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<document id="E7605602536236A9329A58DE057A1EF0" ID-DOI="10.5852/ejt.2024.951.2645" ID-ISSN="2118-9773" ID-Zenodo-Dep="13735747" ID-ZooBank="B17B7AE4-76A6-499C-A5E9-62A14D898484" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725878715189" checkinUser="plazi" docAuthor="Zouhair, Laila, Grootaert, Patrick &amp; Kettani, Kawtar" docDate="2024" docId="314BDF25FF90F23780546D8ADAAFF9D8" docLanguage="en" docName="EJT.2024.951.1-53.pdf" docOrigin="European Journal of Taxonomy 951 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2645/12209" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Platypalpus verbekei Grootaert &amp; Chvala 1992" docType="treatment" docVersion="2" lastPageNumber="49" masterDocId="CD72A75DFFBFF20782546B2CD825FFB2" masterDocTitle="Twelve new species of Platypalpus Macquart (Diptera: Hybotidae) from Morocco, with additional new records" masterLastPageNumber="53" masterPageNumber="1" pageNumber="48" updateTime="1725879277050" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1AE991C9B72E34C3D2D785EF835466CD">Twelve new species of Platypalpus Macquart (Diptera: Hybotidae) from Morocco, with additional new records</mods:title>
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<mods:namePart id="FA39D10763ED019E9ACB46D2EF5B07FC">Zouhair, Laila</mods:namePart>
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<mods:affiliation id="7D6F37F36AE5B68F48E35EF5002F9068">Laboratory of Ecology, Systematics and Conservation of Biodiversity (LESCB), URL-CNRST N ° 18, FS, Abdelmalek Essaadi University, Tétouan, Morocco.</mods:affiliation>
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<mods:namePart id="B012F35F5AD00037ECA73FBC7F2CD4BC">Grootaert, Patrick</mods:namePart>
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<mods:affiliation id="08A03C7C5D07740316E8606D453582E4">Laboratory of Ecology, Systematics and Conservation of Biodiversity (LESCB), URL-CNRST N ° 18, FS, Abdelmalek Essaadi University, Tétouan, Morocco.</mods:affiliation>
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<taxonomicName id="7EE215B0FF90F22880546D8ADC11F973" ID-CoL="4JVDT" authority="Grootaert &amp; Chvala, 1992" authorityName="Grootaert &amp; Chvala" authorityYear="1992" box="[512,1076,1702,1729]" class="Insecta" family="Hybotidae" genus="Platypalpus" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="verbekei">
<emphasis id="8B96B221FF90F22880546D8ADADFF973" bold="true" box="[512,762,1702,1729]" italics="true" pageId="47" pageNumber="48">Platypalpus verbekei</emphasis>
<bibRefCitation id="DD7313C2FF90F22881556D8BDC11F973" author="Grootaert P. &amp; Chvala M." box="[769,1076,1702,1729]" pageId="47" pageNumber="48" pagination="1 - 226" refId="ref24608" refString="Grootaert P. &amp; Chvala M. 1992. Monograph of the genus Platypalpus (Diptera: Empidoidea, Hybotidae) of the Mediterranean region and the Canary Islands. Acta Universitatis Carolinae - Biologica 36 (1 - 2): 1 - 226." type="journal article" year="1992">Grootaert &amp; Chvála, 1992</bibRefCitation>
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<collectingDate id="DD18B11BFF90F22880CE6C16DB0CF8E6" box="[666,809,1850,1876]" pageId="47" pageNumber="48" value="2018-04-08">8 Apr. 2018</collectingDate>
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<materialsCitation id="098A646EFF90F22886A16C16DB65F8C5" collectingDate="2019-01-08" collectionCode="LESCB" collectorName="K. Kettani" country="Morocco" location="Jnane Niche" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Rif">
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<materialsCitation id="098A646EFF90F228810D6C71D9FCF828" collectingDate="2019-12-19" collectionCode="LESCB" collectorName="K. Kettani" country="Morocco" location="Oued Afertane" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Rif">
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<materialsCitation id="098A646EFF90F22883A26CACDD01F828" box="[502,1316,1920,1947]" collectingDate="2021-04-20" collectionCode="LESCB" collectorName="L. Zouhair" country="Morocco" location="Oued Taida" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Rif">
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<collectingDate id="DD18B11BFF90F228809B6CACDB4AF828" box="[719,879,1920,1946]" pageId="47" pageNumber="48" value="2021-04-20">20 Apr. 2021</collectingDate>
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<materialsCitation id="098A646EFF90F22887146CACDB95F80F" collectingDate="2021-04-22" collectionCode="LESCB" collectorName="L. Zouhair" country="Morocco" location="Oued Tanina" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Rif">
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<collectingDate id="DD18B11BFF90F22883306C8FDA24F80F" box="[356,513,1955,1981]" pageId="47" pageNumber="48" value="2021-04-22">22 Apr. 2021</collectingDate>
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<collectorName id="14170BE5FF90F22880C76C88DB37F80F" box="[659,786,1955,1982]" pageId="47" pageNumber="48">L. Zouhair</collectorName>
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<materialsCitation id="098A646EFF90F228819F6C8FDA45F853" collectingDate="2021-05-02" collectionCode="RBINS" collectorName="L. Zouhair" country="Morocco" location="Oued Azoumagh" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Rif">
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<collectorName id="14170BE5FF90F22883116CEBD9E1F852" box="[325,452,1990,2017]" pageId="47" pageNumber="48">L. Zouhair</collectorName>
leg.;
<collectionCode id="DFF3F6F6FF90F228805E6CEBDA45F853" box="[522,608,1991,2017]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="47" pageNumber="48" type="Museum">RBINS</collectionCode>
</materialsCitation>
<materialsCitation id="098A646EFF90F22880296CEBDD54F852" box="[637,1393,1990,2017]" collectingDate="2023-01-05" collectorName="K. Kettani" country="Morocco" location="Briyech" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Rif">
<specimenCount id="AFE4A5BAFF90F22880296CEBDA8BF853" box="[637,686,1991,2017]" count="1" pageId="47" pageNumber="48" type="male">1 ♂</specimenCount>
; beach of
<location id="BC3D38E8FF90F22881786CEADBA8F852" LSID="urn:lsid:plazi:treatment:314BDF25FF90F23780546D8ADAAFF9D8:BC3D38E8FF90F22881786CEADBA8F852" box="[812,909,1990,2016]" country="Morocco" name="Briyech" pageId="47" pageNumber="48" stateProvince="Rif">Briyech</location>
;
<date id="CD5C48F3FF90F22881CE6CEADC0DF853" box="[922,1064,1990,2017]" pageId="47" pageNumber="48" value="2023-01-05">
<collectingDate id="DD18B11BFF90F22881CE6CEADC0DF853" box="[922,1064,1990,2017]" pageId="47" pageNumber="48" value="2023-01-05">5 Jan. 2023</collectingDate>
</date>
;
<collectingMethod id="60A31624FF90F22886626CEBDC8AF853" box="[1078,1199,1991,2017]" pageId="47" pageNumber="48">sweep net</collectingMethod>
;
<collectorName id="14170BE5FF90F22886EA6CEBDD1FF852" box="[1214,1338,1990,2017]" pageId="47" pageNumber="48">K. Kettani</collectorName>
leg.
</materialsCitation>
;
</paragraph>
<caption id="ED9D3EBBFF8FF23782E96F5EDD03FB3E" ID-DOI="http://doi.org/10.5281/zenodo.13735819" ID-Zenodo-Dep="13735819" box="[189,1318,1138,1164]" httpUri="https://zenodo.org/record/13735819/files/figure.png" pageId="48" pageNumber="49" startId="48.[189,232,1138,1164]" targetBox="[189,1399,265,1112]" targetPageId="48">
<paragraph id="B95D6E33FF8FF23782E96F5EDD03FB3E" blockId="48.[189,1318,1138,1164]" box="[189,1318,1138,1164]" pageId="48" pageNumber="49">
<emphasis id="8B96B221FF8FF23782E96F5ED93DFB3E" bold="true" box="[189,280,1138,1164]" pageId="48" pageNumber="49">Fig. 34.</emphasis>
<taxonomicName id="7EE215B0FF8FF23783746F5EDB77FB3E" authority="Grootaert &amp; Chvala, 1992" authorityName="Grootaert &amp; Chvala" authorityYear="1992" box="[288,850,1138,1164]" class="Insecta" family="Hybotidae" genus="Platypalpus" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="verbekei">
<emphasis id="8B96B221FF8FF23783746F5EDA33FB3E" box="[288,534,1138,1164]" italics="true" pageId="48" pageNumber="49">Platypalpus verbekei</emphasis>
<bibRefCitation id="DD7313C2FF8FF237804A6F5EDB77FB3E" author="Grootaert P. &amp; Chvala M." box="[542,850,1138,1164]" pageId="48" pageNumber="49" pagination="1 - 226" refId="ref24608" refString="Grootaert P. &amp; Chvala M. 1992. Monograph of the genus Platypalpus (Diptera: Empidoidea, Hybotidae) of the Mediterranean region and the Canary Islands. Acta Universitatis Carolinae - Biologica 36 (1 - 2): 1 - 226." type="journal article" year="1992">Grootaert &amp; Chvála, 1992</bibRefCitation>
</taxonomicName>
, ♂, habitus (Oued Azoumagh, RBINS).
</paragraph>
</caption>
<paragraph id="B95D6E33FF8FF23782E96FE2DB80FAB9" blockId="48.[189,1398,1230,1292]" pageId="48" pageNumber="49">
<collectionCode id="DFF3F6F6FF8FF23782E96FE2D93DFB5A" box="[189,280,1230,1256]" pageId="48" pageNumber="49">LESCB</collectionCode>
<materialsCitation id="098A646EFF8FF23783666FE2DD0DFB5A" box="[306,1320,1230,1257]" collectingDate="2023-01-09" collectionCode="LESCB" collectorName="Briyech &amp; K. Kettani &amp; High Atlas" pageId="48" pageNumber="49" specimenCount="1" specimenCount-male="1">
<specimenCount id="AFE4A5BAFF8FF23783666FE2D947FB5B" box="[306,354,1230,1257]" count="1" pageId="48" pageNumber="49" type="male">1 ♂</specimenCount>
; beach of
<collectorName id="14170BE5FF8FF237838F6FE2DA19FB5A" box="[475,572,1230,1256]" pageId="48" pageNumber="49">Briyech</collectorName>
;
<date id="CD5C48F3FF8FF237801C6FE2DAF6FB5A" box="[584,723,1230,1257]" pageId="48" pageNumber="49" value="2023-01-09">
<collectingDate id="DD18B11BFF8FF237801C6FE2DAF6FB5A" box="[584,723,1230,1257]" pageId="48" pageNumber="49" value="2023-01-09">9 Jan. 2023</collectingDate>
</date>
;
<collectingMethod id="60A31624FF8FF237808B6FE3DB72FB5B" box="[735,855,1231,1257]" pageId="48" pageNumber="49">sweep net</collectingMethod>
;
<collectorName id="14170BE5FF8FF23781306FE3DBFBFB5A" box="[868,990,1230,1257]" pageId="48" pageNumber="49">K. Kettani</collectorName>
leg.;
<collectionCode id="DFF3F6F6FF8FF23786766FE2DCA4FB5A" box="[1058,1153,1230,1256]" pageId="48" pageNumber="49">LESCB</collectionCode>
.
<emphasis id="8B96B221FF8FF23786F76FE2DD0DFB5A" bold="true" box="[1187,1320,1230,1256]" pageId="48" pageNumber="49">
<collectorName id="14170BE5FF8FF23786F76FE2DD0DFB5A" box="[1187,1320,1230,1256]" pageId="48" pageNumber="49">High Atlas</collectorName>
</emphasis>
</materialsCitation>
<materialsCitation id="098A646EFF8FF23787156FE2DB84FAB9" collectingDate="2017-03-25" collectionCode="LESCB" collectorName="Tahannaout &amp; K. Kettani" pageId="48" pageNumber="49" specimenCount="1" specimenCount-male="1">
<specimenCount id="AFE4A5BAFF8FF23787156FE2DD55FB5B" box="[1345,1392,1230,1257]" count="1" pageId="48" pageNumber="49" type="male">1 ♂</specimenCount>
;
<collectorName id="14170BE5FF8FF23782E96FDDD96CFAB9" box="[189,329,1265,1291]" pageId="48" pageNumber="49">Tahannaout</collectorName>
;
<date id="CD5C48F3FF8FF23783016FDDD9D0FAB9" box="[341,501,1265,1292]" pageId="48" pageNumber="49" value="2017-03-25">
<collectingDate id="DD18B11BFF8FF23783016FDDD9D0FAB9" box="[341,501,1265,1292]" pageId="48" pageNumber="49" value="2017-03-25">25 Mar. 2017</collectingDate>
</date>
;
<collectingMethod id="60A31624FF8FF23780566FDEDA5CFABE" box="[514,633,1266,1292]" pageId="48" pageNumber="49">sweep net</collectingMethod>
;
<collectorName id="14170BE5FF8FF23780D26FDEDADAFAB9" box="[646,767,1265,1292]" pageId="48" pageNumber="49">K. Kettani</collectorName>
leg.;
<collectionCode id="DFF3F6F6FF8FF23781166FDDDB84FAB9" box="[834,929,1265,1291]" pageId="48" pageNumber="49">LESCB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1F83DB8FF8FF23782E96E07DCD0FAC6" pageId="48" pageNumber="49" type="distribution">
<paragraph id="B95D6E33FF8FF23782E96E07DA09FAF7" blockId="48.[189,556,1323,1349]" box="[189,556,1323,1349]" pageId="48" pageNumber="49">
<heading id="E215D95FFF8FF23782E96E07DA09FAF7" bold="true" box="[189,556,1323,1349]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="8B96B221FF8FF23782E96E07DA09FAF7" bold="true" box="[189,556,1323,1349]" pageId="48" pageNumber="49">
Distribution outside
<collectingCountry id="C1F52EA3FF8FF23783EA6E07DA09FAF7" box="[446,556,1323,1349]" name="Morocco" pageId="48" pageNumber="49">Morocco</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="B95D6E33FF8FF23782E96E76DCD0FAC6" blockId="48.[189,1269,1369,1396]" box="[189,1269,1369,1396]" pageId="48" pageNumber="49">
<collectingCountry id="C1F52EA3FF8FF23782E96E76D93DFAC6" box="[189,280,1370,1396]" name="Algeria" pageId="48" pageNumber="49">Algeria</collectingCountry>
,
<collectingCountry id="C1F52EA3FF8FF23783706E76D952FAC6" box="[292,375,1370,1396]" name="France" pageId="48" pageNumber="49">France</collectingCountry>
,
<collectingCountry id="C1F52EA3FF8FF23783D66E76D9E0FAC6" box="[386,453,1370,1396]" name="Spain" pageId="48" pageNumber="49">Spain</collectingCountry>
(
<bibRefCitation id="DD7313C2FF8FF23783806E76DB21FAC6" author="Grootaert P. &amp; Chvala M." box="[468,772,1370,1396]" pageId="48" pageNumber="49" pagination="1 - 226" refId="ref24608" refString="Grootaert P. &amp; Chvala M. 1992. Monograph of the genus Platypalpus (Diptera: Empidoidea, Hybotidae) of the Mediterranean region and the Canary Islands. Acta Universitatis Carolinae - Biologica 36 (1 - 2): 1 - 226." type="journal article" year="1992">Grootaert &amp; Chvála 1992</bibRefCitation>
;
<bibRefCitation id="DD7313C2FF8FF23781456E76DBDCFAC6" author="De Jong Y. &amp; Verbeek M. &amp; Michelsen V. &amp; Bjorn P. D. P. &amp; Los W. &amp; Steeman F. &amp; Bailly N. &amp; Basire C. &amp; Chylarecki P. &amp; Stloukal E. &amp; Hagedorn G. &amp; Wetzel F. T. &amp; Glockler F. &amp; Kroupa A. &amp; Korb G. &amp; Hoffmann A. &amp; Hauser C. &amp; Kohlbecker A. &amp; Muller A. &amp; Penev L." box="[785,1017,1370,1396]" pageId="48" pageNumber="49" refId="ref24274" refString="De Jong Y., Verbeek M., Michelsen V., Bjorn P. D. P., Los W., Steeman F., Bailly N., Basire C., Chylarecki P., Stloukal E., Hagedorn G., Wetzel F. T., Glockler F., Kroupa A., Korb G., Hoffmann A., Hauser C., Kohlbecker A., Muller A., ... &amp; Penev L. 2014. Fauna Europaea - all European animal species on the web. Biodiversity Data Journal 2: e 4034. https: // doi. org / 10.3897 / BDJ. 2. e 4034" type="journal volume" year="2014">
De Jong
<emphasis id="8B96B221FF8FF237812D6E76DB89FAC6" box="[889,940,1370,1396]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. 2014
</bibRefCitation>
;
<bibRefCitation id="DD7313C2FF8FF23786506E76DCCCFAC6" author="Ventura D. &amp; Ghelamallah A. &amp; Bouhraoua R. &amp; Boualem M. &amp; Villar P. J." box="[1028,1257,1369,1396]" pageId="48" pageNumber="49" pagination="39 - 54" refId="ref25772" refString="Ventura D., Ghelamallah A., Bouhraoua R., Boualem M. &amp; Villar P. J. 2015. Platypalpus verbekei Grootaert &amp; Chvala, 1992, nova especie per Algeria i pel Nord d'Africa amb un cataleg de les especies conegudes d'Algeria de la familia Hybotidae (Diptera: Empidoidea: Hybotidae). Butlleti de la Institucio catalana d'Historia natural 79: 39 - 54." type="journal article" year="2015">
Ventura
<emphasis id="8B96B221FF8FF23786326E76DCBFFAC6" box="[1126,1178,1370,1396]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. 2015
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F1F83DB8FF8FF23782E96EB8DAAFF9D8" pageId="48" pageNumber="49" type="discussion">
<paragraph id="B95D6E33FF8FF23782E96EB8D90AFA1C" blockId="48.[189,303,1428,1454]" box="[189,303,1428,1454]" pageId="48" pageNumber="49">
<heading id="E215D95FFF8FF23782E96EB8D90AFA1C" bold="true" box="[189,303,1428,1454]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="8B96B221FF8FF23782E96EB8D90AFA1C" bold="true" box="[189,303,1428,1454]" pageId="48" pageNumber="49">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="B95D6E33FF8FF23782E96EEEDAAFF9D8" blockId="48.[189,1399,1474,1642]" pageId="48" pageNumber="49">
This is the first record of
<taxonomicName id="7EE215B0FF8FF23783B36EEEDAFBFA6E" authorityName="Grootaert &amp; Chvala" authorityYear="1992" box="[487,734,1474,1500]" class="Insecta" family="Hybotidae" genus="Platypalpus" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="verbekei">
<emphasis id="8B96B221FF8FF23783B36EEEDAFBFA6E" box="[487,734,1474,1500]" italics="true" pageId="48" pageNumber="49">Platypalpus verbekei</emphasis>
</taxonomicName>
from
<collectingCountry id="C1F52EA3FF8FF23781726EEFDBB5FA6F" box="[806,912,1475,1501]" name="Morocco" pageId="48" pageNumber="49">Morocco</collectingCountry>
and the second record from North Africa after its report by
<bibRefCitation id="DD7313C2FF8FF23783DC6ECADA53F9B2" author="Ventura D. &amp; Ghelamallah A. &amp; Bouhraoua R. &amp; Boualem M. &amp; Villar P. J." box="[392,630,1509,1536]" pageId="48" pageNumber="49" pagination="39 - 54" refId="ref25772" refString="Ventura D., Ghelamallah A., Bouhraoua R., Boualem M. &amp; Villar P. J. 2015. Platypalpus verbekei Grootaert &amp; Chvala, 1992, nova especie per Algeria i pel Nord d'Africa amb un cataleg de les especies conegudes d'Algeria de la familia Hybotidae (Diptera: Empidoidea: Hybotidae). Butlleti de la Institucio catalana d'Historia natural 79: 39 - 54." type="journal article" year="2015">
Ventura
<emphasis id="8B96B221FF8FF23783B36ECADA3CFA4D" box="[487,537,1509,1535]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. (2015)
</bibRefCitation>
from
<collectingCountry id="C1F52EA3FF8FF23780ED6EC9DB30FA4D" box="[697,789,1509,1535]" name="Algeria" pageId="48" pageNumber="49">Algeria</collectingCountry>
.According to the original description in
<bibRefCitation id="DD7313C2FF8FF23786BE6EC9D94FF991" author="Grootaert P. &amp; Chvala M." pageId="48" pageNumber="49" pagination="1 - 226" refId="ref24608" refString="Grootaert P. &amp; Chvala M. 1992. Monograph of the genus Platypalpus (Diptera: Empidoidea, Hybotidae) of the Mediterranean region and the Canary Islands. Acta Universitatis Carolinae - Biologica 36 (1 - 2): 1 - 226." type="journal article" year="1992">Grootaert &amp; Chvála (1992)</bibRefCitation>
from
<collectingCountry id="C1F52EA3FF8FF23783EE6D25DA28F991" box="[442,525,1545,1571]" name="France" pageId="48" pageNumber="49">France</collectingCountry>
, the Moroccan specimen is entirely compatible with the morphological description and the genitalia illustrations. It is also compatible with the Algerian specimen re-described and illustrated by
<bibRefCitation id="DD7313C2FF8FF23783DB6D63DAA3F9D8" author="Ventura D. &amp; Ghelamallah A. &amp; Bouhraoua R. &amp; Boualem M. &amp; Villar P. J." box="[399,646,1615,1642]" pageId="48" pageNumber="49" pagination="39 - 54" refId="ref25772" refString="Ventura D., Ghelamallah A., Bouhraoua R., Boualem M. &amp; Villar P. J. 2015. Platypalpus verbekei Grootaert &amp; Chvala, 1992, nova especie per Algeria i pel Nord d'Africa amb un cataleg de les especies conegudes d'Algeria de la familia Hybotidae (Diptera: Empidoidea: Hybotidae). Butlleti de la Institucio catalana d'Historia natural 79: 39 - 54." type="journal article" year="2015">
Ventura
<emphasis id="8B96B221FF8FF23783A66D7CDA03F9DB" box="[498,550,1615,1641]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. (2015)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>