treatments-xml/data/87/49/CC/8749CC06FFEBFFD830EFD879FA32FF37.xml

<document id="00EC7200AE403AFB1C8A6C292EC254CC" ID-CLB-Dataset="32712" ID-DOI="10.11646/zootaxa.4290.2.4" ID-GBIF-Dataset="3937c7f9-d611-4ce1-be9d-3d615adb919c" ID-ISSN="1175-5326" ID-Zenodo-Dep="829077" ID-ZooBank="AB23F5B8-8955-4BD6-9A23-9392EC275686" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110787816" checkinUser="plazi" docAuthor="Seifert, Bernhard, Okita, Ichiro &amp; Heinze, Jürgen" docDate="2017" docId="8749CC06FFEBFFD830EFD879FA32FF37" docLanguage="en" docName="zootaxa.4290.2.4.pdf" docOrigin="Zootaxa 4290 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cardiocondyla itsukii Seifert, Okita &amp; Heinze, 2017, sp. nov." docType="treatment" docUuid="30A74350-0849-4AD8-8890-E43EDA9B52C9" docUuidSource="ZooBank" docVersion="11" lastPageNumber="343" masterDocId="7B70B47EFFE4FFCC3078DE4EFFBCFFE2" masterDocTitle="A taxonomic revision of the Cardiocondyla nuda group (Hymenoptera: Formicidae)" masterLastPageNumber="356" masterPageNumber="324" pageNumber="339" updateTime="1698465409454" updateUser="ExternalLinkService">
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<mods:title id="1AB6EA359151441E33ECF37D727E212D">A taxonomic revision of the Cardiocondyla nuda group (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="B69BC2AEC64B09556422206E990D62B9">Seifert, Bernhard</mods:namePart>
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<mods:namePart id="5ADF08045ED9204E3CA12E32E07A28B1">Okita, Ichiro</mods:namePart>
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<mods:date id="5EA613D704F819B0A79C01DEF14F355E">2017</mods:date>
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<paragraph id="0F5F7D10FFEBFFC330EFD879FE41F9B3" blockId="15.[151,509,1591,1651]" box="[151,509,1591,1617]" pageId="15" pageNumber="339">
<heading id="5417CA7CFFEBFFC330EFD879FE41F9B3" bold="true" box="[151,509,1591,1617]" fontSize="11" level="1" pageId="15" pageNumber="339" reason="1">
<emphasis id="3D94A102FFEBFFC330EFD879FE41F9B3" bold="true" box="[151,509,1591,1617]" italics="true" pageId="15" pageNumber="339">
<taxonomicName id="C8E00693FFEBFFC330EFD879FE26F9B3" box="[151,410,1591,1617]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="339" phylum="Arthropoda" rank="species" species="itsukii" status="sp. nov.">Cardiocondyla itsukii</taxonomicName>
<taxonomicNameLabel id="26A71C79FFEBFFC331D9D879FE41F9B3" box="[417,509,1591,1617]" pageId="15" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<tableCitation id="426248ABFFEBFFC330EFD814FF60F991" box="[151,220,1626,1651]" captionStart="TABLE 1" captionStartId="18.[151,239,548,571]" captionText="TABLE 1. Morphometric data of workers of C. mauritanica species complex in the sequence arithmetic mean ± standard deviation [minimum, maximum], n = number of measured individuals." pageId="15" pageNumber="339">Tab. 1</tableCitation>
, 
<figureCitation id="97DB6195FFEBFFC33092D814FED1F991" box="[234,365,1626,1651]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="16.[151,250,1432,1454]" captionStartId-1="17.[151,250,1529,1551]" captionStartId-2="19.[151,250,1380,1402]" captionTargetBox-0="[173,1413,195,1408]" captionTargetBox-1="[151,1436,553,1508]" captionTargetBox-2="[324,1263,193,1346]" captionTargetId-0="figure@16.[173,1413,195,1408]" captionTargetId-1="figure@17.[151,1436,553,1508]" captionTargetId-2="figure@19.[324,1263,193,1358]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="19" captionText-0="FIGURE 10. Head of holotype of Cardiocondyla itsukii sp. nov. in dorsal aspect." captionText-1="FIGURE 11. Lateral aspect of holotype of Cardiocondyla itsukii sp. nov." captionText-2="FIGURE 12. Waist and gaster of holotype of Cardiocondyla itsukii sp. nov. in dorsal aspect." httpUri-0="https://zenodo.org/record/829093/files/figure.png" httpUri-1="https://zenodo.org/record/829095/files/figure.png" httpUri-2="https://zenodo.org/record/829097/files/figure.png" pageId="15" pageNumber="339">
Figs. 10 
<emphasis id="3D94A102FFEBFFC3313CD813FEEDF990" bold="true" box="[324,337,1629,1650]" pageId="15" pageNumber="339">–</emphasis>
12
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</paragraph>
</subSubSection>
<subSubSection id="47FA2E9BFFEBFFC330EFD8ECFB9DF959" box="[151,1057,1698,1723]" pageId="15" pageNumber="339" type="etymology">
<paragraph id="0F5F7D10FFEBFFC330EFD8ECFB9DF959" blockId="15.[151,1436,1698,2011]" box="[151,1057,1698,1723]" pageId="15" pageNumber="339">
<emphasis id="3D94A102FFEBFFC330EFD8ECFEA2F959" bold="true" box="[151,286,1698,1723]" pageId="15" pageNumber="339">Etymology.</emphasis>
The species is dedicated to Itsuki Okita, the son of the junior author.
</paragraph>
</subSubSection>
<subSubSection id="47FA2E9BFFEBFFDD30BFD888FB27FE16" lastPageId="17" lastPageNumber="341" pageId="15" pageNumber="339" type="materials_examined">
<paragraph id="0F5F7D10FFEBFFC330BFD888FB2CF871" blockId="15.[151,1436,1698,2011]" pageId="15" pageNumber="339">
<materialsCitation id="BF88774DFFEBFFC330BFD888FCA6F88D" ID-GBIF-Occurrence="1572325869" collectionCode="SMNG" collectorName="I. Okita &amp; The &amp; Seifert &amp; Okita &amp; These" elevation="12" latitude="34.72297" location="The" longLatPrecision="1" longitude="137.8388" pageId="15" pageNumber="339" specimenCount="14" specimenCount-male="2" specimenCount-worker="4" stateProvince="JAP" typeStatus="holotype">
<emphasis id="3D94A102FFEBFFC330BFD888FEC4F93D" bold="true" box="[199,376,1734,1759]" pageId="15" pageNumber="339">
<typeStatus id="D05BC3B2FFEBFFC330BFD888FEBEF93D" box="[199,258,1734,1759]" pageId="15" pageNumber="339">Type</typeStatus>
material.
</emphasis>
<location id="0A3F2BCBFFEBFFC331F9D888FE11F93D" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFEBFFC331F9D888FE11F93D" box="[385,429,1734,1759]" latitude="34.72297" longLatPrecision="1" longitude="137.8388" name="The" pageId="15" pageNumber="339" stateProvince="JAP">The</location>
<typeStatus id="D05BC3B2FFEBFFC331CFD888FDA5F93D" box="[439,537,1734,1759]" pageId="15" pageNumber="339" type="holotype">holotype</typeStatus>
plus 
<specimenCount id="19E6B699FFEBFFC33223D889FD62F93D" box="[603,734,1735,1759]" pageId="15" pageNumber="339" type="generic" typeStatus="paratypes">8 paratypes</specimenCount>
(
<specimenCount id="19E6B699FFEBFFC33289D889FCDBF93D" box="[753,871,1734,1759]" pageId="15" pageNumber="339" type="worker">4 workers</specimenCount>
, 2 gynes and 
<specimenCount id="19E6B699FFEBFFC33472D889FBD9F93D" box="[1034,1125,1734,1759]" pageId="15" pageNumber="339" type="male">2 males</specimenCount>
) are on three pins labelled &quot; 
<collectingRegion id="CD24B3F2FFEBFFC330D8D8A5FF69F8E1" box="[160,213,1771,1795]" country="Micronesia" name="Yap" pageId="15" pageNumber="339">JAP</collectingRegion>
: 
<geoCoordinate id="6AD41BD7FFEBFFC3309CD8A5FECDF8E0" box="[228,369,1771,1795]" direction="north" orientation="latitude" pageId="15" pageNumber="339" precision="1" value="34.72297">34.72297 N</geoCoordinate>
, 
<geoCoordinate id="6AD41BD7FFEBFFC331F9D8A5FDA9F8E0" box="[385,533,1771,1795]" direction="east" orientation="longitude" pageId="15" pageNumber="339" precision="1" value="137.8388">137.83881 E</geoCoordinate>
/ 
<collectingRegion id="CD24B3F2FFEBFFC3324DD8A4FD21F8E1" box="[565,669,1770,1795]" country="Japan" name="Shizuoka" pageId="15" pageNumber="339">Shizuoka</collectingRegion>
Pref. Iwata-shi, 
<elevation id="84CD9A23FFEBFFC3331DD8A5FC1FF8E0" box="[869,931,1771,1795]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="339" unit="m" value="12.0">
<quantity id="C818D0F5FFEBFFC3331DD8A5FC1FF8E0" box="[869,931,1771,1795]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="339" unit="m" value="12.0">12 m</quantity>
</elevation>
/ street margin in rural land/ leg. 
<collectorName id="A21518C6FFEBFFC3353AD8A5FA20F8E1" box="[1346,1436,1770,1795]" pageId="15" pageNumber="339">I. Okita</collectorName>
2010.09.05-81&quot;. 
<collectorName id="A21518C6FFEBFFC33120D940FE38F8C5" box="[344,388,1806,1831]" pageId="15" pageNumber="339">The</collectorName>
pin with the 
<typeStatus id="D05BC3B2FFEBFFC33251D940FD37F8C5" box="[553,651,1806,1831]" pageId="15" pageNumber="339" type="holotype">holotype</typeStatus>
carries a red 
<typeStatus id="D05BC3B2FFEBFFC3334CD941FCD8F8C5" box="[820,868,1807,1831]" pageId="15" pageNumber="339">type</typeStatus>
label &quot; 
<typeStatus id="D05BC3B2FFEBFFC333C2D940FB99F8C5" box="[954,1061,1806,1831]" pageId="15" pageNumber="339" type="holotype">Holotype</typeStatus>
(top specimen) / &amp; 
<typeStatus id="D05BC3B2FFEBFFC33564D941FA35F8C5" box="[1308,1417,1807,1831]" pageId="15" pageNumber="339" type="paratype">Paratypes</typeStatus>
/ 
<taxonomicName id="C8E00693FFEBFFC330EFD97CFE36F8A9" box="[151,394,1842,1867]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="339" phylum="Arthropoda" rank="species" species="itsukii">
<emphasis id="3D94A102FFEBFFC330EFD97CFE36F8A9" box="[151,394,1842,1867]" italics="true" pageId="15" pageNumber="339">Cardiocondyla itsukii</emphasis>
</taxonomicName>
/ 
<collectorName id="A21518C6FFEBFFC331DCD97CFE53F8A9" box="[420,495,1842,1867]" pageId="15" pageNumber="339">Seifert</collectorName>
&amp; 
<collectorName id="A21518C6FFEBFFC3326ED97CFDEBF8A9" box="[534,599,1842,1867]" pageId="15" pageNumber="339">Okita</collectorName>
&quot; and the two pins with 
<typeStatus id="D05BC3B2FFEBFFC33310D97DFC6FF8A9" box="[872,979,1843,1867]" pageId="15" pageNumber="339" type="paratype">paratypes</typeStatus>
have labels &quot; 
<typeStatus id="D05BC3B2FFEBFFC33414D97DFB60F8A9" box="[1132,1244,1843,1867]" pageId="15" pageNumber="339" type="paratype">Paratypes</typeStatus>
/ 
<taxonomicName id="C8E00693FFEBFFC3348DD97CFF60F88D" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="339" phylum="Arthropoda" rank="species" species="itsukii">
<emphasis id="3D94A102FFEBFFC3348DD97CFF60F88D" italics="true" pageId="15" pageNumber="339">Cardiocondyla itsukii</emphasis>
</taxonomicName>
/ 
<collectorName id="A21518C6FFEBFFC3308CD918FE83F88D" box="[244,319,1878,1903]" pageId="15" pageNumber="339">Seifert</collectorName>
&amp; 
<collectorName id="A21518C6FFEBFFC3311DD918FE1FF88D" box="[357,419,1878,1903]" pageId="15" pageNumber="339">Okita</collectorName>
<emphasis id="3D94A102FFEBFFC331DBD918FE0EF88D" bold="true" box="[419,434,1878,1903]" pageId="15" pageNumber="339">&quot;</emphasis>
. 
<collectorName id="A21518C6FFEBFFC331BAD918FDB9F88D" box="[450,517,1878,1903]" pageId="15" pageNumber="339">These</collectorName>
are deposited at 
<collectionCode id="69F1E5D5FFEBFFC332B1D919FCA6F88D" box="[713,794,1879,1903]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="15" pageNumber="339">SMNG</collectionCode>
</materialsCitation>
. Five additional paratype workers are deposited at CAS. Furthermore, 120 workers, 86 gynes, and 1 ergatoid male are stored in ethanol at MNEHS.
</paragraph>
<paragraph id="0F5F7D10FFEBFFC330BFD9D0FE93F839" blockId="15.[151,1436,1698,2011]" pageId="15" pageNumber="339">
<emphasis id="3D94A102FFEBFFC330BFD9D0FE57F855" bold="true" box="[199,491,1950,1975]" pageId="15" pageNumber="339">All material examined.</emphasis>
A total of 45 nest samples with 
<specimenCount id="19E6B699FFEBFFC333D8D9D1FB88F855" box="[928,1076,1950,1975]" pageId="15" pageNumber="339" type="worker">108 workers</specimenCount>
were subject to NUMOBAT investigation.
</paragraph>
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<paragraph id="0F5F7D10FFF4FFDC30EFDBD6FC6DFA4C" blockId="16.[151,977,1432,1454]" box="[151,977,1432,1454]" pageId="16" pageNumber="340">
<emphasis id="3D94A102FFF4FFDC30EFDBD6FEA3FA4F" bold="true" box="[151,287,1432,1454]" pageId="16" pageNumber="340">FIGURE 10.</emphasis>
Head of holotype of 
<taxonomicName id="C8E00693FFF4FFDC318FDBD6FD6EFA4C" box="[503,722,1432,1454]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="340" phylum="Arthropoda" rank="species" species="itsukii" status="sp. nov.">
<emphasis id="3D94A102FFF4FFDC318FDBD6FD6EFA4C" box="[503,722,1432,1454]" italics="true" pageId="16" pageNumber="340">Cardiocondyla itsukii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26A71C79FFF4FFDC32A0DBD7FC9BFA4C" box="[728,807,1433,1454]" pageId="16" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
in dorsal aspect.
</paragraph>
</caption>
<paragraph id="0F5F7D10FFF4FFDD30BFDB98FB27FE16" blockId="16.[151,1437,1494,2023]" lastBlockId="17.[151,1436,151,501]" lastPageId="17" lastPageNumber="341" pageId="16" pageNumber="340">
<emphasis id="3D94A102FFF4FFDC30BFDB98FE95FA0D" bold="true" box="[199,297,1494,1519]" pageId="16" pageNumber="340">Bhutan:</emphasis>
Puntsholing, 1972.04.0 5, [26.86, 89.39]. 
<materialsCitation id="BF88774DFFF4FFDC3363DB98FF50F9F1" ID-GBIF-Occurrence="1572325872" country="China" location="Hawaii" pageId="16" pageNumber="340" specimenCount="1" stateProvince="Hawaii">
<collectingCountry id="77F73D80FFF4FFDC3363DB98FCD8FA0D" box="[795,868,1494,1519]" name="China" pageId="16" pageNumber="340">
<emphasis id="3D94A102FFF4FFDC3363DB98FCD8FA0D" bold="true" box="[795,868,1494,1519]" pageId="16" pageNumber="340">China</emphasis>
</collectingCountry>
: Nanning, 1966.06.0 2, [22.82, 108.30]. 
<emphasis id="3D94A102FFF4FFDC353EDB98FF50F9F1" bold="true" pageId="16" pageNumber="340">
<collectingRegion id="CD24B3F2FFF4FFDC353EDB98FA27FA0D" box="[1350,1435,1494,1519]" country="United States of America" name="Hawaii" pageId="16" pageNumber="340">Hawaii</collectingRegion>
Islands
</emphasis>
</materialsCitation>
: 
<materialsCitation id="BF88774DFFF4FFDC3083DBB4FC3CF9F1" ID-GBIF-Occurrence="1572325901" box="[251,896,1530,1555]" country="China" location="Hawaii" pageId="16" pageNumber="340" specimenCount="1" stateProvince="Hawaii">
<collectingRegion id="CD24B3F2FFF4FFDC3083DBB4FEF3F9F1" box="[251,335,1530,1555]" country="United States of America" name="Hawaii" pageId="16" pageNumber="340">Hawaii</collectingRegion>
, 2002.04.0 9, No PKS127/130, [19.442, -155.316]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF4FFDC33F2DBB4FEB4F9D5" ID-GBIF-Occurrence="1572325889" country="China" location="Hawaii" pageId="16" pageNumber="340" specimenCount="1" stateProvince="Hawaii">
<collectingRegion id="CD24B3F2FFF4FFDC33F2DBB4FC62F9F1" box="[906,990,1530,1555]" country="United States of America" name="Hawaii" pageId="16" pageNumber="340">Hawaii</collectingRegion>
, 2002.04.xx, No PDK129/131, [20.579, -156.136]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF4FFDC3163D850FC2CF9D5" ID-GBIF-Occurrence="1572325892" box="[283,912,1566,1591]" country="China" location="Hawaii" pageId="16" pageNumber="340" specimenCount="1" stateProvince="Hawaii">
<collectingRegion id="CD24B3F2FFF4FFDC3163D850FECCF9D5" box="[283,368,1566,1591]" country="United States of America" name="Hawaii" pageId="16" pageNumber="340">Hawaii</collectingRegion>
: Mauna Kea, 2001.06.14, [19.726, -155.447]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF4FFDC33DAD850FC63F8ED" ID-GBIF-Occurrence="1572325895" collectorName="Kauai &amp; No HI &amp; Maui &amp; No &amp; Kapalua &amp; Oahu &amp; Waikane Trail &amp; Pachmarhi &amp; Calcutta" country="India" county="Mauna Kea" location="East India" municipality="No" pageId="16" pageNumber="340" specimenCount="1" stateProvince="Hawaii">
<collectingRegion id="CD24B3F2FFF4FFDC33DAD850FC4BF9D5" box="[930,1015,1566,1591]" country="United States of America" name="Hawaii" pageId="16" pageNumber="340">Hawaii</collectingRegion>
: 
<collectingCounty id="E63E059CFFF4FFDC3471D851FB29F9D4" box="[1033,1173,1567,1590]" pageId="16" pageNumber="340">Mauna Kea</collectingCounty>
, 2002.06.13, 
<collectingMunicipality id="EF3BE76AFFF4FFDC3543D851FAE1F9D4" box="[1339,1373,1567,1590]" pageId="16" pageNumber="340">No</collectingMunicipality>
0 49, (GenBank 
<accessionNumber id="10B3E0F3FFF4FFDC316AD80CFE33F9B8" box="[274,399,1602,1626]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ023088" pageId="16" pageNumber="340">
DQ 
<date id="7B5E5BD0FFF4FFDC3143D80CFE33F9B8" box="[315,399,1602,1626]" pageId="16" pageNumber="340" value="88-02-30">023088</date>
</accessionNumber>
), [19.726, -155.447]; 
<collectorName id="A21518C6FFF4FFDC32ECD80CFD66F9B9" box="[660,730,1602,1627]" pageId="16" pageNumber="340">Kauai</collectorName>
, 2006.09.0 4, 
<collectorName id="A21518C6FFF4FFDC330CD80DFC02F9B9" box="[884,958,1602,1627]" pageId="16" pageNumber="340">No HI</collectorName>
54, [22.129, -159.661]; 
<collectorName id="A21518C6FFF4FFDC34B4D80CFAB7F9B9" box="[1228,1291,1602,1627]" pageId="16" pageNumber="340">Maui</collectorName>
: 2001.08.18, 
<collectorName id="A21518C6FFF4FFDC30EFD829FF05F99C" box="[151,185,1639,1662]" pageId="16" pageNumber="340">No</collectorName>
PKS121, [20.681,-156.136]; 
<collectorName id="A21518C6FFF4FFDC3261D828FDEBF99D" box="[537,599,1638,1663]" pageId="16" pageNumber="340">Maui</collectorName>
, 2003.06.16, 
<collectorName id="A21518C6FFF4FFDC3284D829FCA2F99C" box="[764,798,1639,1662]" pageId="16" pageNumber="340">No</collectorName>
173, [20.747, -156.247]; 
<collectorName id="A21518C6FFF4FFDC3420D828FB2AF99D" box="[1112,1174,1638,1663]" pageId="16" pageNumber="340">Maui</collectorName>
, 2003.07.0 3, 
<collectorName id="A21518C6FFF4FFDC3543D829FAE1F99C" box="[1339,1373,1639,1662]" pageId="16" pageNumber="340">No</collectorName>
170, [20.749, -156.241]; 
<collectorName id="A21518C6FFF4FFDC31FFD8C4FE79F941" box="[391,453,1674,1699]" pageId="16" pageNumber="340">Maui</collectorName>
, 2003.xx.xx, 
<collectorName id="A21518C6FFF4FFDC3214D8C5FD32F940" box="[620,654,1675,1698]" pageId="16" pageNumber="340">No</collectorName>
144, [20.751, -156.240]; 
<collectorName id="A21518C6FFF4FFDC33B6D8C4FBB1F941" box="[974,1037,1674,1699]" pageId="16" pageNumber="340">Maui</collectorName>
: 
<collectorName id="A21518C6FFF4FFDC3458D8C4FB3CF941" box="[1056,1152,1674,1699]" pageId="16" pageNumber="340">Kapalua</collectorName>
, 1990.02.23, [21.000, - 156.659]; 
<collectorName id="A21518C6FFF4FFDC316DD8E0FEEAF925" box="[277,342,1710,1735]" pageId="16" pageNumber="340">Oahu</collectorName>
: 
<collectorName id="A21518C6FFF4FFDC3114D8E0FDA4F925" box="[364,536,1710,1735]" pageId="16" pageNumber="340">Waikane Trail</collectorName>
, 1989.09.30, [21.483, -157.856]. 
<collectingCountry id="77F73D80FFF4FFDC33C4D8E0FC40F925" box="[956,1020,1710,1735]" name="India" pageId="16" pageNumber="340">
<emphasis id="3D94A102FFF4FFDC33C4D8E0FC40F925" bold="true" box="[956,1020,1710,1735]" pageId="16" pageNumber="340">India</emphasis>
</collectingCountry>
: Bhopal- 
<quantity id="C818D0F5FFF4FFDC340AD8E0FB6DF925" box="[1138,1233,1710,1735]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="16" pageNumber="340" unit="km" value="130.0">130 km</quantity>
SE, 
<collectorName id="A21518C6FFF4FFDC3565D8E0FA24F925" box="[1309,1432,1710,1735]" pageId="16" pageNumber="340">Pachmarhi</collectorName>
, 2006.06.26, [22.47, 78.43]; 
<collectorName id="A21518C6FFF4FFDC31A9D89CFD8DF909" box="[465,561,1746,1771]" pageId="16" pageNumber="340">Calcutta</collectorName>
, pre 1920 (Rothney-Smith), [22.60, 88.40]; Dehradun- 
<quantity id="C818D0F5FFF4FFDC34E4D89CFB5DF909" box="[1180,1249,1746,1771]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="340" unit="km" value="10.0">10 km</quantity>
SE, 1996.xx.xx, 
<collectorName id="A21518C6FFF4FFDC30EFD8B9FF05F8EC" box="[151,185,1783,1806]" pageId="16" pageNumber="340">No</collectorName>
552, [30.21, 78.13]; 
<location id="0A3F2BCBFFF4FFDC31DDD8B9FDA4F8ED" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC31DDD8B9FDA4F8ED" box="[421,536,1782,1807]" country="India" county="Mauna Kea" municipality="No" name="East India" pageId="16" pageNumber="340" stateProvince="Hawaii">East India</location>
, pre 1920 (Rothney-Smith), [23.6, 88.4]
</materialsCitation>
. 
<materialsCitation id="BF88774DFFF4FFDC3391D8B8FE1EF8D1" ID-GBIF-Occurrence="1572325905" collectingDate="I 6" country="Indonesia" location="Cibodas" pageId="16" pageNumber="340" specimenCount="1" stateProvince="Java">
<collectingCountry id="77F73D80FFF4FFDC3391D8B8FBE2F8ED" box="[1001,1118,1782,1807]" name="Indonesia" pageId="16" pageNumber="340">
<emphasis id="3D94A102FFF4FFDC3391D8B8FBE2F8ED" bold="true" box="[1001,1118,1782,1807]" pageId="16" pageNumber="340">Indonesia</emphasis>
</collectingCountry>
: 
<collectingRegion id="CD24B3F2FFF4FFDC3415D8B9FB1FF8EC" box="[1133,1187,1783,1806]" country="Indonesia" name="Jawa Timur" pageId="16" pageNumber="340">Java</collectingRegion>
: 
<location id="0A3F2BCBFFF4FFDC34D5D8B8FAB0F8ED" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC34D5D8B8FAB0F8ED" box="[1197,1292,1782,1807]" country="Indonesia" name="Cibodas" pageId="16" pageNumber="340" stateProvince="Java">Cibodas</location>
, 2000.01.0 5, No 
<collectingDate id="6B1AA238FFF4FFDC30BED954FF63F8D1" box="[198,223,1818,1843]" pageId="16" pageNumber="340">
<date id="7B5E5BD0FFF4FFDC30BED954FF63F8D1" box="[198,223,1818,1843]" pageId="16" pageNumber="340">I6</date>
</collectingDate>
, [-6.83, 107.68]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF4FFDC31CBD954FED5F805" ID-GBIF-Occurrence="1572325912" country="Japan" county="Dumoga Bone National Park" location="No" municipality="Fukuroi" pageId="16" pageNumber="340" specimenCount="1" stateProvince="Sulawesi" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF4FFDC31CBD954FDA1F8D1" box="[435,541,1818,1843]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="340">Sulawesi</collectingRegion>
: 
<collectingCounty id="E63E059CFFF4FFDC3254D955FC38F8D1" box="[556,900,1818,1843]" pageId="16" pageNumber="340">Dumoga Bone National Park</collectingCounty>
, 1985.xx.xx, [0.58, 123.90]. 
<collectingCountry id="77F73D80FFF4FFDC34A7D954FA94F8D1" box="[1247,1320,1818,1843]" name="Japan" pageId="16" pageNumber="340">
<emphasis id="3D94A102FFF4FFDC34A7D954FA94F8D1" bold="true" box="[1247,1320,1818,1843]" pageId="16" pageNumber="340">Japan</emphasis>
</collectingCountry>
: 
<collectingMunicipality id="EF3BE76AFFF4FFDC3544D954FA24F8D1" box="[1340,1432,1818,1843]" pageId="16" pageNumber="340">Fukuroi</collectingMunicipality>
, 2010.10.24, 
<location id="0A3F2BCBFFF4FFDC3156D971FEECF8B4" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC3156D971FEECF8B4" box="[302,336,1855,1878]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="No" pageId="16" pageNumber="340" stateProvince="Sulawesi">No</location>
88, (GenBank 
<accessionNumber id="10B3E0F3FFF4FFDC3264D970FD24F8B5" box="[540,664,1854,1879]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199504" pageId="16" pageNumber="340">LC199504</accessionNumber>
), [34.791,137.865]; 
<location id="0A3F2BCBFFF4FFDC33E9D971FBA6F8B4" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC33E9D971FBA6F8B4" box="[913,1050,1855,1878]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="Hamamatsu" pageId="16" pageNumber="340" stateProvince="Sulawesi">Hamamatsu</location>
, 2010.08.0 7, 
<location id="0A3F2BCBFFF4FFDC34BFD971FB55F8B4" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC34BFD971FB55F8B4" box="[1223,1257,1855,1878]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="No" pageId="16" pageNumber="340" stateProvince="Sulawesi">No</location>
71 (GenBank 
<accessionNumber id="10B3E0F3FFF4FFDC30EFD92CFEAEF899" box="[151,274,1890,1915]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199505" pageId="16" pageNumber="340">LC199505</accessionNumber>
), [34.716, 137.604]; 
<location id="0A3F2BCBFFF4FFDC3278D92CFDDFF899" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC3278D92CFDDFF899" box="[512,611,1890,1915]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="Iriomote" pageId="16" pageNumber="340" stateProvince="Sulawesi">Iriomote</location>
, 2013.09.18, 
<location id="0A3F2BCBFFF4FFDC3284D92DFCA2F898" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC3284D92DFCA2F898" box="[764,798,1891,1914]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="No" pageId="16" pageNumber="340" stateProvince="Sulawesi">No</location>
199 (GenBank 
<accessionNumber id="10B3E0F3FFF4FFDC33AAD92CFBF3F899" box="[978,1103,1890,1915]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199506" pageId="16" pageNumber="340">LC199506</accessionNumber>
), [24.258, 123.865]; 
<location id="0A3F2BCBFFF4FFDC3543D92CFA24F899" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC3543D92CFA24F899" box="[1339,1432,1890,1915]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="Ishigaki" pageId="16" pageNumber="340" stateProvince="Sulawesi">Ishigaki</location>
, 2013.09.18, 
<location id="0A3F2BCBFFF4FFDC3156D9C9FEECF87C" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC3156D9C9FEECF87C" box="[302,336,1927,1950]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="No" pageId="16" pageNumber="340" stateProvince="Sulawesi">No</location>
197 (GenBank 
<accessionNumber id="10B3E0F3FFF4FFDC325CD9C8FD1CF87D" box="[548,672,1926,1951]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199507" pageId="16" pageNumber="340">LC199507</accessionNumber>
), [24.337, 124.154]; 
<location id="0A3F2BCBFFF4FFDC33D4D9C8FBB6F87D" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC33D4D9C8FBB6F87D" box="[940,1034,1926,1951]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="Iwakuni" pageId="16" pageNumber="340" stateProvince="Sulawesi">Iwakuni</location>
, 2013.08.27, 
<location id="0A3F2BCBFFF4FFDC34C1D9C9FB67F87C" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC34C1D9C9FB67F87C" box="[1209,1243,1927,1950]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="No" pageId="16" pageNumber="340" stateProvince="Sulawesi">No</location>
175 (GenBank 
<accessionNumber id="10B3E0F3FFF4FFDC30EFD9E4FEAEF821" box="[151,274,1962,1987]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199508" pageId="16" pageNumber="340">LC199508</accessionNumber>
), [34.076, 132.210]; 
<location id="0A3F2BCBFFF4FFDC327CD9E4FDD3F821" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC327CD9E4FDD3F821" box="[516,623,1962,1987]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="Iwata-shi" pageId="16" pageNumber="340" stateProvince="Sulawesi">Iwata-shi</location>
, 2010.09.0 5, 
<location id="0A3F2BCBFFF4FFDC3372D9E5FC90F820" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC3372D9E5FC90F820" box="[778,812,1963,1986]" country="Japan" county="Dumoga Bone National Park" municipality="Fukuroi" name="No" pageId="16" pageNumber="340" stateProvince="Sulawesi">No</location>
81 (GenBank 
<accessionNumber id="10B3E0F3FFF4FFDC33AED9E4FBEEF821" box="[982,1106,1962,1987]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199509" pageId="16" pageNumber="340">LC199509</accessionNumber>
), 
<typeStatus id="D05BC3B2FFF4FFDC341FD9E5FB2BF821" box="[1127,1175,1963,1987]" pageId="16" pageNumber="340">type</typeStatus>
<taxonomicName id="C8E00693FFF4FFDC34D8D9E4FA28F821" box="[1184,1428,1962,1987]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="340" phylum="Arthropoda" rank="species" species="itsukii">
<emphasis id="3D94A102FFF4FFDC34D8D9E4FA28F821" box="[1184,1428,1962,1987]" italics="true" pageId="16" pageNumber="340">Cardiocondyla itsukii</emphasis>
</taxonomicName>
, [34.723, 137.839]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF4FFDC3102D980FC50F805" ID-GBIF-Occurrence="1572325903" box="[378,1004,1998,2023]" country="Japan" location="Mt. Unzen" pageId="16" pageNumber="340" specimenCount="1" stateProvince="Nagasaki" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF4FFDC3102D980FE5BF805" box="[378,487,1998,2023]" country="Japan" name="Nagasaki" pageId="16" pageNumber="340">Nagasaki</collectingRegion>
: 
<location id="0A3F2BCBFFF4FFDC318FD981FDCBF804" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF4FFDC318FD981FDCBF804" box="[503,631,1999,2022]" country="Japan" name="Mt. Unzen" pageId="16" pageNumber="340" stateProvince="Nagasaki">Mt. Unzen</location>
, 1983.08.20, [32.760, 130.301]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF4FFDD3385D980FD15FF1A" ID-GBIF-Occurrence="1572325891" country="Japan" county="Mt. Unzen" lastPageId="17" lastPageNumber="341" location="No" municipality="Nakagusuku" pageId="16" pageNumber="340" specimenCount="1" stateProvince="Nagasaki" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF4FFDC3385D980FBD6F805" box="[1021,1130,1998,2023]" country="Japan" name="Nagasaki" pageId="16" pageNumber="340">Nagasaki</collectingRegion>
: 
<collectingCounty id="E63E059CFFF4FFDC3402D981FB4BF804" box="[1146,1271,1999,2022]" pageId="16" pageNumber="340">Mt. Unzen</collectingCounty>
without date, [32.760, 130.301]; 
<collectingMunicipality id="EF3BE76AFFF5FFDD31F0DED9FDABFF52" box="[392,535,151,176]" pageId="17" pageNumber="341">Nakagusuku</collectingMunicipality>
, 2013.06.0 7, 
<location id="0A3F2BCBFFF5FFDD32B0DED6FD56FF4D" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF5FFDD32B0DED6FD56FF4D" box="[712,746,152,175]" country="Japan" county="Mt. Unzen" municipality="Nakagusuku" name="No" pageId="17" pageNumber="341" stateProvince="Nagasaki">No</location>
148 (GenBank 
<accessionNumber id="10B3E0F3FFF5FFDD33BBDED9FBFCFF52" box="[963,1088,151,176]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199510" pageId="17" pageNumber="341">LC199510</accessionNumber>
), [26.275, 127.781]; Nanjo, 2013.06.0 2, No 128 (GenBank 
<accessionNumber id="10B3E0F3FFF5FFDD327EDEF2FD3DFF37" box="[518,641,188,213]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199511" pageId="17" pageNumber="341">LC199511</accessionNumber>
), [26.178, 127.828]; 
<location id="0A3F2BCBFFF5FFDD330FDEF2FBF9FF37" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF5FFDD330FDEF2FBF9FF37" box="[887,1093,188,213]" country="Japan" county="Mt. Unzen" municipality="Nakagusuku" name="Ogasawara Island" pageId="17" pageNumber="341" stateProvince="Nagasaki">Ogasawara Island</location>
, 1972.07.29, [27.10, 142.20]; 
<location id="0A3F2BCBFFF5FFDD30EFDEAEFEDEFF1A" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF5FFDD30EFDEAEFEDEFF1A" box="[151,354,223,248]" country="Japan" county="Mt. Unzen" municipality="Nakagusuku" name="Ogasawara Island" pageId="17" pageNumber="341" stateProvince="Nagasaki">Ogasawara Island</location>
, 1977.02.11, [27.10, 142.20]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF5FFDD32CDDE91FF4DFEFE" ID-GBIF-Occurrence="1572325904" country="Iceland" location="Okinawa" pageId="17" pageNumber="341" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF5FFDD32CDDE91FCA3FF1A" box="[693,799,223,248]" country="Japan" name="Okinawa" pageId="17" pageNumber="341">Okinawa</collectingRegion>
: Ada 
<collectingCountry id="77F73D80FFF5FFDD3327DE91FC1AFF1A" box="[863,934,223,248]" name="Iceland" pageId="17" pageNumber="341">Island</collectingCountry>
, 1994.09.22, (GenBank 
<accessionNumber id="10B3E0F3FFF5FFDD34C1DEAEFA85FF1A" box="[1209,1337,224,248]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ023082" pageId="17" pageNumber="341">
DQ 
<date id="7B5E5BD0FFF5FFDD349ADEAEFA85FF1A" box="[1250,1337,224,248]" pageId="17" pageNumber="341" value="82-02-30">023082</date>
</accessionNumber>
), [26.74, 128.32]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF5FFDD3087DF4AFCE4FEFE" ID-GBIF-Occurrence="1572325893" box="[255,856,260,285]" country="Iceland" location="Okinawa" pageId="17" pageNumber="341" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF5FFDD3087DF4AFED5FEFF" box="[255,361,260,285]" country="Japan" name="Okinawa" pageId="17" pageNumber="341">Okinawa</collectingRegion>
: Izena 
<collectingCountry id="77F73D80FFF5FFDD31C6DF4AFDB9FEFF" box="[446,517,260,285]" name="Iceland" pageId="17" pageNumber="341">Island</collectingCountry>
, 1985.03.31, [26.93, 127.94]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF5FFDD331EDF4AFC6CFEFF" ID-GBIF-Occurrence="1572325899" box="[870,976,260,285]" country="Iceland" location="Okinawa" pageId="17" pageNumber="341" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF5FFDD331EDF4AFC6CFEFF" box="[870,976,260,285]" country="Japan" name="Okinawa" pageId="17" pageNumber="341">Okinawa</collectingRegion>
</materialsCitation>
: 
<materialsCitation id="BF88774DFFF5FFDD33A6DF4AFC11FEA2" ID-GBIF-Occurrence="1572325907" country="Iceland" location="No" municipality="Tahara" pageId="17" pageNumber="341" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF5FFDD33A6DF4AFBFAFEFF" box="[990,1094,260,285]" country="Japan" name="Okinawa" pageId="17" pageNumber="341">Okinawa</collectingRegion>
, 2003.06.26, [26.50, 127.94]; 
<collectingMunicipality id="EF3BE76AFFF5FFDD30EFDF69FF5AFEA2" box="[151,230,295,320]" pageId="17" pageNumber="341">Tahara</collectingMunicipality>
, 2011.08.0 9, 
<location id="0A3F2BCBFFF5FFDD3104DF66FE22FEDD" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF5FFDD3104DF66FE22FEDD" box="[380,414,296,319]" country="Iceland" municipality="Tahara" name="No" pageId="17" pageNumber="341" stateProvince="Okinawa">No</location>
107 (GenBank 
<accessionNumber id="10B3E0F3FFF5FFDD3229DF69FD71FEA2" box="[593,717,295,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199512" pageId="17" pageNumber="341">LC199512</accessionNumber>
), [34.587, 137.026]
</materialsCitation>
; 
<materialsCitation id="BF88774DFFF5FFDD33C0DF69FD19FE6A" ID-GBIF-Occurrence="1572325896" country="Kiribati" location="Enderbury Island" municipality="No" pageId="17" pageNumber="341" specimenCount="1" stateProvince="Wakayama" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF5FFDD33C0DF69FB8BFEA2" box="[952,1079,295,320]" country="Japan" name="Wakayama" pageId="17" pageNumber="341">Wakayama</collectingRegion>
, 2013.08.30, 
<collectingMunicipality id="EF3BE76AFFF5FFDD34B6DF66FB4CFEDD" box="[1230,1264,296,319]" pageId="17" pageNumber="341">No</collectingMunicipality>
186 (GenBank 
<accessionNumber id="10B3E0F3FFF5FFDD30EFDF02FEAEFE87" box="[151,274,332,357]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199513" pageId="17" pageNumber="341">LC199513</accessionNumber>
), [34.288, 135.072]; Yaizu, 2010.10.31, No 94 (GenBank 
<accessionNumber id="10B3E0F3FFF5FFDD33B9DF02FB81FE87" box="[961,1085,332,357]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC199514" pageId="17" pageNumber="341">LC199514</accessionNumber>
), [34.895, 138.347]. 
<collectingCountry id="77F73D80FFF5FFDD354CDF02FA28FE87" box="[1332,1428,332,357]" name="Kiribati" pageId="17" pageNumber="341">
<emphasis id="3D94A102FFF5FFDD354CDF02FA28FE87" bold="true" box="[1332,1428,332,357]" pageId="17" pageNumber="341">Kiribati</emphasis>
</collectingCountry>
: 
<location id="0A3F2BCBFFF5FFDD30EFDF21FEE6FE6A" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF5FFDD30EFDF21FEE6FE6A" box="[151,346,367,392]" country="Kiribati" municipality="No" name="Enderbury Island" pageId="17" pageNumber="341" stateProvince="Wakayama">Enderbury Island</location>
, 2000.05.0 7, [-3.13, -171.09]
</materialsCitation>
. 
<emphasis id="3D94A102FFF5FFDD32D6DF21FD48FE6A" bold="true" box="[686,756,367,392]" pageId="17" pageNumber="341">Nepal</emphasis>
: Pohkara- 
<quantity id="C818D0F5FFF5FFDD3311DF21FC12FE6A" box="[873,942,367,392]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="17" pageNumber="341" unit="km" value="27.0">27 km</quantity>
NW, 1995.11.xx, [28.302, 83.781]: Pohkara vicinity, 1995.11.xx, No 975, [28.20, 83.98]. 
<emphasis id="3D94A102FFF5FFDD32E6DFDAFC98FE4F" bold="true" box="[670,804,404,429]" pageId="17" pageNumber="341">Philippines</emphasis>
: Paloc, 1995.01, [14.580, 120.550]. 
<materialsCitation id="BF88774DFFF5FFDD34B1DFDAFDD1FE32" ID-GBIF-Occurrence="1572325902" collectorName="Le Port" country="Réunion" location="Reunion" pageId="17" pageNumber="341" specimenCount="1" typeStatus="holotype">
<collectingCountry id="77F73D80FFF5FFDD34B1DFDAFA92FE4F" box="[1225,1326,404,429]" name="Réunion" pageId="17" pageNumber="341">
<emphasis id="3D94A102FFF5FFDD34B1DFDAFA92FE4F" bold="true" box="[1225,1326,404,429]" pageId="17" pageNumber="341">Réunion</emphasis>
</collectingCountry>
: 
<collectorName id="A21518C6FFF5FFDD3547DFDBFA24FE4E" box="[1343,1432,405,428]" pageId="17" pageNumber="341">Le Port</collectorName>
, 2007.05.21, No 17714, [-20.938, 55.295]
</materialsCitation>
. 
<emphasis id="3D94A102FFF5FFDD3201DFF9FD40FE32" bold="true" box="[633,764,439,464]" pageId="17" pageNumber="341">Sri Lanka:</emphasis>
Bandarawella, 1988.01.16, [6.824, 80.994]; Nuwara Eliya, 1988.01.16, [6.953, 80.783]. 
<emphasis id="3D94A102FFF5FFDD31A5DF92FDF6FE17" bold="true" box="[477,586,476,501]" pageId="17" pageNumber="341">Thailand</emphasis>
: Koh Chiang, beach, 2004.01.xx, [12.048, 102.302].
</paragraph>
</subSubSection>
<caption id="5B9F2D98FFF5FFDD30EFDBB7FCC2F9ED" box="[151,894,1529,1551]" httpUri="https://zenodo.org/record/829095/files/figure.png" pageId="17" pageNumber="341" targetBox="[151,1436,553,1508]" targetPageId="17">
<paragraph id="0F5F7D10FFF5FFDD30EFDBB7FCC2F9ED" blockId="17.[151,894,1529,1551]" box="[151,894,1529,1551]" pageId="17" pageNumber="341">
<emphasis id="3D94A102FFF5FFDD30EFDBB7FEA2F9EC" bold="true" box="[151,286,1529,1551]" pageId="17" pageNumber="341">FIGURE 11.</emphasis>
Lateral aspect of holotype of 
<taxonomicName id="C8E00693FFF5FFDD3235DBB7FC95F9ED" box="[589,809,1529,1551]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="341" phylum="Arthropoda" rank="species" species="itsukii" status="sp. nov.">
<emphasis id="3D94A102FFF5FFDD3235DBB7FC95F9ED" box="[589,809,1529,1551]" italics="true" pageId="17" pageNumber="341">Cardiocondyla itsukii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26A71C79FFF5FFDD3357DBB4FCC2F9ED" box="[815,894,1530,1551]" pageId="17" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="47FA2E9BFFF5FFDF30BFD878FD8DF9AD" lastPageId="19" lastPageNumber="343" pageId="17" pageNumber="341" type="description">
<paragraph id="0F5F7D10FFF5FFDE30BFD878FD26FE16" blockId="17.[151,1437,1590,2012]" lastBlockId="18.[151,1437,151,501]" lastPageId="18" lastPageNumber="342" pageId="17" pageNumber="341">
<emphasis id="3D94A102FFF5FFDD30BFD878FDA4F9AD" bold="true" box="[199,536,1590,1615]" pageId="17" pageNumber="341">Description of worker caste.</emphasis>
Worker (
<tableCitation id="426248ABFFF5FFDD32FAD878FD76F9AD" box="[642,714,1590,1615]" captionStart="TABLE 1" captionStartId="18.[151,239,548,571]" captionText="TABLE 1. Morphometric data of workers of C. mauritanica species complex in the sequence arithmetic mean ± standard deviation [minimum, maximum], n = number of measured individuals." pageId="17" pageNumber="341">Tab. 1</tableCitation>
, 
<figureCitation id="97DB6195FFF5FFDD32A0D878FCDCF9AD" box="[728,864,1590,1615]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="16.[151,250,1432,1454]" captionStartId-1="17.[151,250,1529,1551]" captionStartId-2="19.[151,250,1380,1402]" captionTargetBox-0="[173,1413,195,1408]" captionTargetBox-1="[151,1436,553,1508]" captionTargetBox-2="[324,1263,193,1346]" captionTargetId-0="figure@16.[173,1413,195,1408]" captionTargetId-1="figure@17.[151,1436,553,1508]" captionTargetId-2="figure@19.[324,1263,193,1358]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="19" captionText-0="FIGURE 10. Head of holotype of Cardiocondyla itsukii sp. nov. in dorsal aspect." captionText-1="FIGURE 11. Lateral aspect of holotype of Cardiocondyla itsukii sp. nov." captionText-2="FIGURE 12. Waist and gaster of holotype of Cardiocondyla itsukii sp. nov. in dorsal aspect." httpUri-0="https://zenodo.org/record/829093/files/figure.png" httpUri-1="https://zenodo.org/record/829095/files/figure.png" httpUri-2="https://zenodo.org/record/829097/files/figure.png" pageId="17" pageNumber="341">
Figs. 10 
<emphasis id="3D94A102FFF5FFDD334BD877FC83F9AC" bold="true" box="[819,831,1593,1614]" pageId="17" pageNumber="341">–</emphasis>
12
</figureCitation>
): Head less elongated than in all other members of the 
<taxonomicName id="C8E00693FFF5FFDD30BBD815FEA2F991" box="[195,286,1626,1651]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="341" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="3D94A102FFF5FFDD30BBD815FEA2F991" box="[195,286,1626,1651]" italics="true" pageId="17" pageNumber="341">C. nuda</emphasis>
</taxonomicName>
group, CL/CW 1.169. Postocular distance moderately large, PoOc/CL 0.444. Eyes relatively small, EYE 0.228. Frontal carinae immediately caudal of the FRS level parallel or very slightly converging (
<figureCitation id="97DB6195FFF5FFDD3541D830FA33F975" box="[1337,1423,1662,1687]" captionStart="FIGURE 10" captionStartId="16.[151,250,1432,1454]" captionTargetBox="[173,1413,195,1408]" captionTargetId="figure@16.[173,1413,195,1408]" captionTargetPageId="16" captionText="FIGURE 10. Head of holotype of Cardiocondyla itsukii sp. nov. in dorsal aspect." httpUri="https://zenodo.org/record/829093/files/figure.png" pageId="17" pageNumber="341">Fig. 10</figureCitation>
). Foveolae on vertex without interspaces, deeply impressed, with 15 
<emphasis id="3D94A102FFF5FFDD33F7D8EBFC27F958" bold="true" box="[911,923,1701,1722]" pageId="17" pageNumber="341">–</emphasis>
22 µm diameter, and with an inner corona (a flat tubercle) of 7 
<emphasis id="3D94A102FFF5FFDD311FD887FECFF93C" bold="true" box="[359,371,1737,1758]" pageId="17" pageNumber="341">–</emphasis>
9 µm diameter having the base of a decumbent pubescence hair in its center. This 
<typeStatus id="D05BC3B2FFF5FFDD3531D886FAC5F902" box="[1353,1401,1736,1760]" pageId="17" pageNumber="341">type</typeStatus>
of sculpture can also be described as a strongly sculptured microreticulum. Longitudinal sculpture on vertex reduced; only frontal laminae, clypeus, and a narrow area on anteromedian vertex finely longitudinally carinulate; a weak semicircular rugosity is found around the antennal fossae. Lateral mesosoma on whole surface regularly and strongly microreticulate; longitudinal sculpture except for 4 
<emphasis id="3D94A102FFF5FFDD3339D917FCF1F88C" bold="true" box="[833,845,1881,1902]" pageId="17" pageNumber="341">–</emphasis>
6 weak and short carinulae on metapleuron entirely absent (
<figureCitation id="97DB6195FFF5FFDD3096D934FE83F871" box="[238,319,1914,1939]" captionStart="FIGURE 11" captionStartId="17.[151,250,1529,1551]" captionTargetBox="[151,1436,553,1508]" captionTargetId="figure@17.[151,1436,553,1508]" captionTargetPageId="17" captionText="FIGURE 11. Lateral aspect of holotype of Cardiocondyla itsukii sp. nov." httpUri="https://zenodo.org/record/829095/files/figure.png" pageId="17" pageNumber="341">Fig. 11</figureCitation>
); dorsal promesonotum with more irregular reticulum the meshes of which have twice the diameter than on lateral mesosoma. Whole surface of petiole and postpetiole more shining, with a very fine microreticulum. The strength of sculpture on mesosoma and waist in particular shows considerable variation within the huge distributional range, without showing regional trends. Cuticular surface of first gaster tergite smooth and shining, a very delicate microreticulum with wide meshes becomes visible at higher microscopic resolution. Pubescence hair length on gaster tergites is the largest within the 
<taxonomicName id="C8E00693FFF6FFDE32A5DEAEFC87FF1A" box="[733,827,223,248]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="342" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="3D94A102FFF6FFDE32A5DEAEFC87FF1A" box="[733,827,223,248]" italics="true" pageId="18" pageNumber="342">C. nuda</emphasis>
</taxonomicName>
group, PLG/CS 7.12% (
<figureCitation id="97DB6195FFF6FFDE3418DE91FB04FF1A" box="[1120,1208,223,248]" captionStart="FIGURE 12" captionStartId="19.[151,250,1380,1402]" captionTargetBox="[324,1263,193,1346]" captionTargetId="figure@19.[324,1263,193,1358]" captionTargetPageId="19" captionText="FIGURE 12. Waist and gaster of holotype of Cardiocondyla itsukii sp. nov. in dorsal aspect." httpUri="https://zenodo.org/record/829097/files/figure.png" pageId="18" pageNumber="342">Fig. 12</figureCitation>
). Metanotal groove distinct but rather flat, MGr/CS 2.16%. Propodeal spines reduced to short dents. Petiolar profile in many specimens with a steeper frontal face compared to 
<taxonomicName id="C8E00693FFF6FFDE321ADF66FD45FEA2" box="[610,761,296,320]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="342" phylum="Arthropoda" rank="species" species="strigifrons">
<emphasis id="3D94A102FFF6FFDE321ADF66FD45FEA2" box="[610,761,296,320]" italics="true" pageId="18" pageNumber="342">C. strigifrons</emphasis>
</taxonomicName>
and 
<taxonomicName id="C8E00693FFF6FFDE334CDF66FC6CFEA2" box="[820,976,295,320]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="342" phylum="Arthropoda" rank="species" species="kagutsuchi">
<emphasis id="3D94A102FFF6FFDE334CDF66FC6CFEA2" box="[820,976,295,320]" italics="true" pageId="18" pageNumber="342">C. kagutsuchi</emphasis>
</taxonomicName>
. Petiole node slightly longer than wide. Postpetiole narrower than in 
<taxonomicName id="C8E00693FFF6FFDE31A1DF03FD3EFE86" box="[473,642,333,356]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="342" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="3D94A102FFF6FFDE31A1DF03FD3EFE86" box="[473,642,333,356]" italics="true" pageId="18" pageNumber="342">C. mauritanica</emphasis>
</taxonomicName>
; in dorsal view with clearly angulate sides and straight anterior margin that is clearly shorter than posterior margin (
<figureCitation id="97DB6195FFF6FFDE32F0DF21FD67FE6A" box="[648,731,367,392]" captionStart="FIGURE 12" captionStartId="19.[151,250,1380,1402]" captionTargetBox="[324,1263,193,1346]" captionTargetId="figure@19.[324,1263,193,1358]" captionTargetPageId="19" captionText="FIGURE 12. Waist and gaster of holotype of Cardiocondyla itsukii sp. nov. in dorsal aspect." httpUri="https://zenodo.org/record/829097/files/figure.png" pageId="18" pageNumber="342">Fig. 12</figureCitation>
); postpetiolar sternite flat, without any protrusions. Color most variable over the huge distributional range: most frequent are morphs with a medium brown mesosoma and waist, a dark brown head and a dark to blackish brown gaster but much lighter or darker color morphs are not rare. For morphometric data of 
<specimenCount id="19E6B699FFF6FFDE31E8DF92FDA7FE17" box="[400,539,476,501]" pageId="18" pageNumber="342" type="worker">108 workers</specimenCount>
see 
<tableCitation id="426248ABFFF6FFDE3236DF92FD28FE16" box="[590,660,476,501]" captionStart="TABLE 1" captionStartId="18.[151,239,548,571]" captionText="TABLE 1. Morphometric data of workers of C. mauritanica species complex in the sequence arithmetic mean ± standard deviation [minimum, maximum], n = number of measured individuals." pageId="18" pageNumber="342">Tab. 1</tableCitation>
.
</paragraph>
<caption id="5B9F2D98FFF6FFDE30EFDC6AFC36FDB9" pageId="18" pageNumber="342">
<paragraph id="0F5F7D10FFF6FFDE30EFDC6AFC36FDB9" blockId="18.[151,1436,547,603]" pageId="18" pageNumber="342">
<emphasis id="3D94A102FFF6FFDE30EFDC6AFEB5FDD9" bold="true" box="[151,265,548,571]" pageId="18" pageNumber="342">TABLE 1.</emphasis>
Morphometric data of workers of 
<taxonomicName id="C8E00693FFF6FFDE320EDC6DFCAAFDD9" box="[630,790,547,571]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="342" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="3D94A102FFF6FFDE320EDC6DFCAAFDD9" box="[630,790,547,571]" italics="true" pageId="18" pageNumber="342">C. mauritanica</emphasis>
</taxonomicName>
species complex in the sequence arithmetic mean ± standard deviation [minimum, maximum], n = number of measured individuals.
</paragraph>
</caption>
<paragraph id="0F5F7D10FFF6FFDE312EDC38FB5AF8C9" pageId="18" pageNumber="342">
<table id="7DE08FB0FFF6003330E7DC38FAA7F8C9" box="[159,1307,630,1835]" gridcols="5" gridrows="17" pageId="18" pageNumber="342">
<tr id="B1D07F52FFF6003330E7DC38FAA7FD49" box="[159,1307,630,683]" gridrow="0" pageId="18" pageNumber="342" rowspan-0="1">
<th id="F201162EFFF60033312EDC38FE5FFD49" box="[342,483,630,683]" gridcol="1" gridrow="0" pageId="18" pageNumber="342">
<taxonomicName id="C8E00693FFF6FFDE312EDC38FE6AFD6E" box="[342,470,630,652]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="342" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="3D94A102FFF6FFDE312EDC38FE6AFD6E" bold="true" box="[342,470,630,652]" italics="true" pageId="18" pageNumber="342">mauritanica</emphasis>
</taxonomicName>
(n=147)
</th>
<th id="F201162EFFF600333211DC38FD49FD49" box="[617,757,630,683]" gridcol="2" gridrow="0" pageId="18" pageNumber="342">
<taxonomicName id="C8E00693FFF6FFDE3211DC38FD6AFD6E" box="[617,726,630,652]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="342" phylum="Arthropoda" rank="species" species="strigifrons">
<emphasis id="3D94A102FFF6FFDE3211DC38FD6AFD6E" bold="true" box="[617,726,630,652]" italics="true" pageId="18" pageNumber="342">strigifrons</emphasis>
</taxonomicName>
(n=44)
</th>
<th id="F201162EFFF600333304DC38FBB4FD49" box="[892,1032,630,683]" gridcol="3" gridrow="0" pageId="18" pageNumber="342">
<taxonomicName id="C8E00693FFF6FFDE3304DC38FC53FD6E" box="[892,1007,630,652]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="342" phylum="Arthropoda" rank="species" species="kagutsuchi">
<emphasis id="3D94A102FFF6FFDE3304DC38FC53FD6E" bold="true" box="[892,1007,630,652]" italics="true" pageId="18" pageNumber="342">kagutsuchi</emphasis>
</taxonomicName>
(n=97)
</th>
<th id="F201162EFFF6003334F7DC38FAA7FD49" box="[1167,1307,630,683]" gridcol="4" gridrow="0" pageId="18" pageNumber="342">
<taxonomicName id="C8E00693FFF6FFDE34F7DC38FB73FD6E" box="[1167,1231,630,652]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="342" phylum="Arthropoda" rank="species" species="itsukii">
<emphasis id="3D94A102FFF6FFDE34F7DC38FB73FD6E" bold="true" box="[1167,1231,630,652]" italics="true" pageId="18" pageNumber="342">itsukii</emphasis>
</taxonomicName>
(n=108)
</th>
</tr>
<tr id="B1D07F52FFF6003330E7DCF0FAA7FD11" box="[159,1307,702,755]" gridrow="1" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DCF0FEA4FD11" box="[159,280,702,755]" gridcol="0" gridrow="1" pageId="18" pageNumber="342">CS [µm]</th>
<td id="F201162EFFF60033312EDCF0FE5FFD11" box="[342,483,702,755]" gridcol="1" gridrow="1" pageId="18" pageNumber="342">514 ± 21 [460,568]</td>
<td id="F201162EFFF600333211DCF0FD49FD11" box="[617,757,702,755]" gridcol="2" gridrow="1" pageId="18" pageNumber="342">530 ± 28 [475,591]</td>
<td id="F201162EFFF600333304DCF0FBB4FD11" box="[892,1032,702,755]" gridcol="3" gridrow="1" pageId="18" pageNumber="342">516 ± 24 [455,565]</td>
<td id="F201162EFFF6003334F7DCF0FAA7FD11" box="[1167,1307,702,755]" gridcol="4" gridrow="1" pageId="18" pageNumber="342">531 ± 20 [477,574]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DD48FAA7FCD9" box="[159,1307,774,827]" gridrow="2" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DD48FEA4FCD9" box="[159,280,774,827]" gridcol="0" gridrow="2" pageId="18" pageNumber="342">CL/CW</th>
<td id="F201162EFFF60033312EDD48FE5FFCD9" box="[342,483,774,827]" gridcol="1" gridrow="2" pageId="18" pageNumber="342">1.182 ± 0.020 [1.126,1.224]</td>
<td id="F201162EFFF600333211DD48FD49FCD9" box="[617,757,774,827]" gridcol="2" gridrow="2" pageId="18" pageNumber="342">1.202 ± 0.024 [1.159,1.256]</td>
<td id="F201162EFFF600333304DD48FBB4FCD9" box="[892,1032,774,827]" gridcol="3" gridrow="2" pageId="18" pageNumber="342">1.198 ± 0.024 [1.144,1.263]</td>
<td id="F201162EFFF6003334F7DD48FAA7FCD9" box="[1167,1307,774,827]" gridcol="4" gridrow="2" pageId="18" pageNumber="342">1.169 ± 0.018 [1.125,1.215]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DD00FAA7FC61" box="[159,1307,846,899]" gridrow="3" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DD00FEA4FC61" box="[159,280,846,899]" gridcol="0" gridrow="3" pageId="18" pageNumber="342">SL/CS</th>
<td id="F201162EFFF60033312EDD00FE5FFC61" box="[342,483,846,899]" gridcol="1" gridrow="3" pageId="18" pageNumber="342">0.813 ± 0.012 [0.787,0.849]</td>
<td id="F201162EFFF600333211DD00FD49FC61" box="[617,757,846,899]" gridcol="2" gridrow="3" pageId="18" pageNumber="342">0.840 ± 0.013 [0.816,0.872]</td>
<td id="F201162EFFF600333304DD00FBB4FC61" box="[892,1032,846,899]" gridcol="3" gridrow="3" pageId="18" pageNumber="342">0.833 ± 0.013 [0.802,0.862]</td>
<td id="F201162EFFF6003334F7DD00FAA7FC61" box="[1167,1307,846,899]" gridcol="4" gridrow="3" pageId="18" pageNumber="342">0.826 ± 0.012 [0.801,0.856]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DDD8FAA7FC29" box="[159,1307,918,971]" gridrow="4" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DDD8FEA4FC29" box="[159,280,918,971]" gridcol="0" gridrow="4" pageId="18" pageNumber="342">PoOc/CL</th>
<td id="F201162EFFF60033312EDDD8FE5FFC29" box="[342,483,918,971]" gridcol="1" gridrow="4" pageId="18" pageNumber="342">0.447 ± 0.008 [0.426,0.467]</td>
<td id="F201162EFFF600333211DDD8FD49FC29" box="[617,757,918,971]" gridcol="2" gridrow="4" pageId="18" pageNumber="342">0.446 ± 0.007 [0.435,0.464]</td>
<td id="F201162EFFF600333304DDD8FBB4FC29" box="[892,1032,918,971]" gridcol="3" gridrow="4" pageId="18" pageNumber="342">0.457 ± 0.009 [0.433,0.484]</td>
<td id="F201162EFFF6003334F7DDD8FAA7FC29" box="[1167,1307,918,971]" gridcol="4" gridrow="4" pageId="18" pageNumber="342">0.444 ± 0.008 [0.428,0.469]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DD90FAA7FBF1" box="[159,1307,990,1043]" gridrow="5" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DD90FEA4FBF1" box="[159,280,990,1043]" gridcol="0" gridrow="5" pageId="18" pageNumber="342">EYE</th>
<td id="F201162EFFF60033312EDD90FE5FFBF1" box="[342,483,990,1043]" gridcol="1" gridrow="5" pageId="18" pageNumber="342">0.232 ± 0.005 [0.222,0.246]</td>
<td id="F201162EFFF600333211DD90FD49FBF1" box="[617,757,990,1043]" gridcol="2" gridrow="5" pageId="18" pageNumber="342">0.234 ± 0.006 [0.219,0.245]</td>
<td id="F201162EFFF600333304DD90FBB4FBF1" box="[892,1032,990,1043]" gridcol="3" gridrow="5" pageId="18" pageNumber="342">0.231 ± 0.005 [0.218,0.245]</td>
<td id="F201162EFFF6003334F7DD90FAA7FBF1" box="[1167,1307,990,1043]" gridcol="4" gridrow="5" pageId="18" pageNumber="342">0.228 ± 0.005 [0.218,0.239]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DA68FAA7FBB9" box="[159,1307,1062,1115]" gridrow="6" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DA68FEA4FBB9" box="[159,280,1062,1115]" gridcol="0" gridrow="6" pageId="18" pageNumber="342">dFOV [µm]</th>
<td id="F201162EFFF60033312EDA68FE5FFBB9" box="[342,483,1062,1115]" gridcol="1" gridrow="6" pageId="18" pageNumber="342">17.7 ± 1.2 [15.0,20.0]</td>
<td id="F201162EFFF600333211DA68FD49FBB9" box="[617,757,1062,1115]" gridcol="2" gridrow="6" pageId="18" pageNumber="342">17.7 ± 1.1 [15.7,20.5]</td>
<td id="F201162EFFF600333304DA68FBB4FBB9" box="[892,1032,1062,1115]" gridcol="3" gridrow="6" pageId="18" pageNumber="342">16.7 ± 1.4 [14.3,21.0]</td>
<td id="F201162EFFF6003334F7DA68FAA7FBB9" box="[1167,1307,1062,1115]" gridcol="4" gridrow="6" pageId="18" pageNumber="342">18.1 ± 1.6 [15.0,21.7]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DA20FAA7FB41" box="[159,1307,1134,1187]" gridrow="7" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DA20FEA4FB41" box="[159,280,1134,1187]" gridcol="0" gridrow="7" pageId="18" pageNumber="342">FRS/CS</th>
<td id="F201162EFFF60033312EDA20FE5FFB41" box="[342,483,1134,1187]" gridcol="1" gridrow="7" pageId="18" pageNumber="342">0.265 ± 0.007 [0.248,0.286]</td>
<td id="F201162EFFF600333211DA20FD49FB41" box="[617,757,1134,1187]" gridcol="2" gridrow="7" pageId="18" pageNumber="342">0.265 ± 0.007 [0.252,0.285]</td>
<td id="F201162EFFF600333304DA20FBB4FB41" box="[892,1032,1134,1187]" gridcol="3" gridrow="7" pageId="18" pageNumber="342">0.261 ± 0.007 [0.246,0.273]</td>
<td id="F201162EFFF6003334F7DA20FAA7FB41" box="[1167,1307,1134,1187]" gridcol="4" gridrow="7" pageId="18" pageNumber="342">0.270 ± 0.009 [0.250,0.290]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DAF8FAA7FB09" box="[159,1307,1206,1259]" gridrow="8" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DAF8FEA4FB09" box="[159,280,1206,1259]" gridcol="0" gridrow="8" pageId="18" pageNumber="342">SPBA/CS</th>
<td id="F201162EFFF60033312EDAF8FE5FFB09" box="[342,483,1206,1259]" gridcol="1" gridrow="8" pageId="18" pageNumber="342">0.268± 0.010 [0.239,0.296]</td>
<td id="F201162EFFF600333211DAF8FD49FB09" box="[617,757,1206,1259]" gridcol="2" gridrow="8" pageId="18" pageNumber="342">0.252 ± 0.011 [0.230,0.280]</td>
<td id="F201162EFFF600333304DAF8FBB4FB09" box="[892,1032,1206,1259]" gridcol="3" gridrow="8" pageId="18" pageNumber="342">0.261 ± 0.012 [0.227,0.301]</td>
<td id="F201162EFFF6003334F7DAF8FAA7FB09" box="[1167,1307,1206,1259]" gridcol="4" gridrow="8" pageId="18" pageNumber="342">0.257 ± 0.013 [0.208,0.283]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DAB0FAA7FAD1" box="[159,1307,1278,1331]" gridrow="9" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DAB0FEA4FAD1" box="[159,280,1278,1331]" gridcol="0" gridrow="9" pageId="18" pageNumber="342">SP/CS</th>
<td id="F201162EFFF60033312EDAB0FE5FFAD1" box="[342,483,1278,1331]" gridcol="1" gridrow="9" pageId="18" pageNumber="342">0.090 ± 0.013 [0.047,0.119]</td>
<td id="F201162EFFF600333211DAB0FD49FAD1" box="[617,757,1278,1331]" gridcol="2" gridrow="9" pageId="18" pageNumber="342">0.061 ± 0.009 [0.042,0.079]</td>
<td id="F201162EFFF600333304DAB0FBB4FAD1" box="[892,1032,1278,1331]" gridcol="3" gridrow="9" pageId="18" pageNumber="342">0.075 ± 0.011 [0.050,0.102]</td>
<td id="F201162EFFF6003334F7DAB0FAA7FAD1" box="[1167,1307,1278,1331]" gridcol="4" gridrow="9" pageId="18" pageNumber="342">0.072 ± 0.011 [0.042,0.096]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DB08FAA7FA99" box="[159,1307,1350,1403]" gridrow="10" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DB08FEA4FA99" box="[159,280,1350,1403]" gridcol="0" gridrow="10" pageId="18" pageNumber="342">PEW/CS</th>
<td id="F201162EFFF60033312EDB08FE5FFA99" box="[342,483,1350,1403]" gridcol="1" gridrow="10" pageId="18" pageNumber="342">0.265 ± 0.013 [0.233,0.298]</td>
<td id="F201162EFFF600333211DB08FD49FA99" box="[617,757,1350,1403]" gridcol="2" gridrow="10" pageId="18" pageNumber="342">0.278 ± 0.012 [0.260,0.318]</td>
<td id="F201162EFFF600333304DB08FBB4FA99" box="[892,1032,1350,1403]" gridcol="3" gridrow="10" pageId="18" pageNumber="342">0.268 ± 0.013 [0.236,0.335]</td>
<td id="F201162EFFF6003334F7DB08FAA7FA99" box="[1167,1307,1350,1403]" gridcol="4" gridrow="10" pageId="18" pageNumber="342">0.271 ± 0.011 [0.247,0.301]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DBC0FAA7FA21" box="[159,1307,1422,1475]" gridrow="11" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DBC0FEA4FA21" box="[159,280,1422,1475]" gridcol="0" gridrow="11" pageId="18" pageNumber="342">PPW/CS</th>
<td id="F201162EFFF60033312EDBC0FE5FFA21" box="[342,483,1422,1475]" gridcol="1" gridrow="11" pageId="18" pageNumber="342">0.485 ± 0.013 [0.449,0.533]</td>
<td id="F201162EFFF600333211DBC0FD49FA21" box="[617,757,1422,1475]" gridcol="2" gridrow="11" pageId="18" pageNumber="342">0.475 ± 0.021 [0.442,0.527]</td>
<td id="F201162EFFF600333304DBC0FBB4FA21" box="[892,1032,1422,1475]" gridcol="3" gridrow="11" pageId="18" pageNumber="342">0.466 ± 0.020 [0.431,0.524]</td>
<td id="F201162EFFF6003334F7DBC0FAA7FA21" box="[1167,1307,1422,1475]" gridcol="4" gridrow="11" pageId="18" pageNumber="342">0.455 ± 0.012 [0.430,0.487]</td>
</tr>
<tr id="B1D07F52FFF6003330E7DB98FAA7F9E9" box="[159,1307,1494,1547]" gridrow="12" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7DB98FEA4F9E9" box="[159,280,1494,1547]" gridcol="0" gridrow="12" pageId="18" pageNumber="342">PEH/CS</th>
<td id="F201162EFFF60033312EDB98FE5FF9E9" box="[342,483,1494,1547]" gridcol="1" gridrow="12" pageId="18" pageNumber="342">0.329 ± 0.009 [0.309,0.350]</td>
<td id="F201162EFFF600333211DB98FD49F9E9" box="[617,757,1494,1547]" gridcol="2" gridrow="12" pageId="18" pageNumber="342">0.313 ± 0.009 [0.293,0.336]</td>
<td id="F201162EFFF600333304DB98FBB4F9E9" box="[892,1032,1494,1547]" gridcol="3" gridrow="12" pageId="18" pageNumber="342">0.311 ± 0.013 [0.283,0.367]</td>
<td id="F201162EFFF6003334F7DB98FAA7F9E9" box="[1167,1307,1494,1547]" gridcol="4" gridrow="12" pageId="18" pageNumber="342">0.314 ± 0.009 [0.296,0.338]</td>
</tr>
<tr id="B1D07F52FFF6003330E7D850FAA7F9B1" box="[159,1307,1566,1619]" gridrow="13" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7D850FEA4F9B1" box="[159,280,1566,1619]" gridcol="0" gridrow="13" pageId="18" pageNumber="342">PPH/CS</th>
<td id="F201162EFFF60033312ED850FE5FF9B1" box="[342,483,1566,1619]" gridcol="1" gridrow="13" pageId="18" pageNumber="342">0.287 ± 0.008 [0.270,0.318]</td>
<td id="F201162EFFF600333211D850FD49F9B1" box="[617,757,1566,1619]" gridcol="2" gridrow="13" pageId="18" pageNumber="342">0.285 ± 0.010 [0.257,0.303]</td>
<td id="F201162EFFF600333304D850FBB4F9B1" box="[892,1032,1566,1619]" gridcol="3" gridrow="13" pageId="18" pageNumber="342">0.280 ± 0.013 [0.250,0.326]</td>
<td id="F201162EFFF6003334F7D850FAA7F9B1" box="[1167,1307,1566,1619]" gridcol="4" gridrow="13" pageId="18" pageNumber="342">0.282 ± 0.009 [0.258,0.307]</td>
</tr>
<tr id="B1D07F52FFF6003330E7D828FAA7F979" box="[159,1307,1638,1691]" gridrow="14" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7D828FEA4F979" box="[159,280,1638,1691]" gridcol="0" gridrow="14" pageId="18" pageNumber="342">sqrtPDG</th>
<td id="F201162EFFF60033312ED828FE5FF979" box="[342,483,1638,1691]" gridcol="1" gridrow="14" pageId="18" pageNumber="342">3.72 ± 0.28 [3.15,4.64]</td>
<td id="F201162EFFF600333211D828FD49F979" box="[617,757,1638,1691]" gridcol="2" gridrow="14" pageId="18" pageNumber="342">3.44 ± 0.16 [3.13,4.03]</td>
<td id="F201162EFFF600333304D828FBB4F979" box="[892,1032,1638,1691]" gridcol="3" gridrow="14" pageId="18" pageNumber="342">3.53 ± 0.19 [3.11,4.00]</td>
<td id="F201162EFFF6003334F7D828FAA7F979" box="[1167,1307,1638,1691]" gridcol="4" gridrow="14" pageId="18" pageNumber="342">3.54 ± 0.20 [3.10,4.15]</td>
</tr>
<tr id="B1D07F52FFF6003330E7D8E0FAA7F901" box="[159,1307,1710,1763]" gridrow="15" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7D8E0FEA4F901" box="[159,280,1710,1763]" gridcol="0" gridrow="15" pageId="18" pageNumber="342">PLG/CS [%]</th>
<td id="F201162EFFF60033312ED8E0FE5FF901" box="[342,483,1710,1763]" gridcol="1" gridrow="15" pageId="18" pageNumber="342">6.38 ± 0.41 [5.43,7.73]</td>
<td id="F201162EFFF600333211D8E0FD49F901" box="[617,757,1710,1763]" gridcol="2" gridrow="15" pageId="18" pageNumber="342">6.76 ± 0.42 [5.90,7.54]</td>
<td id="F201162EFFF600333304D8E0FBB4F901" box="[892,1032,1710,1763]" gridcol="3" gridrow="15" pageId="18" pageNumber="342">6.72 ± 0.37 [5.51,7.91]</td>
<td id="F201162EFFF6003334F7D8E0FAA7F901" box="[1167,1307,1710,1763]" gridcol="4" gridrow="15" pageId="18" pageNumber="342">7.12 ± 0.36 [6.02,7.98]</td>
</tr>
<tr id="B1D07F52FFF6003330E7D8B8FAA7F8C9" box="[159,1307,1782,1835]" gridrow="16" pageId="18" pageNumber="342">
<th id="F201162EFFF6003330E7D8B8FEA4F8C9" box="[159,280,1782,1835]" gridcol="0" gridrow="16" pageId="18" pageNumber="342">MGr/CS [%]</th>
<td id="F201162EFFF60033312ED8B8FE5FF8C9" box="[342,483,1782,1835]" gridcol="1" gridrow="16" pageId="18" pageNumber="342">2.14 ± 0.52 [1.1,3.5]</td>
<td id="F201162EFFF600333211D8B8FD49F8C9" box="[617,757,1782,1835]" gridcol="2" gridrow="16" pageId="18" pageNumber="342">1.71 ± 0.46 [0.8,2.7]</td>
<td id="F201162EFFF600333304D8B8FBB4F8C9" box="[892,1032,1782,1835]" gridcol="3" gridrow="16" pageId="18" pageNumber="342">1.84 ± 0.51 [0.8,3.8]</td>
<td id="F201162EFFF6003334F7D8B8FAA7F8C9" box="[1167,1307,1782,1835]" gridcol="4" gridrow="16" pageId="18" pageNumber="342">2.16 ± 0.54 [0.8,3.3]</td>
</tr>
</table>
</paragraph>
<caption id="5B9F2D98FFF7FFDF30EFDB2AFBFEFA98" box="[151,1090,1380,1402]" httpUri="https://zenodo.org/record/829097/files/figure.png" pageId="19" pageNumber="343" targetBox="[324,1263,193,1346]" targetPageId="19">
<paragraph id="0F5F7D10FFF7FFDF30EFDB2AFBFEFA98" blockId="19.[151,1090,1380,1402]" box="[151,1090,1380,1402]" pageId="19" pageNumber="343">
<emphasis id="3D94A102FFF7FFDF30EFDB2AFEA3FA9B" bold="true" box="[151,287,1380,1402]" pageId="19" pageNumber="343">FIGURE 12.</emphasis>
Waist and gaster of holotype of 
<taxonomicName id="C8E00693FFF7FFDF3210DB2AFCFFFA98" box="[616,835,1380,1402]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="343" phylum="Arthropoda" rank="species" species="itsukii" status="sp. nov.">
<emphasis id="3D94A102FFF7FFDF3210DB2AFCFFFA98" box="[616,835,1380,1402]" italics="true" pageId="19" pageNumber="343">Cardiocondyla itsukii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="26A71C79FFF7FFDF3331DB2BFC24FA98" box="[841,920,1381,1402]" pageId="19" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
in dorsal aspect.
</paragraph>
</caption>
<paragraph id="0F5F7D10FFF7FFDF30BFDBE8FD8DF9AD" blockId="19.[151,1436,1446,2011]" pageId="19" pageNumber="343">
<materialsCitation id="BF88774DFFF7FFDF30BFDBE8FDCFFA01" ID-GBIF-Occurrence="1572325900" collectorName="over East India" country="Iceland" location="East India" pageId="19" pageNumber="343" specimenCount="1" stateProvince="Reunion">
<emphasis id="3D94A102FFF7FFDF30BFDBE8FE1BFA5D" bold="true" box="[199,423,1446,1471]" pageId="19" pageNumber="343">Geographic range.</emphasis>
<taxonomicName id="C8E00693FFF7FFDF31C8DBE9FDABFA5D" box="[432,535,1446,1471]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="343" phylum="Arthropoda" rank="species" species="itsukii">
<emphasis id="3D94A102FFF7FFDF31C8DBE9FDABFA5D" box="[432,535,1446,1471]" italics="true" pageId="19" pageNumber="343">C. itsukii</emphasis>
</taxonomicName>
shows a huge range extending over 
<quantity id="C818D0F5FFF7FFDF33C7DBE9FB8EFA5D" box="[959,1074,1446,1471]" metricMagnitude="7" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="343" unit="km" value="17000.0">17000 km</quantity>
from the 
<collectingCountry id="77F73D80FFF7FFDF34DEDBE8FB56FA5D" box="[1190,1258,1446,1471]" name="Iceland" pageId="19" pageNumber="343">Island</collectingCountry>
<collectingRegion id="CD24B3F2FFF7FFDF348BDBE8FAEEFA5D" box="[1267,1362,1446,1471]" country="France" name="Reunion" pageId="19" pageNumber="343">Reunion</collectingRegion>
in the Indian Ocean, 
<collectorName id="A21518C6FFF7FFDF3138DB85FE4EFA01" box="[320,498,1482,1507]" pageId="19" pageNumber="343">
over 
<location id="0A3F2BCBFFF7FFDF3104DB85FE4EFA01" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF7FFDF3104DB85FE4EFA01" box="[380,498,1482,1507]" country="Iceland" name="East India" pageId="19" pageNumber="343" stateProvince="Reunion">East India</location>
</collectorName>
, Indochina
</materialsCitation>
, 
<materialsCitation id="BF88774DFFF7FFDF32F8DB85FB13FA01" ID-GBIF-Occurrence="1572325910" box="[640,1199,1482,1507]" country="Japan" location="Pacific Islands" pageId="19" pageNumber="343" specimenCount="1" stateProvince="Hawaii">
<collectingCountry id="77F73D80FFF7FFDF32F8DB85FD7CFA00" box="[640,704,1483,1506]" name="Japan" pageId="19" pageNumber="343">Japan</collectingCountry>
and diverse 
<location id="0A3F2BCBFFF7FFDF332EDB84FC41FA01" LSID="urn:lsid:plazi:treatment:8749CC06FFEBFFD830EFD879FA32FF37:0A3F2BCBFFF7FFDF332EDB84FC41FA01" box="[854,1021,1482,1507]" country="Japan" name="Pacific Islands" pageId="19" pageNumber="343" stateProvince="Hawaii">Pacific Islands</location>
east to 
<collectingRegion id="CD24B3F2FFF7FFDF3423DB84FB13FA01" box="[1115,1199,1482,1507]" country="United States of America" name="Hawaii" pageId="19" pageNumber="343">Hawaii</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="BF88774DFFF7FFDF34C5DB84FD8DF9AD" ID-GBIF-Occurrence="1572325930" country="Island Reunion" location="Hawaii" pageId="19" pageNumber="343" specimenCount="1" stateProvince="Hawaii">
The most marginal, isolated populations on 
<collectingCountry id="77F73D80FFF7FFDF31C2DBA0FDD1F9E5" box="[442,621,1518,1543]" isEnumeration="true" pageId="19" pageNumber="343">Island Reunion</collectingCountry>
and 
<collectingRegion id="CD24B3F2FFF7FFDF32CBDBA0FCB8F9E5" box="[691,772,1518,1543]" country="United States of America" name="Hawaii" pageId="19" pageNumber="343">Hawaii</collectingRegion>
most probably have been founded by anthropogenous introduction. Despite showing tramp species properties 
<taxonomicName id="C8E00693FFF7FFDF337DD85DFCD6F9C9" box="[773,874,1554,1579]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="343" phylum="Arthropoda" rank="species" species="itsukii">
<emphasis id="3D94A102FFF7FFDF337DD85DFCD6F9C9" box="[773,874,1554,1579]" italics="true" pageId="19" pageNumber="343">C. itsukii</emphasis>
</taxonomicName>
is rare in the Indo-Malayan Archipelago where 
<taxonomicName id="C8E00693FFF7FFDF35FAD85DFEACF9AD" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="343" phylum="Arthropoda" rank="species" species="kagutsuchi">
<emphasis id="3D94A102FFF7FFDF35FAD85DFEACF9AD" italics="true" pageId="19" pageNumber="343">C. kagutsuchi</emphasis>
</taxonomicName>
and 
<taxonomicName id="C8E00693FFF7FFDF313ED879FE06F9AD" box="[326,442,1591,1615]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="343" phylum="Arthropoda" rank="species" species="strigifrons">
<emphasis id="3D94A102FFF7FFDF313ED879FE06F9AD" box="[326,442,1591,1615]" italics="true" pageId="19" pageNumber="343">strigifrons</emphasis>
</taxonomicName>
dominate.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="47FA2E9BFFF7FFDF30BFD814FE1AF991" box="[199,422,1626,1651]" pageId="19" pageNumber="343" type="diagnosis">
<paragraph id="0F5F7D10FFF7FFDF30BFD814FE1AF991" blockId="19.[151,1436,1446,2011]" box="[199,422,1626,1651]" pageId="19" pageNumber="343">
<emphasis id="3D94A102FFF7FFDF30BFD814FEFDF991" bold="true" box="[199,321,1626,1651]" pageId="19" pageNumber="343">Diagnosis.</emphasis>
See key.
</paragraph>
</subSubSection>
<subSubSection id="47FA2E9BFFF7FFD830BFD830FA32FF37" lastPageId="20" lastPageNumber="344" pageId="19" pageNumber="343" type="biology_ecology">
<paragraph id="0F5F7D10FFF7FFD830BFD830FA32FF37" blockId="19.[151,1436,1446,2011]" lastBlockId="20.[151,1436,151,213]" lastPageId="20" lastPageNumber="344" pageId="19" pageNumber="343">
<emphasis id="3D94A102FFF7FFDF30BFD830FE9BF975" bold="true" box="[199,295,1662,1687]" pageId="19" pageNumber="343">Biology.</emphasis>
On 
<collectingRegion id="CD24B3F2FFF7FFDF3126D830FE13F975" box="[350,431,1662,1687]" country="United States of America" name="Hawaii" pageId="19" pageNumber="343">Hawaii</collectingRegion>
it is one of the very few ants occurring in high densities in primary rain forests with 
<taxonomicName id="C8E00693FFF7FFDF30EFD8ECFE96F959" box="[151,298,1698,1723]" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="343" phylum="Tracheophyta" rank="genus">
<emphasis id="3D94A102FFF7FFDF30EFD8ECFE96F959" box="[151,298,1698,1723]" italics="true" pageId="19" pageNumber="343">Metrosideros</emphasis>
</taxonomicName>
sp. trees and 
<taxonomicName id="C8E00693FFF7FFDF319BD8ECFDF7F959" box="[483,587,1698,1723]" class="Polypodiopsida" family="Cibotiaceae" genus="Cibotium" kingdom="Plantae" order="Cyatheales" pageId="19" pageNumber="343" phylum="Tracheophyta" rank="genus">
<emphasis id="3D94A102FFF7FFDF319BD8ECFDF7F959" box="[483,587,1698,1723]" italics="true" pageId="19" pageNumber="343">Cibotium</emphasis>
</taxonomicName>
ferns and is found also at higher elevations on Mauna Kea volcano (
<bibRefCitation id="6B7100E1FFF7FFDF30E6D888FE7BF93D" author="Krushelnycky" box="[158,455,1734,1759]" pageId="19" pageNumber="343" refString="Krushelnycky, P., Loope, L. L. &amp; Reimer, N. J. (2005) The ecology, policy, and management of ants in Hawaii. Proceedings of the Hawaiian Entomological Society, 37, 1 - 25." type="journal article" year="2005">
Krushelnycky 
<emphasis id="3D94A102FFF7FFDF313DD886FE39F93D" box="[325,389,1734,1759]" italics="true" pageId="19" pageNumber="343">et. al.</emphasis>
2005
</bibRefCitation>
, 
<bibRefCitation id="6B7100E1FFF7FFDF31ABD886FD03F93D" author="Wetterer" box="[467,703,1734,1759]" pageId="19" pageNumber="343" refString="Wetterer, J. K., Banko, P. C., Laniawe, L. P., Slotterback, J. W. &amp; Brenner, G. J. (1998) Nonindigenous ants at high elevations on Mauna Kea, Hawai'i. Pacific Science, 52, 228 - 236." type="journal article" year="1998">
Wetterer 
<emphasis id="3D94A102FFF7FFDF3243D886FDC7F93D" box="[571,635,1734,1759]" italics="true" pageId="19" pageNumber="343">et. al.</emphasis>
1998
</bibRefCitation>
). Biological traits of 
<taxonomicName id="C8E00693FFF7FFDF33D6D889FBAFF93D" box="[942,1043,1734,1759]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="343" phylum="Arthropoda" rank="species" species="itsukii">
<emphasis id="3D94A102FFF7FFDF33D6D889FBAFF93D" box="[942,1043,1734,1759]" italics="true" pageId="19" pageNumber="343">C. itsukii</emphasis>
</taxonomicName>
were studied by 
<bibRefCitation id="6B7100E1FFF7FFDF34AED888FE80F8E1" author="Frohschammer" pageId="19" pageNumber="343" refString="Frohschammer, S. &amp; Heinze, J. (2009) A heritable component in sex ratio and caste determination in a Cardiocondyla ant. Frontiers in Zoology, 6, 27." type="journal article" year="2009">Frohschammer &amp; Heinze (2009)</bibRefCitation>
, 
<bibRefCitation id="6B7100E1FFF7FFDF3131D8A4FD89F8E1" box="[329,565,1770,1795]" pageId="19" pageNumber="343" refString="Heinze, J., Frohschammer, S. &amp; Bernadou, A. (2013) Queen life span and total reproductive success are positively associated in the ant Cardiocondyla cf. kagutsuchi. Behavioral Ecology and Sociobiology, 67, 1555 - 1562. https: // doi. org / 10.1007 / s 00265 - 013 - 1567 - 9" type="journal article">
Heinze 
<emphasis id="3D94A102FFF7FFDF31D9D8A2FE5EF8E1" box="[417,482,1770,1795]" italics="true" pageId="19" pageNumber="343">et. al.</emphasis>
(2013)
</bibRefCitation>
and 
<bibRefCitation id="6B7100E1FFF7FFDF3217D8A4FCF1F8E1" box="[623,845,1770,1795]" pageId="19" pageNumber="343" refString="Okita, I., Terayama, M. &amp; Tsuchida, K. (2015) Cryptic lineages in the Cardiocondyla sl. kagutsuchi Terayama (Hymenoptera: Formicidae) discovered by phylogenetic and morphological approaches. Sociobiology, 62 (3), 401 - 411. https: // doi. org / 10.13102 / sociobiology. v 62 i 3.805" type="journal article">
Okita 
<emphasis id="3D94A102FFF7FFDF32CED8A2FD4BF8E1" box="[694,759,1770,1795]" italics="true" pageId="19" pageNumber="343">et. al.</emphasis>
(2015)
</bibRefCitation>
. Winged males are not known in 
<taxonomicName id="C8E00693FFF7FFDF34B6D8A5FA89F8E1" box="[1230,1333,1770,1795]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="343" phylum="Arthropoda" rank="species" species="itsukii">
<emphasis id="3D94A102FFF7FFDF34B6D8A5FA89F8E1" box="[1230,1333,1770,1795]" italics="true" pageId="19" pageNumber="343">C. itsukii</emphasis>
</taxonomicName>
but only wingless ergatoid males with more than one male usually being present in a colony. The males are in reproductive competition and use the strong shear-shaped mandibles to attack freshly eclosed rivals by crushing their heads or cutting-off appendages but males do not fight when in adult stage. As in 
<taxonomicName id="C8E00693FFF7FFDF3396D919FB27F88D" box="[1006,1179,1879,1903]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="343" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="3D94A102FFF7FFDF3396D919FB27F88D" box="[1006,1179,1879,1903]" italics="true" pageId="19" pageNumber="343">C. mauritanica</emphasis>
</taxonomicName>
, the strictly intranidal mating and polygyny is considered as a preadaptation for a successful tramp species strategy. The total number of sexual offspring of queens is positively associated with their life-span. More fecund queens live longer than less fecund queens and early onset of sexual production does not negatively affect the queen’s life-span. The number of eggs present in colonies increased with queen’s age until shortly before death, indicating negligible reproductive senescence. Several queens produced only males late in their lives, suggesting the occurrence of sperm depletion.
</paragraph>
</subSubSection>
</treatment>
</document>