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<document id="582348E889495453E92F60EC418E353D" ID-DOI="10.5852/ejt.2024.966.2697" ID-ISSN="2118-9773" ID-Zenodo-Dep="14001790" ID-ZooBank="FFE41347-8B55-4CF1-B10B-53EBB4CF0E26" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1730118416961" checkinUser="plazi" docAuthor="Henrard, Arnaud, Griswold, Charles &amp; Jocqué, Rudy" docDate="2024" docId="3157E00AFFCEFFF0FDDD092CFDB7DA9B" docLanguage="en" docName="EJT.2024.966.1-80.pdf" docOrigin="European Journal of Taxonomy 966 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zorodictyna almae Henrard, Griswold &amp; Jocqué, 2024, sp. nov." docType="treatment" docUuid="9F5FEFEF-155C-4C64-862B-3F6F67A52A01" docUuidSource="ZooBank" docVersion="4" lastPageNumber="59" masterDocId="CD6E9872FFF8FFCAFFAA0824FFD5DF0E" masterDocTitle="On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar" masterLastPageNumber="80" masterPageNumber="1" pageNumber="55" updateTime="1730315048679" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2609D322AE9C89DF08BDAB0781C20405">On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar</mods:title>
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<mods:namePart id="412C656726A5AAE5C93A19AB50C5BD7B">Henrard, Arnaud</mods:namePart>
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<mods:namePart id="8A70F8D735F786E06F7310AE1B091016">Griswold, Charles</mods:namePart>
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<mods:affiliation id="5D7ED794B33129C2B15E04CC218F65D6">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA.</mods:affiliation>
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<mods:namePart id="5C8C83E11BAB2BC5ED7B15F7EEF77642">Jocqué, Rudy</mods:namePart>
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<mods:affiliation id="4AC75630501214BFDC67E377F1D3BB44">Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium.</mods:affiliation>
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<paragraph id="B941511CFFCEFFFCFDDD092CFC68DE2D" blockId="54.[371,1217,264,363]" box="[631,957,264,291]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFDDD092CFC68DE2D" box="[631,957,264,291]" centered="true" fontSize="11" level="2" pageId="54" pageNumber="55" reason="2">
<taxonomicName id="7EFE2A9FFFCEFFFCFDDD092CFC8FDE2D" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[631,858,264,291]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFCEFFFCFDDD092CFC8FDE2D" bold="true" box="[631,858,264,291]" italics="true" pageId="54" pageNumber="55">Zorodictyna almae</emphasis>
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<taxonomicNameLabel id="90B93075FFCEFFFCFCC8092DFC68DE2D" box="[866,957,265,291]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="B941511CFFCEFFFCFD680974FCA4DE64" blockId="54.[371,1217,264,363]" box="[706,881,336,363]" pageId="54" pageNumber="55">
<figureCitation id="21C54D99FFCEFFFCFD680974FC9DDE65" box="[706,840,336,363]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="56.[189,231,1928,1954]" captionStartId-1="57.[189,232,1636,1662]" captionTargetBox-0="[189,1398,1121,1890]" captionTargetBox-1="[219,1348,279,1580]" captionTargetId-0="figure-176@56.[189,1398,1121,1890]" captionTargetId-1="figure-12@57.[189,1398,266,1598]" captionTargetPageId-0="56" captionTargetPageId-1="57" captionText-0="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." captionText-1="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi-0="http://doi.org/10.5281/zenodo.14001843" figureDoi-1="http://doi.org/10.5281/zenodo.14001845" httpUri-0="https://zenodo.org/record/14001843/files/figure.png" httpUri-1="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Figs 22–23</figureCitation>
, 
<figureCitation id="21C54D99FFCEFFFCFCFE0974FCA4DE64" box="[852,881,336,362]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="54" pageNumber="55">26</figureCitation>
</paragraph>
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<taxonomicName id="7EFE2A9FFFCEFFFCFF1709B6FE5FDEA3" box="[189,394,402,429]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="8B8A8D0EFFCEFFFCFF1709B6FE9EDEA2" box="[189,331,402,428]" italics="true" pageId="54" pageNumber="55">Zorodictyna</emphasis>
sp. 1
</taxonomicName>
– 
<bibRefCitation id="DD6F2CEDFFCEFFFCFE0D09B6FD45DEA3" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[423,656,402,429]" pageId="54" pageNumber="55" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow 
<emphasis id="8B8A8D0EFFCEFFFCFDBB09B7FD9EDEA2" box="[529,587,402,428]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2015
</bibRefCitation>
.
</paragraph>
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<subSubSection id="F1E40297FFCEFFFCFF1709F0FCDEDCE1" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="B941511CFFCEFFFCFF1709F0FEE3DEE0" blockId="54.[189,310,468,494]" box="[189,310,468,494]" pageId="54" pageNumber="55">
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<emphasis id="8B8A8D0EFFCEFFFCFF1709F0FEE3DEE0" bold="true" box="[189,310,468,494]" pageId="54" pageNumber="55">Diagnosis</emphasis>
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<paragraph id="B941511CFFCEFFFCFF170A27FCDEDCE1" blockId="54.[187,1400,515,1007]" pageId="54" pageNumber="55">
Males of 
<taxonomicName id="7EFE2A9FFFCEFFFCFE8F0A27FE2BDD13" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[293,510,515,541]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFCEFFFCFE8F0A27FE2BDD13" box="[293,510,515,541]" italics="true" pageId="54" pageNumber="55">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCEFFFCFDAE0A20FD89DD10" box="[516,604,516,542]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
are distinguished from 
<taxonomicName id="7EFE2A9FFFCEFFFCFCCF0A20FBA1DD10" authority="(Lenz, 1891)" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[869,1140,515,542]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCEFFFCFCCF0A20FC09DD13" box="[869,988,515,541]" italics="true" pageId="54" pageNumber="55">Z. oswaldi</emphasis>
(
<bibRefCitation id="DD6F2CEDFFCEFFFCFC430A20FBB9DD10" author="Lenz H." box="[1001,1132,516,542]" pageId="54" pageNumber="55" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz, 1891</bibRefCitation>
)
</taxonomicName>
by having a retrobasal cymbial extension (CE) that extends at least ¼ the length of the papal tibia and a thick retrobasal paracymbial process (PP) (
<figureCitation id="21C54D99FFCEFFFCFE510A6FFD52DD6B" box="[507,647,587,613]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23B–C</figureCitation>
) (vs 
<taxonomicName id="7EFE2A9FFFCEFFFCFD6E0A68FCEADD6B" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[708,831,587,613]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCEFFFCFD6E0A68FCEADD6B" box="[708,831,587,613]" italics="true" pageId="54" pageNumber="55">Z. oswaldi</emphasis>
</taxonomicName>
lacks the PP and has a very short CE; 
<bibRefCitation id="DD6F2CEDFFCEFFFCFAA30A6FFE8CDD87" author="Griswold C. E." pageId="54" pageNumber="55" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993: 19–21</bibRefCitation>
). It is distinguished from 
<taxonomicName id="7EFE2A9FFFCEFFFCFD390A4BFC91DD87" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[659,836,623,649]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="silvadavilae" status="sp. nov.">
<emphasis id="8B8A8D0EFFCEFFFCFD390A4BFC91DD87" box="[659,836,623,649]" italics="true" pageId="54" pageNumber="55">Z. silvadavilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCEFFFCFCE10A4BFC7CDD87" box="[843,937,623,649]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
in having the male palp with a nearly triangular basal embolic hook, the embolus lacking a crest, and a retromedian longitudinal ridge that extends for most of the tegulum length (
<figureCitation id="21C54D99FFCEFFFCFD080A92FCE4DDDE" box="[674,817,694,720]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23A–C</figureCitation>
) (vs 
<taxonomicName id="7EFE2A9FFFCEFFFCFCDA0A93FBCBDDDE" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[880,1054,694,720]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="silvadavilae">
<emphasis id="8B8A8D0EFFCEFFFCFCDA0A93FBCBDDDE" box="[880,1054,694,720]" italics="true" pageId="54" pageNumber="55">Z. silvadavilae</emphasis>
</taxonomicName>
has slender, crescentic basal embolic hook, the embolus with subapical distal crest, and the tegular retromedian longitudinal ridge that is weak, short, extending for less than one-half tegulum length, 
<figureCitation id="21C54D99FFCEFFFCFC490AD9FBA6DC16" box="[995,1139,765,792]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="54" pageNumber="55">Fig. 25A–C</figureCitation>
). Females of 
<taxonomicName id="7EFE2A9FFFCEFFFCFABA0ADAFAA2DC19" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1296,1399,765,791]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="almae">
<emphasis id="8B8A8D0EFFCEFFFCFABA0ADAFAA2DC19" box="[1296,1399,765,791]" italics="true" pageId="54" pageNumber="55">Z. almae</emphasis>
</taxonomicName>
are distinguished from 
<taxonomicName id="7EFE2A9FFFCEFFFCFE780B06FD98DC35" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[466,589,801,827]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCEFFFCFE780B06FD98DC35" box="[466,589,801,827]" italics="true" pageId="54" pageNumber="55">Z. oswaldi</emphasis>
</taxonomicName>
by lacking teeth on the lateral lobes (LL) of the epigyne (
<figureCitation id="21C54D99FFCEFFFCFAA90B05FABADC35" box="[1283,1391,801,827]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23D</figureCitation>
) (vs 
<taxonomicName id="7EFE2A9FFFCEFFFCFF4D0B62FEB5DC51" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[231,352,837,863]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCEFFFCFF4D0B62FEB5DC51" box="[231,352,837,863]" italics="true" pageId="54" pageNumber="55">Z. oswaldi</emphasis>
</taxonomicName>
has the LL with teeth; 
<bibRefCitation id="DD6F2CEDFFCEFFFCFDC20B61FCCFDC51" author="Griswold C. E." box="[616,794,837,863]" pageId="54" pageNumber="55" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: fig. 24); females of 
<taxonomicName id="7EFE2A9FFFCEFFFCFBAF0B62FBBEDC51" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1029,1131,837,863]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="almae">
<emphasis id="8B8A8D0EFFCEFFFCFBAF0B62FBBEDC51" box="[1029,1131,837,863]" italics="true" pageId="54" pageNumber="55">Z. almae</emphasis>
</taxonomicName>
are distinguished from 
<taxonomicName id="7EFE2A9FFFCEFFFCFF160B4EFEBFDC8D" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[188,362,873,899]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="silvadavilae">
<emphasis id="8B8A8D0EFFCEFFFCFF160B4EFEBFDC8D" box="[188,362,873,899]" italics="true" pageId="54" pageNumber="55">Z. silvadavilae</emphasis>
</taxonomicName>
by having the lateral lobes of epigyne parallel and the median sector (MS) rectangular (
<figureCitation id="21C54D99FFCEFFFCFF6D0BA8FEE1DCA9" box="[199,308,908,935]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23D</figureCitation>
). The female endogyne has copulatory ducts (CD) larger than spermathecae (
<figureCitation id="21C54D99FFCEFFFCFB150BA8FAF3DCA9" box="[1215,1318,908,935]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23E</figureCitation>
) (vs in 
<taxonomicName id="7EFE2A9FFFCEFFFCFF170B95FEBCDCC4" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[189,361,944,970]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="silvadavilae">
<emphasis id="8B8A8D0EFFCEFFFCFF170B95FEBCDCC4" box="[189,361,944,970]" italics="true" pageId="54" pageNumber="55">Z. silvadavilae</emphasis>
</taxonomicName>
, the LL are converging posteriorly, and the MS is triangular, 
<figureCitation id="21C54D99FFCEFFFCFB980B94FB48DCC5" box="[1074,1181,944,971]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="54" pageNumber="55">Fig. 25D</figureCitation>
, and the endogyne has CD and spermathecae equal in size, 
<figureCitation id="21C54D99FFCEFFFCFD3F0BF0FD2BDCE1" box="[661,766,980,1007]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="54" pageNumber="55">Fig. 25E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFCEFFFCFF170C32FAA2DBA8" pageId="54" pageNumber="55" type="etymology">
<paragraph id="B941511CFFCEFFFCFF170C32FE91DB3E" blockId="54.[189,324,1046,1072]" box="[189,324,1046,1072]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFF170C32FE91DB3E" bold="true" box="[189,324,1046,1072]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="8B8A8D0EFFCEFFFCFF170C32FE91DB3E" bold="true" box="[189,324,1046,1072]" pageId="54" pageNumber="55">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170C61FAA2DBA8" blockId="54.[188,1400,1093,1191]" pageId="54" pageNumber="55">
The species name is a matronym in honor of Alma Saucedo Mejia, participant in the 2011 expedition that produced the female 
<typeStatus id="6645EFBEFFCEFFFCFE1D0C4DFDC9DB8D" box="[439,540,1129,1155]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
(CASENT_9031271) that was a DNA voucher in 
<bibRefCitation id="DD6F2CEDFFCEFFFCFBDD0C4DFAADDB8D" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[1143,1400,1128,1155]" pageId="54" pageNumber="55" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow 
<emphasis id="8B8A8D0EFFCEFFFCFB4F0C4DFAF4DB8D" box="[1253,1313,1129,1155]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
(2015)
</bibRefCitation>
[DNA-DP-Lyc-voucher, Z-1, DP2014d]. She is also the discoverer of many 
<collectingCountry id="C1E9118CFFCEFFFCFB9C0CA8FB11DBA8" box="[1078,1220,1164,1190]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
goblin spiders.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFCEFFFCFF170CEBFE20D85A" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="B941511CFFCEFFFCFF170CEBFEB9DBE6" blockId="54.[189,364,1230,1257]" box="[189,364,1230,1257]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFF170CEBFEB9DBE6" bold="true" box="[189,364,1230,1257]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="8B8A8D0EFFCEFFFCFF170CEBFEB9DBE6" bold="true" box="[189,364,1230,1257]" pageId="54" pageNumber="55">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF400CDAFE89DA16" blockId="54.[189,1398,1278,1376]" box="[234,348,1278,1304]" pageId="54" pageNumber="55">
<emphasis id="8B8A8D0EFFCEFFFCFF400CDAFE89DA16" bold="true" box="[234,348,1278,1304]" pageId="54" pageNumber="55">
<typeStatus id="6645EFBEFFCEFFFCFF400CDAFE89DA16" box="[234,348,1278,1304]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170D05FA9FDA51" blockId="54.[189,1398,1278,1376]" pageId="54" pageNumber="55">
<materialsCitation id="09965B41FFCEFFFCFF170D05FA9FDA51" collectingDate="1997-02-04" collectingDateMax="1997-02-07" collectingDateMin="1997-02-04" collectionCode="FMNH" collectorName="R. F. l'Ankarafantsika &amp; S. Goodman" country="Madagascar" elevation="160" latitude="-16.338333" location="Mahajunga Province" longLatPrecision="127" longitude="46.793335" pageId="54" pageNumber="55" specimenCode="CASENT_9082245" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="C1E9118CFFCEFFFCFF170D05FE47DA35" box="[189,402,1313,1339]" name="Madagascar" pageId="54" pageNumber="55">MADAGASCAR</collectingCountry>
– 
<emphasis id="8B8A8D0EFFCEFFFCFE1D0D05FD1DDA35" bold="true" box="[439,712,1313,1339]" pageId="54" pageNumber="55">
<location id="BC2107C7FFCEFFFCFE1D0D05FD1DDA35" LSID="urn:lsid:plazi:treatment:3157E00AFFCEFFF0FDDD092CFDB7DA9B:BC2107C7FFCEFFFCFE1D0D05FD1DDA35" box="[439,712,1313,1339]" country="Madagascar" latitude="-16.338333" longLatPrecision="127" longitude="46.793335" name="Mahajunga Province" pageId="54" pageNumber="55">Mahajunga Province</location>
</emphasis>
• 
<specimenCount id="AFF89A95FFCEFFFCFD430D06FCD7DA32" box="[745,770,1314,1340]" count="1" pageId="54" pageNumber="55" type="male">♂</specimenCount>
; 
<collectorName id="140B34CAFFCEFFFCFCB80D06FBFCDA35" box="[786,1065,1313,1340]" pageId="54" pageNumber="55">R. F. l’Ankarafantsika</collectorName>
, 
<locationDeviation id="A8F3483EFFCEFFFCFB920D05FAA7DA35" box="[1080,1394,1313,1340]" location="Ampijaroa" pageId="54" pageNumber="55">
<quantity id="7E06FCF9FFCEFFFCFB920D05FBADDA35" box="[1080,1144,1313,1340]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="54" pageNumber="55" unit="km" value="5.0">5 km</quantity>
SSW of Ampijaroa
</locationDeviation>
; 
<geoCoordinate id="DCCA37DBFFCEFFFCFF170D67FEECDA51" box="[189,313,1347,1375]" degrees="16" direction="south" minutes="20.3" orientation="latitude" pageId="54" pageNumber="55" precision="92" value="-16.338333">16°20.3′ S</geoCoordinate>
, 
<geoCoordinate id="DCCA37DBFFCEFFFCFEEC0D67FE13DA51" box="[326,454,1347,1375]" degrees="46" direction="east" minutes="47.6" orientation="longitude" pageId="54" pageNumber="55" precision="92" value="46.793335">46°47.6′ E</geoCoordinate>
; 
<elevation id="32D3B62FFFCEFFFCFE780D61FD89DA51" box="[466,604,1349,1375]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="54" pageNumber="55" unit="m" value="160.0">
<quantity id="7E06FCF9FFCEFFFCFE780D61FDC9DA51" box="[466,540,1349,1375]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="54" pageNumber="55" unit="m" value="160.0">160 m</quantity>
a.s.l.
</elevation>
; 
<date id="CD4077DCFFCEFFFCFDC00D61FCC3DA51" box="[618,790,1349,1375]" pageId="54" pageNumber="55" value="1997-02-04" valueMax="1997-02-07" valueMin="1997-02-04">
<collectingDate id="DD048E34FFCEFFFCFDC00D61FCC3DA51" box="[618,790,1349,1375]" pageId="54" pageNumber="55" value="1997-02-04" valueMax="1997-02-07" valueMin="1997-02-04">4–7 Feb. 1997</collectingDate>
</date>
; 
<collectorName id="140B34CAFFCEFFFCFC880D61FC60DA51" box="[802,949,1349,1375]" pageId="54" pageNumber="55">S. Goodman</collectorName>
leg.; 
<specimenCode id="E958F967FFCEFFFCFC5C0D61FB37DA6E" box="[1014,1250,1349,1376]" pageId="54" pageNumber="55">CASENT_9082245</specimenCode>
; 
<collectionCode id="DFEFC9D9FFCEFFFCFB470D61FA93DA51" box="[1261,1350,1349,1375]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="54" pageNumber="55" type="Museum">FMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF400DA3FEB2DAAF" blockId="54.[188,1399,1415,1691]" box="[234,359,1415,1441]" pageId="54" pageNumber="55">
<emphasis id="8B8A8D0EFFCEFFFCFF400DA3FEB2DAAF" bold="true" box="[234,359,1415,1441]" pageId="54" pageNumber="55">
<typeStatus id="6645EFBEFFCEFFFCFF400DA3FEB2DAAF" box="[234,359,1415,1441]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170D8FFC02D995" blockId="54.[188,1399,1415,1691]" pageId="54" pageNumber="55">
<materialsCitation id="09965B41FFCEFFFCFF170D8FFAA2DACB" box="[189,1399,1450,1477]" collectingDate="1997-02-04" collectingDateMax="1997-02-07" collectingDateMin="1997-02-04" collectionCode="FMNH" collectorName="R. F. l'Ankarafantsika &amp; S. Goodman" country="Madagascar" elevation="160" latitude="-16.338333" location="Mahajunga Province" longLatPrecision="127" longitude="46.793335" pageId="54" pageNumber="55" specimenCode="CASENT_9082245" specimenCount="1" specimenCount-female="1" stateProvince="Mahajunga Province" typeStatus="paratype">
<collectingCountry id="C1E9118CFFCEFFFCFF170D8FFE58DACB" box="[189,397,1451,1477]" name="Madagascar" pageId="54" pageNumber="55">MADAGASCAR</collectingCountry>
– 
<emphasis id="8B8A8D0EFFCEFFFCFE0C0D8EFD7FDACA" bold="true" box="[422,682,1450,1476]" pageId="54" pageNumber="55">
<collectingRegion id="7B3A9FFEFFCEFFFCFE0C0D8EFD7FDACA" box="[422,682,1450,1476]" pageId="54" pageNumber="55">Mahajunga Province</collectingRegion>
</emphasis>
• 
<specimenCount id="AFF89A95FFCEFFFCFD6A0D8FFD38DACB" box="[704,749,1451,1477]" count="1" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
; same data as for holotype; 
<specimenCode id="E958F967FFCEFFFCFB870D8EFACDDACB" box="[1069,1304,1450,1477]" pageId="54" pageNumber="55">CASENT_9082245</specimenCode>
; 
<collectionCode id="DFEFC9D9FFCEFFFCFA880D8FFAA2DACB" box="[1314,1399,1451,1477]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="54" pageNumber="55" type="Museum">FMNH</collectionCode>
</materialsCitation>
• 
<materialsCitation id="09965B41FFCEFFFCFF7A0DEBFAFDD93E" collectingDate="2009-01-26" collectingDateMax="2009-01-28" collectingDateMin="2009-01-26" collectionCode="CAS" collectorName="C. Griswold &amp; A. Saucedo &amp; H. Wood &amp; Polotow D. &amp; Carmichael A. &amp; Griswold C. E." country="Madagascar" county="Ampijoroa" elevation="0" location="near Lake Ravelobe" municipality="National Park Ankarafantsika" pageId="54" pageNumber="55" specimenCode="CASENT_9031271" specimenCount="1" specimenCount-female="1" stateProvince="Mahajunga Province" typeStatus="paratype">
<specimenCount id="AFF89A95FFCEFFFCFF7A0DEBFF2ADAE7" box="[208,255,1487,1513]" count="1" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
; 
<collectingCounty id="50202990FFCEFFFCFEA70DEAFE45DAE6" box="[269,400,1486,1512]" pageId="54" pageNumber="55">Ampijoroa</collectingCounty>
, 
<collectingMunicipality id="5925CB66FFCEFFFCFE340DEAFCD7DAE6" box="[414,770,1486,1512]" pageId="54" pageNumber="55">National Park Ankarafantsika</collectingMunicipality>
, 
<location id="BC2107C7FFCEFFFCFCA50DEBFBD7DAE6" LSID="urn:lsid:plazi:treatment:3157E00AFFCEFFF0FDDD092CFDB7DA9B:BC2107C7FFCEFFFCFCA50DEBFBD7DAE6" box="[783,1026,1486,1513]" country="Madagascar" county="Ampijoroa" municipality="National Park Ankarafantsika" name="near Lake Ravelobe" pageId="54" pageNumber="55" stateProvince="Mahajunga Province">near Lake Ravelobe</location>
; 
<geoCoordinate id="DCCA37DBFFCEFFFCFBA50DE9FB62DAE7" box="[1039,1207,1485,1513]" pageId="54" pageNumber="55">16°17.48′7″ S</geoCoordinate>
, 
<elevation id="32D3B62FFFCEFFFCFB6C0DE9FAA4DAE7" box="[1222,1393,1485,1513]" pageId="54" pageNumber="55">46°48.50′7″ E</elevation>
; 
<date id="CD4077DCFFCEFFFCFF170DD6FE55D902" box="[189,384,1522,1549]" pageId="54" pageNumber="55" value="2009-01-26" valueMax="2009-01-28" valueMin="2009-01-26">
<collectingDate id="DD048E34FFCEFFFCFF170DD6FE55D902" box="[189,384,1522,1549]" pageId="54" pageNumber="55" value="2009-01-26" valueMax="2009-01-28" valueMin="2009-01-26">26–28 Jan. 2009</collectingDate>
</date>
; 
<collectorName id="140B34CAFFCEFFFCFE210DD6FDC9D902" box="[395,540,1522,1548]" pageId="54" pageNumber="55">C. Griswold</collectorName>
, 
<collectorName id="140B34CAFFCEFFFCFD8F0DD6FD7ED902" box="[549,683,1522,1548]" pageId="54" pageNumber="55">A. Saucedo</collectorName>
and 
<collectorName id="140B34CAFFCEFFFCFD490DD7FC9FD902" box="[739,842,1522,1549]" pageId="54" pageNumber="55">H. Wood</collectorName>
leg.; DNA-DP-Lyc-voucher, 
<taxonomicName id="7EFE2A9FFFCEFFFCFB090DD6FABAD902" box="[1187,1391,1522,1549]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="8B8A8D0EFFCEFFFCFB090DD6FAE4D902" box="[1187,1329,1522,1548]" italics="true" pageId="54" pageNumber="55">Zorodictyna</emphasis>
sp. 1
</taxonomicName>
, in 
<collectorName id="140B34CAFFCEFFFCFF710E32FEB6D93E" box="[219,355,1558,1584]" pageId="54" pageNumber="55">Polotow D.</collectorName>
, 
<collectorName id="140B34CAFFCEFFFCFEDA0E32FDCFD93E" box="[368,538,1558,1584]" pageId="54" pageNumber="55">Carmichael A.</collectorName>
&amp; 
<collectorName id="140B34CAFFCEFFFCFDE80E32FD3ED93E" box="[578,747,1558,1584]" pageId="54" pageNumber="55">Griswold C.E.</collectorName>
2015; Z-1; 
<accessionNumber id="A6ADCCFFFFCEFFFCFCD20E32FC3ED93E" box="[888,1003,1558,1584]" pageId="54" pageNumber="55">DP2014d</accessionNumber>
; 
<specimenCode id="E958F967FFCEFFFCFC5D0E32FB37D93E" box="[1015,1250,1558,1584]" pageId="54" pageNumber="55">CASENT_9031271</specimenCode>
; 
<collectionCode id="DFEFC9D9FFCEFFFCFB450E32FAFDD93E" box="[1263,1320,1558,1584]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="09965B41FFCEFFFCFAE80E32FC06D995" collectedFrom="tropical dry forest" collectingDate="2001-04-02" collectingDateMax="2001-04-08" collectingDateMin="2001-04-02" collectingMethod="general collecting" collectionCode="CAS" collectorName="J. J. Rafanomezantsoa" country="Madagascar" elevation="135" latitude="-16.061388" location="Foret de Tsimaloto" longLatPrecision="21" longitude="46.143612" municipality="Parc National d'Ankarafantsika" pageId="54" pageNumber="55" specimenCode="CASENT_9002829" specimenCount="1" specimenCount-female="1" stateProvince="Mahajunga Province" typeStatus="paratype">
<specimenCount id="AFF89A95FFCEFFFCFAE80E32FABAD93E" box="[1346,1391,1558,1584]" count="1" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
; 
<collectingMunicipality id="5925CB66FFCEFFFCFF170E1EFDECD95D" box="[189,569,1593,1620]" pageId="54" pageNumber="55">Parc National d’Ankarafantsika</collectingMunicipality>
, 
<location id="BC2107C7FFCEFFFCFDED0E1EFCF9D95D" LSID="urn:lsid:plazi:treatment:3157E00AFFCEFFF0FDDD092CFDB7DA9B:BC2107C7FFCEFFFCFDED0E1EFCF9D95D" box="[583,812,1593,1620]" country="Madagascar" latitude="-16.061388" longLatPrecision="21" longitude="46.143612" municipality="Parc National d'Ankarafantsika" name="Foret de Tsimaloto" pageId="54" pageNumber="55" stateProvince="Mahajunga Province">Forêt de Tsimaloto</location>
, 
<locationDeviation id="A8F3483EFFCEFFFCFC930E1EFB35D95D" box="[825,1248,1593,1620]" location="Tsaramandroso" pageId="54" pageNumber="55">
<quantity id="7E06FCF9FFCEFFFCFC930E1EFC49D95D" box="[825,924,1593,1620]" metricMagnitude="4" metricUnit="m" metricValue="1.83" pageId="54" pageNumber="55" unit="km" value="18.3">18.3 km</quantity>
46° NE of Tsaramandroso
</locationDeviation>
; 
<geoCoordinate id="DCCA37DBFFCEFFFCFB470E1CFAA4D95A" box="[1261,1393,1592,1620]" degrees="16" direction="south" minutes="3" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="41" value="-16.061388">16°3′41″ S</geoCoordinate>
, 
<geoCoordinate id="DCCA37DBFFCEFFFCFF170E78FE95D976" box="[189,320,1628,1656]" degrees="46" direction="east" minutes="8" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="37" value="46.143612">46°8′37″ E</geoCoordinate>
; 
<elevation id="32D3B62FFFCEFFFCFEE10E79FE07D979" box="[331,466,1629,1656]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="54" pageNumber="55" unit="m" value="135.0">
<quantity id="7E06FCF9FFCEFFFCFEE10E79FE46D976" box="[331,403,1629,1656]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="54" pageNumber="55" unit="m" value="135.0">135 m</quantity>
a.s.l.
</elevation>
; 
<date id="CD4077DCFFCEFFFCFE740E79FD50D979" box="[478,645,1629,1656]" pageId="54" pageNumber="55" value="2001-04-02" valueMax="2001-04-08" valueMin="2001-04-02">
<collectingDate id="DD048E34FFCEFFFCFE740E79FD50D979" box="[478,645,1629,1656]" pageId="54" pageNumber="55" value="2001-04-02" valueMax="2001-04-08" valueMin="2001-04-02">2–8 Apr. 2001</collectingDate>
</date>
; 
<collectorName id="140B34CAFFCEFFFCFD3A0E7AFC59D979" box="[656,908,1629,1656]" pageId="54" pageNumber="55">J.J. Rafanomezantsoa</collectorName>
<emphasis id="8B8A8D0EFFCEFFFCFC380E7AFC1ED979" box="[914,971,1629,1655]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
leg.; 
<collectingMethod id="60BF290BFFCEFFFCFBA00E79FB08D979" box="[1034,1245,1629,1655]" pageId="54" pageNumber="55">general collecting</collectingMethod>
, day, 
<collectedFrom id="0AE212E1FFCEFFFCFAB70E79FEE4D995" pageId="54" pageNumber="55">tropical dry forest</collectedFrom>
; [epigyne illustrated by CEG]; 
<specimenCode id="E958F967FFCEFFFCFD350EA5FC5ED995" box="[671,907,1665,1691]" pageId="54" pageNumber="55">CASENT_9002829</specimenCode>
; 
<collectionCode id="DFEFC9D9FFCEFFFCFC3C0EA5FC06D995" box="[918,979,1665,1691]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170EE7FE2CD9D3" blockId="54.[189,505,1731,1757]" box="[189,505,1731,1757]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFF170EE7FE2CD9D3" bold="true" box="[189,505,1731,1757]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="8B8A8D0EFFCEFFFCFF170EE7FE2CD9D3" bold="true" box="[189,505,1731,1757]" pageId="54" pageNumber="55">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170ED6FE20D85A" blockId="54.[189,1399,1778,1876]" pageId="54" pageNumber="55">
<materialsCitation id="09965B41FFCEFFFCFF170ED6FE24D85A" collectingDate="2009-01-26" collectingDateMax="2009-01-28" collectingDateMin="2009-01-26" collectionCode="CAS" collectorName="C. Griswold &amp; A. Saucedo &amp; H. Wood" country="Madagascar" county="Ampijoroa" latitude="-16.29145" location="Lake Ravelobe" longLatPrecision="1" longitude="46.80845" municipality="National Park Ankarafantsika" pageId="54" pageNumber="55" specimenCode="CASENT_9031270" specimenCount="1" specimenCount-female="1" stateProvince="Mahajanga Province">
<collectingCountry id="C1E9118CFFCEFFFCFF170ED6FE5BD802" box="[189,398,1778,1804]" name="Madagascar" pageId="54" pageNumber="55">MADAGASCAR</collectingCountry>
– 
<emphasis id="8B8A8D0EFFCEFFFCFE070ED6FD67D802" bold="true" box="[429,690,1778,1804]" pageId="54" pageNumber="55">
<collectingRegion id="7B3A9FFEFFCEFFFCFE070ED6FD67D802" box="[429,690,1778,1804]" country="Madagascar" name="Mahajanga" pageId="54" pageNumber="55">Mahajanga Province</collectingRegion>
</emphasis>
• 
<specimenCount id="AFF89A95FFCEFFFCFD670ED6FD2FD803" box="[717,762,1778,1805]" count="1" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
; 
<collectingCounty id="50202990FFCEFFFCFCAD0ED6FC5CD802" box="[775,905,1778,1804]" pageId="54" pageNumber="55">Ampijoroa</collectingCounty>
, 
<collectingMunicipality id="5925CB66FFCEFFFCFC3C0ED6FB20D802" box="[918,1269,1778,1804]" pageId="54" pageNumber="55">National Park Ankarafantsika</collectingMunicipality>
, near 
<location id="BC2107C7FFCEFFFCFA960ED6FEE5D83E" LSID="urn:lsid:plazi:treatment:3157E00AFFCEFFF0FDDD092CFDB7DA9B:BC2107C7FFCEFFFCFA960ED6FEE5D83E" country="Madagascar" county="Ampijoroa" latitude="-16.29145" longLatPrecision="1" longitude="46.80845" municipality="National Park Ankarafantsika" name="Lake Ravelobe" pageId="54" pageNumber="55" stateProvince="Mahajanga Province">Lake Ravelobe</location>
; 
<geoCoordinate id="DCCA37DBFFCEFFFCFE970F30FE0CD83E" box="[317,473,1812,1840]" degrees="16" direction="south" minutes="17.487" orientation="latitude" pageId="54" pageNumber="55" precision="1" value="-16.29145">16°17.487′ S</geoCoordinate>
, 
<geoCoordinate id="DCCA37DBFFCEFFFCFE4D0F30FD53D83E" box="[487,646,1812,1840]" degrees="46" direction="east" minutes="48.507" orientation="longitude" pageId="54" pageNumber="55" precision="1" value="46.80845">46°48.507′ E</geoCoordinate>
; 
<date id="CD4077DCFFCEFFFCFD3E0F32FC88D83E" box="[660,861,1814,1840]" pageId="54" pageNumber="55" value="2009-01-26" valueMax="2009-01-28" valueMin="2009-01-26">
<collectingDate id="DD048E34FFCEFFFCFD3E0F32FC88D83E" box="[660,861,1814,1840]" pageId="54" pageNumber="55" value="2009-01-26" valueMax="2009-01-28" valueMin="2009-01-26">26–28 Jan. 2009</collectingDate>
</date>
; 
<collectorName id="140B34CAFFCEFFFCFCC00F32FC2AD83E" box="[874,1023,1814,1840]" pageId="54" pageNumber="55">C. Griswold</collectorName>
, 
<collectorName id="140B34CAFFCEFFFCFBA00F32FB46D83E" box="[1034,1171,1814,1840]" pageId="54" pageNumber="55">A. Saucedo</collectorName>
and 
<collectorName id="140B34CAFFCEFFFCFB650F32FAEFD83E" box="[1231,1338,1814,1840]" pageId="54" pageNumber="55">H. Wood</collectorName>
leg.; 
<specimenCode id="E958F967FFCEFFFCFF170F1EFE7CD85A" box="[189,425,1850,1876]" pageId="54" pageNumber="55">CASENT_9031270</specimenCode>
; 
<collectionCode id="DFEFC9D9FFCEFFFCFE1E0F1EFE24D85A" box="[436,497,1850,1876]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFCEFFF0FF170F5FFCFDDCDC" lastPageId="58" lastPageNumber="59" pageId="54" pageNumber="55" type="description">
<paragraph id="B941511CFFCEFFFCFF170F5FFE9BD89B" blockId="54.[189,334,1915,1941]" box="[189,334,1915,1941]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFF170F5FFE9BD89B" bold="true" box="[189,334,1915,1941]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="8B8A8D0EFFCEFFFCFF170F5FFE9BD89B" bold="true" box="[189,334,1915,1941]" pageId="54" pageNumber="55">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF400F8FFCD6D8CB" blockId="54.[189,1189,1962,2025]" box="[234,771,1962,1989]" pageId="54" pageNumber="55">
<emphasis id="8B8A8D0EFFCEFFFCFF400F8FFE48D8CB" bold="true" box="[234,413,1963,1989]" pageId="54" pageNumber="55">
Male 
<typeStatus id="6645EFBEFFCEFFFCFE9B0F8FFE48D8CB" box="[305,413,1963,1989]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
</emphasis>
(CASENT_9082245; FMNH)
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170FEAFB70D8E7" blockId="54.[189,1189,1962,2025]" box="[189,1189,1998,2025]" pageId="54" pageNumber="55">MEASUREMENTS. Total length: 21.00. Carapace: length 12.00, width 9.80, height 5.70.</paragraph>
<paragraph id="B941511CFFCFFFFDFF17092DFBBBDE4B" blockId="55.[189,1399,265,326]" pageId="55" pageNumber="56">
COLORATION (
<figureCitation id="21C54D99FFCFFFFDFECC092DFE2DDE2D" box="[358,504,265,291]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="55" pageNumber="56">Fig. 22A–B</figureCitation>
). Dorsal and lateral sides of cephalothorax, abdomen and legs unicolorous grey-brown, venter of sternum, mouthparts, coxae and trochanters yellow white.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170948FB8ADE88" blockId="55.[189,1119,364,391]" box="[189,1119,364,391]" pageId="55" pageNumber="56">
STERNUM (
<figureCitation id="21C54D99FFCFFFFDFE970948FE7DDE88" box="[317,424,364,391]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="55" pageNumber="56">Fig. 22B</figureCitation>
). Oval with slightly sinuous margins; 4.50 wide, 5.60 long.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170989FDF7DEC9" blockId="55.[189,546,428,455]" box="[189,546,428,455]" pageId="55" pageNumber="56">CHILUM. 0.25 high, 0.85 wide.</paragraph>
<paragraph id="B941511CFFCFFFFDFF1709C9FCCADD24" blockId="55.[188,1399,493,554]" pageId="55" pageNumber="56">EYES. ALE 0.28; AME 0.28; PLE 0.35; PME 0.30; ALE–AME 0.13; AME–AME 0.70; PLE–PME 0.45; PME–PME 0.28; MOQ: AW 0.73, PW 0.83, L 0.80.</paragraph>
<paragraph id="B941511CFFCFFFFDFF170A75FDC7DD65" blockId="55.[189,530,592,619]" box="[189,530,592,619]" pageId="55" pageNumber="56">CLYPEUS. With 18 long setae.</paragraph>
<paragraph id="B941511CFFCFFFFDFF170AB5FACFDDC0" blockId="55.[189,1399,657,718]" pageId="55" pageNumber="56">LEGS. Trochanters deeply notched. Unpaired claws smooth, rudimentary on legs III and IV. Strong scopulae beneath all tarsi, apices of metatarsi III and IV, and apical halves of metatarsi I and II.</paragraph>
<caption id="ED810194FFCFFFFDFF170AD1FE4FDC01" box="[189,410,757,783]" inLine="true" pageId="55" pageNumber="56" startId="55.[189,235,757,783]" targetBox="[203,804,799,985]" targetPageId="55">
<paragraph id="B941511CFFCFFFFDFF170AD1FE4FDC01" blockId="55.[189,410,757,783]" box="[189,410,757,783]" pageId="55" pageNumber="56">Leg measurements</paragraph>
</caption>
<paragraph id="B941511CFFCFFFFDFEA00B3BFCF1DCD7" pageId="55" pageNumber="56">
<table id="CBFEA3BCFFCF0035FF610B3BFCF1DCD7" box="[203,804,799,985]" gridcols="7" gridrows="5" pageId="55" pageNumber="56" rowsContinueIn="55.[203,988,1066,1252]">
<tr id="07CE535EFFCF0035FF610B3BFCF1DC37" box="[203,804,799,825]" gridrow="0" pageId="55" pageNumber="56" rowspan-0="1">
<th id="441F3A22FFCF0035FEA00B3BFE99DC37" box="[266,332,799,825]" gridcol="1" gridrow="0" pageId="55" pageNumber="56">F</th>
<th id="441F3A22FFCF0035FEC50B3BFE77DC37" box="[367,418,799,825]" gridcol="2" gridrow="0" pageId="55" pageNumber="56">P</th>
<th id="441F3A22FFCF0035FE6E0B3BFDD3DC37" box="[452,518,799,825]" gridcol="3" gridrow="0" pageId="55" pageNumber="56">T</th>
<th id="441F3A22FFCF0035FD820B3BFDBFDC37" box="[552,618,799,825]" gridcol="4" gridrow="0" pageId="55" pageNumber="56">Mt</th>
<th id="441F3A22FFCF0035FD270B3BFD15DC37" box="[653,704,799,825]" gridcol="5" gridrow="0" pageId="55" pageNumber="56">t</th>
<th id="441F3A22FFCF0035FD480B3BFCF1DC37" box="[738,804,799,825]" gridcol="6" gridrow="0" pageId="55" pageNumber="56">Total</th>
</tr>
<tr id="07CE535EFFCF0035FF610B63FCF1DC6C" box="[203,804,839,866]" gridrow="1" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610B63FF3FDC6C" box="[203,234,839,866]" gridcol="0" gridrow="1" pageId="55" pageNumber="56">I</th>
<td id="441F3A22FFCF0035FEA00B63FE99DC6C" box="[266,332,839,866]" gridcol="1" gridrow="1" pageId="55" pageNumber="56">10.20</td>
<td id="441F3A22FFCF0035FEC50B63FE77DC6C" box="[367,418,839,866]" gridcol="2" gridrow="1" pageId="55" pageNumber="56">5.10</td>
<td id="441F3A22FFCF0035FE6E0B63FDD3DC6C" box="[452,518,839,866]" gridcol="3" gridrow="1" pageId="55" pageNumber="56">11.50</td>
<td id="441F3A22FFCF0035FD820B63FDBFDC6C" box="[552,618,839,866]" gridcol="4" gridrow="1" pageId="55" pageNumber="56">11.10</td>
<td id="441F3A22FFCF0035FD270B63FD15DC6C" box="[653,704,839,866]" gridcol="5" gridrow="1" pageId="55" pageNumber="56">6.30</td>
<td id="441F3A22FFCF0035FD480B63FCF1DC6C" box="[738,804,839,866]" gridcol="6" gridrow="1" pageId="55" pageNumber="56">44.20</td>
</tr>
<tr id="07CE535EFFCF0035FF610B4BFCF1DC84" box="[203,804,879,906]" gridrow="2" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610B4BFF3FDC84" box="[203,234,879,906]" gridcol="0" gridrow="2" pageId="55" pageNumber="56">II</th>
<td id="441F3A22FFCF0035FEA00B4BFE99DC84" box="[266,332,879,906]" gridcol="1" gridrow="2" pageId="55" pageNumber="56">11.00</td>
<td id="441F3A22FFCF0035FEC50B4BFE77DC84" box="[367,418,879,906]" gridcol="2" gridrow="2" pageId="55" pageNumber="56">5.00</td>
<td id="441F3A22FFCF0035FE6E0B4BFDD3DC84" box="[452,518,879,906]" gridcol="3" gridrow="2" pageId="55" pageNumber="56">9.70</td>
<td id="441F3A22FFCF0035FD820B4BFDBFDC84" box="[552,618,879,906]" gridcol="4" gridrow="2" pageId="55" pageNumber="56">9.80</td>
<td id="441F3A22FFCF0035FD270B4BFD15DC84" box="[653,704,879,906]" gridcol="5" gridrow="2" pageId="55" pageNumber="56">5.00</td>
<td id="441F3A22FFCF0035FD480B4BFCF1DC84" box="[738,804,879,906]" gridcol="6" gridrow="2" pageId="55" pageNumber="56">40.50</td>
</tr>
<tr id="07CE535EFFCF0035FF610BB3FCF1DCBC" box="[203,804,919,946]" gridrow="3" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610BB3FF3FDCBC" box="[203,234,919,946]" gridcol="0" gridrow="3" pageId="55" pageNumber="56">III</th>
<td id="441F3A22FFCF0035FEA00BB3FE99DCBC" box="[266,332,919,946]" gridcol="1" gridrow="3" pageId="55" pageNumber="56">9.30</td>
<td id="441F3A22FFCF0035FEC50BB3FE77DCBC" box="[367,418,919,946]" gridcol="2" gridrow="3" pageId="55" pageNumber="56">4.10</td>
<td id="441F3A22FFCF0035FE6E0BB3FDD3DCBC" box="[452,518,919,946]" gridcol="3" gridrow="3" pageId="55" pageNumber="56">7.60</td>
<td id="441F3A22FFCF0035FD820BB3FDBFDCBC" box="[552,618,919,946]" gridcol="4" gridrow="3" pageId="55" pageNumber="56">9.40</td>
<td id="441F3A22FFCF0035FD270BB3FD15DCBC" box="[653,704,919,946]" gridcol="5" gridrow="3" pageId="55" pageNumber="56">4.60</td>
<td id="441F3A22FFCF0035FD480BB3FCF1DCBC" box="[738,804,919,946]" gridcol="6" gridrow="3" pageId="55" pageNumber="56">35.00</td>
</tr>
<tr id="07CE535EFFCF0035FF610B9AFCF1DCD7" box="[203,804,958,985]" gridrow="4" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610B9AFF3FDCD7" box="[203,234,958,985]" gridcol="0" gridrow="4" pageId="55" pageNumber="56">IV</th>
<td id="441F3A22FFCF0035FEA00B9AFE99DCD7" box="[266,332,958,985]" gridcol="1" gridrow="4" pageId="55" pageNumber="56">11.90</td>
<td id="441F3A22FFCF0035FEC50B9AFE77DCD7" box="[367,418,958,985]" gridcol="2" gridrow="4" pageId="55" pageNumber="56">4.70</td>
<td id="441F3A22FFCF0035FE6E0B9AFDD3DCD7" box="[452,518,958,985]" gridcol="3" gridrow="4" pageId="55" pageNumber="56">10.90</td>
<td id="441F3A22FFCF0035FD820B9AFDBFDCD7" box="[552,618,958,985]" gridcol="4" gridrow="4" pageId="55" pageNumber="56">15.20</td>
<td id="441F3A22FFCF0035FD270B9AFD15DCD7" box="[653,704,958,985]" gridcol="5" gridrow="4" pageId="55" pageNumber="56">6.00</td>
<td id="441F3A22FFCF0035FD480B9AFCF1DCD7" box="[738,804,958,985]" gridcol="6" gridrow="4" pageId="55" pageNumber="56">48.70</td>
</tr>
</table>
</paragraph>
<caption id="ED810194FFCFFFFDFF170BDBFEF8DB17" box="[189,301,1023,1049]" inLine="true" pageId="55" pageNumber="56" startId="55.[189,301,1023,1049]" targetBox="[203,988,1066,1252]" targetPageId="55">
<paragraph id="B941511CFFCFFFFDFF170BDBFEF8DB17" blockId="55.[189,301,1023,1049]" box="[189,301,1023,1049]" pageId="55" pageNumber="56">Spination</paragraph>
</caption>
<paragraph id="B941511CFFCFFFFDFEA00C0EFC09DBED" pageId="55" pageNumber="56">
<table id="CBFEA3BCFFCF0035FF610C0EFC09DBEA" box="[203,988,1066,1252]" gridcols="5" gridrows="5" pageId="55" pageNumber="56" rowsContinueFrom="55.[203,804,799,985]">
<tr id="07CE535EFFCF0035FF610C0EFC09DB4A" box="[203,988,1066,1092]" gridrow="0" pageId="55" pageNumber="56" rowspan-0="1">
<th id="441F3A22FFCF0035FEA00C0EFEBBDB4A" box="[266,366,1066,1092]" gridcol="1" gridrow="0" pageId="55" pageNumber="56">F</th>
<th id="441F3A22FFCF0035FE300C0EFE35DB4A" box="[410,480,1066,1092]" gridcol="2" gridrow="0" pageId="55" pageNumber="56">P</th>
<th id="441F3A22FFCF0035FDBD0C0EFD37DB4A" box="[535,738,1066,1092]" gridcol="3" gridrow="0" pageId="55" pageNumber="56">T</th>
<th id="441F3A22FFCF0035FCBD0C0EFC09DB4A" box="[791,988,1066,1092]" gridcol="4" gridrow="0" pageId="55" pageNumber="56">Mt</th>
</tr>
<tr id="07CE535EFFCF0035FF610C76FC09DB63" box="[203,988,1106,1133]" gridrow="1" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610C76FF3FDB63" box="[203,234,1106,1133]" gridcol="0" gridrow="1" pageId="55" pageNumber="56">I</th>
<td id="441F3A22FFCF0035FEA00C76FEBBDB63" box="[266,366,1106,1133]" gridcol="1" gridrow="1" pageId="55" pageNumber="56">pl3d3rl2</td>
<td id="441F3A22FFCF0035FE300C76FE35DB63" box="[410,480,1106,1133]" gridcol="2" gridrow="1" pageId="55" pageNumber="56">pl1rl1</td>
<td id="441F3A22FFCF0035FDBD0C76FD37DB63" box="[535,738,1106,1133]" gridcol="3" gridrow="1" pageId="55" pageNumber="56">pl3d3rl3v2-2-2-2</td>
<td id="441F3A22FFCF0035FCBD0C76FC09DB63" box="[791,988,1106,1133]" gridcol="4" gridrow="1" pageId="55" pageNumber="56">pl4rl4v2-2-3</td>
</tr>
<tr id="07CE535EFFCF0035FF610C5EFC09DB9A" box="[203,988,1146,1172]" gridrow="2" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610C5EFF3FDB9A" box="[203,234,1146,1172]" gridcol="0" gridrow="2" pageId="55" pageNumber="56">II</th>
<td id="441F3A22FFCF0035FEA00C5EFEBBDB9A" box="[266,366,1146,1172]" gridcol="1" gridrow="2" pageId="55" pageNumber="56">pl3d3rl3</td>
<td id="441F3A22FFCF0035FE300C5EFE35DB9A" box="[410,480,1146,1172]" gridcol="2" gridrow="2" pageId="55" pageNumber="56">–</td>
<td id="441F3A22FFCF0035FDBD0C5EFD37DB9A" box="[535,738,1146,1172]" gridcol="3" gridrow="2" pageId="55" pageNumber="56">pl2d3rl2v2-2-2-2</td>
<td id="441F3A22FFCF0035FCBD0C5EFC09DB9A" box="[791,988,1146,1172]" gridcol="4" gridrow="2" pageId="55" pageNumber="56">pl3rl3v2-2-3</td>
</tr>
<tr id="07CE535EFFCF0035FF610C86FC09DBB2" box="[203,988,1186,1212]" gridrow="3" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610C86FF3FDBB2" box="[203,234,1186,1212]" gridcol="0" gridrow="3" pageId="55" pageNumber="56">III</th>
<td id="441F3A22FFCF0035FEA00C86FEBBDBB2" box="[266,366,1186,1212]" gridcol="1" gridrow="3" pageId="55" pageNumber="56">pl3d3rl3</td>
<td id="441F3A22FFCF0035FE300C86FE35DBB2" box="[410,480,1186,1212]" gridcol="2" gridrow="3" pageId="55" pageNumber="56">pl1rl1</td>
<td id="441F3A22FFCF0035FDBD0C86FD37DBB2" box="[535,738,1186,1212]" gridcol="3" gridrow="3" pageId="55" pageNumber="56">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFCF0035FCBD0C86FC09DBB2" box="[791,988,1186,1212]" gridcol="4" gridrow="3" pageId="55" pageNumber="56">pl4d1rl4v2-2-2</td>
</tr>
<tr id="07CE535EFFCF0035FF610CEDFC09DBEA" box="[203,988,1225,1252]" gridrow="4" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610CEDFF3FDBEA" box="[203,234,1225,1252]" gridcol="0" gridrow="4" pageId="55" pageNumber="56">IV</th>
<td id="441F3A22FFCF0035FEA00CEDFEBBDBEA" box="[266,366,1225,1252]" gridcol="1" gridrow="4" pageId="55" pageNumber="56">pl3d3rl3</td>
<td id="441F3A22FFCF0035FE300CEDFE35DBEA" box="[410,480,1225,1252]" gridcol="2" gridrow="4" pageId="55" pageNumber="56">pl1rl1</td>
<td id="441F3A22FFCF0035FDBD0CEDFD37DBEA" box="[535,738,1225,1252]" gridcol="3" gridrow="4" pageId="55" pageNumber="56">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFCF0035FCBD0CEDFC09DBEA" box="[791,988,1225,1252]" gridcol="4" gridrow="4" pageId="55" pageNumber="56">pl4rl4v1-1-1-2-3</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170D2FFC6AD991" blockId="55.[187,1400,1290,1696]" pageId="55" pageNumber="56">
MALE PALP (
<figureCitation id="21C54D99FFCFFFFDFEFA0D2EFE33DA2A" box="[336,486,1290,1317]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="55" pageNumber="56">Fig. 23 A–C</figureCitation>
). Tibia VTA apical, curved and blunt-tipped, RTA triangular, sharply pointed, located behind this dorso-retrolateral excavation to accommodate retrobasal cymbial extension (CE); cymbium oval, apical extremity narrowly rounded, without dorsal patch of chemosensitive setae, with protruding retrolateral paracymbial process (PP), retrobasal cymbial extension (CE) conical, extending at least ¼ the length of papal tibia; subtegulum large, filling most of cymbial concavity and extending to prolateral cymbial margin, with probasal broad ridge (SL) that interlocks with tegulum; tegulum rounded with rectangular process on prolateral margin (TL), which interlocks with subtegular ridge; embolus (E) proapical, broad and thick, with pointed apex, inner margin with twisted longitudinal ridge and triangular basomedian process (EBP) arising from embolus base; median apophysis (MA) strongly sclerotized, short, hooked, curved prolaterad; conductor (C) hyaline, fan shaped, narrow at base; paraconductor (Pc) large, cradling inner margin of E and extending past E apex; retroventral surface of tegulum with deep longitudinal groove that extends for two thirds of tegulum length.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF400EE2FB66D80C" blockId="55.[189,1203,1733,1794]" pageId="55" pageNumber="56">
<emphasis id="8B8A8D0EFFCFFFFDFF400EE2FE68D9EE" bold="true" box="[234,445,1734,1760]" pageId="55" pageNumber="56">
Female 
<typeStatus id="6645EFBEFFCFFFFDFEE70EE2FE68D9EE" box="[333,445,1734,1760]" pageId="55" pageNumber="56" type="paratype">paratype</typeStatus>
</emphasis>
(CASENT_9082245; FMNH; except as otherwise indicated) MEASUREMENTS. Total length: 30.00. Carapace: length 14.00, width 11.30, height 4.40.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170F0DFAD4D86B" blockId="55.[189,1398,1833,1894]" pageId="55" pageNumber="56">
COLOUR (
<figureCitation id="21C54D99FFCFFFFDFE9F0F0DFE1FD84D" box="[309,458,1833,1859]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="55" pageNumber="56">Fig. 22C–D</figureCitation>
). As in male: dorsal and lateral surfaces of cephalothorax, abdomen and legs unicolorous grey-brown; venter of sternum, mouthparts, coxae and trochanters yellow white.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170FA8FBE8D8A8" blockId="55.[189,1085,1932,1958]" box="[189,1085,1932,1958]" pageId="55" pageNumber="56">
STERNUM (
<figureCitation id="21C54D99FFCFFFFDFE970FA8FE7FD8A8" box="[317,426,1932,1958]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="55" pageNumber="56">Fig. 22D</figureCitation>
). Oval with slightly sinuous sides; 4.90 wide, 6.40 long.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170FEAFDF7D8E6" blockId="55.[189,546,1998,2025]" box="[189,546,1998,2025]" pageId="55" pageNumber="56">CHILUM. 0.25 high, 1.08 wide.</paragraph>
<paragraph id="B941511CFFC0FFF2FF17092DFCCADE48" blockId="56.[188,1399,265,327]" pageId="56" pageNumber="57">EYES. ALE 0.33; AME 0.33; PLE 0.31; PME 0.29; ALE–AME 0.20; AME–AME 0.75; PLE–PME 0.55; PME–PME 0.35; MOQ: AW 0.78, PW 0.93, L 0.78.</paragraph>
<paragraph id="B941511CFFC0FFF2FF17094AFDCDDE86" blockId="56.[189,536,366,392]" box="[189,536,366,392]" pageId="56" pageNumber="57">CLYPEUS. With ten long setae.</paragraph>
<paragraph id="B941511CFFC0FFF2FF17098BFA86DEE3" blockId="56.[189,1399,431,493]" pageId="56" pageNumber="57">LEGS. Trochanters deeply notched. Unpaired claws smooth, rudimentary on legs III and IV. Strong scopulae beneath all tarsi, apex of metatarsus III, apical half of metatarsus II and all of metatarsus I.</paragraph>
<paragraph id="B941511CFFC0FFF2FF170A30FE4FDD20" blockId="56.[189,410,532,558]" box="[189,410,532,558]" pageId="56" pageNumber="57">Leg measurements</paragraph>
<paragraph id="B941511CFFC0FFF2FEA00A1AFCF1DDF9" pageId="56" pageNumber="57">
<table id="CBFEA3BCFFC00035FF610A19FCF1DDF9" box="[203,804,573,759]" gridcols="7" gridrows="5" pageId="56" pageNumber="57" rowsContinueIn="56.[203,912,840,1026]">
<tr id="07CE535EFFC00035FF610A19FCF1DD56" box="[203,804,573,600]" gridrow="0" pageId="56" pageNumber="57" rowspan-0="1">
<th id="441F3A22FFC00035FEA00A19FE99DD56" box="[266,332,573,600]" gridcol="1" gridrow="0" pageId="56" pageNumber="57">F</th>
<th id="441F3A22FFC00035FEC50A19FE77DD56" box="[367,418,573,600]" gridcol="2" gridrow="0" pageId="56" pageNumber="57">P</th>
<th id="441F3A22FFC00035FE6E0A19FDD0DD56" box="[452,517,573,600]" gridcol="3" gridrow="0" pageId="56" pageNumber="57">T</th>
<th id="441F3A22FFC00035FD820A19FDBFDD56" box="[552,618,573,600]" gridcol="4" gridrow="0" pageId="56" pageNumber="57">Mt</th>
<th id="441F3A22FFC00035FD270A19FD15DD56" box="[653,704,573,600]" gridcol="5" gridrow="0" pageId="56" pageNumber="57">t</th>
<th id="441F3A22FFC00035FD480A19FCF1DD56" box="[738,804,573,600]" gridcol="6" gridrow="0" pageId="56" pageNumber="57">Total</th>
</tr>
<tr id="07CE535EFFC00035FF610A41FCF1DD8E" box="[203,804,613,640]" gridrow="1" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610A41FF3FDD8E" box="[203,234,613,640]" gridcol="0" gridrow="1" pageId="56" pageNumber="57">I</th>
<td id="441F3A22FFC00035FEA00A41FE99DD8E" box="[266,332,613,640]" gridcol="1" gridrow="1" pageId="56" pageNumber="57">10.30</td>
<td id="441F3A22FFC00035FEC50A41FE77DD8E" box="[367,418,613,640]" gridcol="2" gridrow="1" pageId="56" pageNumber="57">5.20</td>
<td id="441F3A22FFC00035FE6E0A41FDD0DD8E" box="[452,517,613,640]" gridcol="3" gridrow="1" pageId="56" pageNumber="57">11.50</td>
<td id="441F3A22FFC00035FD820A41FDBFDD8E" box="[552,618,613,640]" gridcol="4" gridrow="1" pageId="56" pageNumber="57">11.10</td>
<td id="441F3A22FFC00035FD270A41FD15DD8E" box="[653,704,613,640]" gridcol="5" gridrow="1" pageId="56" pageNumber="57">6.30</td>
<td id="441F3A22FFC00035FD480A41FCF1DD8E" box="[738,804,613,640]" gridcol="6" gridrow="1" pageId="56" pageNumber="57">44.20</td>
</tr>
<tr id="07CE535EFFC00035FF610AA9FCF1DDA6" box="[203,804,653,680]" gridrow="2" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610AA9FF3FDDA6" box="[203,234,653,680]" gridcol="0" gridrow="2" pageId="56" pageNumber="57">II</th>
<td id="441F3A22FFC00035FEA00AA9FE99DDA6" box="[266,332,653,680]" gridcol="1" gridrow="2" pageId="56" pageNumber="57">9.50</td>
<td id="441F3A22FFC00035FEC50AA9FE77DDA6" box="[367,418,653,680]" gridcol="2" gridrow="2" pageId="56" pageNumber="57">4.90</td>
<td id="441F3A22FFC00035FE6E0AA9FDD0DDA6" box="[452,517,653,680]" gridcol="3" gridrow="2" pageId="56" pageNumber="57">7.20</td>
<td id="441F3A22FFC00035FD820AA9FDBFDDA6" box="[552,618,653,680]" gridcol="4" gridrow="2" pageId="56" pageNumber="57">6.80</td>
<td id="441F3A22FFC00035FD270AA9FD15DDA6" box="[653,704,653,680]" gridcol="5" gridrow="2" pageId="56" pageNumber="57">3.80</td>
<td id="441F3A22FFC00035FD480AA9FCF1DDA6" box="[738,804,653,680]" gridcol="6" gridrow="2" pageId="56" pageNumber="57">32.30</td>
</tr>
<tr id="07CE535EFFC00035FF610A91FCF1DDDE" box="[203,804,693,720]" gridrow="3" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610A91FF3FDDDE" box="[203,234,693,720]" gridcol="0" gridrow="3" pageId="56" pageNumber="57">III</th>
<td id="441F3A22FFC00035FEA00A91FE99DDDE" box="[266,332,693,720]" gridcol="1" gridrow="3" pageId="56" pageNumber="57">8.20</td>
<td id="441F3A22FFC00035FEC50A91FE77DDDE" box="[367,418,693,720]" gridcol="2" gridrow="3" pageId="56" pageNumber="57">4.20</td>
<td id="441F3A22FFC00035FE6E0A91FDD0DDDE" box="[452,517,693,720]" gridcol="3" gridrow="3" pageId="56" pageNumber="57">5.80</td>
<td id="441F3A22FFC00035FD820A91FDBFDDDE" box="[552,618,693,720]" gridcol="4" gridrow="3" pageId="56" pageNumber="57">7.20</td>
<td id="441F3A22FFC00035FD270A91FD15DDDE" box="[653,704,693,720]" gridcol="5" gridrow="3" pageId="56" pageNumber="57">3.80</td>
<td id="441F3A22FFC00035FD480A91FCF1DDDE" box="[738,804,693,720]" gridcol="6" gridrow="3" pageId="56" pageNumber="57">29.20</td>
</tr>
<tr id="07CE535EFFC00035FF610AF9FCF1DDF9" box="[203,804,733,759]" gridrow="4" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610AF9FF3FDDF9" box="[203,234,733,759]" gridcol="0" gridrow="4" pageId="56" pageNumber="57">IV</th>
<td id="441F3A22FFC00035FEA00AF9FE99DDF9" box="[266,332,733,759]" gridcol="1" gridrow="4" pageId="56" pageNumber="57">10.90</td>
<td id="441F3A22FFC00035FEC50AF9FE77DDF9" box="[367,418,733,759]" gridcol="2" gridrow="4" pageId="56" pageNumber="57">4.70</td>
<td id="441F3A22FFC00035FE6E0AF9FDD0DDF9" box="[452,517,733,759]" gridcol="3" gridrow="4" pageId="56" pageNumber="57">9.00</td>
<td id="441F3A22FFC00035FD820AF9FDBFDDF9" box="[552,618,733,759]" gridcol="4" gridrow="4" pageId="56" pageNumber="57">12.60</td>
<td id="441F3A22FFC00035FD270AF9FD15DDF9" box="[653,704,733,759]" gridcol="5" gridrow="4" pageId="56" pageNumber="57">4.80</td>
<td id="441F3A22FFC00035FD480AF9FCF1DDF9" box="[738,804,733,759]" gridcol="6" gridrow="4" pageId="56" pageNumber="57">42.00</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFC0FFF0FF170B3AFC47DED6" blockId="56.[189,301,798,824]" lastBlockId="58.[187,1400,265,473]" lastPageId="58" lastPageNumber="59" pageId="56" pageNumber="57">
Spination GENITALIA. Epigyne (
<typeStatus id="6645EFBEFFC2FFF0FE1A092DFDC0DE2D" box="[432,533,265,291]" pageId="58" pageNumber="59" type="paratype">paratype</typeStatus>
CASENT_9002829; CAS) (
<figureCitation id="21C54D99FFC2FFF0FCF7092DFC11DE2D" box="[861,964,265,291]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="58" pageNumber="59">Fig. 23D</figureCitation>
) transverse quadrangular median lobe (ML) plate with smooth margin; copulatory openings (CO) hidden beneath anterolateral corners of ML; lateral lobes (LL) convex, without lateral teeth or projections but with concavities along lateral margins of ML, LL straight and parallel medially, exposing rectangular area of median sector (MS). Endogyne (
<figureCitation id="21C54D99FFC2FFF0FF6D09BEFEFADEBA" box="[199,303,410,436]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="58" pageNumber="59">Fig. 23E</figureCitation>
) with broad, simple, dark copulatory ducts (CD) larger than spermathecae (S), spermathecae with one half twist; fertilization ducts (FD) short and straight.
</paragraph>
<paragraph id="B941511CFFC0FFF2FEA00B6CFC5EDB0C" pageId="56" pageNumber="57">
<table id="CBFEA3BCFFC00035FF610B6CFC45DB0C" box="[203,912,840,1026]" gridcols="5" gridrows="5" pageId="56" pageNumber="57" rowsContinueFrom="56.[203,804,573,759]">
<tr id="07CE535EFFC00035FF610B6CFC45DC6C" box="[203,912,840,866]" gridrow="0" pageId="56" pageNumber="57" rowspan-0="1">
<th id="441F3A22FFC00035FEA00B6CFEBBDC6C" box="[266,366,840,866]" gridcol="1" gridrow="0" pageId="56" pageNumber="57">F</th>
<th id="441F3A22FFC00035FE390B6CFE0CDC6C" box="[403,473,840,866]" gridcol="2" gridrow="0" pageId="56" pageNumber="57">P</th>
<th id="441F3A22FFC00035FE570B6CFD7ADC6C" box="[509,687,840,866]" gridcol="3" gridrow="0" pageId="56" pageNumber="57">T</th>
<th id="441F3A22FFC00035FD750B6CFC45DC6C" box="[735,912,840,866]" gridcol="4" gridrow="0" pageId="56" pageNumber="57">Mt</th>
</tr>
<tr id="07CE535EFFC00035FF610B54FC45DC85" box="[203,912,880,907]" gridrow="1" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610B54FF3FDC85" box="[203,234,880,907]" gridcol="0" gridrow="1" pageId="56" pageNumber="57">I</th>
<td id="441F3A22FFC00035FEA00B54FEBBDC85" box="[266,366,880,907]" gridcol="1" gridrow="1" pageId="56" pageNumber="57">pl1d2</td>
<td id="441F3A22FFC00035FE390B54FE0CDC85" box="[403,473,880,907]" gridcol="2" gridrow="1" pageId="56" pageNumber="57">–</td>
<td id="441F3A22FFC00035FE570B54FD7ADC85" box="[509,687,880,907]" gridcol="3" gridrow="1" pageId="56" pageNumber="57">v2-2-2-2</td>
<td id="441F3A22FFC00035FD750B54FC45DC85" box="[735,912,880,907]" gridcol="4" gridrow="1" pageId="56" pageNumber="57">v2-2-2</td>
</tr>
<tr id="07CE535EFFC00035FF610BBCFC45DCBC" box="[203,912,920,946]" gridrow="2" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610BBCFF3FDCBC" box="[203,234,920,946]" gridcol="0" gridrow="2" pageId="56" pageNumber="57">II</th>
<td id="441F3A22FFC00035FEA00BBCFEBBDCBC" box="[266,366,920,946]" gridcol="1" gridrow="2" pageId="56" pageNumber="57">pl2d2</td>
<td id="441F3A22FFC00035FE390BBCFE0CDCBC" box="[403,473,920,946]" gridcol="2" gridrow="2" pageId="56" pageNumber="57">–</td>
<td id="441F3A22FFC00035FE570BBCFD7ADCBC" box="[509,687,920,946]" gridcol="3" gridrow="2" pageId="56" pageNumber="57">v2-2-2-2</td>
<td id="441F3A22FFC00035FD750BBCFC45DCBC" box="[735,912,920,946]" gridcol="4" gridrow="2" pageId="56" pageNumber="57">v2-2-2</td>
</tr>
<tr id="07CE535EFFC00035FF610BE4FC45DCD4" box="[203,912,960,986]" gridrow="3" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610BE4FF3FDCD4" box="[203,234,960,986]" gridcol="0" gridrow="3" pageId="56" pageNumber="57">III</th>
<td id="441F3A22FFC00035FEA00BE4FEBBDCD4" box="[266,366,960,986]" gridcol="1" gridrow="3" pageId="56" pageNumber="57">pl3d3rl4</td>
<td id="441F3A22FFC00035FE390BE4FE0CDCD4" box="[403,473,960,986]" gridcol="2" gridrow="3" pageId="56" pageNumber="57">pl1rl1</td>
<td id="441F3A22FFC00035FE570BE4FD7ADCD4" box="[509,687,960,986]" gridcol="3" gridrow="3" pageId="56" pageNumber="57">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFC00035FD750BE4FC45DCD4" box="[735,912,960,986]" gridcol="4" gridrow="3" pageId="56" pageNumber="57">pl4d1rl4v2-2-2</td>
</tr>
<tr id="07CE535EFFC00035FF610BC3FC45DB0C" box="[203,912,999,1026]" gridrow="4" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610BC3FF3FDB0C" box="[203,234,999,1026]" gridcol="0" gridrow="4" pageId="56" pageNumber="57">IV</th>
<td id="441F3A22FFC00035FEA00BC3FEBBDB0C" box="[266,366,999,1026]" gridcol="1" gridrow="4" pageId="56" pageNumber="57">pl1d3rl3</td>
<td id="441F3A22FFC00035FE390BC3FE0CDB0C" box="[403,473,999,1026]" gridcol="2" gridrow="4" pageId="56" pageNumber="57">pl1rl1</td>
<td id="441F3A22FFC00035FE570BC3FD7ADB0C" box="[509,687,999,1026]" gridcol="3" gridrow="4" pageId="56" pageNumber="57">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFC00035FD750BC3FC45DB0C" box="[735,912,999,1026]" gridcol="4" gridrow="4" pageId="56" pageNumber="57">pl5rl4v2-1-2-2</td>
</tr>
</table>
</paragraph>
<caption id="ED810194FFC0FFF2FF170FACFBE1D8E6" ID-DOI="http://doi.org/10.5281/zenodo.14001843" ID-Zenodo-Dep="14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="56" pageNumber="57" startId="56.[189,231,1928,1954]" targetBox="[189,1398,1121,1890]" targetPageId="56">
<paragraph id="B941511CFFC0FFF2FF170FACFBE1D8E6" blockId="56.[187,1399,1928,2025]" pageId="56" pageNumber="57">
<emphasis id="8B8A8D0EFFC0FFF2FF170FACFEC3D8AC" bold="true" box="[189,278,1928,1954]" pageId="56" pageNumber="57">Fig. 22.</emphasis>
<taxonomicName id="7EFE2A9FFFC0FFF2FEB10FACFE21D8AC" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[283,500,1928,1954]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC0FFF2FEB10FACFE21D8AC" box="[283,500,1928,1954]" italics="true" pageId="56" pageNumber="57">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC0FFF2FE560FADFD83D8AD" box="[508,598,1929,1955]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
, male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). 
<emphasis id="8B8A8D0EFFC0FFF2FC190F8FFC39D8CB" bold="true" box="[947,1004,1963,1989]" pageId="56" pageNumber="57">A, C</emphasis>
. Dorsal view. 
<emphasis id="8B8A8D0EFFC0FFF2FB3C0F8FFB1BD8CB" bold="true" box="[1174,1230,1963,1989]" pageId="56" pageNumber="57">B, D</emphasis>
. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme.
</paragraph>
</caption>
<caption id="ED810194FFC1FFF3FF170E40FAC7D87A" ID-DOI="http://doi.org/10.5281/zenodo.14001845" ID-Zenodo-Dep="14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="57" pageNumber="58" startId="57.[189,232,1636,1662]" targetBox="[219,1348,279,1580]" targetPageId="57">
<paragraph id="B941511CFFC1FFF3FF170E40FAC7D87A" blockId="57.[187,1400,1636,1909]" pageId="57" pageNumber="58">
<emphasis id="8B8A8D0EFFC1FFF3FF170E40FEC9D970" bold="true" box="[189,284,1636,1662]" pageId="57" pageNumber="58">Fig. 23.</emphasis>
<taxonomicName id="7EFE2A9FFFC1FFF3FE8F0E40FDDDD970" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[293,520,1636,1662]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC1FFF3FE8F0E40FDDDD970" box="[293,520,1636,1662]" italics="true" pageId="57" pageNumber="58">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC1FFF3FDBA0E40FDA5D970" box="[528,624,1636,1662]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
, genitalia drawings; A–C 
<emphasis id="8B8A8D0EFFC1FFF3FC060E40FC68D970" bold="true" box="[940,957,1636,1662]" pageId="57" pageNumber="58">=</emphasis>
male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS). 
<emphasis id="8B8A8D0EFFC1FFF3FCDC0EA3FC5ED9AF" bold="true" box="[886,907,1671,1697]" pageId="57" pageNumber="58">A</emphasis>
. Left palp, prolateral view. 
<emphasis id="8B8A8D0EFFC1FFF3FB140EA3FB04D9AF" bold="true" box="[1214,1233,1671,1697]" pageId="57" pageNumber="58">B</emphasis>
. Idem, ventral view. 
<emphasis id="8B8A8D0EFFC1FFF3FEAF0E8EFECFD9CA" bold="true" box="[261,282,1706,1732]" pageId="57" pageNumber="58">C</emphasis>
. Idem, retrolateral view. 
<emphasis id="8B8A8D0EFFC1FFF3FDE10E8EFD8AD9CA" bold="true" box="[587,607,1706,1732]" pageId="57" pageNumber="58">D</emphasis>
. Epigyne, ventral view. 
<emphasis id="8B8A8D0EFFC1FFF3FC2C0E8EFC4DD9CA" bold="true" box="[902,920,1706,1732]" pageId="57" pageNumber="58">E</emphasis>
. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold.
</paragraph>
</caption>
<paragraph id="B941511CFFC2FFF0FF170A24FEE6DD14" blockId="58.[189,307,512,538]" box="[189,307,512,538]" pageId="58" pageNumber="59">
<heading id="E209E670FFC2FFF0FF170A24FEE6DD14" bold="true" box="[189,307,512,538]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="8B8A8D0EFFC2FFF0FF170A24FEE6DD14" bold="true" box="[189,307,512,538]" pageId="58" pageNumber="59">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF400A14FEFFDD44" blockId="58.[189,920,560,622]" box="[234,298,560,586]" pageId="58" pageNumber="59">
<emphasis id="8B8A8D0EFFC2FFF0FF400A14FEFFDD44" bold="true" box="[234,298,560,586]" pageId="58" pageNumber="59">Male</emphasis>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170A70FC4DDD60" blockId="58.[189,920,560,622]" box="[189,920,596,622]" pageId="58" pageNumber="59">
Variation unknown: only the single 
<typeStatus id="6645EFBEFFC2FFF0FDCB0A70FD1DDD60" box="[609,712,596,622]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
has been studied.
</paragraph>
<paragraph id="B941511CFFC2FFF0FF400AB3FE48DDBF" blockId="58.[187,1400,663,979]" box="[234,413,663,689]" pageId="58" pageNumber="59">
<emphasis id="8B8A8D0EFFC2FFF0FF400AB3FE90DDBF" bold="true" box="[234,325,663,689]" pageId="58" pageNumber="59">Female</emphasis>
(n = 3)
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170A9FFCFDDCDC" blockId="58.[187,1400,663,979]" pageId="58" pageNumber="59">[Female variation from CASENT_9082245, CASENT_9031270 and CASENT_9031271]. Total length = 20.00–30.00, carapace length/width = 1.24–1.30, carapace height/width = 0.39–0.48, PER/carapace width = 0.21–0.24, PER/OAL = 2.90–3.13, PER/AER = 1.35–1.36, OQP/OQA = 1.18–1.23, sternum length/width = 1.20–1.31, palpal coxa length/width = 2.24–2.44, femur I length/carapace width = 0.87– 1.00, metatarsus I length/carapace width = 0.64–0.74, femur IV length/carapace width = 0.96–1.06, palpal tarsus length/carapace width = 0.34–0.41, palpal tarsus length/palpal patella length = 1.46–2.00, palpal tarsus length/palpal tibia length = 1.09–1.57, palpal tarsus length/palpal femur length = 0.76–0.95, palpal tibia length/palpal patella length = 1.08–1.40.</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFC2FFF0FF170BDFFC8DDBF1" pageId="58" pageNumber="59" type="biology_ecology">
<paragraph id="B941511CFFC2FFF0FF170BDFFE40DB1B" blockId="58.[189,405,1019,1045]" box="[189,405,1019,1045]" pageId="58" pageNumber="59">
<heading id="E209E670FFC2FFF0FF170BDFFE40DB1B" bold="true" box="[189,405,1019,1045]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="8B8A8D0EFFC2FFF0FF170BDFFE40DB1B" bold="true" box="[189,405,1019,1045]" pageId="58" pageNumber="59">Note on synonym</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170C0EFBE8DB67" blockId="58.[189,1399,1066,1129]" pageId="58" pageNumber="59">
A specimen of this species (CASENT_9031271) was used as an exemplar in a previously published molecular phylogeny, listed as “ 
<taxonomicName id="7EFE2A9FFFC2FFF0FD9C0C6AFCD4DB67" box="[566,769,1102,1129]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="8B8A8D0EFFC2FFF0FD9C0C6AFD11DB66" box="[566,708,1102,1128]" italics="true" pageId="58" pageNumber="59">Zorodictyna</emphasis>
sp. 1
</taxonomicName>
” by 
<bibRefCitation id="DD6F2CEDFFC2FFF0FC960C6AFBECDB67" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[828,1081,1102,1129]" pageId="58" pageNumber="59" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow 
<emphasis id="8B8A8D0EFFC2FFF0FC0F0C6BFC35DB66" box="[933,992,1102,1128]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
(2015)
</bibRefCitation>
.
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170CB5FEABDBA5" blockId="58.[189,382,1169,1195]" box="[189,382,1169,1195]" pageId="58" pageNumber="59">
<heading id="E209E670FFC2FFF0FF170CB5FEABDBA5" bold="true" box="[189,382,1169,1195]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="8B8A8D0EFFC2FFF0FF170CB5FEABDBA5" bold="true" box="[189,382,1169,1195]" pageId="58" pageNumber="59">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170CE4FC8DDBF1" blockId="58.[187,1398,1216,1279]" pageId="58" pageNumber="59">
The species 
<taxonomicName id="7EFE2A9FFFC2FFF0FEE30CE4FDF0DBD4" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[329,549,1216,1242]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC2FFF0FEE30CE4FDF0DBD4" box="[329,549,1216,1242]" italics="true" pageId="58" pageNumber="59">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC2FFF0FD860CE5FD50DBD5" box="[556,645,1217,1243]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
is recorded from tropical dry forest.Although they have an entire cribellum we know nothing of their use of cribellate silk.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFC2FFF0FF170D03FDB7DA9B" pageId="58" pageNumber="59" type="distribution">
<paragraph id="B941511CFFC2FFF0FF170D03FE83DA4F" blockId="58.[189,342,1319,1345]" box="[189,342,1319,1345]" pageId="58" pageNumber="59">
<heading id="E209E670FFC2FFF0FF170D03FE83DA4F" bold="true" box="[189,342,1319,1345]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="8B8A8D0EFFC2FFF0FF170D03FE83DA4F" bold="true" box="[189,342,1319,1345]" pageId="58" pageNumber="59">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170D73FDB7DA9B" blockId="58.[189,1399,1367,1429]" pageId="58" pageNumber="59">
<taxonomicName id="7EFE2A9FFFC2FFF0FF170D73FE75DA7F" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[189,416,1367,1393]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC2FFF0FF170D73FE75DA7F" box="[189,416,1367,1393]" italics="true" pageId="58" pageNumber="59">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC2FFF0FE060D73FDDFDA7F" box="[428,522,1367,1393]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
is known from National Park Ankarafantsika in Mahajunga Province in northwestern 
<collectingCountry id="C1E9118CFFC2FFF0FEF50D5FFE38DA9B" box="[351,493,1403,1429]" name="Madagascar" pageId="58" pageNumber="59">Madagascar</collectingCountry>
(
<figureCitation id="21C54D99FFC2FFF0FE570D5FFD80DA9B" box="[509,597,1403,1429]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="58" pageNumber="59">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>