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<document id="F6FD5F37A27C2F33A780C4926271FD13" ID-DOI="10.5281/zenodo.184612" ID-GBIF-Dataset="e30cedee-23dc-4989-835c-fee3db1a8d43" ID-ISSN="1175-5326" ID-Zenodo-Dep="184612" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459992082753" checkinUser="plazi" docAuthor="Bravo, Freddy, Chagas, Cinthia &amp; Alves, Veracilda Ribeiro" docDate="2008" docId="7C6D4423FF91FF8DFF0AFABCFAD8FE09" docLanguage="en" docName="zt01916p068.pdf" docOrigin="Zootaxa 1916" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tonnoira robusta Bravo, Alves &amp; Chagas, sp. nov." docType="treatment" docVersion="9" lastPageNumber="67" masterDocId="80543C5BFF93FF89FF9DFFD7FF96FFAB" masterDocTitle="Description of two new species of Tonnoira Enderlein from caves in the Brazilian Amazon and comments about the taxonomic status of Tonnoira plumaria Quate (Diptera, Psychodidae, Psychodinae)" masterLastPageNumber="68" masterPageNumber="63" pageNumber="65" updateTime="1698232670637" updateUser="plazi">
<mods:mods id="015D78C656E256A41CE39604E696A367" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="657AB7319EB893D6C6D0D864F7C8EBA6">
<mods:title id="6F8FAF615C6FCF6804479A75DC1DCEE6">Description of two new species of Tonnoira Enderlein from caves in the Brazilian Amazon and comments about the taxonomic status of Tonnoira plumaria Quate (Diptera, Psychodidae, Psychodinae)</mods:title>
</mods:titleInfo>
<mods:name id="86C450E7E622AF250924DD4FD86A8925" type="personal">
<mods:role id="417DE0A9DA17CF13D055B86F1D1FB742">
<mods:roleTerm id="3299734E268114F4E4A7DAC9811AD91F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C03A951057B7031E80621D9DE63A48D8">Bravo, Freddy</mods:namePart>
</mods:name>
<mods:name id="0DEB2CA1BB43E9C9912501EC12BC1662" type="personal">
<mods:role id="A54FCF62897CD839A2CF5DCC61A71D04">
<mods:roleTerm id="1197AD16128D395F915D330DEBD9167D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="137CF1BDDF1819DD2702A4D8B27865CE">Chagas, Cinthia</mods:namePart>
</mods:name>
<mods:name id="6C71FC5CCFD72F8878EB4649E2D4D1E7" type="personal">
<mods:role id="814C11CC8AF0DCC5EA75882283056EC8">
<mods:roleTerm id="3DA042D7F0E394783BC0E020B5CE81C8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B27D1548E6333547EDE3608B4B8E440A">Alves, Veracilda Ribeiro</mods:namePart>
</mods:name>
<mods:typeOfResource id="E4551FA47A3E2D739437DB0F85470467">text</mods:typeOfResource>
<mods:relatedItem id="1D3D2FBE56EE7A925F7A55F1C611504C" type="host">
<mods:titleInfo id="E1CBC45E7025E2DB4CB58F306E6C793F">
<mods:title id="583C9A981D1417D6E731852F44560331">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8AACCFA920E8CCA779E380E09A1A45B0">
<mods:date id="6E91A065AB732C45E221E9A7DD2DD0AB">2008</mods:date>
<mods:detail id="04B61F8405BC863EF2783F61559BF35E" type="volume">
<mods:number id="366F546F71A37B017A02F346A5DE1A8A">1916</mods:number>
</mods:detail>
<mods:extent id="4DC31454BF016AAF91997C8C0A0E992A" unit="page">
<mods:start id="07B98F8F3BE96BEFC5B1F2146CD2D528">63</mods:start>
<mods:end id="491EFDC4F1009A5810CEA67A2786BFDA">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6F5B0B3F8E091E605FCFC048AA6BF841">journal article</mods:classification>
<mods:identifier id="1030AA580619EAE4868B8C3D58E5F35A" type="DOI">10.5281/zenodo.184612</mods:identifier>
<mods:identifier id="BC83B4019E13CEB3207453D99D160CEA" type="GBIF-Dataset">e30cedee-23dc-4989-835c-fee3db1a8d43</mods:identifier>
<mods:identifier id="037C52CD1B6522BDD647D697955E71A9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="84AA8BF6D41D0FC8E6F1209BC58869AB" type="Zenodo-Dep">184612</mods:identifier>
</mods:mods>
<treatment id="7C6D4423FF91FF8DFF0AFABCFAD8FE09" ID-DOI="http://doi.org/10.5281/zenodo.6232949" ID-GBIF-Taxon="119380041" ID-Zenodo-Dep="6232949" LSID="urn:lsid:plazi:treatment:7C6D4423FF91FF8DFF0AFABCFAD8FE09" httpUri="http://treatment.plazi.org/id/7C6D4423FF91FF8DFF0AFABCFAD8FE09" lastPageId="4" lastPageNumber="67" pageId="2" pageNumber="65">
<subSubSection id="BCDEA6BEFF91FF8BFF0AFABCFE88FA03" pageId="2" pageNumber="65" type="nomenclature">
<paragraph id="F47BF535FF91FF8BFF0AFABCFD69FA2F" blockId="2.[151,767,1386,1448]" box="[151,767,1386,1413]" pageId="2" pageNumber="65">
<heading id="AF334259FF91FF8BFF0AFABCFD69FA2F" bold="true" box="[151,767,1386,1413]" fontSize="11" level="1" pageId="2" pageNumber="65" reason="1">
<emphasis id="C6B02927FF91FF8BFF0AFABCFD69FA2F" bold="true" box="[151,767,1386,1413]" pageId="2" pageNumber="65">
<taxonomicName id="33C48EB6FF91FF8BFF0AFABCFD02FA2F" ID-CoL="7CHSZ" authority="Bravo, Alves &amp; Chagas" authorityName="Bravo, Alves &amp; Chagas" box="[151,660,1386,1413]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="C6B02927FF91FF8BFF0AFABCFEFEFA2E" bold="true" box="[151,360,1387,1413]" italics="true" pageId="2" pageNumber="65">Tonnoira robusta</emphasis>
Bravo, Alves &amp; Chagas
</taxonomicName>
,
<taxonomicNameLabel id="DD83945CFF91FF8BFD3CFABDFD69FA2F" box="[673,767,1386,1412]" pageId="2" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F47BF535FF91FF8BFF0AFA59FE88FA03" blockId="2.[151,767,1386,1448]" box="[151,286,1422,1448]" pageId="2" pageNumber="65">
<figureCitation id="6CFFE9B0FF91FF8BFF0AFA59FE88FA03" box="[151,286,1422,1448]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Figs. 11–20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BCDEA6BEFF91FF8DFF0AFA09FAD8FE09" lastPageId="4" lastPageNumber="67" pageId="2" pageNumber="65" type="materials_examined">
<paragraph id="F47BF535FF91FF8BFF0AFA09FB81F933" blockId="2.[151,1436,1502,2013]" pageId="2" pageNumber="65">
<typeStatus id="2B7F4B97FF91FF8BFF0AFA09FF47FA53" box="[151,209,1502,1528]" pageId="2" pageNumber="65">Type</typeStatus>
material.
<collectingCountry id="8CD3B5A5FF91FF8BFEC8FA09FE54FA53" box="[341,450,1502,1528]" name="Brazil" pageId="2" pageNumber="65">BRAZIL</collectingCountry>
, Amazonas, Presidente Figueiredo, Gruta Refúgio do Maragua,
<geoCoordinate id="91F093F2FF91FF8BFB71FA09FA0AFA53" box="[1260,1436,1502,1528]" direction="south" orientation="latitude" pageId="2" pageNumber="65" precision="1" value="-2.0507333">02°03´02.64”S</geoCoordinate>
<geoCoordinate id="91F093F2FF91FF8BFF0AF9D1FEC4F98B" box="[151,338,1542,1568]" direction="west" orientation="longitude" pageId="2" pageNumber="65" precision="1" value="-59.96329">59°57´47.85”W</geoCoordinate>
,
<date id="807AD3F5FF91FF8BFEC2F9D1FE72F98B" box="[351,484,1542,1568]" pageId="2" pageNumber="65" value="2006-04-01">01.IV.2006</date>
, Alves, V. col.,
<typeStatus id="2B7F4B97FF91FF8BFD3FF9D1FC9EF98B" box="[674,776,1542,1568]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
male (
<collectionCode id="92D56DF0FF91FF8BFCC7F9D1FC35F98B" box="[858,931,1542,1568]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="65">INPA</collectionCode>
); 1
<typeStatus id="2B7F4B97FF91FF8BFC4DF9D1FBA3F98B" box="[976,1077,1542,1568]" pageId="2" pageNumber="65" type="paratype">paratype</typeStatus>
male, same locality, date, and collector as
<typeStatus id="2B7F4B97FF91FF8BFEB2F9F9FE03F9E3" box="[303,405,1582,1608]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
(
<collectionCode id="92D56DF0FF91FF8BFE37F9F9FDB3F9E3" box="[426,549,1582,1608]" pageId="2" pageNumber="65">MZUEFS</collectionCode>
); 2
<typeStatus id="2B7F4B97FF91FF8BFDF8F9F9FD5CF9E3" box="[613,714,1582,1608]" pageId="2" pageNumber="65" type="paratype">paratype</typeStatus>
males, same locality and collector as
<typeStatus id="2B7F4B97FF91FF8BFB34F9F9FA85F9E3" box="[1193,1299,1582,1608]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
,
<date id="807AD3F5FF91FF8BFAB9F9F9FA0AF9E3" box="[1316,1436,1582,1608]" pageId="2" pageNumber="65" value="2006-05-06">06.V.2006</date>
(
<collectionCode id="92D56DF0FF91FF8BFF02F981FF70F9DB" box="[159,230,1622,1648]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="65">INPA</collectionCode>
,
<collectionCode id="92D56DF0FF91FF8BFF6CF981FEFCF9DB" box="[241,362,1622,1648]" pageId="2" pageNumber="65">MZUEFS</collectionCode>
); 2
<typeStatus id="2B7F4B97FF91FF8BFE0EF981FE6EF9DB" box="[403,504,1622,1648]" pageId="2" pageNumber="65" type="paratype">paratype</typeStatus>
males, same locality and collector as
<typeStatus id="2B7F4B97FF91FF8BFC2CF981FB8CF9DB" box="[945,1050,1622,1648]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
,
<date id="807AD3F5FF91FF8BFBBBF981FB36F9DB" box="[1062,1184,1622,1648]" pageId="2" pageNumber="65" value="2006-02-09">09.II.2006</date>
(
<collectionCode id="92D56DF0FF91FF8BFB32F981FB60F9DB" box="[1199,1270,1622,1648]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="65">INPA</collectionCode>
,
<collectionCode id="92D56DF0FF91FF8BFA9CF981FAECF9DB" box="[1281,1402,1622,1648]" pageId="2" pageNumber="65">MZUEFS</collectionCode>
); 1
<typeStatus id="2B7F4B97FF91FF8BFF0AF9A9FF6AF933" box="[151,252,1662,1688]" pageId="2" pageNumber="65" type="paratype">paratype</typeStatus>
female, same locality and collector as
<typeStatus id="2B7F4B97FF91FF8BFD5EF9A9FCBBF933" box="[707,813,1662,1688]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
,
<date id="807AD3F5FF91FF8BFCA5F9A9FC24F933" box="[824,946,1662,1688]" pageId="2" pageNumber="65" value="2006-02-09">09.II.2006</date>
(
<collectionCode id="92D56DF0FF91FF8BFC5CF9A9FB9CF933" box="[961,1034,1662,1688]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="65">INPA</collectionCode>
).
</paragraph>
<paragraph id="F47BF535FF91FF8DFF5BF971FBFBFE81" blockId="2.[151,1436,1502,2013]" lastBlockId="4.[151,1436,152,418]" lastPageId="4" lastPageNumber="67" pageId="2" pageNumber="65">
Description. Male. Head (
<figureCitation id="6CFFE9B0FF91FF8BFD9EF971FDCBF96B" box="[515,605,1702,1728]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 11</figureCitation>
): eyes separated by 0.5 facet diameters (
<figureCitation id="6CFFE9B0FF91FF8BFBC8F971FB39F96B" box="[1109,1199,1702,1728]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 11</figureCitation>
); interocular suture inverted Y shaped (
<figureCitation id="6CFFE9B0FF91FF8BFE1AF919FE40F943" box="[391,470,1742,1768]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig.11</figureCitation>
). Antenna: scape cylindrical 1.7X length of pedicel (
<figureCitation id="6CFFE9B0FF91FF8BFBC3F919FB2FF943" box="[1118,1209,1742,1768]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 12</figureCitation>
); pedicel spherical (
<figureCitation id="6CFFE9B0FF91FF8BFF02F921FF6CF8BB" box="[159,250,1782,1808]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 12</figureCitation>
); flagellum with 14 flagellomeres; all flagellomeres flask-shaped (
<figureCitation id="6CFFE9B0FF91FF8BFBBDF921FB22F8BB" box="[1056,1204,1782,1808]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Figs. 12, 13</figureCitation>
); first flagellomere same length as second (
<figureCitation id="6CFFE9B0FF91FF8BFE31F8C9FD95F893" box="[428,515,1822,1848]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 12</figureCitation>
); apiculus present in flagellomere 14 (
<figureCitation id="6CFFE9B0FF91FF8BFC58F8C9FB8AF893" box="[965,1052,1822,1848]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 13</figureCitation>
); ascoids not visible. Palpus formula = 1.0:2.1:1.9:2.2; last palpomere striated (
<figureCitation id="6CFFE9B0FF91FF8BFD4DF891FCBFF8CB" box="[720,809,1862,1888]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 14</figureCitation>
). Wing (
<figureCitation id="6CFFE9B0FF91FF8BFC0FF891FC7AF8CB" box="[914,1004,1862,1888]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 15</figureCitation>
): Sc short; vein R2+3 obsolete, radial fork free; radial fork basal; medial fork complete; R5 ending at wing apex. Male terminalia: hypandrium narrow, diademe-like without ventral expansion (
<figureCitation id="6CFFE9B0FF91FF8BFD23F84CFC81F81E" box="[702,791,1947,1973]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 16</figureCitation>
); gonocoxite 0.6X length of the gonostylus (
<figureCitation id="6CFFE9B0FF91FF8BFAA9F84CFA1BF81E" box="[1332,1421,1947,1973]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 16</figureCitation>
); gonostyli convergent, with base larger than apex (
<figureCitation id="6CFFE9B0FF91FF8BFD43F814FCA3F876" box="[734,821,1987,2013]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 16</figureCitation>
); alveoli present along entire surface (
<figureCitation id="6CFFE9B0FF91FF8BFB68F814FADAF876" box="[1269,1356,1987,2013]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="2" pageNumber="65">Fig. 16</figureCitation>
); epandrium rectangular; foramen not observed (
<figureCitation id="6CFFE9B0FF90FF8AFD51F878FCB0F862" box="[716,806,1967,1993]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="3" pageNumber="66">Fig. 17</figureCitation>
); cercopod long, 2.2X length of epandrium, slightly curved (
<figureCitation id="6CFFE9B0FF90FF8AFF6AF800FEDBF85A" box="[247,333,2007,2033]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="3" pageNumber="66">Fig. 18</figureCitation>
), with 2 tenacula, one apical and the other subapical (
<figureCitation id="6CFFE9B0FF90FF8AFC55F800FB89F85A" box="[968,1055,2007,2033]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="3" pageNumber="66">Fig. 19</figureCitation>
); sternite 10 smaller than tergite 10 (
<figureCitation id="6CFFE9B0FF97FF8DFF5AFF4FFEB4FF19" box="[199,290,152,178]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="4" pageNumber="67">Fig. 17</figureCitation>
); aedeagus asymmetrical, bipartite with left branch smaller than right, curved, pointed at apex (
<figureCitation id="6CFFE9B0FF97FF8DFF02FF17FF60FF71" box="[159,246,192,218]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="4" pageNumber="67">Fig. 16</figureCitation>
); right branch of aedeagus with rounded apex (fig. 16); with two asymmetrical parameres ending near the apex of the aedeagus apex (
<figureCitation id="6CFFE9B0FF97FF8DFD9BFF3FFDCBFEA9" box="[518,605,232,258]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="4" pageNumber="67">Fig. 16</figureCitation>
); aedeagal apodeme trapezoidal, 0.75x length of aedeagus, wider anteriorly than posteriorly (
<figureCitation id="6CFFE9B0FF97FF8DFE32FEC7FD90FE81" box="[431,518,272,298]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="4" pageNumber="67">Fig. 16</figureCitation>
); gonocoxal apodeme not fused at midline (
<figureCitation id="6CFFE9B0FF97FF8DFB95FEC7FBF6FE81" box="[1032,1120,272,298]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="4" pageNumber="67">Fig. 16</figureCitation>
).
</paragraph>
<caption id="A0BBA5BDFF90FF8AFF0AF90EFCDDF8D0" httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="3" pageNumber="66" targetBox="[172,1431,187,1706]" targetPageId="3">
<paragraph id="F47BF535FF90FF8AFF0AF90EFCDDF8D0" blockId="3.[151,1437,1753,1915]" pageId="3" pageNumber="66">
<emphasis id="C6B02927FF90FF8AFF0AF90EFECDF95A" bold="true" box="[151,347,1753,1777]" pageId="3" pageNumber="66">FIGURES 11–20.</emphasis>
<taxonomicName id="33C48EB6FF90FF8AFEFFF90DFC89F95B" authority="Bravo, Chagas &amp; Alves" authorityName="Bravo, Chagas &amp; Alves" box="[354,799,1753,1777]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="C6B02927FF90FF8AFEFFF90DFD8FF95A" box="[354,537,1754,1777]" italics="true" pageId="3" pageNumber="66">Tonnoira robusta</emphasis>
Bravo, Chagas &amp; Alves
</taxonomicName>
,
<taxonomicNameLabel id="DD83945CFF90FF8AFCB4F90EFCEDF95B" box="[809,891,1753,1776]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
Male (11–19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d= distance between the base of subgentital plate and apex of apical lobes; S10= sternite 10; T10= tergite 10.
</paragraph>
</caption>
<paragraph id="F47BF535FF97FF8DFF5BFEEFFAD8FE09" blockId="4.[151,1436,152,418]" pageId="4" pageNumber="67">
Female (
<figureCitation id="6CFFE9B0FF97FF8DFEADFEEFFE1AFEF9" box="[304,396,312,338]" captionStart="FIGURES 11 – 20" captionStartId="3.[151,269,1753,1777]" captionTargetBox="[172,1431,187,1706]" captionTargetId="figure@3.[151,1436,170,1727]" captionTargetPageId="3" captionText="FIGURES 11 – 20. Tonnoira robusta Bravo, Chagas &amp; Alves, sp. nov. Male (11 – 19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d = distance between the base of subgentital plate and apex of apical lobes; S 10 = sternite 10; T 10 = tergite 10." httpUri="https://zenodo.org/record/184614/files/figure.png" pageId="4" pageNumber="67">Fig. 20</figureCitation>
). Similar to male except as follows: apical lobes of subgenital plate of females long, rounded, and separated by a shallow apical concavity. Cerci blade-like, 2.2X the total length of genital plate from the anterior border at middle to the lobe apices (distance d). Genital complex small, as illustrated.
</paragraph>
</subSubSection>
</treatment>
</document>
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