1445 lines
158 KiB
XML
1445 lines
158 KiB
XML
<document id="E2405CDEA175AA870349EAAD83FD9D02" ID-CLB-Dataset="45372" ID-DOI="10.5281/zenodo.210618" ID-GBIF-Dataset="48843c3a-a5bc-4b60-9952-aa3802558f1e" ID-ISSN="1175-5326" ID-Zenodo-Dep="210618" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460444333248" checkinUser="plazi" docAuthor="David, Patrick & Vogel, Gernot" docDate="2012" docId="A123915FFF96F62EFEB5D25A2AB0F9A5" docLanguage="en" docName="zt03201p068.pdf" docOrigin="Zootaxa 3201" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Oligodon wagneri David & Vogel, 2012, sp. nov." docType="treatment" docVersion="10" lastPageNumber="65" masterDocId="5D1AE927FF97F629FE22D7602831FF8A" masterDocTitle="A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from Pulau Nias, Indonesia" masterLastPageNumber="68" masterPageNumber="58" pageNumber="59" updateTime="1698333216876" updateUser="ExternalLinkService">
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<mods:mods id="6375CE4BD8BFACAA3F3BE83C2406DF05" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0D98187EBDCE45AB43958DB6AC40BEB1">
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<mods:title id="5143AC8314A3803668565BC1C4B34727">A new species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae) from Pulau Nias, Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name id="CF48242FBA26E81CAFC0D4207D9A800F" type="personal">
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<mods:roleTerm id="6C5D7258B50C77F73238D8524A40A2A3">Author</mods:roleTerm>
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<mods:namePart id="E84A86847DF53AA18F49A9CDE25A623A">David, Patrick</mods:namePart>
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</mods:name>
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<mods:name id="BA020FDE86419EC8DF860FE19C2CA0C2" type="personal">
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<mods:role id="87A308ACD1A880048862D98B12E2A940">
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<mods:roleTerm id="3075B6132A15494C8FDE3C5CF8DC8E4E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="471C4009A55E6FA7630407E0A65A8A35">Vogel, Gernot</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E9540779FA35D994454B6F2355C9EE20">text</mods:typeOfResource>
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<mods:titleInfo id="063CB03CF8EE6C91AB67E93BD28CED0B">
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<mods:title id="D1F3454462FE0EA29D79522BB2DE4820">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3CB7E373B3E5D3FE3DACE02D8DBCB394">
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<mods:date id="A706A2311FEB4CC4F287FFC6E4E98FAB">2012</mods:date>
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<mods:detail id="8489EE2B834425B7E15B70E4A981BF3A" type="volume">
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<mods:number id="1B73F6BE4276399A0A9B57428A440534">3201</mods:number>
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<mods:start id="C30B7AB52049F79F6DB80F91DB402A40">58</mods:start>
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<mods:end id="A04A529A1E9FED9C6F26D392EE745A46">68</mods:end>
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<mods:classification id="71867E2CCC4CC35EC8D5E813C52B091E">journal article</mods:classification>
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<mods:identifier id="33EA5820FA5B6D5DB3B0E65E96A2B082" type="CLB-Dataset">45372</mods:identifier>
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<mods:identifier id="B553759D74C1B57ED6A9D76D47326ED5" type="DOI">10.5281/zenodo.210618</mods:identifier>
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<mods:identifier id="F51D5B30DF84AEED6B514C74C46E51A7" type="GBIF-Dataset">48843c3a-a5bc-4b60-9952-aa3802558f1e</mods:identifier>
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<mods:identifier id="DF34743ECC5C66B24E87C8D3157EC13D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BC5DD7C9D038F521EBB3A350B3D7116B" type="Zenodo-Dep">210618</mods:identifier>
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<treatment id="A123915FFF96F62EFEB5D25A2AB0F9A5" ID-DOI="http://doi.org/10.5281/zenodo.6178297" ID-GBIF-Taxon="119509046" ID-Zenodo-Dep="6178297" LSID="urn:lsid:plazi:treatment:A123915FFF96F62EFEB5D25A2AB0F9A5" httpUri="http://treatment.plazi.org/id/A123915FFF96F62EFEB5D25A2AB0F9A5" lastPageId="7" lastPageNumber="65" pageId="1" pageNumber="59">
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<subSubSection id="619073C2FF96F628FEB5D25A2923FAFF" pageId="1" pageNumber="59" type="nomenclature">
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<paragraph id="29352049FF96F628FEB5D25A29E4FADE" blockId="1.[151,469,1338,1397]" box="[151,469,1338,1364]" pageId="1" pageNumber="59">
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<heading id="727D9725FF96F628FEB5D25A29E4FADE" bold="true" box="[151,469,1338,1364]" fontSize="11" level="1" pageId="1" pageNumber="59" reason="1">
|
||
<emphasis id="1BFEFC5BFF96F628FEB5D25A29E4FADE" bold="true" box="[151,469,1338,1364]" pageId="1" pageNumber="59">
|
||
<taxonomicName id="EE8A5BCAFF96F628FEB5D25A2941FADE" box="[151,368,1338,1364]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="59" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF96F628FEB5D25A2941FADE" bold="true" box="[151,368,1338,1364]" italics="true" pageId="1" pageNumber="59">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="00CD4120FF96F628FF55D25A29E4FADE" box="[375,469,1338,1364]" pageId="1" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="29352049FF96F628FEB5D23D2923FAFF" blockId="1.[151,469,1338,1397]" box="[151,274,1373,1397]" pageId="1" pageNumber="59">
|
||
(
|
||
<figureCitation id="B1B13CCCFF96F628FEBDD23D2938FAFF" box="[159,265,1373,1397]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,249,1561,1584]" captionStartId-1="3.[151,249,1008,1031]" captionStartId-2="3.[151,249,1759,1782]" captionTargetBox-0="[171,1422,1255,1530]" captionTargetBox-1="[151,1434,447,986]" captionTargetBox-2="[151,1434,1110,1737]" captionTargetId-0="figure@2.[151,1436,1234,1541]" captionTargetId-1="figure@3.[151,1436,447,986]" captionTargetId-2="figure@3.[151,1436,1110,1737]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURE 1. Holotype of Oligodon wagneri sp. nov. (MNHN 1891.0233, juvenile male). Dorsal view and lateral view of head, left side. Drawing by Dick Visser." captionText-1="FIGURE 2. Holotype of Oligodon wagneri sp. nov. (MNHN 1891.0233, juvenile male). General view. Photograph by Patrick David." captionText-2="FIGURE 3. Holotype of Oligodon wagneri sp. nov. (MNHN 1891.0233, juvenile male). General ventral view. Photograph by Patrick David." httpUri-0="https://zenodo.org/record/210619/files/figure.png" httpUri-1="https://zenodo.org/record/210620/files/figure.png" httpUri-2="https://zenodo.org/record/210621/files/figure.png" pageId="1" pageNumber="59">Figs. 1–3</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="619073C2FF96F628FEB5D2C52A66F982" pageId="1" pageNumber="59" type="materials_examined">
|
||
<paragraph id="29352049FF96F628FEB5D2C52A66F982" blockId="1.[151,1437,1445,2021]" pageId="1" pageNumber="59">
|
||
<emphasis id="1BFEFC5BFF96F628FEB5D2C52935FA37" bold="true" box="[151,260,1445,1469]" pageId="1" pageNumber="59">
|
||
<typeStatus id="F6319EEBFF96F628FEB5D2C52935FA37" box="[151,260,1445,1469]" pageId="1" pageNumber="59" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectionCode id="4F9BB88CFF96F628FF30D2C62956FA37" LSID="urn:lsid:biocol.org:col:34988" box="[274,359,1446,1469]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="59">MNHN</collectionCode>
|
||
1891.0233, juvenile male, from “Poulo Nias”, i.e.
|
||
<collectingCountry id="519D60D9FF96F628FD88D2C52BDBFA37" box="[938,1002,1445,1469]" name="Iceland" pageId="1" pageNumber="59">Pulau</collectingCountry>
|
||
Nias (Nias Island), Nias Archipelago, Sumatera Utara Province, off the northern part of the west coast of Sumatra,
|
||
<collectingCountry id="519D60D9FF96F628FA3CD2A92CBFFA6B" box="[1054,1166,1481,1505]" name="Indonesia" pageId="1" pageNumber="59">Indonesia</collectingCountry>
|
||
, collected by Mr. Jules Claine and deposited on
|
||
<date id="5D340689FF96F628FF89D2902A62F982" box="[427,595,1517,1544]" pageId="1" pageNumber="59" value="1891-07-23">July 23rd, 1891</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="619073C2FF96F628FEEBD1732DB3F94B" pageId="1" pageNumber="59" type="diagnosis">
|
||
<paragraph id="29352049FF96F628FEEBD1732DB3F94B" blockId="1.[151,1437,1445,2021]" pageId="1" pageNumber="59">
|
||
<emphasis id="1BFEFC5BFF96F628FEEBD173290DF9A6" bold="true" box="[201,316,1555,1580]" pageId="1" pageNumber="59">Diagnosis</emphasis>
|
||
. A species of the genus
|
||
<taxonomicName id="EE8A5BCAFF96F628FC69D1742A84F9A6" box="[587,693,1556,1580]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="59" phylum="Chordata" rank="genus">
|
||
<emphasis id="1BFEFC5BFF96F628FC69D1742A84F9A6" box="[587,693,1556,1580]" italics="true" pageId="1" pageNumber="59">Oligodon</emphasis>
|
||
</taxonomicName>
|
||
characterized by the combination of: (1) 9 maxillary teeth, the last three strongly enlarged; (2) hemipenes not forked, thin and long, reaching 18th SC, covered with spines basally and flounces distally; (3) 15–15–15 DSR; (4) anal plate entire, (5) internasals and loreals present; (6) 7 supralabials, 3rd–4th entering orbit; (7) dorsal pattern made of 14 narrow, white crossbars on the body and 4 on the tail, alternating with three irregular white reticulations; and (7) venter heavily covered with subrectangular dark brown blotches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="619073C2FF96F62EFEEBD1AD2AF7FCBD" lastPageId="7" lastPageNumber="65" pageId="1" pageNumber="59" type="description">
|
||
<paragraph id="29352049FF96F628FEEBD1AD2C2CF8DB" blockId="1.[151,1437,1445,2021]" pageId="1" pageNumber="59">
|
||
<emphasis id="1BFEFC5BFF96F628FEEBD1AD29D2F96C" bold="true" box="[201,483,1741,1766]" pageId="1" pageNumber="59">
|
||
Description of
|
||
<typeStatus id="F6319EEBFF96F628FF5ED1AE29D2F96C" box="[380,483,1742,1766]" pageId="1" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Body cylindrical and robust but elongate; head short, ovoid, depressed, barely distinct from thick neck; snout long, slightly pointed, protracted beyond lower jaw, about 1.8 times as long as eye diameter; large, crescentic nostril piercing laterally on central part of nasal; eye average, its diameter about 1.5 times distance between eye-lip, round pupil; tail average, tapering progressively.
|
||
</paragraph>
|
||
<paragraph id="29352049FF96F62BFEEBD03F2924FC96" blockId="1.[151,1437,1445,2021]" lastBlockId="2.[151,1436,151,1159]" lastPageId="2" lastPageNumber="60" pageId="1" pageNumber="59">
|
||
<emphasis id="1BFEFC5BFF96F628FEEBD03F295AF8FC" box="[201,363,1887,1910]" italics="true" pageId="1" pageNumber="59">Measurements</emphasis>
|
||
. SVL:
|
||
<quantity id="EE728DACFF96F628FFE1D03D2A12F8FF" box="[451,547,1885,1909]" metricMagnitude="-1" metricUnit="m" metricValue="1.97" pageId="1" pageNumber="59" unit="mm" value="197.0">197 mm</quantity>
|
||
; TaL:
|
||
<quantity id="EE728DACFF96F628FC4BD03D2A8AF8FF" box="[617,699,1885,1909]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="59" unit="mm" value="38.0">38 mm</quantity>
|
||
; TL:
|
||
<quantity id="EE728DACFF96F628FCD5D03D2B66F8FF" box="[759,855,1885,1909]" metricMagnitude="-1" metricUnit="m" metricValue="2.35" pageId="1" pageNumber="59" unit="mm" value="235.0">235 mm</quantity>
|
||
; ratio TaL/TL: 0.162; HL:
|
||
<quantity id="EE728DACFF96F628FAA0D03D2CD8F8FF" box="[1154,1257,1885,1909]" metricMagnitude="-3" metricUnit="m" metricValue="9.05" pageId="1" pageNumber="59" unit="mm" value="9.05">9.05 mm</quantity>
|
||
; SnL:
|
||
<quantity id="EE728DACFF96F628FB10D03D2DA9F8FF" box="[1330,1432,1885,1909]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="1" pageNumber="59" unit="mm" value="2.85">2.85 mm</quantity>
|
||
.
|
||
<emphasis id="1BFEFC5BFF96F628FEB5D0E32930F810" box="[151,257,1923,1946]" italics="true" pageId="1" pageNumber="59">Dentition</emphasis>
|
||
. Maxillary teeth: left maxilla with 9 teeth under the formula: 6 subequal teeth + 3 strongly enlarged, blade-like teeth, without diastema.
|
||
<emphasis id="1BFEFC5BFF96F628FC05D0C92AAEF84A" box="[551,671,1961,1984]" italics="true" pageId="1" pageNumber="59">Hemipenis</emphasis>
|
||
. In situ, the hemipenis is thin, not forked and reaches the 18th SC; proximal half of the organ spinose, the distal spines slightly shorter; distal half of the organ densely covered with smooth flounces; sulcus spermaticus simple, extending up to the tip of the organ between papillae; two short papillae extend on a length of 2 or 3 SC from the end of the organ.
|
||
<emphasis id="1BFEFC5BFF95F62BFD76D7DC2BCAFF5E" box="[852,1019,188,212]" italics="true" pageId="2" pageNumber="60">Body scalation</emphasis>
|
||
. DSR: 15–15–15 scales, all smooth; scale of the outer dorsal scale row enlarged. 156 VEN (+ 1 preventral); 41 SC, all paired; anal entire; terminal caudal scale pointing.
|
||
<emphasis id="1BFEFC5BFF95F62BFF48D6642A24FE96" box="[362,533,260,284]" italics="true" pageId="2" pageNumber="60">Head scalation</emphasis>
|
||
. Head scalation complete, including 2 internasals, 2 prefrontals, 2 supraoculars, 1 frontal, and 2 parietals. Rostral large, wider than high, well visible from above, inserting deeply and broadly between internasals on about one half of their length; 1 / 1 relatively large, pentagonal nasals, divided in its lower half by a suture, most of its area being occupied by the nostril; internasals pentagonal, much wider than long, 0.8 times as long as prefrontals; the short suture between internasals is 0.7 times as long as the suture between prefrontals; prefrontals short, much wider than long, their suture only 0.2 times as long as frontal; supraoculars subrectangular, narrow, about 2.2 times as long as broad; frontal, large, hexagonal, long and wide, about 1.3 times longer than wide; parietals very large, extending on about 43% of HL, longer than wide, about 1.1 times longer than frontal, abruptly truncated posteriorly with a straight posterior margin; no nuchal scale behind parietal; 1 / 1 small loreal scale, subrectangular, 1.2 times longer than high; 7 / 7 SL, 1st–2nd in contact with nasal, 2nd–3rd in contact with loreal, 3rd–4th SL entering orbit, 5th–6th SL largest; 1 / 1 narrow preoculars, in contact with prefrontals but not reaching the frontal; no subocular; 2 / 2 small postoculars, of equal size; 1 + 2 temporals on each side, anterior one rather small and elongate, upper posterior one elongate and very large; 8 / 8
|
||
<collectingCountry id="519D60D9FF95F62BFDBDD5DD2B8DFD5E" box="[927,956,701,724]" name="Iceland" pageId="2" pageNumber="60">IL</collectingCountry>
|
||
, 1st
|
||
<collectingCountry id="519D60D9FF95F62BFDC4D5DD2C30FD5E" box="[998,1025,701,724]" name="Iceland" pageId="2" pageNumber="60">IL</collectingCountry>
|
||
in contact each with the other, 1st–4th
|
||
<collectingCountry id="519D60D9FF95F62BFEB5D5812883FD72" box="[151,178,737,760]" name="Iceland" pageId="2" pageNumber="60">IL</collectingCountry>
|
||
in contact with anterior chin shields; mental small; anterior chin shields 2.1 times as long as the very short posterior ones.
|
||
</paragraph>
|
||
<paragraph id="29352049FF95F62BFEEBD4482A66FB0D" blockId="2.[151,1436,151,1159]" pageId="2" pageNumber="60">
|
||
<emphasis id="1BFEFC5BFF95F62BFEEBD4482990FCCA" box="[201,417,808,832]" italics="true" pageId="2" pageNumber="60">Colour and pattern</emphasis>
|
||
. Body and tail chestnut-brown, with irregular darker areas on the upper part of the dorsum (the paler posterior part of the body is due to the discoloration in preservative); numerous irregular cream dots throughout the body, usually arranged along vertical or oblique discontinuous lines of dots every 2 or 3 DSR, forming discontinuous, wavy crossbars on the sides of the body from the 1st DSR up to the vertebral row where these dots are larger than on the sides; a series of 14 on the body + 4 on the tail white transverse crossbars, short but wide, no more than 1 or 1.5 DSR long but 3–4 DSR wide; these vertebral crossbars are more or less broadly edged with dark brown; the vertebral crossbars are separated from each other by two or three irregular cream reticulations formed by large cream dots issued from the lateral lines of dots. The tail is as the body but without reticulations between the narrow dorsal crossbars and with a series of minute white dots on every 2 or 3 scales of the first scale row above the subcaudals, on each side.
|
||
</paragraph>
|
||
<caption id="7DF570C1FF95F62BFEB5D17929DEF9C5" httpUri="https://zenodo.org/record/210619/files/figure.png" pageId="2" pageNumber="60" targetBox="[171,1422,1255,1530]" targetPageId="2">
|
||
<paragraph id="29352049FF95F62BFEB5D17929DEF9C5" blockId="2.[151,1436,1561,1615]" pageId="2" pageNumber="60">
|
||
<emphasis id="1BFEFC5BFF95F62BFEB5D1792923F9A5" bold="true" box="[151,274,1561,1584]" pageId="2" pageNumber="60">FIGURE 1.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="EE8A5BCAFF95F62BFFBAD17A2A60F9BA" box="[408,593,1562,1584]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="60" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF95F62BFFBAD17A2A60F9BA" box="[408,593,1562,1584]" italics="true" pageId="2" pageNumber="60">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF95F62BFC75D17A2A99F9A5" bold="true" box="[599,680,1562,1583]" pageId="2" pageNumber="60">
|
||
<taxonomicNameLabel id="00CD4120FF95F62BFC75D17A2A99F9A5" box="[599,680,1562,1583]" pageId="2" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MNHN 1891.0233, juvenile male). Dorsal view and lateral view of head, left side. Drawing by Dick Visser.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="29352049FF95F62AFEEBD1192AF8FE00" blockId="2.[151,1436,1654,2016]" lastBlockId="3.[151,1436,151,394]" lastPageId="3" lastPageNumber="61" pageId="2" pageNumber="60">Head very pale grey above and on the sides; rostral, areas around nasals, 1st SL and anterior half of 2nd SL dark greyish-brown, with a paler area on the tip of the rostral and two pale grey blotches at the edge between the 1st SL and rostral; an anterior dark greyish-brown crossband extending on the greatest part of prefrontals and anterior half of the frontal between each eye, downwards up to the posterior half of 3rd SL, 4th SL and anterior half of 5th SL; a short dark greyish-brown triangular blotch on the upper rear corner of each eye, not in contact with the anterior crossband; a large, vertical, irregular dark greyish-brown bar on the posterior half of 6th SL; a short, semicircular or peer-shaped brown blotch on the frontal just before the parietals; a broad, anteriorly truncated dark greyish-brown occipital stripe covers much of the parietals and extends obliquely posteriorly in widening downwards on each side across the temporals to the corner of the mouth then, downwards and backwards, connects with the chestnut-brown colour of the body near the level of 1st and 2nd ventrals on the neck; the branches of this stripe are separated from the anterior transversal band by a narrow, oblique area of pale grey background colour; another very pale grey streak, parallel to the previous one, separates on each side the posterior greyish-brown occipital stripe from the brown background colour of the body, each branch of the pale area looking like a nuchal streak extending obliquely downwards and backwards from the nuchal scale and tips of parietals and reaching the 3rd and 4th ventrals; the two lateral branches do not connect each other on the occiput and are separated by a greyish-brown area. The chin and throat are cream, heavily and irregularly covered with numerous, dark brown irregular blotches especially on chin shields, on the infralabials and on the middle of the throat.</paragraph>
|
||
<caption id="7DF570C1FF94F62AFEB5D49028EDFBAF" httpUri="https://zenodo.org/record/210620/files/figure.png" pageId="3" pageNumber="61" targetBox="[151,1434,447,986]" targetPageId="3">
|
||
<paragraph id="29352049FF94F62AFEB5D49028EDFBAF" blockId="3.[151,1436,1008,1061]" pageId="3" pageNumber="61">
|
||
<emphasis id="1BFEFC5BFF94F62AFEB5D4902923FB8F" bold="true" box="[151,274,1008,1031]" pageId="3" pageNumber="61">FIGURE 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="EE8A5BCAFF94F62AFFB8D4902A62FB8C" box="[410,595,1008,1030]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="61" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF94F62AFFB8D4902A62FB8C" box="[410,595,1008,1030]" italics="true" pageId="3" pageNumber="61">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF94F62AFC78D4912A9BFB8C" bold="true" box="[602,682,1009,1030]" pageId="3" pageNumber="61">
|
||
<taxonomicNameLabel id="00CD4120FF94F62AFC78D4912A9BFB8C" box="[602,682,1009,1030]" pageId="3" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MNHN 1891.0233, juvenile male). General view. Photograph by Patrick David.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7DF570C1FF94F62AFEB5D1BF291BF89E" httpUri="https://zenodo.org/record/210621/files/figure.png" pageId="3" pageNumber="61" targetBox="[151,1434,1110,1737]" targetPageId="3">
|
||
<paragraph id="29352049FF94F62AFEB5D1BF291BF89E" blockId="3.[151,1436,1759,1812]" pageId="3" pageNumber="61">
|
||
<emphasis id="1BFEFC5BFF94F62AFEB5D1BF2922F97E" bold="true" box="[151,275,1759,1782]" pageId="3" pageNumber="61">FIGURE 3.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="EE8A5BCAFF94F62AFFB8D1BF2A62F97F" box="[410,595,1759,1781]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="61" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF94F62AFFB8D1BF2A62F97F" box="[410,595,1759,1781]" italics="true" pageId="3" pageNumber="61">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="00CD4120FF94F62AFC78D1802A9AF97F" box="[602,683,1760,1781]" pageId="3" pageNumber="61" rank="species">
|
||
<emphasis id="1BFEFC5BFF94F62AFC78D1802A9AF97F" bold="true" box="[602,683,1760,1781]" pageId="3" pageNumber="61">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(MNHN 1891.0233, juvenile male). General ventral view. Photograph by Patrick David.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="29352049FF94F62DFEEBD05E2D6BFF7D" blockId="3.[151,1436,1851,2029]" lastBlockId="4.[151,1437,151,2023]" lastPageId="4" lastPageNumber="62" pageId="3" pageNumber="61">The venter is cream, with numerous rectangular dark brown blotches on each ventral; on 1st–4th VEN, the dark blotches are narrow and restricted to the outer parts of ventrals, producing a short, longitudinal cream streak, followed by an area up to 28th VEN where the brown colour covers much of each ventral and leaving only small, irregular cream blotches on ventrals, after VEN 28th, the venter is cream with rectangular blotches on every ventral, either covering one outer half the scale or both tips, the scheme irregularly alternating every two ventrals; sometimes two large blotches on the same ventral, making the scale nearly entirely dark brown. The ventral surface of the tail is as venter but dark brown rectangular blotches are limited to the outer tips of subcaudals, at the exception of a larger blotch on the outer part of every two or three subcaudals, no dot or streak in the middle of the tail.</paragraph>
|
||
<paragraph id="29352049FF93F62DFEEBD6632B77FE96" blockId="4.[151,1437,151,2023]" box="[201,838,259,284]" pageId="4" pageNumber="62">
|
||
<emphasis id="1BFEFC5BFF93F62DFEEBD6632906FE96" bold="true" box="[201,311,259,284]" pageId="4" pageNumber="62">Variation</emphasis>
|
||
. This species is known only from its
|
||
<typeStatus id="F6319EEBFF93F62DFCFFD6642B73FE96" box="[733,834,260,284]" pageId="4" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="29352049FF93F62DFEEBD64729DEFE26" blockId="4.[151,1437,151,2023]" pageId="4" pageNumber="62">
|
||
<emphasis id="1BFEFC5BFF93F62DFEEBD647296CFECA" bold="true" box="[201,349,295,320]" pageId="4" pageNumber="62">Comparison</emphasis>
|
||
. Our comparison of
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC6FD6482B2CFECA" box="[589,797,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFC6FD6482B2CFECA" box="[589,797,296,320]" italics="true" pageId="4" pageNumber="62">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFD04D6482BB3FECA" bold="true" box="[806,898,296,320]" pageId="4" pageNumber="62">
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFD04D6482BB3FECA" box="[806,898,296,320]" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
to other species of the genus is based on data available in
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFF3DD62D29E6FEEE" box="[287,471,332,356]" pageId="4" pageNumber="62" refString="De Rooij, N. (1917) The reptiles of the Indo-Australian Archipelago. II. Ophidia. E. J. Brill, Leyden, xvi + 334 pp." type="book">De Rooij (1917)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFFC1D62C2A4EFEEE" author="Taylor" box="[483,639,332,356]" pageId="4" pageNumber="62" refString="Taylor, E. H. (1922) The snakes of the Philippine Islands. Bureau of Printing, Manila, 312 pp., Pls. 1 - 37." type="book chapter" year="1922">Taylor (1922)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFCA8D62C2B11FEEE" author="Smith" box="[650,800,332,356]" pageId="4" pageNumber="62" refString="Smith, M. A. (1943) The fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Vol. III, Serpentes. Taylor & Francis, London, xii + 583 pp." type="book" year="1943">Smith (1943)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFD09D62C2BE4FEEE" author="Leviton" box="[811,981,332,356]" pageId="4" pageNumber="62" refString="Leviton, A. E. (1963) Contributions to a review of Philippine snakes, I. The snakes of the genus Oligodon. The Philippine Journal of Science, 91 [1962] (4), 459 - 484." type="journal article" year="1963">Leviton (1963)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFDC3D62D2CBAFEEE" author="Wagner" box="[993,1163,332,357]" pageId="4" pageNumber="62" refString="Wagner, F. W. (1976) A reassessment of infrageneric groupings and relationships of the Asian Colubrid snake genus Oligodon. Louisiana State University, Seminar 511, Baton Rouge, 37 unnumbered pp." type="journal article" year="1976">Wagner (1976)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFAB4D62C28D2FE0D" author="Manthey" pageId="4" pageNumber="62" refString="Manthey, U. & Grossmann, W. (1997) Amphibien & Reptilien Sudostasiens. Natur und Tier-Verlag, Munster, 512 pp." type="book" year="1997">Manthey & Grossmann (1997)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFECDD60F29EEFE0D" box="[239,479,367,392]" pageId="4" pageNumber="62" refString="Pauwels, O. S. G., Wallach, V., David, P. & Chanhome, L. (2002) A new species of Oligodon Fitzinger, 1826 (Serpentes, Colubridae) from southern Peninsular Thailand. Natural History Journal of Chulalongkorn University, Bangkok, 2 (2), 7 - 18." type="journal article">
|
||
Pauwels
|
||
<emphasis id="1BFEFC5BFF93F62DFF71D61129B5FE02" box="[339,388,368,392]" italics="true" pageId="4" pageNumber="62">et al</emphasis>
|
||
. (2002)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFFC8D6102A5BFE0D" author="Das" box="[490,618,367,391]" pageId="4" pageNumber="62" refString="Das, I. (2010) A field guide to the Reptiles of South-east Asia. New Holland Publishers (UK) Ltd, London-Cape Town-Sydney- Auckland, 376 pp." type="book" year="2010">Das (2010)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFC86D60F2BF4FE0D" author="Tillack" box="[676,965,367,391]" pageId="4" pageNumber="62" refString="Tillack, F. & Gunther, R. (2010) Revision of the species of Oligodon from Sumatra and adjacent islands, with comments on the taxonomic status of Oligodon subcarinatus (Gunther, 1872) and Oligodon annulifer (Boulenger, 1893) from Borneo (Reptilia, Squamata, Colubridae). Russian Journal of Herpetology, 16 [2009] (4), 265 - 294." type="journal article" year="2010">Tillack & Günther (2010)</bibRefCitation>
|
||
, as well as on specimens deposited in collections (see in the Appendix).
|
||
</paragraph>
|
||
<paragraph id="29352049FF93F62DFEEBD6D72BA5FDD5" blockId="4.[151,1437,151,2023]" pageId="4" pageNumber="62">
|
||
The number of DSR around the neck (one head length behind the head) and at midbody is a major diagnostic character in the genus
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFB8D6BC2A35FE7E" box="[410,516,476,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="genus">
|
||
<emphasis id="1BFEFC5BFF93F62DFFB8D6BC2A35FE7E" box="[410,516,476,500]" italics="true" pageId="4" pageNumber="62">Oligodon</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFC60D6BC2B3EFE7E" author="David" box="[578,783,476,500]" pageId="4" pageNumber="62" refString="David, P., Vogel, G. & Van Rooijen, J. (2008) A revision of the Oligodon taeniatus (Gunther, 1861) (Squamata: Colubridae) group, with the description of three new species from the Indochinese Region. Zootaxa, 1965, 1 - 45." type="journal article" year="2008">
|
||
David
|
||
<emphasis id="1BFEFC5BFF93F62DFCADD6BD2AF3FE7E" box="[655,706,476,500]" italics="true" pageId="4" pageNumber="62">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
). This number is usually constant within a given species. Previously, this character has been regarded as variable in some species but most of these are currently regarded as complexes, such as
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF5ED5442ACFFDB6" authority="Cantor, 1839" authorityName="Cantor" authorityYear="1839" box="[380,766,548,572]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="cyclurus">
|
||
<emphasis id="1BFEFC5BFF93F62DFF5ED5442A7EFDB6" box="[380,591,548,572]" italics="true" pageId="4" pageNumber="62">Oligodon cyclurus</emphasis>
|
||
(Cantor, 1839)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFD1DD5442C3DFDB6" author="David" box="[831,1036,548,572]" pageId="4" pageNumber="62" refString="David, P., Das, I. & Vogel, G. (2011) On some taxonomic and nomenclatural problems in Indian species of the genus Oligodon Fitzinger, 1826 (Squamata: Colubridae). Zootaxa, 2799, 1 - 14." type="journal article" year="2011">
|
||
David
|
||
<emphasis id="1BFEFC5BFF93F62DFDAED5452BF1FDB6" box="[908,960,548,572]" italics="true" pageId="4" pageNumber="62">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
). An exception might be
|
||
<taxonomicName id="EE8A5BCAFF93F62DFB10D54429DEFDD5" authority="Schlegel, 1837" authorityName="Schlegel" authorityYear="1837" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="purpurascens">
|
||
<emphasis id="1BFEFC5BFF93F62DFB10D544291EFDEA" italics="true" pageId="4" pageNumber="62">Oligodon purpurascens</emphasis>
|
||
(Schlegel, 1837)
|
||
</taxonomicName>
|
||
which has 19 or 21 DSR at midbody.
|
||
</paragraph>
|
||
<paragraph id="29352049FF93F62DFEEBD50C2CFBFBF6" blockId="4.[151,1437,151,2023]" pageId="4" pageNumber="62">
|
||
Among the about 70 species recognized in the genus
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD07D50C2BBEFD0E" box="[805,911,620,644]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="genus">
|
||
<emphasis id="1BFEFC5BFF93F62DFD07D50C2BBEFD0E" box="[805,911,620,644]" italics="true" pageId="4" pageNumber="62">Oligodon</emphasis>
|
||
</taxonomicName>
|
||
, only 26 have 15 DSR either constantly along the length of body or at least at midbody. Some morphological data for these 26 species are summarized in Table 1. All other species of the genus have 13, 17, 19, 21 or rarely 23 rows at midbody or throughout the body. Of the 26 species that have 15 DSR at midbody, 24 display the condition consistently:
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA3FD5B82D43FD65" authority="Gunther, 1862" authorityName="Gunther" authorityYear="1862" box="[1053,1394,727,752]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="brevicauda">
|
||
<emphasis id="1BFEFC5BFF93F62DFA3FD5B82CF1FD7A" box="[1053,1216,728,752]" italics="true" pageId="4" pageNumber="62">O. brevicauda</emphasis>
|
||
Günther, 1862
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFBA3D5B829EEFC9E" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="calamarius">
|
||
<emphasis id="1BFEFC5BFF93F62DFBA3D5B82924FC9E" italics="true" pageId="4" pageNumber="62">O. calamarius</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFCED59D2B32FC9E" authority="Gray, 1834" authorityName="Gray" authorityYear="1834" box="[492,771,764,788]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="1BFEFC5BFF93F62DFFCED59D2A58FC9E" box="[492,617,764,788]" italics="true" pageId="4" pageNumber="62">O. dorsalis</emphasis>
|
||
(Gray, 1834)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD32D59D2C61FC9E" authority="Wall, 1910" authorityName="Wall" authorityYear="1910" box="[784,1104,764,788]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="erythrorachis">
|
||
<emphasis id="1BFEFC5BFF93F62DFD32D59D2BFBFC9E" box="[784,970,764,788]" italics="true" pageId="4" pageNumber="62">O. erythrorachis</emphasis>
|
||
Wall, 1910
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA7ED59D2DA9FC9E" authority="Boulenger, 1893" authorityName="Boulenger" authorityYear="1893" box="[1116,1432,764,788]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="1BFEFC5BFF93F62DFA7ED59D2CE3FC9E" box="[1116,1234,765,788]" italics="true" pageId="4" pageNumber="62">O. everetti</emphasis>
|
||
Boulenger, 1893
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEB5D44029CEFCBD" authority="Boulenger, 1918" authorityName="Boulenger" authorityYear="1918" box="[151,511,799,824]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="hamptoni">
|
||
<emphasis id="1BFEFC5BFF93F62DFEB5D440291AFCB2" box="[151,299,800,824]" italics="true" pageId="4" pageNumber="62">O. hamptoni</emphasis>
|
||
Boulenger, 1918
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC31D4402BBEFCBD" authority="Boulenger, 1914" authorityName="Boulenger" authorityYear="1914" box="[531,911,799,824]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFC31D4402A98FCB2" box="[531,681,800,824]" italics="true" pageId="4" pageNumber="62">O. inornatus</emphasis>
|
||
(Boulenger, 1914)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD81D440296BFCD6" authority="Pauwels, Wallach, David & Chanhome, 2002" authorityName="Pauwels, Wallach, David & Chanhome" authorityYear="2002" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="jintakunei">
|
||
<emphasis id="1BFEFC5BFF93F62DFD81D4402C0FFCB2" box="[931,1086,800,824]" italics="true" pageId="4" pageNumber="62">O. jintakunei</emphasis>
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFA6FD47F296BFCD6" author="Pauwels" pageId="4" pageNumber="62" refString="Pauwels, O. S. G., Wallach, V., David, P. & Chanhome, L. (2002) A new species of Oligodon Fitzinger, 1826 (Serpentes, Colubridae) from southern Peninsular Thailand. Natural History Journal of Chulalongkorn University, Bangkok, 2 (2), 7 - 18." type="journal article" year="2002">Pauwels, Wallach, David & Chanhome, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF47D4252ADFFCD6" authority="Angel & Bourret, 1933" authorityName="Angel & Bourret" authorityYear="1933" box="[357,750,836,860]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="lacroixi">
|
||
<emphasis id="1BFEFC5BFF93F62DFF47D42529EFFCD6" box="[357,478,836,860]" italics="true" pageId="4" pageNumber="62">O. lacroixi</emphasis>
|
||
Angel & Bourret, 1933
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFCDBD4252CF0FCD6" authority="Zheng & Huang, 1978" authorityName="Zheng & Huang" authorityYear="1978" box="[761,1217,836,861]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="lungshenensis">
|
||
<emphasis id="1BFEFC5BFF93F62DFCDBD4252B88FCD6" box="[761,953,836,860]" italics="true" pageId="4" pageNumber="62">O. lungshenensis</emphasis>
|
||
Zheng & Huang, 1978
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFAEED42528E3FCF5" authority="Wall, 1909" authorityName="Wall" authorityYear="1909" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="melaneus">
|
||
<emphasis id="1BFEFC5BFF93F62DFAEED4252D68FCD6" box="[1228,1369,836,860]" italics="true" pageId="4" pageNumber="62">O. melaneus</emphasis>
|
||
Wall, 1909
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEFDD4082A29FCF5" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" box="[223,536,871,896]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="1BFEFC5BFF93F62DFEFDD408295AFC0A" box="[223,363,872,896]" italics="true" pageId="4" pageNumber="62">O. modestus</emphasis>
|
||
Günther, 1864
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC04D4082BF2FCF5" authority="Whitaker & Dattari, 1982" authorityName="Whitaker & Dattari" authorityYear="1982" box="[550,963,871,896]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="nikhili">
|
||
<emphasis id="1BFEFC5BFF93F62DFC04D4082AA3FC0A" box="[550,658,872,896]" italics="true" pageId="4" pageNumber="62">O. nikhili</emphasis>
|
||
Whitaker & Dattari, 1982
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFDF2D4082D24FCF5" authority="Gunther, 1873" authorityName="Gunther" authorityYear="1873" box="[976,1301,871,896]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="notospilus">
|
||
<emphasis id="1BFEFC5BFF93F62DFDF2D4082C56FC0A" box="[976,1127,872,896]" italics="true" pageId="4" pageNumber="62">O. notospilus</emphasis>
|
||
Günther, 1873
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFB00D40829B4FC2E" authority="Van Denburgh, 1909" authorityName="Van Denburgh" authorityYear="1909" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="ornatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFB00D4082DAAFC0A" box="[1314,1435,872,896]" italics="true" pageId="4" pageNumber="62">O. ornatus</emphasis>
|
||
Van Denburgh, 1909
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFB0D4ED2A48FC2E" authority="Roux" authorityName="Roux" box="[402,633,908,932]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="petronellae">
|
||
<emphasis id="1BFEFC5BFF93F62DFFB0D4ED2A04FC2E" box="[402,565,908,932]" italics="true" pageId="4" pageNumber="62">O. petronellae</emphasis>
|
||
Roux
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFCA0D4ED2AA9FC2E" box="[642,664,909,932]" italics="true" pageId="4" pageNumber="62">in</emphasis>
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFC82D4ED2B63FC2E" author="De" box="[672,850,908,932]" pageId="4" pageNumber="62" refString="De Rooij, N. (1917) The reptiles of the Indo-Australian Archipelago. II. Ophidia. E. J. Brill, Leyden, xvi + 334 pp." type="book" year="1917">De Rooij, 1917</bibRefCitation>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD7DD4ED2C89FC2E" authority="Werner, 1913" authorityName="Werner" authorityYear="1913" box="[863,1208,908,932]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="praefrontalis">
|
||
<emphasis id="1BFEFC5BFF93F62DFD7DD4ED2C25FC2E" box="[863,1044,908,932]" italics="true" pageId="4" pageNumber="62">O. praefrontalis</emphasis>
|
||
Werner, 1913
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFAE6D4ED28E3FC4D" authority="Jan, 1862" authorityName="Jan" authorityYear="1862" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="propinquus">
|
||
<emphasis id="1BFEFC5BFF93F62DFAE6D4ED2D56FC2E" box="[1220,1383,909,932]" italics="true" pageId="4" pageNumber="62">O. propinquus</emphasis>
|
||
Jan, 1862
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEFDD4D02B27FC4D" authority="Dumeril, Bibron & Dumeril, 1854" authorityName="Dumeril, Bibron & Dumeril" authorityYear="1854" box="[223,790,943,968]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="sublineatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFEFDD4D029B2FC42" box="[223,387,944,968]" italics="true" pageId="4" pageNumber="62">O. sublineatus</emphasis>
|
||
Duméril, Bibron & Duméril, 1854
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD01D4D02C42FC4D" authority="Jerdon, 1853" authorityName="Jerdon" authorityYear="1853" box="[803,1139,943,968]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="taeniolatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFD01D4D02BF2FC42" box="[803,963,944,968]" italics="true" pageId="4" pageNumber="62">O. taeniolatus</emphasis>
|
||
(Jerdon, 1853)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFAA3D4D028EBFC66" authority="Boulenger, 1888" authorityName="Boulenger" authorityYear="1888" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFAA3D4D02D3FFC42" box="[1153,1294,944,968]" italics="true" pageId="4" pageNumber="62">O. torquatus</emphasis>
|
||
(Boulenger, 1888)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFECBD4B52A79FC66" authority="Gunther, 1865" authorityName="Gunther" authorityYear="1865" box="[233,584,980,1004]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="1BFEFC5BFF93F62DFECBD4B529B7FC66" box="[233,390,980,1004]" italics="true" pageId="4" pageNumber="62">O. vertebralis</emphasis>
|
||
(Günther, 1865)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFCAAD4B52BEFFC66" authority="Bleeker, 1860" authorityName="Bleeker" authorityYear="1860" box="[648,990,980,1004]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="waandersi">
|
||
<emphasis id="1BFEFC5BFF93F62DFCAAD4B52B13FC66" box="[648,802,980,1004]" italics="true" pageId="4" pageNumber="62">O. waandersi</emphasis>
|
||
(Bleeker, 1860)
|
||
</taxonomicName>
|
||
, as well as
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA45D4B42D06FC66" box="[1127,1335,980,1004]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFA45D4B42D06FC66" box="[1127,1335,980,1004]" italics="true" pageId="4" pageNumber="62">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFB62D4B42DADFC66" bold="true" box="[1344,1436,980,1004]" pageId="4" pageNumber="62">
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFB62D4B42DADFC66" box="[1344,1436,980,1004]" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Exceptionally, two species, both occurring in Sumatra and its region, have usually 17 and unusually 15 DSR at midbody:
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF2BD37D2A1DFBBE" authority="Boie, 1827" authorityName="Boie" authorityYear="1827" box="[265,556,1052,1076]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFF2BD37D299AFBBE" box="[265,427,1052,1076]" italics="true" pageId="4" pageNumber="62">O. bitorquatus</emphasis>
|
||
Boie, 1827
|
||
</taxonomicName>
|
||
(rare value recorded only in one of the
|
||
<typeStatus id="F6319EEBFF93F62DFDC4D37D2C79FBBF" box="[998,1096,1053,1077]" pageId="4" pageNumber="62" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA4FD37D299AFBDD" authority="Dumeril, Bibron & Dumeril, 1854" authorityName="Dumeril, Bibron & Dumeril" authorityYear="1854" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="subquadratum">
|
||
<emphasis id="1BFEFC5BFF93F62DFA4FD37D2D00FBBE" box="[1133,1329,1052,1076]" italics="true" pageId="4" pageNumber="62">O. subquadratum</emphasis>
|
||
Duméril, Bibron & Duméril, 1854
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFCBD3202B10FBDD" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" box="[489,801,1087,1112]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFFCBD3202A59FBD2" box="[489,616,1088,1112]" italics="true" pageId="4" pageNumber="62">O. signatus</emphasis>
|
||
(Günther, 1864)
|
||
</taxonomicName>
|
||
(value recorded in the
|
||
<typeStatus id="F6319EEBFF93F62DFA00D35F2CB5FBDD" box="[1058,1156,1087,1111]" pageId="4" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA8AD3202D5DFBD2" box="[1192,1388,1088,1112]" class="Squamata" family="Colubridae" genus="Simotes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="annulifer">
|
||
<emphasis id="1BFEFC5BFF93F62DFA8AD3202D5DFBD2" box="[1192,1388,1088,1112]" italics="true" pageId="4" pageNumber="62">Simotes annulifer</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF93F62DFB51D3202940FBF6" authority="Van Lidth" authorityName="Van Lidth" class="Squamata" family="Colubridae" genus="Simotes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="62" phylum="Chordata" rank="variety" species="annulifer" variety="annulata">
|
||
var.
|
||
<emphasis id="1BFEFC5BFF93F62DFEB5D30428CAFBF6" box="[151,251,1124,1148]" italics="true" pageId="4" pageNumber="62">annulata</emphasis>
|
||
Van Lidth
|
||
</taxonomicName>
|
||
de Jeude, 1922; synonymized with
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD21D3052BB3FBF6" box="[771,898,1125,1148]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFD21D3052B26FBF6" box="[771,791,1125,1148]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFD07D3052BB3FBF6" box="[805,898,1125,1148]" italics="true" pageId="4" pageNumber="62">signatus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFD8ED3042C8FFBF6" author="Tillack" box="[940,1214,1124,1148]" pageId="4" pageNumber="62" refString="Tillack, F. & Gunther, R. (2010) Revision of the species of Oligodon from Sumatra and adjacent islands, with comments on the taxonomic status of Oligodon subcarinatus (Gunther, 1872) and Oligodon annulifer (Boulenger, 1893) from Borneo (Reptilia, Squamata, Colubridae). Russian Journal of Herpetology, 16 [2009] (4), 265 - 294." type="journal article" year="2010">Tillack & Günther 2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="29352049FF93F62DFEEBD3E72AD1FA6E" blockId="4.[151,1437,151,2023]" pageId="4" pageNumber="62">
|
||
The 26 species with 15 DSR at midbody can be divided into those with a maximum of 15 DSR around the neck or those with 17 DSR around the neck with a reduction to 15 DSR before midbody. Some species fall into both categories (
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEDAD3B0295CFB62" box="[248,365,1232,1256]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="1BFEFC5BFF93F62DFEDAD3B0295CFB62" box="[248,365,1232,1256]" italics="true" pageId="4" pageNumber="62">O. everetti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF59D3B02A39FB62" box="[379,520,1232,1256]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFF59D3B02A39FB62" box="[379,520,1232,1256]" italics="true" pageId="4" pageNumber="62">O. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC34D3B02A86FB62" box="[534,695,1232,1256]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="sublineatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFC34D3B02A1BFB6D" box="[534,554,1232,1255]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFC1AD3B02A86FB62" box="[568,695,1232,1256]" italics="true" pageId="4" pageNumber="62">sublineatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFCD6D3B02BA2FB62" box="[756,915,1232,1256]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="taeniolatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFCD6D3B02BA2FB62" box="[756,915,1232,1256]" italics="true" pageId="4" pageNumber="62">O. taeniolatus</emphasis>
|
||
</taxonomicName>
|
||
). The first category (15–15–15 DSR) includes
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEB5D3952924FA86" box="[151,277,1269,1292]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFEB5D3952924FA86" box="[151,277,1269,1292]" italics="true" pageId="4" pageNumber="62">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFF3ED3942944FA86" bold="true" box="[284,373,1268,1292]" pageId="4" pageNumber="62">
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFF3ED3942944FA86" box="[284,373,1268,1292]" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and the 18 following species:
|
||
<taxonomicName id="EE8A5BCAFF93F62DFCE8D3952B5BFA86" box="[714,874,1268,1292]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="brevicauda">
|
||
<emphasis id="1BFEFC5BFF93F62DFCE8D3952B5BFA86" box="[714,874,1268,1292]" italics="true" pageId="4" pageNumber="62">O. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD55D3952C26FA86" box="[887,1047,1268,1292]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="calamarius">
|
||
<emphasis id="1BFEFC5BFF93F62DFD55D3952C26FA86" box="[887,1047,1268,1292]" italics="true" pageId="4" pageNumber="62">O. calamarius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA07D3952C90FA86" box="[1061,1185,1268,1292]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="1BFEFC5BFF93F62DFA07D3952C90FA86" box="[1061,1185,1268,1292]" italics="true" pageId="4" pageNumber="62">O. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA8DD3952D14FA86" box="[1199,1317,1269,1292]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="1BFEFC5BFF93F62DFA8DD3952D14FA86" box="[1199,1317,1269,1292]" italics="true" pageId="4" pageNumber="62">O. everetti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFB10D39528F2FABA" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="hamptoni">
|
||
<emphasis id="1BFEFC5BFF93F62DFB10D39528F2FABA" italics="true" pageId="4" pageNumber="62">O. hamptoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEF0D2782951FABA" box="[210,352,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFEF0D2782951FABA" box="[210,352,1304,1328]" italics="true" pageId="4" pageNumber="62">O. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF52D2782A35FABA" box="[368,516,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="jintakunei">
|
||
<emphasis id="1BFEFC5BFF93F62DFF52D2782A35FABA" box="[368,516,1304,1328]" italics="true" pageId="4" pageNumber="62">O. jintakunei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC31D2782ABCFABA" box="[531,653,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="lacroixi">
|
||
<emphasis id="1BFEFC5BFF93F62DFC31D2782ABCFABA" box="[531,653,1304,1328]" italics="true" pageId="4" pageNumber="62">O. lacroixi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFCBED2782B16FABA" box="[668,807,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="1BFEFC5BFF93F62DFCBED2782B16FABA" box="[668,807,1304,1328]" italics="true" pageId="4" pageNumber="62">O. modestus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD15D2782BFFFABA" box="[823,974,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="notospilus">
|
||
<emphasis id="1BFEFC5BFF93F62DFD15D2782BFFFABA" box="[823,974,1304,1328]" italics="true" pageId="4" pageNumber="62">O. notospilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFDFFD2782C67FABA" box="[989,1110,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="ornatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFDFFD2782C67FABA" box="[989,1110,1304,1328]" italics="true" pageId="4" pageNumber="62">O. ornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA44D2782D36FABA" box="[1126,1287,1304,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="petronellae">
|
||
<emphasis id="1BFEFC5BFF93F62DFA44D2782D36FABA" box="[1126,1287,1304,1328]" italics="true" pageId="4" pageNumber="62">O. petronellae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFB35D278288FFADE" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="propinquus">
|
||
<emphasis id="1BFEFC5BFF93F62DFB35D278288FFADE" italics="true" pageId="4" pageNumber="62">O. propinquus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEEDD25D2963FADE" box="[207,338,1341,1364]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFEEDD25D2963FADE" box="[207,338,1341,1364]" italics="true" pageId="4" pageNumber="62">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
(value recorded in the
|
||
<typeStatus id="F6319EEBFF93F62DFC44D25C2AF9FADE" box="[614,712,1340,1364]" pageId="4" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="EE8A5BCAFF93F62DFCD6D25D2B8CFADE" box="[756,957,1340,1364]" class="Squamata" family="Colubridae" genus="Simotes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="annulifer">
|
||
<emphasis id="1BFEFC5BFF93F62DFCD6D25D2B8CFADE" box="[756,957,1340,1364]" italics="true" pageId="4" pageNumber="62">Simotes annulifer</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF93F62DFDE5D25D2CEAFADE" authority="Van Lidth" authorityName="Van Lidth" box="[967,1243,1340,1364]" class="Squamata" family="Colubridae" genus="Simotes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="4" pageNumber="62" phylum="Chordata" rank="variety" species="annulifer" variety="annulata">
|
||
var.
|
||
<emphasis id="1BFEFC5BFF93F62DFDD9D25C2C6EFADE" box="[1019,1119,1340,1364]" italics="true" pageId="4" pageNumber="62">annulata</emphasis>
|
||
Van Lidth
|
||
</taxonomicName>
|
||
de Jeude, 1922; synonymized with O.
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFB7D20129C3FAF2" box="[405,498,1377,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFFB7D20129C3FAF2" box="[405,498,1377,1400]" italics="true" pageId="4" pageNumber="62">signatus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFC03D2002B09FAF2" author="Tillack" box="[545,824,1376,1400]" pageId="4" pageNumber="62" refString="Tillack, F. & Gunther, R. (2010) Revision of the species of Oligodon from Sumatra and adjacent islands, with comments on the taxonomic status of Oligodon subcarinatus (Gunther, 1872) and Oligodon annulifer (Boulenger, 1893) from Borneo (Reptilia, Squamata, Colubridae). Russian Journal of Herpetology, 16 [2009] (4), 265 - 294." type="journal article" year="2010">Tillack & Günther 2010</bibRefCitation>
|
||
),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD6CD2002BC0FAF2" box="[846,1009,1376,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="sublineatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFD6CD2002BC0FAF2" box="[846,1009,1376,1400]" italics="true" pageId="4" pageNumber="62">O. sublineatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA20D2002CA0FAF2" box="[1026,1169,1376,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFA20D2002CA0FAF2" box="[1026,1169,1376,1400]" italics="true" pageId="4" pageNumber="62">O. torquatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA83D2002D0FFAF2" box="[1185,1342,1376,1400]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="1BFEFC5BFF93F62DFA83D2002D0FFAF2" box="[1185,1342,1376,1400]" italics="true" pageId="4" pageNumber="62">O. vertebralis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFBA3D200293DFA16" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="waandersi">
|
||
<emphasis id="1BFEFC5BFF93F62DFBA3D200293DFA16" italics="true" pageId="4" pageNumber="62">O. waandersi</emphasis>
|
||
</taxonomicName>
|
||
. By its formula of 15–15–15 DSR,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFCB8D2E52B29FA16" box="[666,792,1413,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFCB8D2E52B29FA16" box="[666,792,1413,1436]" italics="true" pageId="4" pageNumber="62">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFD3CD2E42B46FA16" bold="true" box="[798,887,1412,1436]" pageId="4" pageNumber="62">
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFD3CD2E42B46FA16" box="[798,887,1412,1436]" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA28D2E52C9DFA16" box="[1034,1196,1412,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFA28D2E52C9DFA16" box="[1034,1196,1412,1436]" italics="true" pageId="4" pageNumber="62">O. bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA98D2E52D42FA16" box="[1210,1395,1412,1436]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="erythrorachis">
|
||
<emphasis id="1BFEFC5BFF93F62DFA98D2E52D42FA16" box="[1210,1395,1412,1436]" italics="true" pageId="4" pageNumber="62">O. erythrorachis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFBA3D2E52930FA4A" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="melaneus">
|
||
<emphasis id="1BFEFC5BFF93F62DFBA3D2E52930FA4A" italics="true" pageId="4" pageNumber="62">O. melaneus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF62D2C8299AFA4A" box="[320,427,1448,1472]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="nikhili">
|
||
<emphasis id="1BFEFC5BFF93F62DFF62D2C8299AFA4A" box="[320,427,1448,1472]" italics="true" pageId="4" pageNumber="62">O. nikhili</emphasis>
|
||
</taxonomicName>
|
||
which always have 17 DSR around the neck. The complete dorsal scale row formulas of
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEB5D2AD2967FA6E" box="[151,342,1484,1508]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="lungshenensis">
|
||
<emphasis id="1BFEFC5BFF93F62DFEB5D2AD2967FA6E" box="[151,342,1484,1508]" italics="true" pageId="4" pageNumber="62">O. lungshenensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFAED2AD2A71FA6E" box="[396,576,1484,1508]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="praefrontalis">
|
||
<emphasis id="1BFEFC5BFF93F62DFFAED2AD2A71FA6E" box="[396,576,1484,1508]" italics="true" pageId="4" pageNumber="62">O. praefrontalis</emphasis>
|
||
</taxonomicName>
|
||
are unknown.
|
||
</paragraph>
|
||
<paragraph id="29352049FF93F62DFEEBD28F2AC0F885" blockId="4.[151,1437,151,2023]" pageId="4" pageNumber="62">
|
||
The structure and length of the hemipenes are major diagnostic characters in the genus
|
||
<taxonomicName id="EE8A5BCAFF93F62DFABAD2902D33F982" box="[1176,1282,1520,1544]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="genus">
|
||
<emphasis id="1BFEFC5BFF93F62DFABAD2902D33F982" box="[1176,1282,1520,1544]" italics="true" pageId="4" pageNumber="62">Oligodon</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFB33D28F2DA6F98D" author="Smith" box="[1297,1431,1519,1543]" pageId="4" pageNumber="62" refString="Smith, M. A. (1943) The fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Vol. III, Serpentes. Taylor & Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4D1B5DB8FF93F62DFEB5D1752901F9A6" author="Wagner" box="[151,304,1556,1581]" pageId="4" pageNumber="62" refString="Wagner, F. W. (1976) A reassessment of infrageneric groupings and relationships of the Asian Colubrid snake genus Oligodon. Louisiana State University, Seminar 511, Baton Rouge, 37 unnumbered pp." type="journal article" year="1976">Wagner 1976</bibRefCitation>
|
||
).
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF66D1742A20F9A6" box="[324,529,1556,1580]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFF66D1742A20F9A6" box="[324,529,1556,1580]" italics="true" pageId="4" pageNumber="62">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFC3AD1742A43F9A6" bold="true" box="[536,626,1556,1580]" pageId="4" pageNumber="62">
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFC3AD1742A43F9A6" box="[536,626,1556,1580]" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD25D1752BB5F9A6" box="[775,900,1556,1580]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="1BFEFC5BFF93F62DFD25D1752BB5F9A6" box="[775,900,1556,1580]" italics="true" pageId="4" pageNumber="62">O. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFDB0D1752C05F9A6" box="[914,1076,1556,1580]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="sublineatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFDB0D1752B97F9A6" box="[914,934,1557,1580]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFD96D1742C05F9A6" box="[948,1076,1556,1580]" italics="true" pageId="4" pageNumber="62">sublineatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA49D1752D3BF9A6" box="[1131,1290,1556,1580]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="taeniolatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFA49D1752C4EF9A6" box="[1131,1151,1557,1580]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFAAFD1742D3BF9A6" box="[1165,1290,1556,1580]" italics="true" pageId="4" pageNumber="62">taeniolatus</emphasis>
|
||
</taxonomicName>
|
||
by its unforked (vs. forked) hemipenes. Among the species with unforked hemipenes,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFDFDD1582C6FF9DA" box="[991,1118,1592,1616]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFDFDD1582C6FF9DA" box="[991,1118,1592,1616]" italics="true" pageId="4" pageNumber="62">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFA46D1582C8FF9DA" bold="true" box="[1124,1214,1592,1616]" pageId="4" pageNumber="62">
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFA46D1582C8FF9DA" box="[1124,1214,1592,1616]" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs by its hemipenes being longer (reaching the 18th SC) than in
|
||
<taxonomicName id="EE8A5BCAFF93F62DFCFFD1002BB1F9FD" box="[733,896,1631,1655]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="calamarius">
|
||
<emphasis id="1BFEFC5BFF93F62DFCFFD1002BB1F9FD" box="[733,896,1631,1655]" italics="true" pageId="4" pageNumber="62">O. calamarius</emphasis>
|
||
</taxonomicName>
|
||
(up to 9–10th),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA1AD1002C89F9FD" box="[1080,1208,1631,1655]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="1BFEFC5BFF93F62DFA1AD1002C89F9FD" box="[1080,1208,1631,1655]" italics="true" pageId="4" pageNumber="62">O. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
(11th),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFB2FD1002DADF9FD" box="[1293,1436,1631,1655]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="hamptoni">
|
||
<emphasis id="1BFEFC5BFF93F62DFB2FD1002DADF9FD" box="[1293,1436,1631,1655]" italics="true" pageId="4" pageNumber="62">O. hamptoni</emphasis>
|
||
</taxonomicName>
|
||
(11th),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEFDD1E6295DF914" box="[223,364,1670,1694]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFEFDD1E6295DF914" box="[223,364,1670,1694]" italics="true" pageId="4" pageNumber="62">O. inornatus</emphasis>
|
||
</taxonomicName>
|
||
(10–12th),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFC5D1E62A6FF914" box="[487,606,1670,1694]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="ornatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFFC5D1E62A6FF914" box="[487,606,1670,1694]" italics="true" pageId="4" pageNumber="62">O. ornatus</emphasis>
|
||
</taxonomicName>
|
||
(9th),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC83D1E62B72F914" box="[673,835,1670,1694]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="propinquus">
|
||
<emphasis id="1BFEFC5BFF93F62DFC83D1E62A84F917" box="[673,693,1670,1693]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFCE6D1E72B72F914" box="[708,835,1671,1694]" italics="true" pageId="4" pageNumber="62">propinquus</emphasis>
|
||
</taxonomicName>
|
||
(5th),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFDA6D1E62C16F917" box="[900,1063,1669,1693]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="sublineatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFDA6D1E62C16F917" box="[900,1063,1669,1693]" italics="true" pageId="4" pageNumber="62">O. sublineatus</emphasis>
|
||
</taxonomicName>
|
||
(13th),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA54D1E62D24F917" box="[1142,1301,1669,1693]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="taeniolatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFA54D1E62D24F917" box="[1142,1301,1669,1693]" italics="true" pageId="4" pageNumber="62">O. taeniolatus</emphasis>
|
||
</taxonomicName>
|
||
(9–11th),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFBA3D1E62930F94E" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFBA3D1E62930F94E" italics="true" pageId="4" pageNumber="62">O. torquatus</emphasis>
|
||
</taxonomicName>
|
||
(8th), and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF56D1CD2A3DF94E" box="[372,524,1708,1732]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="waandersi">
|
||
<emphasis id="1BFEFC5BFF93F62DFF56D1CD2A3DF94E" box="[372,524,1708,1732]" italics="true" pageId="4" pageNumber="62">O. waandersi</emphasis>
|
||
</taxonomicName>
|
||
(5–7th). In contrast, hemipenes are shorter in
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA2FD1CD2CBDF94E" box="[1037,1164,1709,1732]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFA2FD1CD2CBDF94E" box="[1037,1164,1709,1732]" italics="true" pageId="4" pageNumber="62">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFAB1D1CD2CDFF94F" box="[1171,1262,1709,1733]" pageId="4" pageNumber="62" rank="species">
|
||
<emphasis id="1BFEFC5BFF93F62DFAB1D1CD2CDFF94F" bold="true" box="[1171,1262,1709,1733]" pageId="4" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
than in
|
||
<taxonomicName id="EE8A5BCAFF93F62DFB68D1CD28F5F961" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="1BFEFC5BFF93F62DFB68D1CD28F5F961" italics="true" pageId="4" pageNumber="62">O. everetti</emphasis>
|
||
</taxonomicName>
|
||
(up to 35th SC) and in
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFE7D1B42A74F961" box="[453,581,1748,1771]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFFE7D1B42A74F961" box="[453,581,1748,1771]" italics="true" pageId="4" pageNumber="62">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
(25–26th). Hemipenes length in
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD92D1B42C1FF961" box="[944,1070,1748,1771]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFD92D1B42C1FF961" box="[944,1070,1748,1771]" italics="true" pageId="4" pageNumber="62">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFA17D1B32CBEF961" bold="true" box="[1077,1167,1747,1771]" pageId="4" pageNumber="62">
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFA17D1B32CBEF961" box="[1077,1167,1747,1771]" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is more or less equal to that of
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEC4D19829B9F885" box="[230,392,1783,1807]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFEC4D19828CBF885" box="[230,250,1784,1807]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFF2AD19729B9F885" box="[264,392,1783,1807]" italics="true" pageId="4" pageNumber="62">bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
, O.
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF9BD1972A12F885" box="[441,547,1783,1807]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="melaneus">
|
||
<emphasis id="1BFEFC5BFF93F62DFF9BD1972A12F885" box="[441,547,1783,1807]" italics="true" pageId="4" pageNumber="62">melaneus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC42D1982ADBF885" box="[608,746,1783,1807]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="1BFEFC5BFF93F62DFC42D1982A45F885" box="[608,628,1784,1807]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFCA0D1972ADBF885" box="[642,746,1783,1807]" italics="true" pageId="4" pageNumber="62">modestus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="29352049FF93F62DFEEBD07B2C0EF86D" blockId="4.[151,1437,151,2023]" pageId="4" pageNumber="62">
|
||
With 9 maxillary teeth,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFF4D07B2A95F8B9" box="[470,676,1819,1843]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFFF4D07B2A95F8B9" box="[470,676,1819,1843]" italics="true" pageId="4" pageNumber="62">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFC8ED07B2B37F8B9" bold="true" box="[684,774,1819,1843]" pageId="4" pageNumber="62">
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFC8ED07B2B37F8B9" box="[684,774,1819,1843]" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has more than
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD96D07C2C64F8B9" box="[948,1109,1819,1843]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="brevicauda">
|
||
<emphasis id="1BFEFC5BFF93F62DFD96D07C2C64F8B9" box="[948,1109,1819,1843]" italics="true" pageId="4" pageNumber="62">O. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
(7–8),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA8AD07C2D78F8B9" box="[1192,1353,1819,1843]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="calamarius">
|
||
<emphasis id="1BFEFC5BFF93F62DFA8AD07C2D78F8B9" box="[1192,1353,1819,1843]" italics="true" pageId="4" pageNumber="62">O. calamarius</emphasis>
|
||
</taxonomicName>
|
||
(7),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFBA3D07C28C0F8DD" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="1BFEFC5BFF93F62DFBA3D07C28C0F8DD" italics="true" pageId="4" pageNumber="62">O. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
(6–7),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF61D02029CCF8DD" box="[323,509,1855,1879]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="erythrorachis">
|
||
<emphasis id="1BFEFC5BFF93F62DFF61D02029CCF8DD" box="[323,509,1855,1879]" italics="true" pageId="4" pageNumber="62">O. erythrorachis</emphasis>
|
||
</taxonomicName>
|
||
(7–8),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC6DD0202AF4F8DD" box="[591,709,1856,1879]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="1BFEFC5BFF93F62DFC6DD0202A52F8DD" box="[591,611,1856,1879]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFC50D0202AF4F8DD" box="[626,709,1856,1879]" italics="true" pageId="4" pageNumber="62">everetti</emphasis>
|
||
</taxonomicName>
|
||
(7–8),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD35D0202B95F8DD" box="[791,932,1855,1879]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="hamptoni">
|
||
<emphasis id="1BFEFC5BFF93F62DFD35D0202B95F8DD" box="[791,932,1855,1879]" italics="true" pageId="4" pageNumber="62">O. hamptoni</emphasis>
|
||
</taxonomicName>
|
||
(7–8),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFDD7D0202CB9F8DD" box="[1013,1160,1855,1879]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="jintakunei">
|
||
<emphasis id="1BFEFC5BFF93F62DFDD7D0202CB9F8DD" box="[1013,1160,1855,1879]" italics="true" pageId="4" pageNumber="62">O. jintakunei</emphasis>
|
||
</taxonomicName>
|
||
(6),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFA9CD0202D7AF8DD" box="[1214,1355,1855,1879]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="melaneus">
|
||
<emphasis id="1BFEFC5BFF93F62DFA9CD0202D7AF8DD" box="[1214,1355,1855,1879]" italics="true" pageId="4" pageNumber="62">O. melaneus</emphasis>
|
||
</taxonomicName>
|
||
(7),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFBA3D02028CEF8F1" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="1BFEFC5BFF93F62DFBA3D02028CEF8F1" italics="true" pageId="4" pageNumber="62">O. modestus</emphasis>
|
||
</taxonomicName>
|
||
(7–8),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFF70D00429D9F8F1" box="[338,488,1891,1915]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="notospilus">
|
||
<emphasis id="1BFEFC5BFF93F62DFF70D00429D9F8F1" box="[338,488,1891,1915]" italics="true" pageId="4" pageNumber="62">O. notospilus</emphasis>
|
||
</taxonomicName>
|
||
(7–8),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC19D0042A82F8F1" box="[571,691,1892,1915]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="ornatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFC19D0042A82F8F1" box="[571,691,1892,1915]" italics="true" pageId="4" pageNumber="62">O. ornatus</emphasis>
|
||
</taxonomicName>
|
||
(6–8),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD24D0042B96F8F1" box="[774,935,1891,1915]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="petronellae">
|
||
<emphasis id="1BFEFC5BFF93F62DFD24D0042B96F8F1" box="[774,935,1891,1915]" italics="true" pageId="4" pageNumber="62">O. petronellae</emphasis>
|
||
</taxonomicName>
|
||
(7–8),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFDD8D0042C48F8F1" box="[1018,1145,1892,1915]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFDD8D0042C3FF8F1" box="[1018,1038,1892,1915]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFA3ED0042C48F8F1" box="[1052,1145,1892,1915]" italics="true" pageId="4" pageNumber="62">signatus</emphasis>
|
||
</taxonomicName>
|
||
(7–8),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFAEED0042D5EF8F1" box="[1228,1391,1891,1915]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="sublineatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFAEED0042D5EF8F1" box="[1228,1391,1891,1915]" italics="true" pageId="4" pageNumber="62">O. sublineatus</emphasis>
|
||
</taxonomicName>
|
||
(6– 8), and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFEC9D0E829B8F815" box="[235,393,1927,1951]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="taeniolatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFEC9D0E829B8F815" box="[235,393,1927,1951]" italics="true" pageId="4" pageNumber="62">O. taeniolatus</emphasis>
|
||
</taxonomicName>
|
||
(6–7). The numbers can be considered equal or nearly equal with those of
|
||
<taxonomicName id="EE8A5BCAFF93F62DFAECD0E82D5EF815" box="[1230,1391,1927,1951]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFAECD0E82D5EF815" box="[1230,1391,1927,1951]" italics="true" pageId="4" pageNumber="62">O. bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
(6– 9),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFE9ED0CC2905F849" box="[188,308,1963,1987]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="lacroixi">
|
||
<emphasis id="1BFEFC5BFF93F62DFE9ED0CC2905F849" box="[188,308,1963,1987]" italics="true" pageId="4" pageNumber="62">O. lacroixi</emphasis>
|
||
</taxonomicName>
|
||
(8–12),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFFB6D0CC2A07F849" box="[404,566,1964,1987]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="propinquus">
|
||
<emphasis id="1BFEFC5BFF93F62DFFB6D0CC2A07F849" box="[404,566,1964,1987]" italics="true" pageId="4" pageNumber="62">O. propinquus</emphasis>
|
||
</taxonomicName>
|
||
(9),
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC49D0CC2B37F849" box="[619,774,1963,1987]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="1BFEFC5BFF93F62DFC49D0CC2A4EF849" box="[619,639,1964,1987]" italics="true" pageId="4" pageNumber="62">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF93F62DFCAFD0CB2B37F849" box="[653,774,1963,1987]" italics="true" pageId="4" pageNumber="62">vertebralis</emphasis>
|
||
</taxonomicName>
|
||
(8–9), and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFDAAD0CC2C2EF849" box="[904,1055,1963,1987]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="waandersi">
|
||
<emphasis id="1BFEFC5BFF93F62DFDAAD0CC2C2EF849" box="[904,1055,1963,1987]" italics="true" pageId="4" pageNumber="62">O. waandersi</emphasis>
|
||
</taxonomicName>
|
||
(8–9). In contrast,
|
||
<taxonomicName id="EE8A5BCAFF93F62DFAD7D0CC2D42F849" box="[1269,1395,1964,1987]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF93F62DFAD7D0CC2D42F849" box="[1269,1395,1964,1987]" italics="true" pageId="4" pageNumber="62">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF93F62DFB58D0CB28F9F86D" bold="true" pageId="4" pageNumber="62">
|
||
<taxonomicNameLabel id="00CD4120FF93F62DFB58D0CB28F9F86D" pageId="4" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has fewer maxillary teeth than
|
||
<taxonomicName id="EE8A5BCAFF93F62DFC0BD0B02A86F86D" box="[553,695,2000,2023]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFC0BD0B02A86F86D" box="[553,695,2000,2023]" italics="true" pageId="4" pageNumber="62">O. inornatus</emphasis>
|
||
</taxonomicName>
|
||
(11–12) and
|
||
<taxonomicName id="EE8A5BCAFF93F62DFD69D0B02BE9F86D" box="[843,984,2000,2023]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1BFEFC5BFF93F62DFD69D0B02BE9F86D" box="[843,984,2000,2023]" italics="true" pageId="4" pageNumber="62">O. torquatus</emphasis>
|
||
</taxonomicName>
|
||
(15–16).
|
||
</paragraph>
|
||
<paragraph id="29352049FF91F62FFEEBD7F82DB2FF7D" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="64">
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEEBD7F829A4FF3A" box="[201,405,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF91F62FFEEBD7F829A4FF3A" box="[201,405,152,176]" italics="true" pageId="6" pageNumber="64">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="00CD4120FF91F62FFFBED7F829C4FF3A" box="[412,501,152,176]" pageId="6" pageNumber="64" rank="species">
|
||
<emphasis id="1BFEFC5BFF91F62FFFBED7F829C4FF3A" bold="true" box="[412,501,152,176]" pageId="6" pageNumber="64">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="EE8A5BCAFF91F62FFCAAD7F82B16FF3A" box="[648,807,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="brevicauda">
|
||
<emphasis id="1BFEFC5BFF91F62FFCAAD7F82B16FF3A" box="[648,807,152,176]" italics="true" pageId="6" pageNumber="64">O. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD16D7F82BE5FF3A" box="[820,980,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="calamarius">
|
||
<emphasis id="1BFEFC5BFF91F62FFD16D7F82BE5FF3A" box="[820,980,152,176]" italics="true" pageId="6" pageNumber="64">O. calamarius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFDC0D7F82C6CFF3A" box="[994,1117,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="1BFEFC5BFF91F62FFDC0D7F82C6CFF3A" box="[994,1117,152,176]" italics="true" pageId="6" pageNumber="64">O. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFA4ED7F82D14FF3A" box="[1132,1317,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="erythrorachis">
|
||
<emphasis id="1BFEFC5BFF91F62FFA4ED7F82D14FF3A" box="[1132,1317,152,176]" italics="true" pageId="6" pageNumber="64">O. erythrorachis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFB11D7F828F2FF5E" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="hamptoni">
|
||
<emphasis id="1BFEFC5BFF91F62FFB11D7F828F2FF5E" italics="true" pageId="6" pageNumber="64">O. hamptoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEF0D7DD2950FF5E" box="[210,353,189,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFEF0D7DD2950FF5E" box="[210,353,189,212]" italics="true" pageId="6" pageNumber="64">O. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFF53D7DD2A35FF5E" box="[369,516,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="jintakunei">
|
||
<emphasis id="1BFEFC5BFF91F62FFF53D7DD2A35FF5E" box="[369,516,188,212]" italics="true" pageId="6" pageNumber="64">O. jintakunei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFC36D7DD2ABFFF5E" box="[532,654,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="lacroixi">
|
||
<emphasis id="1BFEFC5BFF91F62FFC36D7DD2ABFFF5E" box="[532,654,188,212]" italics="true" pageId="6" pageNumber="64">O. lacroixi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFCBFD7DD2B1AFF5E" box="[669,811,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="melaneus">
|
||
<emphasis id="1BFEFC5BFF91F62FFCBFD7DD2B1AFF5E" box="[669,811,188,212]" italics="true" pageId="6" pageNumber="64">O. melaneus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD19D7DD2BF7FF5E" box="[827,966,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="1BFEFC5BFF91F62FFD19D7DD2BF7FF5E" box="[827,966,188,212]" italics="true" pageId="6" pageNumber="64">O. modestus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFDF4D7DD2C73FF5E" box="[982,1090,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="nikhili">
|
||
<emphasis id="1BFEFC5BFF91F62FFDF4D7DD2C73FF5E" box="[982,1090,188,212]" italics="true" pageId="6" pageNumber="64">O. nikhili</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFA73D7DD2CD9FF5E" box="[1105,1256,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="notospilus">
|
||
<emphasis id="1BFEFC5BFF91F62FFA73D7DD2CD9FF5E" box="[1105,1256,188,212]" italics="true" pageId="6" pageNumber="64">O. notospilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFADAD7DD2D40FF5E" box="[1272,1393,189,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="ornatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFADAD7DD2D40FF5E" box="[1272,1393,189,212]" italics="true" pageId="6" pageNumber="64">O. ornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFBA3D7DD2919FF72" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="praefrontalis">
|
||
<emphasis id="1BFEFC5BFF91F62FFBA3D7DD2919FF72" italics="true" pageId="6" pageNumber="64">O. praefrontalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFF14D78029E6FF72" box="[310,471,224,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="sublineatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFF14D78029E6FF72" box="[310,471,224,248]" italics="true" pageId="6" pageNumber="64">O. sublineatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFFC4D7802AB5FF72" box="[486,644,224,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="taeniolatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFFC4D78029CBFF7D" box="[486,506,224,247]" italics="true" pageId="6" pageNumber="64">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF91F62FFC2AD7802AB5FF72" box="[520,644,224,248]" italics="true" pageId="6" pageNumber="64">taeniolatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFCB0D7802B2FFF72" box="[658,798,224,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFCB0D7802B2FFF72" box="[658,798,224,248]" italics="true" pageId="6" pageNumber="64">O. torquatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD7ED7802BC2FF72" box="[860,1011,224,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="waandersi">
|
||
<emphasis id="1BFEFC5BFF91F62FFD7ED7802BC2FF72" box="[860,1011,224,248]" italics="true" pageId="6" pageNumber="64">O. waandersi</emphasis>
|
||
</taxonomicName>
|
||
by its entire (undivided) anal plate.
|
||
</paragraph>
|
||
<paragraph id="29352049FF91F62FFEEBD6642A1EFE26" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="64">
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEEBD66429A4FE96" box="[201,405,260,284]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF91F62FFEEBD66429A4FE96" box="[201,405,260,284]" italics="true" pageId="6" pageNumber="64">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF91F62FFFB9D66429C5FE96" bold="true" box="[411,500,260,284]" pageId="6" pageNumber="64">
|
||
<taxonomicNameLabel id="00CD4120FF91F62FFFB9D66429C5FE96" box="[411,500,260,284]" pageId="6" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a complete head scalation complement, and thus differs from
|
||
<taxonomicName id="EE8A5BCAFF91F62FFAF6D6652D42FE96" box="[1236,1395,260,284]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="brevicauda">
|
||
<emphasis id="1BFEFC5BFF91F62FFAF6D6652D42FE96" box="[1236,1395,260,284]" italics="true" pageId="6" pageNumber="64">O. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFBA3D6652930FECA" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="hamptoni">
|
||
<emphasis id="1BFEFC5BFF91F62FFBA3D6652930FECA" italics="true" pageId="6" pageNumber="64">O. hamptoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFF32D6482995FECA" box="[272,420,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="jintakunei">
|
||
<emphasis id="1BFEFC5BFF91F62FFF32D6482995FECA" box="[272,420,296,320]" italics="true" pageId="6" pageNumber="64">O. jintakunei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFF96D6482A5BFECA" box="[436,618,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="praefrontalis">
|
||
<emphasis id="1BFEFC5BFF91F62FFF96D6482A5BFECA" box="[436,618,296,320]" italics="true" pageId="6" pageNumber="64">O. praefrontalis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EE8A5BCAFF91F62FFC89D6482B08FECA" box="[683,825,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="hamptoni">
|
||
<emphasis id="1BFEFC5BFF91F62FFC89D6482B08FECA" box="[683,825,296,320]" italics="true" pageId="6" pageNumber="64">O. hamptoni</emphasis>
|
||
</taxonomicName>
|
||
which lack internasals. Furthermore, the loreal scale, present in
|
||
<taxonomicName id="EE8A5BCAFF91F62FFF2FD62D29BDFEEE" box="[269,396,333,356]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF91F62FFF2FD62D29BDFEEE" box="[269,396,333,356]" italics="true" pageId="6" pageNumber="64">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF91F62FFFB1D62C29DFFEEE" bold="true" box="[403,494,332,356]" pageId="6" pageNumber="64">
|
||
<taxonomicNameLabel id="00CD4120FF91F62FFFB1D62C29DFFEEE" box="[403,494,332,356]" pageId="6" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, is absent in
|
||
<taxonomicName id="EE8A5BCAFF91F62FFCA1D62D2B12FEEE" box="[643,803,332,356]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="brevicauda">
|
||
<emphasis id="1BFEFC5BFF91F62FFCA1D62D2B12FEEE" box="[643,803,332,356]" italics="true" pageId="6" pageNumber="64">O. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD10D62D2BDDFEEE" box="[818,1004,332,356]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="erythrorachis">
|
||
<emphasis id="1BFEFC5BFF91F62FFD10D62D2BDDFEEE" box="[818,1004,332,356]" italics="true" pageId="6" pageNumber="64">O. erythrorachis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFDD9D62D2C45FEEE" box="[1019,1140,332,356]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="lacroixi">
|
||
<emphasis id="1BFEFC5BFF91F62FFDD9D62D2C45FEEE" box="[1019,1140,332,356]" italics="true" pageId="6" pageNumber="64">O. lacroixi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFAA1D62D2D3FFEEE" box="[1155,1294,332,356]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="1BFEFC5BFF91F62FFAA1D62D2D3FFEEE" box="[1155,1294,332,356]" italics="true" pageId="6" pageNumber="64">O. modestus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFB3ED62D2DA5FEEE" box="[1308,1428,333,356]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="ornatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFB3ED62D2DA5FEEE" box="[1308,1428,333,356]" italics="true" pageId="6" pageNumber="64">O. ornatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEE5D610294DFE02" box="[199,380,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="praefrontalis">
|
||
<emphasis id="1BFEFC5BFF91F62FFEE5D610294DFE02" box="[199,380,368,392]" italics="true" pageId="6" pageNumber="64">O. praefrontalis</emphasis>
|
||
</taxonomicName>
|
||
. The loreal is either present or absent in
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD72D6102BEFFE02" box="[848,990,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="hamptoni">
|
||
<emphasis id="1BFEFC5BFF91F62FFD72D6102BEFFE02" box="[848,990,368,392]" italics="true" pageId="6" pageNumber="64">O. hamptoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EE8A5BCAFF91F62FFA34D6102C9FFE02" box="[1046,1198,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="waandersi">
|
||
<emphasis id="1BFEFC5BFF91F62FFA34D6102C9FFE02" box="[1046,1198,368,392]" italics="true" pageId="6" pageNumber="64">O. waandersi</emphasis>
|
||
</taxonomicName>
|
||
. It is usually present but may be absent in
|
||
<taxonomicName id="EE8A5BCAFF91F62FFFA7D6F52A19FE26" box="[389,552,404,428]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFFA7D6F52A19FE26" box="[389,552,404,428]" italics="true" pageId="6" pageNumber="64">O. bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="29352049FF91F62FFEE5D6D82D18FD9D" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="64">
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEE5D6D829A2FE5A" box="[199,403,440,464]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF91F62FFEE5D6D829A2FE5A" box="[199,403,440,464]" italics="true" pageId="6" pageNumber="64">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF91F62FFFBBD6D829C3FE5A" bold="true" box="[409,498,440,464]" pageId="6" pageNumber="64">
|
||
<taxonomicNameLabel id="00CD4120FF91F62FFFBBD6D829C3FE5A" box="[409,498,440,464]" pageId="6" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has more ventral scales than
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD1BD6D82BE9FE5A" box="[825,984,440,464]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="calamarius">
|
||
<emphasis id="1BFEFC5BFF91F62FFD1BD6D82BE9FE5A" box="[825,984,440,464]" italics="true" pageId="6" pageNumber="64">O. calamarius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFDC4D6D82C6BFE5A" box="[998,1114,440,464]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="1BFEFC5BFF91F62FFDC4D6D82C6BFE5A" box="[998,1114,440,464]" italics="true" pageId="6" pageNumber="64">O. everetti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFA4AD6D82CE0FE5A" box="[1128,1233,440,464]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="nikhili">
|
||
<emphasis id="1BFEFC5BFF91F62FFA4AD6D82CE0FE5A" box="[1128,1233,440,464]" italics="true" pageId="6" pageNumber="64">O. nikhili</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFAFCD6D82D42FE5A" box="[1246,1395,440,464]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="notospilus">
|
||
<emphasis id="1BFEFC5BFF91F62FFAFCD6D82D42FE5A" box="[1246,1395,440,464]" italics="true" pageId="6" pageNumber="64">O. notospilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFBA3D6D82924FE7E" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="propinquus">
|
||
<emphasis id="1BFEFC5BFF91F62FFBA3D6D82924FE7E" italics="true" pageId="6" pageNumber="64">O. propinquus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFF06D6BD2992FE7E" box="[292,419,477,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFF06D6BD2909FE7E" box="[292,312,477,500]" italics="true" pageId="6" pageNumber="64">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF91F62FFF64D6BD2992FE7E" box="[326,419,477,500]" italics="true" pageId="6" pageNumber="64">signatus</emphasis>
|
||
</taxonomicName>
|
||
(males only), and
|
||
<taxonomicName id="EE8A5BCAFF91F62FFC54D6BD2B23FE7E" box="[630,786,476,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="1BFEFC5BFF91F62FFC54D6BD2B23FE7E" box="[630,786,476,500]" italics="true" pageId="6" pageNumber="64">O. vertebralis</emphasis>
|
||
</taxonomicName>
|
||
, and fewer than
|
||
<taxonomicName id="EE8A5BCAFF91F62FFDEDD6BD2C5EFE7E" box="[975,1135,476,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="brevicauda">
|
||
<emphasis id="1BFEFC5BFF91F62FFDEDD6BD2C5EFE7E" box="[975,1135,476,500]" italics="true" pageId="6" pageNumber="64">O. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFA5FD6BD2CCBFE7E" box="[1149,1274,476,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="1BFEFC5BFF91F62FFA5FD6BD2CCBFE7E" box="[1149,1274,476,500]" italics="true" pageId="6" pageNumber="64">O. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFB2AD6BD2DA4FE7E" box="[1288,1429,476,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="hamptoni">
|
||
<emphasis id="1BFEFC5BFF91F62FFB2AD6BD2DA4FE7E" box="[1288,1429,476,500]" italics="true" pageId="6" pageNumber="64">O. hamptoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEB5D5602915FD92" box="[151,292,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFEB5D5602915FD92" box="[151,292,512,536]" italics="true" pageId="6" pageNumber="64">O. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFF10D56029F4FD92" box="[306,453,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="jintakunei">
|
||
<emphasis id="1BFEFC5BFF91F62FFF10D56029F4FD92" box="[306,453,512,536]" italics="true" pageId="6" pageNumber="64">O. jintakunei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFFF1D5602A7AFD92" box="[467,587,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="lacroixi">
|
||
<emphasis id="1BFEFC5BFF91F62FFFF1D5602A7AFD92" box="[467,587,512,536]" italics="true" pageId="6" pageNumber="64">O. lacroixi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFC7BD5602B29FD92" box="[601,792,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="lungshenensis">
|
||
<emphasis id="1BFEFC5BFF91F62FFC7BD5602B29FD92" box="[601,792,512,536]" italics="true" pageId="6" pageNumber="64">O. lungshenensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD04D5602BEBFD92" box="[806,986,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="praefrontalis">
|
||
<emphasis id="1BFEFC5BFF91F62FFD04D5602BEBFD92" box="[806,986,512,536]" italics="true" pageId="6" pageNumber="64">O. praefrontalis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EE8A5BCAFF91F62FFA35D5602C87FD92" box="[1047,1206,512,536]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="taeniolatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFA35D5602C87FD92" box="[1047,1206,512,536]" italics="true" pageId="6" pageNumber="64">O. taeniolatus</emphasis>
|
||
</taxonomicName>
|
||
(Table 1).
|
||
</paragraph>
|
||
<paragraph id="29352049FF91F62FFEEBD5442CB6FD0E" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="64">
|
||
Among species present in Sunda Islands and neighbouring areas,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD80D5452C11FDB6" box="[930,1056,549,572]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF91F62FFD80D5452C11FDB6" box="[930,1056,549,572]" italics="true" pageId="6" pageNumber="64">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF91F62FFA04D5442C4EFDB6" bold="true" box="[1062,1151,548,572]" pageId="6" pageNumber="64">
|
||
<taxonomicNameLabel id="00CD4120FF91F62FFA04D5442C4EFDB6" box="[1062,1151,548,572]" pageId="6" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="EE8A5BCAFF91F62FFB33D5452889FDEA" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFB33D5452D14FDB6" box="[1297,1317,549,572]" italics="true" pageId="6" pageNumber="64">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF91F62FFB11D5442889FDEA" italics="true" pageId="6" pageNumber="64">bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEE4D528290DFDEA" box="[198,316,584,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="1BFEFC5BFF91F62FFEE4D528290DFDEA" box="[198,316,584,608]" italics="true" pageId="6" pageNumber="64">O. everetti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFF69D52829EFFDEA" box="[331,478,584,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="jintakunei">
|
||
<emphasis id="1BFEFC5BFF91F62FFF69D52829EFFDEA" box="[331,478,584,608]" italics="true" pageId="6" pageNumber="64">O. jintakunei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFFCFD5282A5CFDEA" box="[493,621,584,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFFCFD5282A5CFDEA" box="[493,621,584,608]" italics="true" pageId="6" pageNumber="64">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
by a shorter tail, i.e. a lower ratio Tail length / Total length in males. In contrast, the new species differs from
|
||
<taxonomicName id="EE8A5BCAFF91F62FFC63D50D2AC4FD0E" box="[577,757,620,644]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="praefrontalis">
|
||
<emphasis id="1BFEFC5BFF91F62FFC63D50D2AC4FD0E" box="[577,757,620,644]" italics="true" pageId="6" pageNumber="64">O. praefrontalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD09D50D2BF3FD0E" box="[811,962,620,644]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="waandersi">
|
||
<emphasis id="1BFEFC5BFF91F62FFD09D50D2BF3FD0E" box="[811,962,620,644]" italics="true" pageId="6" pageNumber="64">O. waandersi</emphasis>
|
||
</taxonomicName>
|
||
by its longer tail.
|
||
</paragraph>
|
||
<paragraph id="29352049FF91F62FFEEBD5F02A93FCBD" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="64">
|
||
In the region of Sumatra, only
|
||
<taxonomicName id="EE8A5BCAFF91F62FFC04D5F02AF6FD22" box="[550,711,656,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="petronellae">
|
||
<emphasis id="1BFEFC5BFF91F62FFC04D5F02A0BFD2D" box="[550,570,656,679]" italics="true" pageId="6" pageNumber="64">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF91F62FFC6BD5F02AF6FD22" box="[585,711,656,680]" italics="true" pageId="6" pageNumber="64">petronellae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EE8A5BCAFF91F62FFCF4D5F02B66FD22" box="[726,855,656,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFCF4D5F02B66FD22" box="[726,855,656,680]" italics="true" pageId="6" pageNumber="64">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
and, possibly
|
||
<taxonomicName id="EE8A5BCAFF91F62FFDD9D5F02C9EFD22" box="[1019,1199,656,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="praefrontalis">
|
||
<emphasis id="1BFEFC5BFF91F62FFDD9D5F02C9EFD22" box="[1019,1199,656,680]" italics="true" pageId="6" pageNumber="64">O. praefrontalis</emphasis>
|
||
</taxonomicName>
|
||
may have 15–15–15 DSR. In
|
||
<taxonomicName id="EE8A5BCAFF91F62FFED9D5D5294BFD46" box="[251,378,693,716]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFED9D5D5294BFD46" box="[251,378,693,716]" italics="true" pageId="6" pageNumber="64">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
, this occurrence is rare and has been met only in the
|
||
<typeStatus id="F6319EEBFF91F62FFDF7D5D42C06FD46" box="[981,1079,692,716]" pageId="6" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="EE8A5BCAFF91F62FFA7FD5D52D12FD46" box="[1117,1315,692,716]" class="Squamata" family="Colubridae" genus="Simotes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="annulifer">
|
||
<emphasis id="1BFEFC5BFF91F62FFA7FD5D52D12FD46" box="[1117,1315,692,716]" italics="true" pageId="6" pageNumber="64">Simotes annulifer</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFB08D5D528F2FD7A" class="Squamata" family="Colubridae" genus="Simotes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="64" phylum="Chordata" rank="variety" species="annulifer" variety="annulata">
|
||
var.
|
||
<emphasis id="1BFEFC5BFF91F62FFB79D5D528F2FD7A" italics="true" pageId="6" pageNumber="64">annulata</emphasis>
|
||
</taxonomicName>
|
||
. The description of
|
||
<taxonomicName id="EE8A5BCAFF91F62FFF81D5B82A67FD7A" box="[419,598,728,752]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="praefrontalis">
|
||
<emphasis id="1BFEFC5BFF91F62FFF81D5B82986FD65" box="[419,439,728,751]" italics="true" pageId="6" pageNumber="64">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF91F62FFFE6D5B82A67FD7A" box="[452,598,728,752]" italics="true" pageId="6" pageNumber="64">praefrontalis</emphasis>
|
||
</taxonomicName>
|
||
did not mention the complete scale row formula, and the
|
||
<typeStatus id="F6319EEBFF91F62FFAF5D5B72D08FD65" box="[1239,1337,727,751]" pageId="6" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
and sole specimen of this species is lost. A summary of morphological characters of the nine species of
|
||
<taxonomicName id="EE8A5BCAFF91F62FFAEAD59C2D03FC9E" box="[1224,1330,764,788]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="genus">
|
||
<emphasis id="1BFEFC5BFF91F62FFAEAD59C2D03FC9E" box="[1224,1330,764,788]" italics="true" pageId="6" pageNumber="64">Oligodon</emphasis>
|
||
</taxonomicName>
|
||
recorded from the region of Sumatra appears in
|
||
<tableCitation id="640815F2FF91F62FFC6AD47F2AAFFCBD" box="[584,670,799,823]" captionStart="TABLE 2" captionStartId="6.[151,235,872,893]" captionText="TABLE 2. Main characters of Oligodon species recorded from the region of Sumatra. Exceptional values are placed in brackets; values for VEN, SC and relative tail length in males only given in square brackets where these differ from known species ranges. See Materials and Methods for abbreviations, with the addition of: Anal: D—divided; E—entire." pageId="6" pageNumber="64">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="7DF570C1FF91F62FFEB5D4082C8AFC31" pageId="6" pageNumber="64">
|
||
<paragraph id="29352049FF91F62FFEB5D4082C8AFC31" blockId="6.[151,1436,872,1205]" pageId="6" pageNumber="64">
|
||
<emphasis id="1BFEFC5BFF91F62FFEB5D4082935FCF7" bold="true" box="[151,260,872,893]" pageId="6" pageNumber="64">TABLE 2.</emphasis>
|
||
Main characters of
|
||
<taxonomicName id="EE8A5BCAFF91F62FFFF3D4082A01FCF4" box="[465,560,872,894]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="genus">
|
||
<emphasis id="1BFEFC5BFF91F62FFFF3D4082A01FCF4" box="[465,560,872,894]" italics="true" pageId="6" pageNumber="64">Oligodon</emphasis>
|
||
</taxonomicName>
|
||
species recorded from the region of Sumatra. Exceptional values are placed in brackets; values for VEN, SC and relative tail length in males only given in square brackets where these differ from known species ranges. See Materials and Methods for abbreviations, with the addition of: Anal: D—divided; E—entire.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="29352049FF91F62FFEBDD4B62CEDFC66" blockId="6.[151,1436,872,1205]" box="[159,1244,982,1004]" pageId="6" pageNumber="64">Species Hemipenes Hemipenes DSR MT Anal VEN</paragraph>
|
||
<paragraph id="29352049FF91F62FFC0ED4952A93FB81" blockId="6.[151,1436,872,1205]" box="[556,674,1013,1035]" pageId="6" pageNumber="64">length (SC)</paragraph>
|
||
<paragraph id="29352049FF91F62FFEBDD37F2CFFFBBE" blockId="6.[151,1436,872,1205]" box="[159,1230,1054,1077]" pageId="6" pageNumber="64">
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD37F2920FBBF" box="[159,273,1055,1077]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF91F62FFEBDD37F2920FBBF" box="[159,273,1055,1077]" italics="true" pageId="6" pageNumber="64">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF91F62FFF35D37F2959FBBE" bold="true" box="[279,360,1055,1076]" pageId="6" pageNumber="64">
|
||
<taxonomicNameLabel id="00CD4120FF91F62FFF35D37F2959FBBE" box="[279,360,1055,1076]" pageId="6" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Not forked 18
|
||
<date id="5D340689FF91F62FFCFBD37E2BE5FBBE" box="[729,980,1054,1076]" pageId="6" pageNumber="64" value="1915-09-15" valueMax="1915-09-15" valueMin="1915-09-15">
|
||
15–15–
|
||
<geoCoordinate id="4CBE468EFF91F62FFD04D37E2BE5FBBE" box="[806,980,1054,1076]" direction="east" orientation="longitude" pageId="6" pageNumber="64" precision="925" value="15.15">15 9</geoCoordinate>
|
||
</date>
|
||
E 156
|
||
</paragraph>
|
||
<paragraph id="29352049FF91F62EFEBDD32C2B3EFF5E" blockId="6.[151,1436,872,1205]" lastBlockId="7.[151,1436,151,1583]" lastPageId="7" lastPageNumber="65" pageId="6" pageNumber="64">
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD32C29AFFBE8" authority="Not" authorityName="Not" box="[159,414,1100,1122]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFEBDD32C2903FBE8" box="[159,306,1100,1122]" italics="true" pageId="6" pageNumber="64">O. bitorquatus</emphasis>
|
||
Not
|
||
</taxonomicName>
|
||
forked 15–16 17–(15)
|
||
<date id="5D340689FF91F62FFD0AD32B2BDCFBEB" box="[808,1005,1099,1121]" pageId="6" pageNumber="64" value="1917-06-15" valueMax="1917-09-15" valueMin="1917-06-15">17–15 6–9</date>
|
||
E 136–165 [136–160]
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD31629AFFB01" authority="Not" authorityName="Not" box="[159,414,1141,1163]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="octolineatus">
|
||
<emphasis id="1BFEFC5BFF91F62FFEBDD316290BFB01" box="[159,314,1141,1163]" italics="true" pageId="6" pageNumber="64">O. octolineatus</emphasis>
|
||
Not
|
||
</taxonomicName>
|
||
forked 25–27
|
||
<date id="5D340689FF91F62FFCFBD3162BC8FB01" box="[729,1017,1142,1163]" pageId="6" pageNumber="64" value="1917-08-17" valueMax="1917-10-17" valueMin="1917-08-17">17–17–17 8–10</date>
|
||
E 155–197 [155–175]
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD3FF29AFFB3E" authority="Not" authorityName="Not" box="[159,414,1183,1205]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="petronellae">
|
||
<emphasis id="1BFEFC5BFF91F62FFEBDD3FF2901FB3F" box="[159,304,1183,1205]" italics="true" pageId="6" pageNumber="64">O. petronellae</emphasis>
|
||
Not
|
||
</taxonomicName>
|
||
forked?
|
||
<date id="5D340689FF91F62FFCFBD3FE2BDDFB3E" box="[729,1004,1182,1204]" pageId="6" pageNumber="64" value="1915-07-15" valueMax="1915-08-15" valueMin="1915-07-15">15–15–15 7–8</date>
|
||
E 144–163 [144–155]
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEEBD0D529A7F847" box="[201,406,1973,1997]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF91F62FFEEBD0D529A7F847" box="[201,406,1973,1997]" italics="true" pageId="6" pageNumber="64">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF91F62FFFBFD0D529C6F847" bold="true" box="[413,503,1973,1997]" pageId="6" pageNumber="64">
|
||
<taxonomicNameLabel id="00CD4120FF91F62FFFBFD0D529C6F847" box="[413,503,1973,1997]" pageId="6" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar superficially to
|
||
<taxonomicName id="EE8A5BCAFF91F62FFD7ED0D52BCCF847" box="[860,1021,1973,1997]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="petronellae">
|
||
<emphasis id="1BFEFC5BFF91F62FFD7ED0D52B41F846" box="[860,880,1973,1996]" italics="true" pageId="6" pageNumber="64">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF91F62FFD5DD0D52BCCF847" box="[895,1021,1973,1997]" italics="true" pageId="6" pageNumber="64">petronellae</emphasis>
|
||
</taxonomicName>
|
||
. However it differs readily from this species by its dorsal pattern of pale, transverse vertebral blotches or crossbars vs. elongate vertebral blotches anteriorly, turning to rhombic blotches posteriorly; furthermore, the dorsal surface of
|
||
<taxonomicName id="EE8A5BCAFF90F62EFA1DD7F82C8EFF3A" box="[1087,1215,152,176]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFA1DD7F82C8EFF3A" box="[1087,1215,152,176]" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFAEAD7F82D15FF3A" bold="true" box="[1224,1316,152,176]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFAEAD7F82D15FF3A" box="[1224,1316,152,176]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is heavily dotted with pale dots which are absent in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFC4AD7DD2B39FF5E" box="[616,776,188,212]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="petronellae">
|
||
<emphasis id="1BFEFC5BFF90F62EFC4AD7DD2A4DFF5E" box="[616,636,189,212]" italics="true" pageId="7" pageNumber="65">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF90F62EFCA8D7DC2B39FF5E" box="[650,776,188,212]" italics="true" pageId="7" pageNumber="65">petronellae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="29352049FF91F62FFEBDD3A92C9EF8E5" pageId="6" pageNumber="64">
|
||
<table id="5B8AD2E9FF9109D6FEB5D3A82DADF8FA" box="[151,1436,1224,1904]" gridcols="4" gridrows="5" pageId="6" pageNumber="64">
|
||
<tr id="97BA220BFF9109D6FEB5D3A82DADFA09" box="[151,1436,1224,1411]" gridrow="0" pageId="6" pageNumber="64">
|
||
<th id="D46B4B77FF9109D6FEB5D3A82979FA09" box="[151,328,1224,1411]" gridcol="0" gridrow="0" pageId="6" pageNumber="64">
|
||
<emphasis id="1BFEFC5BFF91F62FFEBDD3A92917FA09" italics="true" pageId="6" pageNumber="64">
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD3A92970FB54" box="[159,321,1224,1246]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="praefrontalis">O. praefrontalis</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD3922974FA8D" box="[159,325,1265,1287]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="pulcherrimus">O. pulcherrimus</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD27B2979FABA" box="[159,328,1307,1328]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="purpurascens">O. purpurascens</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD2242923FAD0" box="[159,274,1348,1370]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="signatus">O. signatus</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD20E2917FA09" box="[159,294,1389,1411]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="trilineatus">O. trilineatus</taxonomicName>
|
||
</emphasis>
|
||
</th>
|
||
<th id="D46B4B77FF9109D6FF5AD3A82A5AFA09" box="[376,619,1224,1411]" gridcol="1" gridrow="0" pageId="6" pageNumber="64">???? Not forked 14–16 Not forked 25–26 Not forked 23</th>
|
||
<th id="D46B4B77FF9109D6FC92D3A82C38FA09" box="[688,1033,1224,1411]" gridcol="2" gridrow="0" pageId="6" pageNumber="64">?–15–?? 17–17–15 8–9 19-21–19-21–15-17 9-10 17–17–15 7–8 17–17–15 7–8</th>
|
||
<th id="D46B4B77FF9109D6FA13D3A82DADFA09" box="[1073,1436,1224,1411]" gridcol="3" gridrow="0" pageId="6" pageNumber="64">D 193 D 152–176 [152–157] E 159–209 [159–181] E 144–160 [144–150] E 146–162 [146–149]</th>
|
||
</tr>
|
||
<tr id="97BA220BFF9109D6FEB5D2CC2DADFA48" box="[151,1436,1452,1474]" gridrow="1" pageId="6" pageNumber="64" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="D46B4B77FF9109D6FEB5D2CC2979FA48" box="[151,328,1452,1474]" gridcol="0" gridrow="1" pageId="6" pageNumber="64">continued.</th>
|
||
</tr>
|
||
<tr id="97BA220BFF9109D6FEB5D2BE2DADFA7E" box="[151,1436,1502,1524]" gridrow="2" pageId="6" pageNumber="64">
|
||
<th id="D46B4B77FF9109D6FEB5D2BE2979FA7E" box="[151,328,1502,1524]" gridcol="0" gridrow="2" pageId="6" pageNumber="64">Species</th>
|
||
<td id="D46B4B77FF9109D6FF5AD2BE2A5AFA7E" box="[376,619,1502,1524]" gridcol="1" gridrow="2" pageId="6" pageNumber="64">SC</td>
|
||
<td id="D46B4B77FF9109D6FC92D2BE2C38FA7E" box="[688,1033,1502,1524]" gridcol="2" gridrow="2" pageId="6" pageNumber="64">SL InN Lor Ate</td>
|
||
<td id="D46B4B77FF9109D6FA13D2BE2DADFA7E" box="[1073,1436,1502,1524]" gridcol="3" gridrow="2" pageId="6" pageNumber="64">Ratio TaL/TL</td>
|
||
</tr>
|
||
<tr id="97BA220BFF9109D6FEB5D16B2DADF9AB" box="[151,1436,1547,1569]" gridrow="3" pageId="6" pageNumber="64">
|
||
<th id="D46B4B77FF9109D6FEB5D16B2979F9AB" box="[151,328,1547,1569]" gridcol="0" gridrow="3" pageId="6" pageNumber="64">
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD16C2920F9AB" box="[159,273,1548,1569]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="wagneri">
|
||
<emphasis id="1BFEFC5BFF91F62FFEBDD16C2920F9AB" box="[159,273,1548,1569]" italics="true" pageId="6" pageNumber="64">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="D46B4B77FF9109D6FF5AD16B2A5AF9AB" box="[376,619,1547,1569]" gridcol="1" gridrow="3" pageId="6" pageNumber="64">41</td>
|
||
<td id="D46B4B77FF9109D6FC92D16B2C38F9AB" box="[688,1033,1547,1569]" gridcol="2" gridrow="3" pageId="6" pageNumber="64">7 2 1 1</td>
|
||
<td id="D46B4B77FF9109D6FA13D16B2DADF9AB" box="[1073,1436,1547,1569]" gridcol="3" gridrow="3" pageId="6" pageNumber="64">0.162</td>
|
||
</tr>
|
||
<tr id="97BA220BFF9109D6FEB5D1582DADF8FA" box="[151,1436,1592,1904]" gridrow="4" pageId="6" pageNumber="64">
|
||
<th id="D46B4B77FF9109D6FEB5D1582979F8FA" box="[151,328,1592,1904]" gridcol="0" gridrow="4" pageId="6" pageNumber="64">
|
||
<emphasis id="1BFEFC5BFF91F62FFEBDD1592917F8FA" italics="true" pageId="6" pageNumber="64">
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD1592903F9C4" box="[159,306,1592,1614]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="bitorquatus">O. bitorquatus</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD103290BF9F2" box="[159,314,1634,1656]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="octolineatus">O. octolineatus</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD1EC2901F92B" box="[159,304,1675,1697]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="petronellae">O. petronellae</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD1D52970F941" box="[159,321,1717,1739]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="praefrontalis">O. praefrontalis</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD1BF2974F97E" box="[159,325,1758,1780]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="pulcherrimus">O. pulcherrimus</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD0682979F897" box="[159,328,1800,1821]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="purpurascens">O. purpurascens</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD0512923F8CD" box="[159,274,1841,1863]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="signatus">O. signatus</taxonomicName>
|
||
<taxonomicName id="EE8A5BCAFF91F62FFEBDD03B2917F8FA" box="[159,294,1882,1904]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="trilineatus">O. trilineatus</taxonomicName>
|
||
</emphasis>
|
||
</th>
|
||
<td id="D46B4B77FF9109D6FF5AD1582A5AF8FA" box="[376,619,1592,1904]" gridcol="1" gridrow="4" pageId="6" pageNumber="64">29–44 [38–44] 40–62 [51–62] 30–42 [35–42] 37 38–53 [39–46] 40–60 [50–56] 44–59 [48–59] 39–61 [54–61]</td>
|
||
<td id="D46B4B77FF9109D6FC92D1582C38F8FA" box="[688,1033,1592,1904]" gridcol="2" gridrow="4" pageId="6" pageNumber="64">(6) 7 2 (0) 1 1 (5) 6 (7) 2 1 1–2 7 2 1 1–2 7 0 0 1 (6) 7 (0)2 1 1 (6) 7–8 2 1 1–2 7 2 1 1 7 2 1 1(2)</td>
|
||
<td id="D46B4B77FF9109D6FA13D1582DADF8FA" box="[1073,1436,1592,1904]" gridcol="3" gridrow="4" pageId="6" pageNumber="64">0.127–0.195 [0.172–0.195] 0.131–0.224 [0.181–0.224] 0.122–0.178 [0.150–0.178] 0.129 0.114–0.212 [0.160–0.212] 0.115–0.188 [0.152–0.188] 0.181–0.249 [0.181–0.242] 0.198–0.244</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="29352049FF90F62EFEEBD7802AC5FE45" blockId="7.[151,1436,151,1583]" pageId="7" pageNumber="65">
|
||
<taxonomicName id="EE8A5BCAFF90F62EFEEBD78029A7FF72" box="[201,406,224,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFEEBD78029A7FF72" box="[201,406,224,248]" italics="true" pageId="7" pageNumber="65">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFFBDD78029C8FF72" bold="true" box="[415,505,224,248]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFFBDD78029C8FF72" box="[415,505,224,248]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
generally differs from
|
||
<taxonomicName id="EE8A5BCAFF90F62EFD23D7802BB0FF72" box="[769,897,224,248]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFD23D7802BB0FF72" box="[769,897,224,248]" italics="true" pageId="7" pageNumber="65">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
by its dorsal scale formula, 15–15–15 DSR vs. usually 17–15–15. When
|
||
<taxonomicName id="EE8A5BCAFF90F62EFF9ED6652A0DFE96" box="[444,572,261,284]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFF9ED6652A0DFE96" box="[444,572,261,284]" italics="true" pageId="7" pageNumber="65">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
has 15–15–15 DSR,
|
||
<taxonomicName id="EE8A5BCAFF90F62EFD11D6652B82FE96" box="[819,947,261,284]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFD11D6652B82FE96" box="[819,947,261,284]" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFD99D6642C27FE96" bold="true" box="[955,1046,260,284]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFD99D6642C27FE96" box="[955,1046,260,284]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs by (1) a much shorter tail, with a ratio TaL/TL of 0.162 vs.
|
||
<quantity id="EE728DACFF90F62EFC37D6472AF1FEB5" box="[533,704,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="5.461" metricValueMax="6.3246" metricValueMin="4.5973999999999995" pageId="7" pageNumber="65" unit="in" value="0.215" valueMax="0.249" valueMin="0.181">0.181–0.249 in</quantity>
|
||
<taxonomicName id="EE8A5BCAFF90F62EFCEBD6482B7BFECA" box="[713,842,296,320]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFCEBD6482B7BFECA" box="[713,842,296,320]" italics="true" pageId="7" pageNumber="65">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
, (2) more ventrals, 156 vs.
|
||
<quantity id="EE728DACFF90F62EFAAED6472D3CFEB5" box="[1164,1293,295,319]" metricMagnitude="0" metricUnit="m" metricValue="3.7338" metricValueMax="3.81" metricValueMin="3.6576" pageId="7" pageNumber="65" unit="in" value="147.0" valueMax="150.0" valueMin="144.0">144–150 in</quantity>
|
||
males of
|
||
<taxonomicName id="EE8A5BCAFF90F62EFBA3D64828C5FEEE" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFBA3D64828C5FEEE" italics="true" pageId="7" pageNumber="65">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
, (3) a dorsal pattern of pale, transversal vertebral blotches or crossbars in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFA14D62D2C84FEEE" box="[1078,1205,333,356]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri">
|
||
<emphasis id="1BFEFC5BFF90F62EFA14D62D2C84FEEE" box="[1078,1205,333,356]" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
vs. pale, long crossbars, the first ones shaped as long, narrow chevrons, the posterior ones as narrow crossbars in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFA9FD6102D0CFE02" box="[1213,1341,368,392]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFA9FD6102D0CFE02" box="[1213,1341,368,392]" italics="true" pageId="7" pageNumber="65">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
, and (4) a heavily ornate venter with large, dark brown blotches in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFD67D6F52BF6FE26" box="[837,967,405,428]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFD67D6F52BF6FE26" box="[837,967,405,428]" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFDF3D6F42C1FFE26" bold="true" box="[977,1070,404,428]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFDF3D6F42C1FFE26" box="[977,1070,404,428]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
vs. venter uniform with a dark brown blotch on the tip of each ventral in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFC4FD6D82ADDFE5A" box="[621,748,440,464]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="signatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFC4FD6D82ADDFE5A" box="[621,748,440,464]" italics="true" pageId="7" pageNumber="65">O. signatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="29352049FF90F62EFEEBD6BC2AF7FCBD" blockId="7.[151,1436,151,1583]" pageId="7" pageNumber="65">
|
||
Lastly,
|
||
<taxonomicName id="EE8A5BCAFF90F62EFF3FD6BC29DDFE7E" box="[285,492,476,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri">
|
||
<emphasis id="1BFEFC5BFF90F62EFF3FD6BC29DDFE7E" box="[285,492,476,500]" italics="true" pageId="7" pageNumber="65">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
is most similar in colour pattern to
|
||
<taxonomicName id="EE8A5BCAFF90F62EFDAFD6BD2C00FE7E" box="[909,1073,476,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFDAFD6BD2B90FE7E" box="[909,929,477,500]" italics="true" pageId="7" pageNumber="65">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF90F62EFD93D6BC2C00FE7E" box="[945,1073,476,500]" italics="true" pageId="7" pageNumber="65">bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="EE8A5BCAFF90F62EFA95D6BD2D06FE7E" box="[1207,1335,477,500]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFA95D6BD2D06FE7E" box="[1207,1335,477,500]" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFB62D6BC2DADFE7E" bold="true" box="[1344,1436,476,500]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFB62D6BC2DADFE7E" box="[1344,1436,476,500]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from this latter species by (1) the number of dorsal scale rows, 15–15–15 vs. 17–17–15 or exceptionally 17– 15–
|
||
<quantity id="EE728DACFF90F62EFEE0D54428CBFDB6" box="[194,250,548,572]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="7" pageNumber="65" unit="in" value="15.0">15 in</quantity>
|
||
<taxonomicName id="EE8A5BCAFF90F62EFF23D5452992FDB6" box="[257,419,548,572]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFF23D5452992FDB6" box="[257,419,548,572]" italics="true" pageId="7" pageNumber="65">O. bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
, (2) a shorter tail, with a ratio TaL/TL equal to 0.162 vs.
|
||
<quantity id="EE728DACFF90F62EFA06D5442CFCFDB6" box="[1060,1229,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="4.6609" metricValueMax="4.953" metricValueMin="4.3688" pageId="7" pageNumber="65" unit="in" value="0.1835" valueMax="0.195" valueMin="0.172">0.172–0.195 in</quantity>
|
||
males of
|
||
<taxonomicName id="EE8A5BCAFF90F62EFB18D54528D3FDEA" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFB18D54528D3FDEA" italics="true" pageId="7" pageNumber="65">O. bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
, (3) vertebral blotches transversally elongate or appearing as narrow crossbars in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFAACD5282D3EFDEA" box="[1166,1295,584,608]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFAACD5282D3EFDEA" box="[1166,1295,584,608]" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFB35D5282D42FDEA" bold="true" box="[1303,1395,584,608]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFB35D5282D42FDEA" box="[1303,1395,584,608]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
vs. rounded vertebral blotches anteriorly in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFC75D50D2AC8FD0E" box="[599,761,620,644]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFC75D50D2AC8FD0E" box="[599,761,620,644]" italics="true" pageId="7" pageNumber="65">O. bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
, (4) one or two enlarged nuchal scales behind parietals in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFBA3D50D2926FD22" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFBA3D50D2926FD22" italics="true" pageId="7" pageNumber="65">O. bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
, absent in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFFACD5F02A3DFD22" box="[398,524,656,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFFACD5F02A3DFD22" box="[398,524,656,680]" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFC30D5F02A5BFD22" bold="true" box="[530,618,656,680]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFC30D5F02A5BFD22" box="[530,618,656,680]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and (5) hemipenis spinose on its proximal half in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFABCD5F02D2AFD22" box="[1182,1307,656,680]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri">
|
||
<emphasis id="1BFEFC5BFF90F62EFABCD5F02D2AFD22" box="[1182,1307,656,680]" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
vs. on only its first proximal quarter in
|
||
<taxonomicName id="EE8A5BCAFF90F62EFFCBD5D52AA0FD46" box="[489,657,692,716]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFFCBD5D529CCFD46" box="[489,509,693,716]" italics="true" pageId="7" pageNumber="65">O</emphasis>
|
||
.
|
||
<emphasis id="1BFEFC5BFF90F62EFC33D5D42AA0FD46" box="[529,657,692,716]" italics="true" pageId="7" pageNumber="65">bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore,
|
||
<taxonomicName id="EE8A5BCAFF90F62EFD66D5D52BDCFD46" box="[836,1005,692,716]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="bitorquatus">
|
||
<emphasis id="1BFEFC5BFF90F62EFD66D5D52BDCFD46" box="[836,1005,692,716]" italics="true" pageId="7" pageNumber="65">O. bitorquatus</emphasis>
|
||
</taxonomicName>
|
||
is known from the islands of Java, Panaitan, Sumbawa, and, in Sumatra, only from Lampung Province (Bandar Lampung; Lake Ranau on the border between the provinces of Sumatera Selatan and Lampung; Gunung Rajabasa; David & Vogel, unpublished), all substantially further south than the island of Nias.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="619073C2FF90F62EFEEBD42429CBFC66" pageId="7" pageNumber="65" type="etymology">
|
||
<paragraph id="29352049FF90F62EFEEBD42429CBFC66" blockId="7.[151,1436,151,1583]" pageId="7" pageNumber="65">
|
||
<emphasis id="1BFEFC5BFF90F62EFEEBD4242979FCD6" bold="true" box="[201,328,836,860]" pageId="7" pageNumber="65">Etymology</emphasis>
|
||
. This new species is named in honour of Mr. Frederick W. Wagner, who noticed the distinct specific status of this new species and for his contribution to the systematics of the genus
|
||
<taxonomicName id="EE8A5BCAFF90F62EFA41D4082CFCFC0A" box="[1123,1229,872,896]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="genus">
|
||
<emphasis id="1BFEFC5BFF90F62EFA41D4082CFCFC0A" box="[1123,1229,872,896]" italics="true" pageId="7" pageNumber="65">Oligodon</emphasis>
|
||
</taxonomicName>
|
||
. Although unpublished, his works (
|
||
<bibRefCitation id="4D1B5DB8FF90F62EFF46D4EC29CFFC2E" author="Wagner" box="[356,510,908,932]" pageId="7" pageNumber="65" refString="Wagner, F. W. (1975) A revision of the Asian colubrid snakes Oligodon cinereus (Gunther), Oligodon joynsoni (Smith), and Oligodon cyclurus (Cantor). Unpublished MS thesis, Louisiana State University, Baton Rouge (USA), 97 pp." type="book" year="1975">Wagner 1975</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4D1B5DB8FF90F62EFC28D4EC2A72FC2E" author="Wagner" box="[522,579,908,932]" pageId="7" pageNumber="65" refString="Wagner, F. W. (1976) A reassessment of infrageneric groupings and relationships of the Asian Colubrid snake genus Oligodon. Louisiana State University, Seminar 511, Baton Rouge, 37 unnumbered pp." type="journal article" year="1976">1976</bibRefCitation>
|
||
) are good bases to the taxonomy of this difficult genus. As common names, we suggest Wagner’s kukri Snake (English),
|
||
<taxonomicName id="EE8A5BCAFF90F62EFCBCD4CF2B39FC4D" box="[670,776,943,967]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="genus">Oligodon</taxonomicName>
|
||
de Wagner (French), Wagners Kukrinatter (
|
||
<collectingCountry id="519D60D9FF90F62EFB26D4CF2D53FC4D" box="[1284,1378,943,967]" name="Germany" pageId="7" pageNumber="65">German</collectingCountry>
|
||
) and Kepel Nias (Bahasa
|
||
<collectingCountry id="519D60D9FF90F62EFF5ED4B429DFFC66" box="[380,494,980,1004]" name="Indonesia" pageId="7" pageNumber="65">Indonesia</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="619073C2FF90F62EFEEBD4972AB0F9A5" pageId="7" pageNumber="65" type="distribution">
|
||
<paragraph id="29352049FF90F62EFEEBD4972B8BFAF0" blockId="7.[151,1436,151,1583]" pageId="7" pageNumber="65">
|
||
<emphasis id="1BFEFC5BFF90F62EFEEBD497296AFB9A" bold="true" box="[201,347,1015,1040]" pageId="7" pageNumber="65">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="EE8A5BCAFF90F62EFF4BD4982A06FB9A" box="[361,567,1016,1040]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFF4BD4982A06FB9A" box="[361,567,1016,1040]" italics="true" pageId="7" pageNumber="65">Oligodon wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFC1DD4982AABFB9A" bold="true" box="[575,666,1016,1040]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFC1DD4982AABFB9A" box="[575,666,1016,1040]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="F6319EEBFF90F62EFD91D4992BD2FB9B" box="[947,995,1017,1041]" pageId="7" pageNumber="65">type</typeStatus>
|
||
locality in Nias Island, Nias Archipelago, off the west coast of Sumatra,
|
||
<collectingCountry id="519D60D9FF90F62EFC16D37C2A95FBBE" box="[564,676,1052,1076]" name="Indonesia" pageId="7" pageNumber="65">Indonesia</collectingCountry>
|
||
, without precise locality data. We considered the possibility of an erroneous
|
||
<typeStatus id="F6319EEBFF90F62EFF2FD320290CFBD2" box="[269,317,1088,1112]" pageId="7" pageNumber="65">type</typeStatus>
|
||
locality. The
|
||
<typeStatus id="F6319EEBFF90F62EFFFBD3202A38FBD2" box="[473,521,1088,1112]" pageId="7" pageNumber="65">type</typeStatus>
|
||
specimen was collected by Mr Jules Claine (
|
||
<date id="5D340689FF90F62EFA2AD35F2CBBFBDD" box="[1032,1162,1087,1111]" pageId="7" pageNumber="65" value="1856" valueMax="1938">1856–1938</date>
|
||
), a French diplomat and traveller, and an accomplished photographer. He travelled extensively in Sumatra,
|
||
<collectingCountry id="519D60D9FF90F62EFA6FD3042CF0FBF6" box="[1101,1217,1124,1148]" name="Singapore" pageId="7" pageNumber="65">Singapore</collectingCountry>
|
||
, Pinang Island and West
|
||
<collectingCountry id="519D60D9FF90F62EFEF4D3E7290DFB15" box="[214,316,1159,1183]" name="Malaysia" pageId="7" pageNumber="65">Malaysia</collectingCountry>
|
||
in
|
||
<date id="5D340689FF90F62EFF43D3E729EFFB15" box="[353,478,1159,1183]" pageId="7" pageNumber="65" value="1889" valueMax="1890">1889-1890</date>
|
||
. A description of his trips in
|
||
<collectingCountry id="519D60D9FF90F62EFD05D3E72BBFFB15" box="[807,910,1159,1183]" name="Malaysia" pageId="7" pageNumber="65">Malaysia</collectingCountry>
|
||
can be found in
|
||
<bibRefCitation id="4D1B5DB8FF90F62EFA6ED3E72CDBFB15" author="Claine" box="[1100,1258,1159,1183]" pageId="7" pageNumber="65" refString="Claine, J. (1892) Un an en Malaisie. Le Tour du Monde, 63, 369 - 400." type="journal article" year="1892">Claine (1892)</bibRefCitation>
|
||
. Claine became the French consul in
|
||
<collectingCountry id="519D60D9FF90F62EFFA0D3CD2A36FB4E" box="[386,519,1196,1220]" name="Puerto Rico" pageId="7" pageNumber="65">Puerto Rico</collectingCountry>
|
||
in 1892 and seemingly never visited again the Indo-Malayan Region. There is no mention of a visit in Nias Island in
|
||
<bibRefCitation id="4D1B5DB8FF90F62EFC06D3AF2A8EFB6D" author="Claine" box="[548,703,1231,1255]" pageId="7" pageNumber="65" refString="Claine, J. (1892) Un an en Malaisie. Le Tour du Monde, 63, 369 - 400." type="journal article" year="1892">Claine (1892)</bibRefCitation>
|
||
but the collection deposited in the Paris Museum of Natural History on
|
||
<date id="5D340689FF90F62EFED0D39729ABFA85" box="[242,410,1268,1295]" pageId="7" pageNumber="65" value="1891-07-23">July 23rd, 1891</date>
|
||
contains
|
||
<specimenCount id="3F8CEBC0FF90F62EFC28D3972A95FA85" box="[522,676,1271,1295]" pageId="7" pageNumber="65" type="generic">15 specimens</specimenCount>
|
||
of reptile species indeed present in Nias or in the adjacent part of Sumatra, plus the
|
||
<typeStatus id="F6319EEBFF90F62EFF47D27A29F6FAB8" box="[357,455,1306,1330]" pageId="7" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="EE8A5BCAFF90F62EFFCDD27B2A5EFAB9" box="[495,623,1307,1331]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFFCDD27B2A5EFAB9" box="[495,623,1307,1331]" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFC55D27B2AE3FAB9" bold="true" box="[631,722,1307,1331]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFC55D27B2AE3FAB9" box="[631,722,1307,1331]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
There remains a possibility that some of these specimens may have been collected on Penang Island, off the west coast of West
|
||
<collectingCountry id="519D60D9FF90F62EFD4DD25F2BE8FADD" box="[879,985,1343,1367]" name="Malaysia" pageId="7" pageNumber="65">Malaysia</collectingCountry>
|
||
. Nevertheless, the characters of
|
||
<taxonomicName id="EE8A5BCAFF90F62EFB60D22028F5FAF1" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="wagneri" status="sp. nov.">
|
||
<emphasis id="1BFEFC5BFF90F62EFB60D22028F5FAF1" italics="true" pageId="7" pageNumber="65">O. wagneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BFEFC5BFF90F62EFEE9D2032914FAF1" bold="true" box="[203,293,1379,1403]" pageId="7" pageNumber="65">
|
||
<taxonomicNameLabel id="00CD4120FF90F62EFEE9D2032914FAF1" box="[203,293,1379,1403]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
do not agree with any known species from West
|
||
<collectingCountry id="519D60D9FF90F62EFD6ED2022B87FAF0" box="[844,950,1378,1402]" name="Malaysia" pageId="7" pageNumber="65">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="29352049FF90F62EFEEBD2E72AB0F9A5" blockId="7.[151,1436,151,1583]" pageId="7" pageNumber="65">
|
||
<emphasis id="1BFEFC5BFF90F62EFEEBD2E729B0FA15" bold="true" box="[201,385,1414,1439]" pageId="7" pageNumber="65">Natural history</emphasis>
|
||
. There are no data about the
|
||
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|
||
. Nias Island is a large oceanic island off the northern part of the west coast of Sumatra (ca.
|
||
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|
||
,
|
||
<geoCoordinate id="4CBE468EFF90F62EFC91D2CA2B23FA48" box="[691,786,1450,1474]" direction="east" orientation="longitude" pageId="7" pageNumber="65" precision="925" value="97.55">97°33’E</geoCoordinate>
|
||
). Its highest point reaches
|
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|
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a.s.l. This island, which receives about
|
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|
||
of precipitation per year, is subject to the equatorial climate. In spite of these favourable natural circumstances, only patches of lowland rainforests and inland swamps are still present (
|
||
<bibRefCitation id="4D1B5DB8FF90F62EFA99D2922DBEF980" author="Collins" box="[1211,1423,1522,1547]" pageId="7" pageNumber="65" refString="Collins, N. M., Sayer, J. A. & Whitmore, T. C. (Eds.) (1991) The conservation atlas of tropical forests - Asia and the Pacific. Simon & Schuster, New York, 256 pp." type="book" year="1991">
|
||
Collins
|
||
<emphasis id="1BFEFC5BFF90F62EFB36D2942D74F981" box="[1300,1349,1523,1547]" italics="true" pageId="7" pageNumber="65">et al</emphasis>
|
||
. 1991
|
||
</bibRefCitation>
|
||
); most of the original forest has been cleared.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |