpUri="https://zenodo.org/record/12718837/files/figure.png" pageId="14" pageNumber="599">Fig. 6c</figureCitation>
). Dorsal and mid-dorsal series of plates with moderate developed keels. Median series bearing lateral line and without keels. Mid-ventral series angled to fifth first plates, without keel posteriorly. Ventral series smooth angled ventrally towards posterior portion of caudal peduncle. Ventral surface of head covered with platelets, with exception to region beneath lower lip. Abdomen covered with minute platelets in specimens larger than 
<quantity id="E2609B26055D1B11BAF462F8FB480D4D" box="[1161,1233,1009,1033]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="599" unit="mm" value="80.0">80 mm</quantity>
SL, with exception of very small areas around pectoral- and pelvic-fin insertions and at urogenital opening. Preanal plate present. Median series of plates with 26-28 (mode 27*), three predorsal plates, plates between dorsal and adipose fins 7*-9 (mode 7), plates between adipose and caudal fins 7*-8 (mode 7) and plates below dorsal-fin base 7*-8 (mode 7).
</paragraph>
<paragraph id="252736C3055D1B11BD2B65D8FAE60FED" blockId="14.[816,1460,177,1705]" pageId="14" pageNumber="599">Dorsal fin II,7, its origin at vertical through midpoint between pectoral and pelvic fins or slightly posterior to that point. Dorsal-fin distal margin straight; dorsal-fin posterior rays not reaching adipose-fin spine. Adipose spine compressed and curved inward. Pectoral fin I,6, its distal border straight. Pectoral-fin spine slightly curved inward, covered with moderately developed odontodes, more developed on distal portion in larger specimens. Tip of adpressed pectoral fin reaching to one-third of adpressed pelvic-fin spine. Pelvic fin i,5, its distal border straight to slightly convex; its adpressed unbranched ray surpassing anal-fin origin. Anal fin i,4, its tip reaching sixth plate after its origin. Unbranched and fifth branched rays smaller than second, third, and fourth rays; third branched ray generally longest. Caudal fin i,14,i; distal margin falcate, with ventral lobe slightly longer than dorsal one.</paragraph>
<paragraph id="252736C3055D1B10BD4D67D8FDC60C4D" blockId="14.[816,1459,1745,1961]" lastBlockId="15.[129,772,1041,1289]" lastPageId="15" lastPageNumber="600" pageId="14" pageNumber="599">
<emphasis id="17ECEAD1055D1B11BD4D67D8FC440FAD" bold="true" box="[816,989,1745,1769]" pageId="14" pageNumber="599">Color in alcohol.</emphasis>
Ground color generally yellowish-brown, some specimens tan. One faint stripe between mid-dorsal and mid-ventral keels. Color of fins equal to ground color of trunk, sometimes lighter; some specimens with ventral lobe of caudal fin slightly darker than dorsal lobe. Region around dorsal-fin base darker, extending medially along dorsal portion of caudal peduncle, usually forming inconspicuous dark stripe on dorsal region; anal fin color pattern similar to that of dorsal fin. Spots at least in some part of body and fins in all specimens. Body generally with sparse and small spots, concentrated on anterior portion of body; some specimens have few or even lack spots altogether. All fins generally covered by small spots, closely spaced; spots diffuse or absent in some specimens. Color pattern of juvenile specimens similar to adults; diffuse spots on body and fins in some juvenile specimens.
</paragraph>
<caption id="71E7664B055C1B10BEFC6293FBE20A96" ID-DOI="http://doi.org/10.5281/zenodo.12718855" ID-Zenodo-Dep="12718855" httpUri="https://zenodo.org/record/12718855/files/figure.png" pageId="15" pageNumber="600" startId="15.[129,168,922,946]" targetBox="[129,1452,176,902]" targetPageId="15" targetType="figure">
<paragraph id="252736C3055C1B10BEFC6293FBE20A96" blockId="15.[129,1452,922,978]" pageId="15" pageNumber="600">
<emphasis id="17ECEAD1055C1B10BEFC6293FF480AF6" bold="true" box="[129,209,922,946]" pageId="15" pageNumber="600">Fig. 14.</emphasis>
Color pattern of live specimen of 
<taxonomicName id="E2984D40055C1B10BC536292FC320AF6" authority=", ZUFMS-PIS" authorityName="ZUFMS-PIS" box="[558,939,922,946]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055C1B10BC536292FC810AF6" box="[558,792,922,946]" italics="true" pageId="15" pageNumber="600">Hypostomus khimaera</emphasis>
, ZUFMS-PIS
</taxonomicName>
3738, 10.6 mm SL, rio Aquidauana, Mato Grosso do Sul, Brazil, showing the dark tan stripe along midline of flank and diffuse spots on the body.
</paragraph>
</caption>
<paragraph id="252736C3055C1B10BEFC6439FE2A0CCD" blockId="15.[129,771,1328,1417]" pageId="15" pageNumber="600">
<emphasis id="17ECEAD1055C1B10BEFC6439FE910C0C" bold="true" box="[129,264,1328,1352]" pageId="15" pageNumber="600">Color in life.</emphasis>
Color in life is similar to that in alcohol preserved specimens, except for more visible stripes and more yellowish-brown ground color (
<figureCitation id="BDA32A46055C1B10BF276478FE3F0CCD" box="[346,422,1393,1417]" captionStart="Fig" captionStartId="15.[129,168,922,946]" captionTargetBox="[129,1452,176,902]" captionTargetId="figure-474@15.[129,1452,176,902]" captionTargetPageId="15" captionText="Fig. 14. Color pattern of live specimen of Hypostomus khimaera, ZUFMS-PIS 3738, 10.6 mm SL, rioAquidauana, Mato Grosso do Sul, Brazil, showing the dark tan stripe along midline of flank and diffuse spots on the body." figureDoi="http://doi.org/10.5281/zenodo.12718855" httpUri="https://zenodo.org/record/12718855/files/figure.png" pageId="15" pageNumber="600">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="252736C3055C1B10BEFC64B9FD7A0C8D" blockId="15.[129,739,1456,1481]" box="[129,739,1456,1481]" pageId="15" pageNumber="600">
<emphasis id="17ECEAD1055C1B10BEFC64B9FEC70C8C" bold="true" box="[129,350,1456,1480]" pageId="15" pageNumber="600">Sexual dimorphism.</emphasis>
No sexual dimorphism was observed.
</paragraph>
</subSubSection>
<subSubSection id="6D826548055C1B10BEFC64F9FDC80E2D" pageId="15" pageNumber="600" type="distribution">
<paragraph id="252736C3055C1B10BEFC64F9FE4C0F2D" blockId="15.[129,771,1520,1641]" pageId="15" pageNumber="600">
<emphasis id="17ECEAD1055C1B10BEFC64F9FE880F4C" bold="true" box="[129,273,1520,1544]" pageId="15" pageNumber="600">Distribution.</emphasis>
<taxonomicName id="E2984D40055C1B10BF6764F8FD900F4D" box="[282,521,1521,1545]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055C1B10BF6764F8FD900F4D" box="[282,521,1521,1545]" italics="true" pageId="15" pageNumber="600">Hypostomus khimaera</emphasis>
</taxonomicName>
is known from several localities of rio 
<collectingCountry id="5D8F7653055C1B10BF546718FE120F6D" box="[297,395,1553,1577]" name="Paraguay" pageId="15" pageNumber="600">Paraguay</collectingCountry>
basin, such as rios Cuiabá, Manso, Quilombo, Coxipó, Coxim, baía de Chacororé, rios Vermelho and Aquidauana basins (
<figureCitation id="BDA32A46055C1B10BFFA6758FE5E0F2D" box="[391,455,1617,1641]" captionStart="Fig" captionStartId="9.[129,168,1872,1896]" captionTargetBox="[310,1270,1114,1837]" captionTargetId="figure-36@9.[309,1273,1110,1841]" captionTargetPageId="9" captionText="Fig. 9. Type locality of Hypostomus basilisko (yellow star), Hypostomus cochliodon (dark star) and Hypostomus khimaera (red star). The dark triangles represent areas of co-occurrence of Hypostomus cochliodon and Hypostomus khimaera; dark diamonds represent area of occurrence of Hypostomus cochliodon in the upper rio Paraná basin." figureDoi="http://doi.org/10.5281/zenodo.12718845" httpUri="https://zenodo.org/record/12718845/files/figure.png" pageId="15" pageNumber="600">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="252736C3055C1B10BEFC6799FDC80E2D" blockId="15.[129,772,1680,1897]" pageId="15" pageNumber="600">
<emphasis id="17ECEAD1055C1B10BEFC6799FEA40FED" bold="true" box="[129,317,1680,1705]" pageId="15" pageNumber="600">Ecological notes.</emphasis>
<taxonomicName id="E2984D40055C1B10BF3E6798FDA90FED" box="[323,560,1681,1705]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055C1B10BF3E6798FDA90FED" box="[323,560,1681,1705]" italics="true" pageId="15" pageNumber="600">Hypostomus khimaera</emphasis>
</taxonomicName>
occurs syntopically with 
<taxonomicName id="E2984D40055C1B10BEC867B8FED00F8D" authorityName="Armbruster" authorityYear="2003" box="[181,329,1713,1737]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="cochliodon">
<emphasis id="17ECEAD1055C1B10BEC867B8FED00F8D" box="[181,329,1713,1737]" italics="true" pageId="15" pageNumber="600">H. cochliodon</emphasis>
</taxonomicName>
in all aforementioned localities. However, in ichthyological surveys of headwater streams of rio Aquidauana basin, 
<taxonomicName id="E2984D40055C1B10BF3567F8FE530E4D" box="[328,458,1777,1801]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055C1B10BF3567F8FE530E4D" box="[328,458,1777,1801]" italics="true" pageId="15" pageNumber="600">H. khimaera</emphasis>
</taxonomicName>
was more commonly found in small streams than in larger streams. In the rio Aquidauana basin streams, 
<taxonomicName id="E2984D40055C1B10BF616638FE060E0D" box="[284,415,1841,1865]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055C1B10BF616638FE060E0D" box="[284,415,1841,1865]" italics="true" pageId="15" pageNumber="600">H. khimaera</emphasis>
</taxonomicName>
was collected generally along the margins of deeper sites with sandy bottoms.
</paragraph>
</subSubSection>
<subSubSection id="6D826548055C1B10BEFC6698FBF10C0C" pageId="15" pageNumber="600" type="etymology">
<paragraph id="252736C3055C1B10BEFC6698FBF10C0C" blockId="15.[129,771,1937,1961]" lastBlockId="15.[809,1452,1040,1352]" pageId="15" pageNumber="600">
<emphasis id="17ECEAD1055C1B10BEFC6698FF600EED" bold="true" box="[129,249,1937,1961]" pageId="15" pageNumber="600">Etymology.</emphasis>
The epithet 
<taxonomicName id="E2984D40055C1B10BF136698FE540EED" box="[366,461,1937,1961]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055C1B10BF136698FE540EED" box="[366,461,1937,1961]" italics="true" pageId="15" pageNumber="600">khimaera</emphasis>
</taxonomicName>
derives from the Greek, cimaira or 
<emphasis id="17ECEAD1055C1B10BD396519FC3A0D6C" box="[836,931,1040,1064]" italics="true" pageId="15" pageNumber="600">khímaira</emphasis>
, a mythological creature with hybrid body, formed essentially by three animals, a lion, a snake and a goat. The new species belongs to the 
<taxonomicName id="E2984D40055C1B10BA306558FAD50D2C" authorityName="Armbruster" authorityYear="2003" box="[1101,1356,1104,1128]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="cochliodon">
<emphasis id="17ECEAD1055C1B10BA306558FAD50D2C" box="[1101,1356,1104,1128]" italics="true" pageId="15" pageNumber="600">Hypostomus cochliodon</emphasis>
</taxonomicName>
group, a group commonly associated with relatively few, spoon-shape teeth, but has, instead, numerous teeth that are not spoon-shaped, very similar to that found in 
<taxonomicName id="E2984D40055C1B10BAD165B8FAF60D8C" baseAuthorityName="Eigenmann" baseAuthorityYear="1922" box="[1196,1391,1200,1224]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="plecostomoides">
<emphasis id="17ECEAD1055C1B10BAD165B8FAF60D8C" box="[1196,1391,1200,1224]" italics="true" pageId="15" pageNumber="600">H. plecostomoides</emphasis>
</taxonomicName>
or 
<taxonomicName id="E2984D40055C1B10BBEF65B8FCF70DAC" authorityName="Hollanda Carvalho &amp; Weber" authorityYear="2005" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="soniae">
<emphasis id="17ECEAD1055C1B10BBEF65B8FCF70DAC" italics="true" pageId="15" pageNumber="600">H. soniae</emphasis>
</taxonomicName>
. However, its general body morphology is similar to 
<taxonomicName id="E2984D40055C1B10BBEF65D8FC070C4C" authorityName="Armbruster" authorityYear="2003" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="cochliodon">
<emphasis id="17ECEAD1055C1B10BBEF65D8FC070C4C" italics="true" pageId="15" pageNumber="600">H. cochliodon</emphasis>
</taxonomicName>
. The epithet 
<taxonomicName id="E2984D40055C1B10BA4D65F9FB0D0C4C" box="[1072,1172,1264,1288]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="species" species="khimaera">
<emphasis id="17ECEAD1055C1B10BA4D65F9FB0D0C4C" box="[1072,1172,1264,1288]" italics="true" pageId="15" pageNumber="600">khimaera</emphasis>
</taxonomicName>
makes an allusion due to the new species possess features of conspicuously distinct species. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="6D826548055C1B10BD546479FB690EEC" pageId="15" pageNumber="600" type="discussion">
<paragraph id="252736C3055C1B10BD546479FB690EEC" blockId="15.[809,1452,1392,1960]" pageId="15" pageNumber="600">
<emphasis id="17ECEAD1055C1B10BD546479FC010CCC" bold="true" box="[809,920,1392,1416]" pageId="15" pageNumber="600">Remarks.</emphasis>
Some morphological differences were observed between the specimens from the rio Cuiabá and those from streams of rio 
<collectingCountry id="5D8F7653055C1B10BDC064B8FB860C8D" box="[957,1055,1457,1481]" name="Paraguay" pageId="15" pageNumber="600">Paraguay</collectingCountry>
basin. Characters such as mesial cusp morphology (round to oblong), development of lateral cusp (moderate to well developed), development of keels (moderate to well developed) and roughness of the body plates (moderate to well developed odontodes on lateral body plates) usually show some degree of variation even among specimens collected at the same site. Due to the presence of such variations, only the specimens from the rio Salobo were designated as 
<typeStatus id="FA238861055C1B10BD5467B8FC0B0F8D" box="[809,914,1713,1737]" pageId="15" pageNumber="600" type="paratype">paratypes</typeStatus>
. However, some degree of morphological variation is more common among loricariid populations inhabiting small to medium rivers than in populations of species inhabiting channels of larger rivers (C. H. Zawadzki, pers. obs.). Probably, because the sedentary characteristics of most of the species of 
<emphasis id="17ECEAD1055C1B10BD546658FC2B0E2C" box="[809,946,1873,1896]" italics="true" pageId="15" pageNumber="600">
<taxonomicName id="E2984D40055C1B10BD546658FC370E2C" box="[809,942,1873,1896]" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="600" phylum="Chordata" rank="genus">Hypostomus</taxonomicName>
,
</emphasis>
any degree of geographical isolation can result in gradual levels of morphological variation from each small stream population to the neighboring ones.
</paragraph>
</subSubSection>
</treatment>
</document>