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<paragraph id="B941511CFFC9FFFBFDF30E30FC0ED920" blockId="49.[601,987,1555,1618]" box="[601,987,1555,1582]" pageId="49" pageNumber="50">
<heading id="E209E670FFC9FFFBFDF30E30FC0ED920" box="[601,987,1555,1582]" centered="true" fontSize="11" level="2" pageId="49" pageNumber="50" reason="2">
Genus
<taxonomicName id="7EFE2A9FFFC9FFFBFD010E37FC0ED920" ID-CoL="648PK" authority="Strand, 1907" authorityName="Strand" authorityYear="1907" box="[683,987,1555,1582]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFC9FFFBFD010E37FCEBD920" bold="true" box="[683,830,1555,1582]" italics="true" pageId="49" pageNumber="50">Zorodictyna</emphasis>
<bibRefCitation id="DD6F2CEDFFC9FFFBFCEF0E30FC0ED920" author="Strand E." box="[837,987,1556,1582]" pageId="49" pageNumber="50" pagination="725 - 748" refId="ref38669" refString="Strand E. 1907. Diagnosen neuer Spinnen aus Madagaskar und Sansibar. Zoologischer Anzeiger 31: 725 - 748." type="journal article" year="1907">Strand, 1907</bibRefCitation>
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<subSubSection id="F1E40297FFC9FFFBFD720E1CFC8ED95F" box="[728,859,1591,1618]" pageId="49" pageNumber="50" type="description">
<paragraph id="B941511CFFC9FFFBFD720E1CFC8ED95F" blockId="49.[601,987,1555,1618]" box="[728,859,1591,1618]" pageId="49" pageNumber="50">
<figureCitation id="21C54D99FFC9FFFBFD720E1CFC8ED95F" box="[728,859,1591,1618]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="56.[189,231,1928,1954]" captionStartId-1="57.[189,232,1636,1662]" captionStartId-2="59.[189,234,1894,1920]" captionStartId-3="61.[189,231,1752,1778]" captionStartId-4="63.[189,231,1857,1883]" captionTargetBox-0="[189,1398,1121,1890]" captionTargetBox-1="[219,1348,279,1580]" captionTargetBox-2="[189,1398,1046,1855]" captionTargetBox-3="[211,1360,510,1693]" captionTargetBox-4="[189,1398,1005,1819]" captionTargetId-0="figure-176@56.[189,1398,1121,1890]" captionTargetId-1="figure-12@57.[189,1398,266,1598]" captionTargetId-2="figure-180@59.[189,1398,1046,1855]" captionTargetId-3="figure-97@61.[189,1398,507,1714]" captionTargetId-4="figure-208@63.[189,1398,1005,1819]" captionTargetPageId-0="56" captionTargetPageId-1="57" captionTargetPageId-2="59" captionTargetPageId-3="61" captionTargetPageId-4="63" captionText-0="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; AB = male holotype (CASENT_9082245; FMNH); CD = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." captionText-1="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." captionText-2="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." captionText-3="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." captionText-4="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi-0="http://doi.org/10.5281/zenodo.14001843" figureDoi-1="http://doi.org/10.5281/zenodo.14001845" figureDoi-2="http://doi.org/10.5281/zenodo.14001847" figureDoi-3="http://doi.org/10.5281/zenodo.14001849" figureDoi-4="http://doi.org/10.5281/zenodo.14001851" httpUri-0="https://zenodo.org/record/14001843/files/figure.png" httpUri-1="https://zenodo.org/record/14001845/files/figure.png" httpUri-2="https://zenodo.org/record/14001847/files/figure.png" httpUri-3="https://zenodo.org/record/14001849/files/figure.png" httpUri-4="https://zenodo.org/record/14001851/files/figure.png" pageId="49" pageNumber="50">Figs 2226</figureCitation>
</paragraph>
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<subSubSection id="F1E40297FFC9FFFBFF170E5EFEB5D9E9" pageId="49" pageNumber="50" type="type_taxon">
<paragraph id="B941511CFFC9FFFBFF170E5EFE8DD99A" blockId="49.[189,344,1658,1684]" box="[189,344,1658,1684]" pageId="49" pageNumber="50">
<heading id="E209E670FFC9FFFBFF170E5EFE8DD99A" bold="true" box="[189,344,1658,1684]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="8B8A8D0EFFC9FFFBFF170E5EFE8DD99A" bold="true" box="[189,344,1658,1684]" pageId="49" pageNumber="50">
<typeStatus id="6645EFBEFFC9FFFBFF170E5EFF2FD99A" box="[189,250,1658,1684]" pageId="49" pageNumber="50">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC9FFFBFF170E8DFEB5D9E9" blockId="49.[187,1397,1705,1768]" pageId="49" pageNumber="50">
<taxonomicName id="7EFE2A9FFFC9FFFBFF170E8DFDB6D9CD" ID-CoL="7GH4W" authority="Strand, 1907" authorityName="Strand" authorityYear="1907" box="[189,611,1705,1731]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="8B8A8D0EFFC9FFFBFF170E8DFE19D9CD" box="[189,460,1705,1731]" italics="true" pageId="49" pageNumber="50">Zorodictyna intermedia</emphasis>
<bibRefCitation id="DD6F2CEDFFC9FFFBFE7A0E8DFDB6D9CD" author="Strand E." box="[464,611,1705,1731]" pageId="49" pageNumber="50" pagination="725 - 748" refId="ref38669" refString="Strand E. 1907. Diagnosen neuer Spinnen aus Madagaskar und Sansibar. Zoologischer Anzeiger 31: 725 - 748." type="journal article" year="1907">Strand, 1907</bibRefCitation>
</taxonomicName>
(= junior synonym of
<taxonomicName id="7EFE2A9FFFC9FFFBFCCA0E8DFB3CD9CD" ID-CoL="7GH58" authority="(Lenz, 1891)" authorityName="" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[864,1257,1705,1731]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFC9FFFBFCCA0E8DFB99D9CD" box="[864,1100,1705,1731]" italics="true" pageId="49" pageNumber="50">Zorodictyna oswaldi</emphasis>
(
<bibRefCitation id="DD6F2CEDFFC9FFFBFBF00E8DFB0AD9CD" author="Lenz H." box="[1114,1247,1705,1731]" pageId="49" pageNumber="50" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz, 1891</bibRefCitation>
)
</taxonomicName>
, see below), by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFC9FFF8FF170F2BFC62DE65" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="B941511CFFC9FFFBFF170F2BFEE3D827" blockId="49.[189,310,1807,1833]" box="[189,310,1807,1833]" pageId="49" pageNumber="50">
<heading id="E209E670FFC9FFFBFF170F2BFEE3D827" bold="true" box="[189,310,1807,1833]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="8B8A8D0EFFC9FFFBFF170F2BFEE3D827" bold="true" box="[189,310,1807,1833]" pageId="49" pageNumber="50">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC9FFF8FF170F1AFC62DE65" blockId="49.[187,1401,1854,2025]" lastBlockId="50.[189,1399,265,363]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
Cribellate udubids having an entire or vestigial cribellum and a strong scopulae beneath leg tarsi and most of length of metatarsi. The male palp (
<figureCitation id="21C54D99FFC9FFFBFDD20F46FD32D873" box="[632,743,1890,1917]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="49" pageNumber="50">Figs 23C</figureCitation>
,
<figureCitation id="21C54D99FFC9FFFBFD580F46FCF1D873" box="[754,804,1890,1917]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="49" pageNumber="50">25C</figureCitation>
) is provided with a short paracymbium retrolateral process (PP) (absent in
<taxonomicName id="7EFE2A9FFFC9FFFBFE700FA2FCADD8AF" authority="(Lenz, 1891))" authorityName="" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[474,888,1926,1953]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFC9FFFBFE700FA2FD19D8AE" box="[474,716,1926,1952]" italics="true" pageId="49" pageNumber="50">Zorodictyna oswaldi</emphasis>
(
<bibRefCitation id="DD6F2CEDFFC9FFFBFD740FA3FCB0D8AF" author="Lenz H." box="[734,869,1927,1953]" pageId="49" pageNumber="50" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz, 1891</bibRefCitation>
))
</taxonomicName>
and the female epigyne has a median lobe (ML) broad, nearly rectangular, not extending to posterior margin of epigyne (
<figureCitation id="21C54D99FFC9FFFBFBEA0F8EFB7AD8CA" box="[1088,1199,1962,1988]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="49" pageNumber="50">Figs 23D</figureCitation>
,
<figureCitation id="21C54D99FFC9FFFBFB130F8EFB3AD8CB" box="[1209,1263,1962,1989]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="49" pageNumber="50">25D</figureCitation>
). The genus is distinguished from
<taxonomicName id="7EFE2A9FFFC9FFFBFE6F0FEAFDC0D8E6" authorityName="Simon" authorityYear="1880" box="[453,533,1998,2024]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFC9FFFBFE6F0FEAFDC0D8E6" box="[453,533,1998,2024]" italics="true" pageId="49" pageNumber="50">Uduba</emphasis>
</taxonomicName>
males in lacking a threadlike embolus and a knob-like process between palpal patella and tibia and from
<taxonomicName id="7EFE2A9FFFCAFFF8FDEE092DFD41DE2D" authorityName="Simon" authorityYear="1880" box="[580,660,265,291]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFCAFFF8FDEE092DFD41DE2D" box="[580,660,265,291]" italics="true" pageId="50" pageNumber="51">Uduba</emphasis>
</taxonomicName>
females by having compact spermathecae rather than multiple loops. It is distinguished from
<taxonomicName id="7EFE2A9FFFCAFFF8FD8F0909FD44DE49" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[549,657,301,327]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8B8A8D0EFFCAFFF8FD8F0909FD44DE49" box="[549,657,301,327]" italics="true" pageId="50" pageNumber="51">Tabiboka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCAFFF8FD330909FCD1DE49" box="[665,772,301,327]" pageId="50" pageNumber="51" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="7EFE2A9FFFCAFFF8FC94090AFC7FDE49" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[830,938,302,327]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8B8A8D0EFFCAFFF8FC94090AFC7FDE49" box="[830,938,302,327]" italics="true" pageId="50" pageNumber="51">Zorascar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCAFFF8FC1A0909FBC9DE49" box="[944,1052,301,327]" pageId="50" pageNumber="51" rank="genus">gen. nov.</taxonomicNameLabel>
by having a strong tarsal and metatarsal scopulae and having a cribellum rather than a colulus.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFCAFFF9FF1709B7FC10DA52" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="description">
<paragraph id="B941511CFFCAFFF8FF1709B7FE9BDEA3" blockId="50.[189,334,403,429]" box="[189,334,403,429]" pageId="50" pageNumber="51">
<heading id="E209E670FFCAFFF8FF1709B7FE9BDEA3" bold="true" box="[189,334,403,429]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="8B8A8D0EFFCAFFF8FF1709B7FE9BDEA3" bold="true" box="[189,334,403,429]" pageId="50" pageNumber="51">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCAFFF8FF1709E6FD0CDD46" blockId="50.[189,1400,450,585]" pageId="50" pageNumber="51">
BODY. Large spiders (males: 13.0024.00; females: 19.0031.00) with smooth teguments of carapace, sternum and legs (
<figureCitation id="21C54D99FFCAFFF8FE3D09C2FE26DD0E" box="[407,499,486,512]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; AB = male holotype (CASENT_9082245; FMNH); CD = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="50" pageNumber="51">Figs 22</figureCitation>
,
<figureCitation id="21C54D99FFCAFFF8FDAA09C2FDF4DD0E" box="[512,545,486,512]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="50" pageNumber="51">24</figureCitation>
). Carapace piriform, longer than wide (L/W: males: 1.121.50; females: 1.241.41), with thoracic fovea linear; with sparse cover of short, dark setae; profile higher in ocular and thoracic area and abruptly sloping posteriorly.
</paragraph>
<paragraph id="B941511CFFCAFFF8FF170A55FEC6DDDC" blockId="50.[189,1400,624,722]" pageId="50" pageNumber="51">
COLORATION (
<figureCitation id="21C54D99FFCAFFF8FEC90A54FE68DD85" box="[355,445,624,651]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; AB = male holotype (CASENT_9082245; FMNH); CD = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="50" pageNumber="51">Figs 22</figureCitation>
,
<figureCitation id="21C54D99FFCAFFF8FE630A55FE3FDD85" box="[457,490,625,651]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="50" pageNumber="51">24</figureCitation>
). Cephalothorax brown dorsally, yellow-brown ventrally; abdomen dark with obscure pattern (
<taxonomicName id="7EFE2A9FFFCAFFF8FE2C0AB1FE3BDDA0" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[390,494,660,686]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFCAFFF8FE2C0AB1FE3BDDA0" box="[390,494,660,686]" italics="true" pageId="50" pageNumber="51">Z. almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCAFFF8FE530AB1FD82DDA1" box="[505,599,661,687]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="7EFE2A9FFFCAFFF8FDCF0AB1FCC6DDA0" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[613,787,660,686]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="silvadavilae" status="sp. nov.">
<emphasis id="8B8A8D0EFFCAFFF8FDCF0AB1FCC6DDA0" box="[613,787,660,686]" italics="true" pageId="50" pageNumber="51">Z. silvadavilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCAFFF8FCB10AB1FCACDDA1" box="[795,889,661,687]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
) or with bold markings (
<taxonomicName id="7EFE2A9FFFCAFFF8FB0F0AB1FEDBDDDC" authority="(Lenz, 1891))" authorityName="" baseAuthorityName="Lenz" baseAuthorityYear="1891" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCAFFF8FB0F0AB1FAF7DDA0" box="[1189,1314,660,686]" italics="true" pageId="50" pageNumber="51">Z. oswaldi</emphasis>
(
<bibRefCitation id="DD6F2CEDFFCAFFF8FA9E0AB1FF2FDDDC" author="Lenz H." pageId="50" pageNumber="51" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz, 1891</bibRefCitation>
))
</taxonomicName>
.
</paragraph>
<paragraph id="B941511CFFCAFFF8FF170ADFFBAEDC37" blockId="50.[189,1398,762,825]" pageId="50" pageNumber="51">
EYES (
<figureCitation id="21C54D99FFCAFFF8FEAD0ADEFE4CDC1B" box="[263,409,762,789]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; AB = male holotype (CASENT_9082245; FMNH); CD = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="50" pageNumber="51">Figs 22A, C</figureCitation>
,
<figureCitation id="21C54D99FFCAFFF8FE0F0ADFFE2BDC1B" box="[421,510,763,789]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="50" pageNumber="51">24A, C</figureCitation>
). In two nearly straight rows: ALE larger than AME, PLE and PME similar to ALE; tapetum of secondary eyes canoe shaped, shiny. Clypeus with long setae.
</paragraph>
<paragraph id="B941511CFFCAFFF8FF170B45FAADDB21" blockId="50.[188,1400,865,1071]" pageId="50" pageNumber="51">
PROSOMA. Chilum double, about three times or more as wide as high. Chelicerae: paturon conical, with some long setae medially, three teeth on both margins, with retrolateral thick setae at fang base; fang with serrate texture on proximal edge. Labium (
<figureCitation id="21C54D99FFCAFFF8FD400B8DFCAEDCCD" box="[746,891,937,963]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; AB = male holotype (CASENT_9082245; FMNH); CD = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="50" pageNumber="51">Figs 22B, D</figureCitation>
,
<figureCitation id="21C54D99FFCAFFF8FC2D0B8DFC0EDCCD" box="[903,987,937,963]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="50" pageNumber="51">24B, D</figureCitation>
) rectangular. Endites (
<figureCitation id="21C54D99FFCAFFF8FB480B8DFAA6DCCD" box="[1250,1395,937,963]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; AB = male holotype (CASENT_9082245; FMNH); CD = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="50" pageNumber="51">Figs 22B, D</figureCitation>
,
<figureCitation id="21C54D99FFCAFFF8FF160BE9FEC7DCE9" box="[188,274,973,999]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="50" pageNumber="51">24B, D</figureCitation>
) roughly rectangular, with retrolateral margins sinuous, distal part slightly wider than proximal part, with subdistal serrula present as simple row. Sternum (
<figureCitation id="21C54D99FFCAFFF8FC2B0BD4FBC3DB05" box="[897,1046,1008,1035]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; AB = male holotype (CASENT_9082245; FMNH); CD = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="50" pageNumber="51">Figs 22B, D</figureCitation>
,
<figureCitation id="21C54D99FFCAFFF8FB890BD5FBACDB05" box="[1059,1145,1009,1035]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="50" pageNumber="51">24B, D</figureCitation>
) oval, slightly longer than wide, length 1.221.33 times width, with slightly curved margins, not extending between coxae IV.
</paragraph>
<paragraph id="B941511CFFCAFFF8FF170C73FE5DDABA" blockId="50.[188,1400,1111,1461]" pageId="50" pageNumber="51">
LEGS (
<figureCitation id="21C54D99FFCAFFF8FEA30C73FEB6DB7F" box="[265,355,1111,1137]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; AB = male holotype (CASENT_9082245; FMNH); CD = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="50" pageNumber="51">Figs 22</figureCitation>
,
<figureCitation id="21C54D99FFCAFFF8FEC50C73FE45DB7F" box="[367,400,1111,1137]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="50" pageNumber="51">24</figureCitation>
). Slender. Formula 4123 or 1423. Three tarsal claws, unpaired claw smooth, small to absent, superior claws with six to eight teeth; without claw tufts or tenent setae; scopulae present beneath all tarsi and distal parts of metatarsi. Trichobothria in two or more rows on tarsi, in three or more on metatarsi, all equal in length; distal and proximal plates of bothria well differentiated, proximal plates with transverse ridges (
<bibRefCitation id="DD6F2CEDFFCAFFF8FE640CC2FD51DA0F" author="Griswold C. E." box="[462,644,1254,1281]" pageId="50" pageNumber="51" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: fig. 62). Tarsal organ with tear-drop or keyhole-shaped opening (
<bibRefCitation id="DD6F2CEDFFCAFFF8FF6F0D2EFEA9DA2B" author="Griswold C. E." box="[197,380,1290,1317]" pageId="50" pageNumber="51" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: fig. 63), close to tip of tarsus. Spines numerous, with four pairs of ventral spines on tibia I, not overlapping, and three pairs of ventral spines on metatarsus. Calamistrum present, oval in basal third of metatarsus. Male tibial crack (autospasy suture) present in males as a suture through the bases of leg tibia just distad to the basalmost pair of ventral spines (
<figureCitation id="21C54D99FFCAFFF8FC690D52FBFEDA9F" box="[963,1067,1398,1425]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="50" pageNumber="51">Fig. 24B</figureCitation>
, arrow). Trochanters distally notched to entire.
</paragraph>
<paragraph id="B941511CFFCAFFF8FF170DF9FEEBD9FC" blockId="50.[187,1399,1500,1779]" pageId="50" pageNumber="51">
ABDOMEN (
<figureCitation id="21C54D99FFCAFFF8FEEA0DF8FE42DAF9" box="[320,407,1500,1527]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; AB = male holotype (CASENT_9082245; FMNH); CD = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="50" pageNumber="51">Figs 22</figureCitation>
,
<figureCitation id="21C54D99FFCAFFF8FE0B0DF9FE17DAF9" box="[417,450,1501,1527]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="50" pageNumber="51">24</figureCitation>
). Oval, without scuta. Epiandrous glands absent. Cribellum entire. Spinnerets:ALS and PLS large, conical, biarticulate; PMS one-segmented, small. Females with two ALS major ampullate gland spigots and field of about 1020 piriform gland spigots and tartipores; PMS with two anterior minor ampullate gland spigots and posterior cylindrical gland spigots and aciniform gland spigots; PLS with few cylindrical gland spigots and aciniform gland spigots, apically with unique spigot, probably “modified” or “pseudoflagelliform” gland spigot; males lack cylindrical gland spigots and have “modified” gland spigot and second of each pair of major ampullate gland and minor ampullate gland spigots spigot reduced to nubbins.
</paragraph>
<paragraph id="B941511CFFCAFFF9FF170F3FFBA7DEBF" blockId="50.[187,1401,1818,2025]" lastBlockId="51.[189,1401,265,434]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
MALE PALP (
<figureCitation id="21C54D99FFCAFFF8FEE60F3EFE09D83B" box="[332,476,1818,1845]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="50" pageNumber="51">Figs 23AC</figureCitation>
,
<figureCitation id="21C54D99FFCAFFF8FE4C0F3EFDEAD83B" box="[486,575,1818,1845]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="50" pageNumber="51">25AC</figureCitation>
;
<bibRefCitation id="DD6F2CEDFFCAFFF8FDE30F3FFD18D83B" author="Lenz H." box="[585,717,1819,1845]" pageId="50" pageNumber="51" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">LENZ 1891</bibRefCitation>
:
<figureCitation id="21C54D99FFCAFFF8FD7D0F3FFC93D83A" box="[727,838,1818,1845]" captionStart="Fig" captionStartId="19.[188,230,1787,1813]" captionTargetBox="[188,1399,479,1749]" captionTargetId="figure-89@19.[188,1399,479,1749]" captionTargetPageId="19" captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AC = 0.43 mm; DF = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick." figureDoi="http://doi.org/10.5281/zenodo.14001804" httpUri="https://zenodo.org/record/14001804/files/figure.png" pageId="50" pageNumber="51">FIG. 6bc</figureCitation>
;
<bibRefCitation id="DD6F2CEDFFCAFFF8FCFA0F3EFC2BD83A" author="Lehtinen P. T." box="[848,1022,1818,1844]" pageId="50" pageNumber="51" pagination="199 - 468" refId="ref37164" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen 1967</bibRefCitation>
: fig. 78;
<bibRefCitation id="DD6F2CEDFFCAFFF8FBF50F3EFAC4D83B" author="Griswold C. E." box="[1119,1297,1818,1845]" pageId="50" pageNumber="51" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: figs 19 21). Tibia with large or stout RTA, inserted medially, provided with basal concavity, with short rounded or tapered VTA; cymbium oval with rounded distal extremity, without dorsal patch of short setae, with rounded paracymbial retrolateral process (PP) (absent in
<taxonomicName id="7EFE2A9FFFCAFFF8FCC10FA2FB88D8AE" authorityName="" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[875,1117,1926,1952]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCAFFF8FCC10FA2FB88D8AE" box="[875,1117,1926,1952]" italics="true" pageId="50" pageNumber="51">Zorodictyna oswaldi</emphasis>
</taxonomicName>
) and conical retrobasal extension (CE); bulbus provided with retrobasal subtegulum lobe (locking lobe, SL), and tegulum rounded with well-developed prolateral conical protrusion at base of embolus (corresponding locking lobe, TL); embolus (E) with fixed base, originating prolateraly, directed clockwise, short and lamellate (
<figureCitation id="21C54D99FFCBFFF9FB48092DFAA7DE2D" box="[1250,1394,265,291]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="51" pageNumber="52">Figs 23AB</figureCitation>
,
<figureCitation id="21C54D99FFCBFFF9FF170908FECCDE49" box="[189,281,300,327]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="51" pageNumber="52">25AB</figureCitation>
); median apophysis (MA) hook-shaped, situated opposite to E; conductor (C) hyaline, well developed, fan-shaped, medially extending from narrow base to wide flattened tip; paraconductor (Pc) as narrow, finger-like membranous extension inserted between tegulum and base of MA with sharp triangular dorso-prolateral process (EBP) arising near bases of embolus and paraconductor.
</paragraph>
<paragraph id="B941511CFFCBFFF9FF1709FDFD5DDD8F" blockId="51.[187,1399,473,642]" pageId="51" pageNumber="52">
GENITALIA. Epigyne (
<figureCitation id="21C54D99FFCBFFF9FE1609FDFDFADEFD" box="[444,559,473,499]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="51" pageNumber="52">Figs 23D</figureCitation>
,
<figureCitation id="21C54D99FFCBFFF9FD9709FDFDA6DEFA" box="[573,627,473,500]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="51" pageNumber="52">25D</figureCitation>
;
<bibRefCitation id="DD6F2CEDFFCBFFF9FD2B09FEFCD6DEFD" author="Lenz H." box="[641,771,473,500]" pageId="51" pageNumber="52" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz 1891</bibRefCitation>
: fig. 8b;
<bibRefCitation id="DD6F2CEDFFCBFFF9FCC509FDFBCADEFD" author="Lehtinen P. T." box="[879,1055,473,499]" pageId="51" pageNumber="52" pagination="199 - 468" refId="ref37164" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen 1967</bibRefCitation>
: figs 72, 76;
<bibRefCitation id="DD6F2CEDFFCBFFF9FB1709FDFAA7DEFD" author="Griswold C. E." box="[1213,1394,473,499]" pageId="51" pageNumber="52" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: figs 2224) as single, shield-shaped plate (ML) with convex median and lateral folds; lateral sector convex or with tooth. Endogyne (
<figureCitation id="21C54D99FFCBFFF9FE480A04FD9BDD34" box="[482,590,544,570]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="51" pageNumber="52">Figs 23E</figureCitation>
,
<figureCitation id="21C54D99FFCBFFF9FDF20A04FD59DD35" box="[600,652,544,571]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="51" pageNumber="52">25E</figureCitation>
): copulatory ducts (CD) very short, twisted; spermatheca (S) one or two lobate, head of spermatheca provided with one or more pores; no evidence of enlarged Bennetts Gland pore through spermathecal base.
</paragraph>
<paragraph id="B941511CFFCBFFF9FF170A8DFE59DDCD" blockId="51.[189,396,681,707]" box="[189,396,681,707]" pageId="51" pageNumber="52">
<heading id="E209E670FFCBFFF9FF170A8DFE59DDCD" bold="true" box="[189,396,681,707]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="8B8A8D0EFFCBFFF9FF170A8DFE59DDCD" bold="true" box="[189,396,681,707]" pageId="51" pageNumber="52">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCBFFF9FF170AFCFEB2DC18" blockId="51.[189,1398,728,790]" pageId="51" pageNumber="52">
<taxonomicName id="7EFE2A9FFFCBFFF9FF170AFCFE67DDFC" authorityName="" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[189,434,728,754]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCBFFF9FF170AFCFE67DDFC" box="[189,434,728,754]" italics="true" pageId="51" pageNumber="52">Zorodictyna oswaldi</emphasis>
</taxonomicName>
(
<typeStatus id="6645EFBEFFCBFFF9FE690AFCFE2DDDFC" box="[451,504,728,754]" pageId="51" pageNumber="52">type</typeStatus>
species,
<specimenCount id="AFF89A95FFCBFFF9FDCC0AFCFD54DDFC" box="[614,641,728,754]" count="1" pageId="51" pageNumber="52" type="male"></specimenCount>
<specimenCount id="AFF89A95FFCBFFF9FD2B0AFCFD41DDFC" box="[641,660,728,754]" count="1" pageId="51" pageNumber="52" type="female"></specimenCount>
);
<taxonomicName id="7EFE2A9FFFCBFFF9FD040AFCFC5BDDFC" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[686,910,728,754]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFCBFFF9FD040AFCFC5BDDFC" box="[686,910,728,754]" italics="true" pageId="51" pageNumber="52">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCBFFF9FC320AFDFC20DDFD" box="[920,1013,729,755]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCount id="AFF89A95FFCBFFF9FBA20AFCFBF7DDFC" box="[1032,1058,728,754]" count="1" pageId="51" pageNumber="52" type="male"></specimenCount>
<specimenCount id="AFF89A95FFCBFFF9FB880AFCFBE1DDFC" box="[1058,1076,728,754]" count="1" pageId="51" pageNumber="52" type="female"></specimenCount>
);
<taxonomicName id="7EFE2A9FFFCBFFF9FBE50AFCFAA3DDFC" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1103,1398,728,754]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="silvadavilae" status="sp. nov.">
<emphasis id="8B8A8D0EFFCBFFF9FBE50AFCFAA3DDFC" box="[1103,1398,728,754]" italics="true" pageId="51" pageNumber="52">Zorodictyna silvadavilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCBFFF9FF150AD8FECCDC18" box="[191,281,764,790]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCount id="AFF89A95FFCBFFF9FE810AD8FE90DC18" box="[299,325,764,790]" count="1" pageId="51" pageNumber="52" type="male"></specimenCount>
<specimenCount id="AFF89A95FFCBFFF9FEEF0AD8FE82DC18" box="[325,343,764,790]" count="1" pageId="51" pageNumber="52" type="female"></specimenCount>
).
</paragraph>
<paragraph id="B941511CFFCBFFF9FF170B1AFE32DC59" blockId="51.[189,487,829,856]" box="[189,487,829,856]" pageId="51" pageNumber="52">
<heading id="E209E670FFCBFFF9FF170B1AFE32DC59" bold="true" box="[189,487,829,856]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="8B8A8D0EFFCBFFF9FF170B1AFE32DC59" bold="true" box="[189,487,829,856]" pageId="51" pageNumber="52">
Note on the
<typeStatus id="6645EFBEFFCBFFF9FEFE0B1AFE5FDC56" box="[340,394,830,856]" pageId="51" pageNumber="52">type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCBFFF9FF170B48FD14DBC9" blockId="51.[187,1399,876,1223]" pageId="51" pageNumber="52">
The first species name used in
<taxonomicName id="7EFE2A9FFFCBFFF9FD8E0B48FD67DC88" authorityName="Strand" authorityYear="1907" box="[548,690,876,902]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFCBFFF9FD8E0B48FD67DC88" box="[548,690,876,902]" italics="true" pageId="51" pageNumber="52">Zorodictyna</emphasis>
</taxonomicName>
was
<taxonomicName id="7EFE2A9FFFCBFFF9FD450B48FB76DC89" authority="Strand, 1907" authorityName="Strand" authorityYear="1907" box="[751,1187,876,903]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="8B8A8D0EFFCBFFF9FD450B48FBD6DC88" box="[751,1027,876,902]" italics="true" pageId="51" pageNumber="52">Zorodictyna intermedia</emphasis>
<bibRefCitation id="DD6F2CEDFFCBFFF9FBA10B48FB76DC89" author="Strand E." box="[1035,1187,876,903]" pageId="51" pageNumber="52" pagination="725 - 748" refId="ref38669" refString="Strand E. 1907. Diagnosen neuer Spinnen aus Madagaskar und Sansibar. Zoologischer Anzeiger 31: 725 - 748." type="journal article" year="1907">Strand, 1907</bibRefCitation>
</taxonomicName>
, which is the
<typeStatus id="6645EFBEFFCBFFF9FAEE0B49FAA2DC89" box="[1348,1399,877,903]" pageId="51" pageNumber="52">type</typeStatus>
species by monotypy. However, there is good reason to consider Z.
<taxonomicName id="7EFE2A9FFFCBFFF9FC5A0BB4FAF2DCA5" authority="Strand, 1907 a" authorityName="Strand" authorityYear="1907" box="[1008,1319,912,939]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="8B8A8D0EFFCBFFF9FC5A0BB4FBA5DCA4" box="[1008,1136,912,938]" italics="true" pageId="51" pageNumber="52">intermedia</emphasis>
<bibRefCitation id="DD6F2CEDFFCBFFF9FBD20BB4FAC4DCA4" author="Strand E." box="[1144,1297,912,938]" pageId="51" pageNumber="52" pagination="725 - 748" refId="ref38669" refString="Strand E. 1907. Diagnosen neuer Spinnen aus Madagaskar und Sansibar. Zoologischer Anzeiger 31: 725 - 748." type="journal article" year="1907">Strand, 1907</bibRefCitation>
a
</taxonomicName>
junior synonym of
<taxonomicName id="7EFE2A9FFFCBFFF9FEE20B90FD71DCC0" authority="Simon, 1906" authorityName="Simon" authorityYear="1906" box="[328,676,948,974]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="inhonesta">
<emphasis id="8B8A8D0EFFCBFFF9FEE20B90FDD8DCC0" box="[328,525,948,974]" italics="true" pageId="51" pageNumber="52">Uduba inhonesta</emphasis>
<bibRefCitation id="DD6F2CEDFFCBFFF9FDB80B90FD71DCC0" author="Simon E." box="[530,676,948,974]" pageId="51" pageNumber="52" pagination="284 - 308" refId="ref38537" refString="Simon E. 1906. Etude sur les araignees de la section des cribellates. Annales de la Societe entomologique de Belgique 50: 284 - 308. https: // doi. org / 10.5962 / bhl. part. 19947" type="journal article" year="1906">Simon, 1906</bibRefCitation>
</taxonomicName>
(now
<taxonomicName id="7EFE2A9FFFCBFFF9FD400B90FCACDCC0" authorityName="Simon" authorityYear="1906" box="[746,889,948,974]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="inhonesta">
Z.
<emphasis id="8B8A8D0EFFCBFFF9FCA30B90FCACDCC0" box="[777,889,948,974]" italics="true" pageId="51" pageNumber="52">inhonesta</emphasis>
</taxonomicName>
), which was already recognized by
<bibRefCitation id="DD6F2CEDFFCBFFF9FABA0B90FED8DCFF" author="Lehtinen P. T." pageId="51" pageNumber="52" pagination="199 - 468" refId="ref37164" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen (1967)</bibRefCitation>
. Furthermore, we have carefully examined the
<typeStatus id="6645EFBEFFCBFFF9FCEA0BFCFCA6DCFC" box="[832,883,984,1010]" pageId="51" pageNumber="52">type</typeStatus>
specimens of
<taxonomicName id="7EFE2A9FFFCBFFF9FB890BFCFAA2DCFF" authority="Lenz, 1891" authorityName="Lenz" authorityYear="1891" box="[1059,1399,983,1010]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCBFFF9FB890BFCFB3CDCFF" box="[1059,1257,983,1010]" italics="true" pageId="51" pageNumber="52">Agroeca oswaldi</emphasis>
<bibRefCitation id="DD6F2CEDFFCBFFF9FB5B0BFCFAA2DCFF" author="Lenz H." box="[1265,1399,983,1010]" pageId="51" pageNumber="52" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz, 1891</bibRefCitation>
</taxonomicName>
(now
<taxonomicName id="7EFE2A9FFFCBFFF9FEAB0BD8FEAEDB1B" authorityName="Lenz" authorityYear="1891" box="[257,379,1019,1045]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCBFFF9FEAB0BD8FEAEDB1B" box="[257,379,1019,1045]" italics="true" pageId="51" pageNumber="52">Z. oswaldi</emphasis>
</taxonomicName>
) and
<taxonomicName id="7EFE2A9FFFCBFFF9FE140BD8FD9ADB1B" authorityName="Simon" authorityYear="1906" box="[446,591,1019,1045]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="inhonesta">
<emphasis id="8B8A8D0EFFCBFFF9FE140BD8FD9ADB1B" box="[446,591,1019,1045]" italics="true" pageId="51" pageNumber="52">Z. inhonesta</emphasis>
</taxonomicName>
: the distinction observed by
<bibRefCitation id="DD6F2CEDFFCBFFF9FC090BDFFB9BDB1B" author="Lehtinen P. T." box="[931,1102,1019,1045]" pageId="51" pageNumber="52" pagination="199 - 468" refId="ref37164" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen 1967</bibRefCitation>
was artifactual, resulting from a mating plug lodged in the epigyne of the
<typeStatus id="6645EFBEFFCBFFF9FCA90C3BFCE3DB37" box="[771,822,1055,1081]" pageId="51" pageNumber="52">type</typeStatus>
of
<taxonomicName id="7EFE2A9FFFCBFFF9FCF50C3BFC08DB36" authorityName="Lenz" authorityYear="1891" box="[863,989,1054,1081]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="59" phylum="Arthropoda" rank="species" species="oswaldi">
Z.
<emphasis id="8B8A8D0EFFCBFFF9FC280C3AFC08DB36" box="[898,989,1054,1080]" italics="true" pageId="51" pageNumber="52">oswaldi</emphasis>
</taxonomicName>
in contrast with the clean epigyne of the
<typeStatus id="6645EFBEFFCBFFF9FEA60C67FEEADB53" box="[268,319,1091,1117]" pageId="51" pageNumber="52">type</typeStatus>
of
<taxonomicName id="7EFE2A9FFFCBFFF9FEC30C66FD0DDB52" authority="Simon, 1906" authorityName="Simon" authorityYear="1906" box="[361,728,1090,1116]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="inhonesta">
<emphasis id="8B8A8D0EFFCBFFF9FEC30C66FDE1DB52" box="[361,564,1090,1116]" italics="true" pageId="51" pageNumber="52">Uduba inhonesta</emphasis>
<bibRefCitation id="DD6F2CEDFFCBFFF9FD970C66FD0DDB52" author="Simon E." box="[573,728,1090,1116]" pageId="51" pageNumber="52" pagination="284 - 308" refId="ref38537" refString="Simon E. 1906. Etude sur les araignees de la section des cribellates. Annales de la Societe entomologique de Belgique 50: 284 - 308. https: // doi. org / 10.5962 / bhl. part. 19947" type="journal article" year="1906">Simon, 1906</bibRefCitation>
</taxonomicName>
. We have established that
<taxonomicName id="7EFE2A9FFFCBFFF9FBBE0C67FA8FDB52" authority="(Simon, 1906)" baseAuthorityName="Simon" baseAuthorityYear="1906" box="[1044,1370,1090,1116]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="inhonesta">
<emphasis id="8B8A8D0EFFCBFFF9FBBE0C67FB72DB52" box="[1044,1191,1090,1116]" italics="true" pageId="51" pageNumber="52">Z. inhonesta</emphasis>
(
<bibRefCitation id="DD6F2CEDFFCBFFF9FB1D0C66FA84DB52" author="Simon E." box="[1207,1361,1090,1116]" pageId="51" pageNumber="52" pagination="284 - 308" refId="ref38537" refString="Simon E. 1906. Etude sur les araignees de la section des cribellates. Annales de la Societe entomologique de Belgique 50: 284 - 308. https: // doi. org / 10.5962 / bhl. part. 19947" type="journal article" year="1906">Simon, 1906</bibRefCitation>
)
</taxonomicName>
is itself a junior synonym of
<taxonomicName id="7EFE2A9FFFCBFFF9FE5C0C42FDA5DB8E" authorityName="Lenz" authorityYear="1891" box="[502,624,1126,1152]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCBFFF9FE5C0C42FDA5DB8E" box="[502,624,1126,1152]" italics="true" pageId="51" pageNumber="52">Z. oswaldi</emphasis>
</taxonomicName>
. Therefore, the
<typeStatus id="6645EFBEFFCBFFF9FC860C42FC8ADB8E" box="[812,863,1126,1152]" pageId="51" pageNumber="52">type</typeStatus>
species of
<taxonomicName id="7EFE2A9FFFCBFFF9FC4E0C42FAC5DB8E" authority="Strand, 1907" authorityName="Strand" authorityYear="1907" box="[996,1296,1126,1152]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFCBFFF9FC4E0C42FBA7DB8E" box="[996,1138,1126,1152]" italics="true" pageId="51" pageNumber="52">Zorodictyna</emphasis>
<bibRefCitation id="DD6F2CEDFFCBFFF9FBD30C42FAC5DB8E" author="Strand E." box="[1145,1296,1126,1152]" pageId="51" pageNumber="52" pagination="725 - 748" refId="ref38669" refString="Strand E. 1907. Diagnosen neuer Spinnen aus Madagaskar und Sansibar. Zoologischer Anzeiger 31: 725 - 748." type="journal article" year="1907">Strand, 1907</bibRefCitation>
</taxonomicName>
is also a junior synonym of
<taxonomicName id="7EFE2A9FFFCBFFF9FE370CAEFDC2DBAD" authorityName="Lenz" authorityYear="1891" box="[413,535,1161,1188]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCBFFF9FE370CAEFE78DBAD" box="[413,429,1162,1187]" italics="true" pageId="51" pageNumber="52">Z</emphasis>
.
<emphasis id="8B8A8D0EFFCBFFF9FE160CADFDC2DBAD" box="[444,535,1161,1187]" italics="true" pageId="51" pageNumber="52">oswaldi</emphasis>
</taxonomicName>
. For more complete details concerning the synonymy, see the description of
<taxonomicName id="7EFE2A9FFFCBFFF9FF770C89FDB8DBC9" authority="(Lenz, 1891)" authorityName="" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[221,621,1197,1223]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCBFFF9FF770C89FE18DBC9" box="[221,461,1197,1223]" italics="true" pageId="51" pageNumber="52">Zorodictyna oswaldi</emphasis>
(
<bibRefCitation id="DD6F2CEDFFCBFFF9FE770C89FDB1DBC9" author="Lenz H." box="[477,612,1197,1223]" pageId="51" pageNumber="52" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz, 1891</bibRefCitation>
)
</taxonomicName>
below.
</paragraph>
<paragraph id="B941511CFFCBFFF9FF170CCAFEF9DA06" blockId="51.[189,300,1262,1288]" box="[189,300,1262,1288]" pageId="51" pageNumber="52">
<heading id="E209E670FFCBFFF9FF170CCAFEF9DA06" bold="true" box="[189,300,1262,1288]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="8B8A8D0EFFCBFFF9FF170CCAFEF9DA06" bold="true" box="[189,300,1262,1288]" pageId="51" pageNumber="52">Affinities</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCBFFF9FF170D3AFC10DA52" blockId="51.[188,1399,1309,1372]" pageId="51" pageNumber="52">
From the molecular analyses (
<figureCitation id="21C54D99FFCBFFF9FDB30D39FD8ADA36" box="[537,607,1309,1336]" captionStart="Fig" captionStartId="9.[189,233,1185,1211]" captionTargetBox="[193,1394,271,1139]" captionTargetId="figure-11@9.[189,1398,265,1148]" captionTargetPageId="9" captionText="Fig. 2. Phylogenetic reconstruction based on the maximum likelihood (MLi) analysis performed on the COI16S dataset, showing the relationships between Zorascar gen. nov. and other relatives of the family Udubidae Griswold &amp; Polotow, 2015 and Zoropsidae Bertkau,1882. The tree is congruent in its topology (for supported nodes) with the Bayesian analyses of the same dataset. Nodal support values (MLi Bootstraps and posterior probabilities (PPr) are indicated as following = MLi/PPr). Values below 50/0.5 are not shown or indicated by hyphens." figureDoi="http://doi.org/10.5281/zenodo.14001792" httpUri="https://zenodo.org/record/14001792/files/figure.png" pageId="51" pageNumber="52">Fig. 2</figureCitation>
),
<taxonomicName id="7EFE2A9FFFCBFFF9FDD80D39FCD5DA39" authorityName="Strand" authorityYear="1907" box="[626,768,1309,1335]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFCBFFF9FDD80D39FCD5DA39" box="[626,768,1309,1335]" italics="true" pageId="51" pageNumber="52">Zorodictyna</emphasis>
</taxonomicName>
appears closely related to species in the
<taxonomicName id="7EFE2A9FFFCBFFF9FB790D39FA90DA39" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[1235,1349,1309,1335]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="family">Udubidae</taxonomicName>
that have been identified as
<taxonomicName id="7EFE2A9FFFCBFFF9FE7A0D65FDE9DA55" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[464,572,1345,1371]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8B8A8D0EFFCBFFF9FE7A0D65FDE9DA55" box="[464,572,1345,1371]" italics="true" pageId="51" pageNumber="52">Tabiboka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCBFFF9FDEE0D66FD7ADA52" box="[580,687,1346,1372]" pageId="51" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="7EFE2A9FFFCBFFF9FD4D0D66FC86DA55" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[743,851,1346,1371]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8B8A8D0EFFCBFFF9FD4D0D66FC86DA55" box="[743,851,1346,1371]" italics="true" pageId="51" pageNumber="52">Zorascar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCBFFF9FCF00D66FC10DA52" box="[858,965,1346,1372]" pageId="51" pageNumber="52" rank="genus">nov. gen.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFCBFFF9FF170DA7FC61DAC2" box="[189,948,1411,1484]" pageId="51" pageNumber="52" type="distribution">
<paragraph id="B941511CFFCBFFF9FF170DA7FE83DA93" blockId="51.[189,342,1411,1437]" box="[189,342,1411,1437]" pageId="51" pageNumber="52">
<heading id="E209E670FFCBFFF9FF170DA7FE83DA93" bold="true" box="[189,342,1411,1437]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="8B8A8D0EFFCBFFF9FF170DA7FE83DA93" bold="true" box="[189,342,1411,1437]" pageId="51" pageNumber="52">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCBFFF9FF170D96FC61DAC2" blockId="51.[189,948,1458,1484]" box="[189,948,1458,1484]" pageId="51" pageNumber="52">
Species of the genus
<taxonomicName id="7EFE2A9FFFCBFFF9FE1F0D96FD96DAC2" authorityName="Strand" authorityYear="1907" box="[437,579,1458,1484]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFCBFFF9FE1F0D96FD96DAC2" box="[437,579,1458,1484]" italics="true" pageId="51" pageNumber="52">Zorodictyna</emphasis>
</taxonomicName>
occur in
<collectingCountry id="C1E9118CFFCBFFF9FD1B0D96FCEADAC2" box="[689,831,1458,1484]" name="Madagascar" pageId="51" pageNumber="52">Madagascar</collectingCountry>
(
<figureCitation id="21C54D99FFCBFFF9FCE50D96FC72DAC2" box="[847,935,1458,1484]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="51" pageNumber="52">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>