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<document id="5F0EFD3F00BA2E884ABC4BABFDAAE8C7" ID-CLB-Dataset="56053" ID-DOI="10.11646/zootaxa.5125.4.1" ID-GBIF-Dataset="cd04ba2d-dd59-46a6-ba04-641fe11e8ed5" ID-ISSN="1175-5326" ID-Zenodo-Dep="6450835" ID-ZooBank="CD46676E-5304-4550-B1DB-8EA6EF30AF62" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649745307548" checkinUser="plazi" docAuthor="Tungpairojwong, Nisarat, Phlai-Ngam, Sirikamon &amp; Jacobus, Luke M." docDate="2022" docId="311AC57CF326832836B34917FED5976A" docLanguage="en" docName="zootaxa.5125.4.1.pdf" docOrigin="Zootaxa 5125 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acentrella fimbriata Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="367" masterDocId="CD23BD04F324833836244B51FFA8962B" masterDocTitle="A new species of Acentrella Bengtsson, 1912 (Ephemeroptera: Baetidae) from Thailand" masterLastPageNumber="378" masterPageNumber="351" pageNumber="353" updateTime="1699319985581" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F56C4183093BF47811353C900B81C2E9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2CC196F63235DFE1936FF012BCB20859">
<mods:title id="50E00DFAB3E6CAF5EA60FC234DEFD8F5">A new species of Acentrella Bengtsson, 1912 (Ephemeroptera: Baetidae) from Thailand</mods:title>
</mods:titleInfo>
<mods:name id="64FF1A44B30316F73A6AA6590157C4FC" type="personal">
<mods:role id="96DFE031C104A80C8903BDB748FD69C5">
<mods:roleTerm id="6F33C6566B47E7064752671EE5035DB0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC7C517AD6A1FD6F8B69F8CC066A6FED">Tungpairojwong, Nisarat</mods:namePart>
<mods:affiliation id="49405984FAF64CEF330F0310714BE600">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand. &amp; Applied Taxonomic Research Center, Khon Kaen University, Khon Kaen 40002, Thailand. &amp; Center of Excellence on Biodiversity Tab Nilaniti Building 2 nd Fl., Department of Biology, Faculty of Science, Chulalongkorn University Phayathai Road, Pathumwan Bangkok 10330, Thailand.</mods:affiliation>
</mods:name>
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<mods:role id="61F22BF8BA11E934A935E71A0049E3A1">
<mods:roleTerm id="9CD12D996AE61B0396600E14A05A4E58">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B82C325B78D5029DEE43A753CCB7941A">Phlai-Ngam, Sirikamon</mods:namePart>
<mods:affiliation id="4B85C16E337BD2D6D13981409999620E">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand. &amp; Center of Excellence on Biodiversity Tab Nilaniti Building 2 nd Fl., Department of Biology, Faculty of Science, Chulalongkorn University Phayathai Road, Pathumwan Bangkok 10330, Thailand.</mods:affiliation>
</mods:name>
<mods:name id="6AC54E5ADB22BCD09CD160FC0D9931A5" type="personal">
<mods:role id="9DBA6E9D658548ABEEF21F23003A38FF">
<mods:roleTerm id="29FB0A02A82D67E23A94E09E86666C40">Author</mods:roleTerm>
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<mods:namePart id="481D77B0BE31F7CBD53361010DDB02FD">Jacobus, Luke M.</mods:namePart>
<mods:nameIdentifier id="3B971AE7A7760DD56654B929B4B18BF1" type="ORCID">0000-0002-7201-3922</mods:nameIdentifier>
<mods:affiliation id="2ABD9BD50FBB327AACD1EA161E71D891">Division of Science, Indiana University Purdue University Columbus, 4601 Central Ave., Columbus, Indiana, 47203, USA. lukemjacobus @ alumni. purdue. edu; https: // orcid. org / 0000 - 0002 - 7201 - 3922</mods:affiliation>
<mods:nameIdentifier id="02AA4A28E3FFA30D9FB8B93C0463F147" type="email">lukemjacobus@alumni.purdue.edu</mods:nameIdentifier>
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<mods:title id="5FFDE3CEF992003FCAAD8EBC4DEF07F3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3BA03A29054D555A2C8B6EEB15281FD6">
<mods:date id="DBA0F1C8E2E2866C64E0B3FCEC22841F">2022</mods:date>
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<mods:number id="1081471C8C481DDAB2DD6D6F9A6D3FFC">2022-04-12</mods:number>
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<mods:number id="4DF0DDA913EF8B1A1E43BD6095407CF5">5125</mods:number>
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<mods:number id="338A5042CA6C95E6F83CB0DE415A6A1B">4</mods:number>
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<mods:start id="2E74B07DFDD1BA3E9287537F0996040E">351</mods:start>
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<mods:identifier id="8E67CB82266D1D7396A39B8CB9D69467" type="DOI">10.11646/zootaxa.5125.4.1</mods:identifier>
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<treatment id="311AC57CF326832836B34917FED5976A" ID-DOI="http://doi.org/10.5281/zenodo.6450839" ID-GBIF-Taxon="194568190" ID-Zenodo-Dep="6450839" LSID="urn:lsid:plazi:treatment:311AC57CF326832836B34917FED5976A" httpUri="http://treatment.plazi.org/id/311AC57CF326832836B34917FED5976A" lastPageId="16" lastPageNumber="367" pageId="2" pageNumber="353">
<subSubSection id="F1A927E1F326833A36B34917FE59944A" box="[151,497,582,609]" pageId="2" pageNumber="353" type="nomenclature">
<paragraph id="B90C746AF326833A36B34917FE59944A" blockId="2.[151,497,582,645]" box="[151,497,582,609]" pageId="2" pageNumber="353">
<heading id="E244C306F326833A36B34917FE59944A" bold="true" box="[151,497,582,609]" fontSize="11" level="1" pageId="2" pageNumber="353" reason="1">
<emphasis id="8BC7A878F326833A36B34917FE59944A" bold="true" box="[151,497,582,609]" pageId="2" pageNumber="353">
<taxonomicName id="7EB30FE9F326833A36B34917FE24944A" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[151,396,582,609]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F326833A36B34917FE24944A" bold="true" box="[151,396,582,609]" italics="true" pageId="2" pageNumber="353">Acentrella fimbriata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90F41503F326833A37B04916FE59944A" box="[404,497,583,609]" pageId="2" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F1A927E1F326833A36B3493AFEB094AE" box="[151,280,619,645]" pageId="2" pageNumber="353" type="description">
<paragraph id="B90C746AF326833A36B3493AFEB094AE" blockId="2.[151,497,582,645]" box="[151,280,619,645]" pageId="2" pageNumber="353">
(
<figureCitation id="218868EFF326833A36BB493AFEA794AE" box="[159,271,619,645]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="4.[151,250,1943,1967]" captionStartId-1="5.[151,250,1751,1775]" captionStartId-10="14.[151,250,1865,1889]" captionStartId-2="6.[151,250,1943,1967]" captionStartId-3="7.[151,250,1943,1967]" captionStartId-4="8.[151,250,1943,1967]" captionStartId-5="9.[151,250,1788,1812]" captionStartId-6="10.[151,250,1910,1934]" captionStartId-7="11.[151,250,1785,1809]" captionStartId-8="12.[151,250,1882,1906]" captionStartId-9="13.[151,250,1855,1879]" captionTargetBox-0="[174,1413,181,1919]" captionTargetBox-1="[151,1436,181,1728]" captionTargetBox-10="[152,1436,182,1841]" captionTargetBox-2="[194,1391,193,1907]" captionTargetBox-3="[192,1392,190,1903]" captionTargetBox-4="[186,1403,191,1901]" captionTargetBox-5="[151,1435,182,1764]" captionTargetBox-6="[151,1436,562,1885]" captionTargetBox-7="[152,1436,182,1761]" captionTargetBox-8="[152,1436,183,1857]" captionTargetBox-9="[152,1435,182,1829]" captionTargetId-0="figure-19@4.[174,1413,181,1919]" captionTargetId-1="figure-19@5.[151,1436,181,1728]" captionTargetId-10="figure-19@14.[151,1436,181,1841]" captionTargetId-2="figure-19@6.[180,1407,181,1919]" captionTargetId-3="figure-19@7.[181,1406,181,1919]" captionTargetId-4="figure-19@8.[177,1410,181,1919]" captionTargetId-5="figure-19@9.[151,1435,181,1764]" captionTargetId-6="figure-205@10.[151,1436,561,1886]" captionTargetId-7="figure-19@11.[151,1436,181,1761]" captionTargetId-8="figure-19@12.[151,1436,181,1858]" captionTargetId-9="figure-19@13.[151,1436,181,1831]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="14" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="FIGURE 1. Acentrella fimbriata sp. nov., female nymph, dorsal habitus. Scale bar = 1 mm." captionText-1="FIGURE 2. Acentrella fimbriata sp. nov., nymph, dorsal views: (A) head and pronotum; (B) thorax; (C) abdomen; (D) abdominal terga V–VII; (E) abdominal gills. Scale bars: A = 0.2 mm; B–E = 0.5 mm." captionText-10="FIGURE 11. Acentrella fimbriata sp. nov., male subimago: (A) dorsal habitus; (B) head and anterior thorax, lateral view; (C) abdomen, dorsal view; (D) forewing; (E) genitalia. Scale bars: A, D = 1 mm; B–C = 0.5 mm; E = 0.05 mm." captionText-2="FIGURE 3. Acentrella fimbriata sp. nov. nymph, head structures: (A) head capsule, frontal view; (B) antenna; (C) labrum, with different setae emphasized on each side; (D) left (angulate) mandible; (E) right (planate) mandible; (F) incisors of left mandible; (G) incisors of right mandible. Scale bars: A–E = 0.1 mm; F–G = 0.02 mm." captionText-3="FIGURE 4. Acentrella fimbriata sp. nov., nymph, mouthparts: (A) maxilla; (B) apex of galae-lacinia; (C) maxillary palp; (D) labium; (E) glossa; (F) paraglossa; (G) labial palp; (H) hypopharynx. Scale bars: A, C = 0.2 mm; B, H = 0.05 mm; D–F = 0.1 mm." captionText-4="FIGURE 5. Acentrella fimbriata sp. nov., nymph, thoracic and abdominal structures: (A) small posterolateral projection on metanotum; (B) foreleg; (C) tarsal claw; (D) abdominal tergum V; (E) abdominal gills I–VII; (F) gill V, enlarged view with details; (G) paraproct; (H) cerci and median filament. Scale bars: A, B, D, E, F = 0.2 mm; C, G = 0.02 mm; H = 0.1 mm." captionText-5="FIGURE 6. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs: (A) head; (B) setae on scape and pedicel of antenna; (C) long feathered bristles on dorsal margin of femur; (D) dense, irregular row of long, simple setae on ventral margin of femur; (E) villopore; (F) tarsal claw. Scale bars: A = 500 µm; B, E, F = 100 µm; C = 200 µm; D = 300 µm." captionText-6="FIGURE 7. Acentrella fimbriata sp. nov., nymph, scanning electron micrograph: abdomen, ventral view. Scale bar = 0.5 mm" captionText-7="FIGURE 8. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A) sterna 3–5; (B) sterna 5–7; (C) sterna 7–10. Scale bars: A, B = 400 µm; C = 500 µm" captionText-8="FIGURE 9. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A–C) details of sternum 8; (D–E) details of sternum 9; (F) paraproct. Scale bars: A = 200 µm; B, D, F = 50 µm; C = 30 µm; E = 20 µm." captionText-9="FIGURE 10. Acentrella fimbriata sp. nov., scanning electron micrographs, A–D nymph, E–F egg: (A) abdominal terga IV–VI; (B) setae and scales on abdominal tergum V; (C) gill with long setae; (D) median filament; (E) egg, general view; (F) egg, longitudinal tissue of inner layer. Scale bars: A = 400 µm; B, D = 100 µm; C = 0.5 mm; E = 50 µm; F = 10 µm." figureDoi-0="http://doi.org/10.5281/zenodo.6450837" figureDoi-1="http://doi.org/10.5281/zenodo.6450843" figureDoi-10="http://doi.org/10.5281/zenodo.6450861" figureDoi-2="http://doi.org/10.5281/zenodo.6450845" figureDoi-3="http://doi.org/10.5281/zenodo.6450847" figureDoi-4="http://doi.org/10.5281/zenodo.6450849" figureDoi-5="http://doi.org/10.5281/zenodo.6450851" figureDoi-6="http://doi.org/10.5281/zenodo.6450853" figureDoi-7="http://doi.org/10.5281/zenodo.6450855" figureDoi-8="http://doi.org/10.5281/zenodo.6450857" figureDoi-9="http://doi.org/10.5281/zenodo.6450859" httpUri-0="https://zenodo.org/record/6450837/files/figure.png" httpUri-1="https://zenodo.org/record/6450843/files/figure.png" httpUri-10="https://zenodo.org/record/6450861/files/figure.png" httpUri-2="https://zenodo.org/record/6450845/files/figure.png" httpUri-3="https://zenodo.org/record/6450847/files/figure.png" httpUri-4="https://zenodo.org/record/6450849/files/figure.png" httpUri-5="https://zenodo.org/record/6450851/files/figure.png" httpUri-6="https://zenodo.org/record/6450853/files/figure.png" httpUri-7="https://zenodo.org/record/6450855/files/figure.png" httpUri-8="https://zenodo.org/record/6450857/files/figure.png" httpUri-9="https://zenodo.org/record/6450859/files/figure.png" pageId="2" pageNumber="353">Figs 1–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F1A927E1F326833A36B349E2FD0E95C6" pageId="2" pageNumber="353" type="materials_examined">
<paragraph id="B90C746AF326833A36B349E2FC1A94DA" blockId="2.[151,1437,690,2013]" pageId="2" pageNumber="353">
<materialsCitation id="09DB7E37F326833A36B349E2FC1A94DA" ID-GBIF-Occurrence="3750826315" collectingDate="2019-12-23" collectionCode="PERC" county="Final" elevation="693" latitude="19.157091" location="Bo Kluea district" longLatPrecision="1" longitude="101.15844" municipality="Mang River" pageId="2" pageNumber="353" specimenCount="1" stateProvince="Nan Province" typeStatus="holotype">
<emphasis id="8BC7A878F326833A36B349E2FEA394E6" bold="true" box="[151,267,691,717]" pageId="2" pageNumber="353">
<typeStatus id="6608CAC8F326833A36B349E2FEAE94E6" box="[151,262,691,717]" pageId="2" pageNumber="353" type="holotype">Holotype</typeStatus>
.
</emphasis>
Final nymphal instar in ethanol [
<collectionCode id="DFA2ECAFF326833A345A49E3FD6494E6" box="[638,716,690,717]" collectionName="PERC" pageId="2" pageNumber="353">PERC</collectionCode>
],
<collectingRegion id="7B77BA88F326833A34FA49E2FCD294E6" box="[734,890,691,717]" country="Thailand" name="Nan" pageId="2" pageNumber="353">Nan Province</collectingRegion>
,
<collectingMunicipality id="5968EE10F326833A35A249E2FBA694E6" box="[902,1038,691,717]" pageId="2" pageNumber="353">Mang River</collectingMunicipality>
,
<location id="BC6C22B1F326833A323D49E2FB7194E6" LSID="urn:lsid:plazi:treatment:311AC57CF326832836B34917FED5976A:BC6C22B1F326833A323D49E2FB7194E6" box="[1049,1241,691,717]" county="Final" latitude="19.157091" longLatPrecision="1" longitude="101.15844" municipality="Mang River" name="Bo Kluea district" pageId="2" pageNumber="353" stateProvince="Nan Province">Bo Kluea district</location>
(
<geoCoordinate id="DC8712ADF326833A32CC49E3FA3094E6" box="[1256,1432,690,717]" degrees="19" direction="north" minutes="09" orientation="latitude" pageId="2" pageNumber="353" precision="1" seconds="25.53" value="19.157091">19°09’25.53”N</geoCoordinate>
,
<geoCoordinate id="DC8712ADF326833A36B34986FEF994DA" box="[151,337,727,753]" degrees="101" direction="east" minutes="09" orientation="longitude" pageId="2" pageNumber="353" precision="1" seconds="30.37" value="101.15844">101°09’30.37”E</geoCoordinate>
, alt.
<elevation id="329E9359F326833A37A24986FDA194D9" box="[390,521,727,754]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="2" pageNumber="353" unit="m" value="693.0">
<quantity id="7E4BD98FF326833A37A24986FE6594DA" box="[390,461,727,753]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="2" pageNumber="353" unit="m" value="693.0">693 m</quantity>
a.s.l.
</elevation>
),
<date id="CD0D52AAF326833A343B4986FD0A94DA" box="[543,674,727,753]" pageId="2" pageNumber="353" value="2019-12-23">
<collectingDate id="DD49AB42F326833A343B4986FD0A94DA" box="[543,674,727,753]" pageId="2" pageNumber="353" value="2019-12-23">23.xii.2019</collectingDate>
</date>
, Sirikamon Phlai-ngam.
</materialsCitation>
</paragraph>
<paragraph id="B90C746AF326833A36E349AAFC79953E" blockId="2.[151,1437,690,2013]" box="[199,977,762,789]" pageId="2" pageNumber="353">
<materialsCitation id="09DB7E37F326833A36E349AAFC65953E" ID-GBIF-Occurrence="3750826306" box="[199,973,762,789]" collectingDate="2019-12-23" collectionCode="KKU-AIC" county="Final" elevation="693" latitude="19.157091" location="Bo Kluea district" longLatPrecision="1" longitude="101.15844" municipality="Mang River" pageId="2" pageNumber="353" specimenCount="1" stateProvince="Nan" typeStatus="paratype">
<emphasis id="8BC7A878F326833A36E349AAFE93953E" bold="true" box="[199,315,763,789]" pageId="2" pageNumber="353">
<typeStatus id="6608CAC8F326833A36E349AAFE9E953E" box="[199,310,763,789]" pageId="2" pageNumber="353" type="paratype">Paratype</typeStatus>
.
</emphasis>
One nymph in ethanol [
<collectionCode id="DFA2ECAFF326833A346D49ABFD6D953E" box="[585,709,762,789]" pageId="2" pageNumber="353">KKU-AIC</collectionCode>
] same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="B90C746AF326833A36E3484FFD0E95C6" blockId="2.[151,1437,690,2013]" pageId="2" pageNumber="353">
<emphasis id="8BC7A878F326833A36E3484FFE6C9513" bold="true" box="[199,452,798,824]" pageId="2" pageNumber="353">Additional materials.</emphasis>
<materialsCitation id="09DB7E37F326833A37ED484EFBE99512" ID-GBIF-Occurrence="3750826313" box="[457,1089,798,825]" collectingDate="2015-02-02" collectingDateMax="2020-03-06" collectingDateMin="2015-02-02" collectionCode="KKU-AIC" county="Khun Korn" elevation="610" latitude="19.520039" location="Chiang Rai Province" longLatPrecision="1" longitude="99.65349" municipality="Pha Kong cave" pageId="2" pageNumber="353" specimenCount="1" stateProvince="Chiang Rai Province">
<collectingRegion id="7B77BA88F326833A37ED484EFD1D9512" box="[457,693,799,825]" country="Thailand" name="Chiang Rai" pageId="2" pageNumber="353">Chiang Rai Province</collectingRegion>
: 20 nymphs in ethanol [
<collectionCode id="DFA2ECAFF326833A359C484FFB9F9512" box="[952,1079,798,825]" pageId="2" pageNumber="353">KKU-AIC</collectionCode>
],
</materialsCitation>
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one nymph in ethanol [
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],
<location id="BC6C22B1F326833A36B34812FECF9576" LSID="urn:lsid:plazi:treatment:311AC57CF326832836B34917FED5976A:BC6C22B1F326833A36B34812FECF9576" box="[151,359,835,861]" county="Khun Korn" latitude="19.520039" longLatPrecision="1" longitude="99.65349" municipality="Pha Kong cave" name="Khun Korn stream" pageId="2" pageNumber="353" stateProvince="Chiang Rai">Khun Korn stream</location>
(
<geoCoordinate id="DC8712ADF326833A37514813FD8D9576" box="[373,549,834,861]" degrees="19" direction="north" minutes="59" orientation="latitude" pageId="2" pageNumber="353" precision="1" seconds="19.80" value="19.988834">19°59’19.80”N</geoCoordinate>
,
<geoCoordinate id="DC8712ADF326833A340B4812FD739576" box="[559,731,835,861]" degrees="99" direction="east" minutes="37" orientation="longitude" pageId="2" pageNumber="353" precision="1" seconds="19.80" value="99.62217">99°37’19.80”E</geoCoordinate>
, alt.
<elevation id="329E9359F326833A352B4812FC389575" box="[783,912,835,862]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="353" unit="m" value="610.0">
<quantity id="7E4BD98FF326833A352B4812FCFD9576" box="[783,853,835,861]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="353" unit="m" value="610.0">610 m</quantity>
a.s.l.
</elevation>
),
<date id="CD0D52AAF326833A35824813FBA49576" box="[934,1036,834,861]" pageId="2" pageNumber="353" value="2015-02-03">
<collectingDate id="DD49AB42F326833A35824813FBA49576" box="[934,1036,834,861]" pageId="2" pageNumber="353" value="2015-02-03">3.ii.2015</collectingDate>
</date>
,
<location id="BC6C22B1F326833A32334812FABF9576" LSID="urn:lsid:plazi:treatment:311AC57CF326832836B34917FED5976A:BC6C22B1F326833A32334812FABF9576" box="[1047,1303,835,861]" county="Khun Korn" latitude="19.520039" longLatPrecision="1" longitude="99.65349" municipality="Pha Kong cave" name="Sirikamon Phlai-ngam" pageId="2" pageNumber="353" stateProvince="Chiang Rai">Sirikamon Phlai-ngam</location>
; 12 nymphs in ethanol [
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],
</materialsCitation>
<materialsCitation id="09DB7E37F326833A37944836FD0095E2" ID-GBIF-Occurrence="3750826301" collectingDate="2020-03-06" collectingDateMax="2020-03-06" collectingDateMin="2015-02-02" collectionCode="KKU-AIC, PERC" county="Khun Korn" elevation="610" latitude="19.520039" location="Phan district" longLatPrecision="1" longitude="99.65349" municipality="Pha Kong cave" pageId="2" pageNumber="353" specimenCount="1" stateProvince="Chiang Rai">
one reared male subimago in ethanol, one nymph in ethanol [
<collectionCode id="DFA2ECAFF326833A32A14837FAAC95AA" box="[1157,1284,870,897]" pageId="2" pageNumber="353">KKU-AIC</collectionCode>
], Khun Korn stream (
<geoCoordinate id="DC8712ADF326833A36D448DBFE08958E" box="[240,416,906,933]" degrees="19" direction="north" minutes="59" orientation="latitude" pageId="2" pageNumber="353" precision="1" seconds="19.80" value="19.988834">19°59’19.80”N</geoCoordinate>
,
<geoCoordinate id="DC8712ADF326833A378E48DAFDFD958E" box="[426,597,907,933]" degrees="99" direction="east" minutes="37" orientation="longitude" pageId="2" pageNumber="353" precision="1" seconds="19.80" value="99.62217">99°37’19.80”E</geoCoordinate>
, alt.
<elevation id="329E9359F326833A34AC48DAFCA0958D" box="[648,776,907,934]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="353" unit="m" value="610.0">
<quantity id="7E4BD98FF326833A34AC48DAFD66958E" box="[648,718,907,933]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="353" unit="m" value="610.0">610 m</quantity>
a.s.l.
</elevation>
),
<date id="CD0D52AAF326833A353A48DAFC23958E" box="[798,907,907,933]" pageId="2" pageNumber="353" value="2020-03-06">
<collectingDate id="DD49AB42F326833A353A48DAFC23958E" box="[798,907,907,933]" pageId="2" pageNumber="353" value="2020-03-06">6.iii.2020</collectingDate>
</date>
, Sirikamon Phlai-ngam; five nymphs in ethanol [
<collectionCode id="DFA2ECAFF326833A36B948FFFEB195E2" box="[157,281,942,969]" pageId="2" pageNumber="353">KKU-AIC</collectionCode>
] and one nymph in ethanol [
<collectionCode id="DFA2ECAFF326833A347448FFFD3695E2" box="[592,670,942,969]" collectionName="PERC" pageId="2" pageNumber="353">PERC</collectionCode>
],
</materialsCitation>
<materialsCitation id="09DB7E37F326833A348B48FEFD0B95C6" ID-GBIF-Occurrence="3750826309" collectingDate="2015-02-02" collectingDateMax="2020-03-06" collectingDateMin="2015-02-02" collectionCode="KKU-AIC, PERC" county="Khun Korn" elevation="287" latitude="19.520039" location="Phan district" longLatPrecision="1" longitude="99.65349" municipality="Pha Kong cave" pageId="2" pageNumber="353" specimenCount="1" stateProvince="Chiang Rai">
<collectingMunicipality id="5968EE10F326833A348B48FEFCF095E2" box="[687,856,943,969]" pageId="2" pageNumber="353">Pha Kong cave</collectingMunicipality>
,
<location id="BC6C22B1F326833A354648FEFC4695E2" LSID="urn:lsid:plazi:treatment:311AC57CF326832836B34917FED5976A:BC6C22B1F326833A354648FEFC4695E2" box="[866,1006,943,969]" county="Khun Korn" latitude="19.520039" longLatPrecision="1" longitude="99.65349" municipality="Pha Kong cave" name="Phan district" pageId="2" pageNumber="353" stateProvince="Chiang Rai">Phan district</location>
(
<geoCoordinate id="DC8712ADF326833A35D848FEFB0395E2" box="[1020,1195,943,969]" degrees="19" direction="north" minutes="31" orientation="latitude" pageId="2" pageNumber="353" precision="1" seconds="12.14" value="19.520039">19°31’12.14”N</geoCoordinate>
,
<geoCoordinate id="DC8712ADF326833A329248FFFAC795E2" box="[1206,1391,942,969]" degrees="099" direction="east" minutes="39" orientation="longitude" pageId="2" pageNumber="353" precision="1" seconds="12.54" value="99.65349">099°39’12.54”E</geoCoordinate>
, alt.
<elevation id="329E9359F326833A36B34882FEB295C5" box="[151,282,979,1006]" metricMagnitude="2" metricUnit="m" metricValue="2.87" pageId="2" pageNumber="353" unit="m" value="287.0">
<quantity id="7E4BD98FF326833A36B34882FF7695C6" box="[151,222,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="2.87" pageId="2" pageNumber="353" unit="m" value="287.0">287 m</quantity>
a.s.l.
</elevation>
),
<date id="CD0D52AAF326833A37144883FE3E95C6" box="[304,406,978,1005]" pageId="2" pageNumber="353" value="2015-02-02">
<collectingDate id="DD49AB42F326833A37144883FE3E95C6" box="[304,406,978,1005]" pageId="2" pageNumber="353" value="2015-02-02">2.ii.2015</collectingDate>
</date>
, Sirikamon Phlai-ngam
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1A927E1F326833236E348A7FC9997FA" lastPageId="10" lastPageNumber="361" pageId="2" pageNumber="353" type="materials_examined">
<paragraph id="B90C746AF326833A36E348A7FECC923B" blockId="2.[151,1437,690,2013]" box="[199,356,1014,1040]" pageId="2" pageNumber="353">
<emphasis id="8BC7A878F326833A36E348A7FECC923B" bold="true" box="[199,356,1014,1040]" pageId="2" pageNumber="353">Descriptions.</emphasis>
</paragraph>
<paragraph id="B90C746AF326833A36E34F4BFC319256" blockId="2.[151,1437,690,2013]" pageId="2" pageNumber="353">
<emphasis id="8BC7A878F326833A36E34F4BFE13921E" bold="true" box="[199,443,1050,1077]" italics="true" pageId="2" pageNumber="353">Final nymphal instar</emphasis>
(
<figureCitation id="218868EFF326833A37ED4F4AFDA1921E" box="[457,521,1051,1077]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[174,1413,181,1919]" captionTargetId="figure-19@4.[174,1413,181,1919]" captionTargetPageId="4" captionText="FIGURE 1. Acentrella fimbriata sp. nov., female nymph, dorsal habitus. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450837" httpUri="https://zenodo.org/record/6450837/files/figure.png" pageId="2" pageNumber="353">Fig. 1</figureCitation>
). Body length
<quantity id="7E4BD98FF326833A348A4F4BFC9E921E" box="[686,822,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.2" metricValueMin="4.3" pageId="2" pageNumber="353" unit="mm" value="4.75" valueMax="5.2" valueMin="4.3">4.3–5.2 mm</quantity>
, head width
<quantity id="7E4BD98FF326833A35E54F4AFB9B921E" box="[961,1075,1051,1077]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="2" pageNumber="353" unit="mm" value="0.95" valueMax="1.0" valueMin="0.9">0.9–1 mm</quantity>
, head length
<quantity id="7E4BD98FF326833A32E04F4BFAE4921E" box="[1220,1356,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="353" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
, antennal length 1.0–
<quantity id="7E4BD98FF326833A37644F6EFE309272" box="[320,408,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="353" unit="mm" value="1.2">1.2 mm</quantity>
, lengths of forefemora, midfemora and hindfemora
<quantity id="7E4BD98FF326833A35C64F6FFB219272" box="[994,1161,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" metricValueMax="6.5" metricValueMin="6.3" pageId="2" pageNumber="353" unit="mm" value="0.64" valueMax="0.65" valueMin="0.63">0.63–0.65 mm</quantity>
. Length of cerci
<quantity id="7E4BD98FF326833A336C4F6EFF6E9256" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="2" pageNumber="353" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
. Body short and ventrally flattened. Thorax apparently enlarged.
</paragraph>
<paragraph id="B90C746AF326833B36E34FD7FB8096FE" blockId="2.[151,1437,690,2013]" lastBlockId="3.[151,1437,151,1977]" lastPageId="3" lastPageNumber="354" pageId="2" pageNumber="353">
<emphasis id="8BC7A878F326833A36E34FD7FEAD928A" bold="true" box="[199,261,1158,1185]" italics="true" pageId="2" pageNumber="353">Head</emphasis>
. Capsule light brown with lighter and darker areas on vertex; frontal suture yellowish with brown markings extending from behind lateral ocelli to occiput. Head with hypognathous orientation; width greater than length. Compound eyes black; ocelli dark brown; orange turbinate eyes of male subimago visible through cuticle of final nymphal instar. Margins of capsule relatively densely covered with fine, long setae, with greatest concentrations on genae and clypeus. Long and translucent scales scattered along capsule margins (
<figureCitation id="218868EFF326833A32044E46FBD5931A" box="[1056,1149,1303,1329]" captionStart="FIGURE 2" captionStartId="5.[151,250,1751,1775]" captionTargetBox="[151,1436,181,1728]" captionTargetId="figure-19@5.[151,1436,181,1728]" captionTargetPageId="5" captionText="FIGURE 2. Acentrella fimbriata sp. nov., nymph, dorsal views: (A) head and pronotum; (B) thorax; (C) abdomen; (D) abdominal terga V–VII; (E) abdominal gills. Scale bars: A = 0.2 mm; B–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6450843" httpUri="https://zenodo.org/record/6450843/files/figure.png" pageId="2" pageNumber="353">Figs 2A</figureCitation>
,
<figureCitation id="218868EFF326833A32AD4E46FB05931A" box="[1161,1197,1303,1329]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[194,1391,193,1907]" captionTargetId="figure-19@6.[180,1407,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Acentrella fimbriata sp. nov. nymph, head structures: (A) head capsule, frontal view; (B) antenna; (C) labrum, with different setae emphasized on each side; (D) left (angulate) mandible; (E) right (planate) mandible; (F) incisors of left mandible; (G) incisors of right mandible. Scale bars: A–E = 0.1 mm; F–G = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6450845" httpUri="https://zenodo.org/record/6450845/files/figure.png" pageId="2" pageNumber="353">3A</figureCitation>
,
<figureCitation id="218868EFF326833A329D4E46FB779319" box="[1209,1247,1303,1330]" captionStart="FIGURE 6" captionStartId="9.[151,250,1788,1812]" captionTargetBox="[151,1435,182,1764]" captionTargetId="figure-19@9.[151,1435,181,1764]" captionTargetPageId="9" captionText="FIGURE 6. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs: (A) head; (B) setae on scape and pedicel of antenna; (C) long feathered bristles on dorsal margin of femur; (D) dense, irregular row of long, simple setae on ventral margin of femur; (E) villopore; (F) tarsal claw. Scale bars: A = 500 µm; B, E, F = 100 µm; C = 200 µm; D = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.6450851" httpUri="https://zenodo.org/record/6450851/files/figure.png" pageId="2" pageNumber="353">6A</figureCitation>
). Antenna length approximately 2 times head width; scape and pedicel brownish, slightly paler than flagellum; scape broader than pedicel, subequal in length. Inner margins of scape and pedicel densely covered with long, fine, simple setae (
<figureCitation id="218868EFF326833A334E4E0EFF1293B6" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[194,1391,193,1907]" captionTargetId="figure-19@6.[180,1407,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Acentrella fimbriata sp. nov. nymph, head structures: (A) head capsule, frontal view; (B) antenna; (C) labrum, with different setae emphasized on each side; (D) left (angulate) mandible; (E) right (planate) mandible; (F) incisors of left mandible; (G) incisors of right mandible. Scale bars: A–E = 0.1 mm; F–G = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6450845" httpUri="https://zenodo.org/record/6450845/files/figure.png" pageId="2" pageNumber="353">Figs 3B</figureCitation>
,
<figureCitation id="218868EFF326833A36ED4ED2FF4593B5" box="[201,237,1411,1438]" captionStart="FIGURE 6" captionStartId="9.[151,250,1788,1812]" captionTargetBox="[151,1435,182,1764]" captionTargetId="figure-19@9.[151,1435,181,1764]" captionTargetPageId="9" captionText="FIGURE 6. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs: (A) head; (B) setae on scape and pedicel of antenna; (C) long feathered bristles on dorsal margin of femur; (D) dense, irregular row of long, simple setae on ventral margin of femur; (E) villopore; (F) tarsal claw. Scale bars: A = 500 µm; B, E, F = 100 µm; C = 200 µm; D = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.6450851" httpUri="https://zenodo.org/record/6450851/files/figure.png" pageId="2" pageNumber="353">6B</figureCitation>
). Labrum (
<figureCitation id="218868EFF326833A374B4ED2FE6F93B6" box="[367,455,1411,1437]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[194,1391,193,1907]" captionTargetId="figure-19@6.[180,1407,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Acentrella fimbriata sp. nov. nymph, head structures: (A) head capsule, frontal view; (B) antenna; (C) labrum, with different setae emphasized on each side; (D) left (angulate) mandible; (E) right (planate) mandible; (F) incisors of left mandible; (G) incisors of right mandible. Scale bars: A–E = 0.1 mm; F–G = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6450845" httpUri="https://zenodo.org/record/6450845/files/figure.png" pageId="2" pageNumber="353">Fig. 3C</figureCitation>
) approximately 1.5 times wider than long, broadly rounded anteriorly; anteromedian notch shallow, each side with one regular row of brush-like setae, arranged in pairs along anterior margin; dorsal surface with one median pair of long, simple setae, and lateral row of six smaller setae extending to margin; dorsal surface otherwise with scattered simple, hairlike setae. Left (angulate) mandible (
<figureCitation id="218868EFF326833A32164EBEFB159022" box="[1074,1213,1519,1545]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[194,1391,193,1907]" captionTargetId="figure-19@6.[180,1407,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Acentrella fimbriata sp. nov. nymph, head structures: (A) head capsule, frontal view; (B) antenna; (C) labrum, with different setae emphasized on each side; (D) left (angulate) mandible; (E) right (planate) mandible; (F) incisors of left mandible; (G) incisors of right mandible. Scale bars: A–E = 0.1 mm; F–G = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6450845" httpUri="https://zenodo.org/record/6450845/files/figure.png" pageId="2" pageNumber="353">Figs 3D, 3F</figureCitation>
) incisors not fused, with seven bluntly pointed teeth; prostheca comb-shaped; margin between mola and prostheca smooth, without setae; molar surface with one enlarged and blunt tooth at innermost edge, followed by about seven smaller molar teeth. Right (planate) mandible (
<figureCitation id="218868EFF326833A37DA4D0AFD2D905E" box="[510,645,1627,1653]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[194,1391,193,1907]" captionTargetId="figure-19@6.[180,1407,181,1919]" captionTargetPageId="6" captionText="FIGURE 3. Acentrella fimbriata sp. nov. nymph, head structures: (A) head capsule, frontal view; (B) antenna; (C) labrum, with different setae emphasized on each side; (D) left (angulate) mandible; (E) right (planate) mandible; (F) incisors of left mandible; (G) incisors of right mandible. Scale bars: A–E = 0.1 mm; F–G = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6450845" httpUri="https://zenodo.org/record/6450845/files/figure.png" pageId="2" pageNumber="353">Figs 3E, 3G</figureCitation>
) incisors not fused, with seven bluntly pointed teeth; prostheca slender, with numerous sharp teeth; edge between mola and prostheca smooth, without setae; molar area with numerous small, round teeth. Maxilla (
<figureCitation id="218868EFF326833A37F84DF2FDF39096" box="[476,603,1699,1725]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[192,1392,190,1903]" captionTargetId="figure-19@7.[181,1406,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Acentrella fimbriata sp. nov., nymph, mouthparts: (A) maxilla; (B) apex of galae-lacinia; (C) maxillary palp; (D) labium; (E) glossa; (F) paraglossa; (G) labial palp; (H) hypopharynx. Scale bars: A, C = 0.2 mm; B, H = 0.05 mm; D–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450847" httpUri="https://zenodo.org/record/6450847/files/figure.png" pageId="2" pageNumber="353">Figs 4A–B</figureCitation>
) with four large teeth on galea-lacinia, rounded at tips; row of five long, complex setae, and row of eight smaller simple setae along margin. Maxillary palp 2-segmented, with scattered small, blunt setae; proximal segment length approximately 3.0 times width, longer than distal segment; articulation between segments distinctly narrowed; distal segment with distinct, nipple-like tip at apex (
<figureCitation id="218868EFF326833A32BC4C5EFB5D9102" box="[1176,1269,1807,1833]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[192,1392,190,1903]" captionTargetId="figure-19@7.[181,1406,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Acentrella fimbriata sp. nov., nymph, mouthparts: (A) maxilla; (B) apex of galae-lacinia; (C) maxillary palp; (D) labium; (E) glossa; (F) paraglossa; (G) labial palp; (H) hypopharynx. Scale bars: A, C = 0.2 mm; B, H = 0.05 mm; D–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450847" httpUri="https://zenodo.org/record/6450847/files/figure.png" pageId="2" pageNumber="353">Figs 4C</figureCitation>
). Labium (
<figureCitation id="218868EFF326833A334B4C5EFF169166" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[192,1392,190,1903]" captionTargetId="figure-19@7.[181,1406,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Acentrella fimbriata sp. nov., nymph, mouthparts: (A) maxilla; (B) apex of galae-lacinia; (C) maxillary palp; (D) labium; (E) glossa; (F) paraglossa; (G) labial palp; (H) hypopharynx. Scale bars: A, C = 0.2 mm; B, H = 0.05 mm; D–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450847" httpUri="https://zenodo.org/record/6450847/files/figure.png" pageId="2" pageNumber="353">Fig. 4D</figureCitation>
): Glossae (
<figureCitation id="218868EFF326833A371D4C62FE269166" box="[313,398,1843,1869]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[192,1392,190,1903]" captionTargetId="figure-19@7.[181,1406,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Acentrella fimbriata sp. nov., nymph, mouthparts: (A) maxilla; (B) apex of galae-lacinia; (C) maxillary palp; (D) labium; (E) glossa; (F) paraglossa; (G) labial palp; (H) hypopharynx. Scale bars: A, C = 0.2 mm; B, H = 0.05 mm; D–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450847" httpUri="https://zenodo.org/record/6450847/files/figure.png" pageId="2" pageNumber="353">Fig. 4E</figureCitation>
) with five acute, serrate setae apically, and row of six acute, robust setae along inner margin, paraglossae (
<figureCitation id="218868EFF326833A370F4C06FE28915A" box="[299,384,1879,1905]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[192,1392,190,1903]" captionTargetId="figure-19@7.[181,1406,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Acentrella fimbriata sp. nov., nymph, mouthparts: (A) maxilla; (B) apex of galae-lacinia; (C) maxillary palp; (D) labium; (E) glossa; (F) paraglossa; (G) labial palp; (H) hypopharynx. Scale bars: A, C = 0.2 mm; B, H = 0.05 mm; D–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450847" httpUri="https://zenodo.org/record/6450847/files/figure.png" pageId="2" pageNumber="353">Fig. 4F</figureCitation>
) width approximately 1.3 times width of glossae, and length approximately 1.3 times length of glossae; apical part of paraglossa with 3 regular rows of long, serrate setae, and row of six submarginal acute setae. Labial palp (
<figureCitation id="218868EFF326833A37484CCEFE6C9192" box="[364,452,1951,1977]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[192,1392,190,1903]" captionTargetId="figure-19@7.[181,1406,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Acentrella fimbriata sp. nov., nymph, mouthparts: (A) maxilla; (B) apex of galae-lacinia; (C) maxillary palp; (D) labium; (E) glossa; (F) paraglossa; (G) labial palp; (H) hypopharynx. Scale bars: A, C = 0.2 mm; B, H = 0.05 mm; D–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450847" httpUri="https://zenodo.org/record/6450847/files/figure.png" pageId="2" pageNumber="353">Fig. 4G</figureCitation>
) 3-segmented, articulation between second and third segments indistinct; third segment rounded; ventral surface with sparse hairlike setae on first and second segments, but longer stout setae on third segment. Hypopharynx (
<figureCitation id="218868EFF327833B37A24BC6FE75969A" box="[390,477,151,177]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[192,1392,190,1903]" captionTargetId="figure-19@7.[181,1406,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Acentrella fimbriata sp. nov., nymph, mouthparts: (A) maxilla; (B) apex of galae-lacinia; (C) maxillary palp; (D) labium; (E) glossa; (F) paraglossa; (G) labial palp; (H) hypopharynx. Scale bars: A, C = 0.2 mm; B, H = 0.05 mm; D–F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450847" httpUri="https://zenodo.org/record/6450847/files/figure.png" pageId="3" pageNumber="354">Fig. 4H</figureCitation>
) nearly subrectangular; lingua apically rounded, covered with fine setae especially on apex; superlingua slightly rounded with small hair-like setae along apical margin.
</paragraph>
<paragraph id="B90C746AF327833B36E34B8FFB1694AD" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="354">
<emphasis id="8BC7A878F327833B36E34B8FFEB596D2" bold="true" box="[199,285,222,249]" italics="true" pageId="3" pageNumber="354">Thorax</emphasis>
(
<figureCitation id="218868EFF327833B370B4B8EFE2F96D2" box="[303,391,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1751,1775]" captionTargetBox="[151,1436,181,1728]" captionTargetId="figure-19@5.[151,1436,181,1728]" captionTargetPageId="5" captionText="FIGURE 2. Acentrella fimbriata sp. nov., nymph, dorsal views: (A) head and pronotum; (B) thorax; (C) abdomen; (D) abdominal terga V–VII; (E) abdominal gills. Scale bars: A = 0.2 mm; B–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6450843" httpUri="https://zenodo.org/record/6450843/files/figure.png" pageId="3" pageNumber="354">Fig. 2B</figureCitation>
). Coloration light brown to brownish with dark brown patterns. Surface of pronotum (
<figureCitation id="218868EFF327833B334B4B8EFF139736" captionStart="FIGURE 2" captionStartId="5.[151,250,1751,1775]" captionTargetBox="[151,1436,181,1728]" captionTargetId="figure-19@5.[151,1436,181,1728]" captionTargetPageId="5" captionText="FIGURE 2. Acentrella fimbriata sp. nov., nymph, dorsal views: (A) head and pronotum; (B) thorax; (C) abdomen; (D) abdominal terga V–VII; (E) abdominal gills. Scale bars: A = 0.2 mm; B–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6450843" httpUri="https://zenodo.org/record/6450843/files/figure.png" pageId="3" pageNumber="354">Fig. 2A</figureCitation>
) brownish with indistinct brown spots. Mesonotum light brown with submedian dark brown stripes and brown longitudinal stripes on half of anterior margin. Metanotum brown, with tiny posterolateral projection on each side. Plural sclerite brown. Sterna mostly pale. Furcasternal projections present on prosternum, but absent on meso- and metasterna. Legs (
<figureCitation id="218868EFF327833B374F4A3EFE6197A2" box="[363,457,366,393]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[186,1403,191,1901]" captionTargetId="figure-19@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Acentrella fimbriata sp. nov., nymph, thoracic and abdominal structures: (A) small posterolateral projection on metanotum; (B) foreleg; (C) tarsal claw; (D) abdominal tergum V; (E) abdominal gills I–VII; (F) gill V, enlarged view with details; (G) paraproct; (H) cerci and median filament. Scale bars: A, B, D, E, F = 0.2 mm; C, G = 0.02 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450849" httpUri="https://zenodo.org/record/6450849/files/figure.png" pageId="3" pageNumber="354">Figs 5B</figureCitation>
) yellow-brown but slightly paler than thorax; tarsal claws darker brownish. Femora broad, subequal to length of tibiae; dorsal margins of femora each with one regular row of long feathered bristlelike setae; length of longest setae approximately 2/3 width of respective femur (
<figureCitation id="218868EFF327833B32054AE6FBD297F9" box="[1057,1146,439,466]" captionStart="FIGURE 6" captionStartId="9.[151,250,1788,1812]" captionTargetBox="[151,1435,182,1764]" captionTargetId="figure-19@9.[151,1435,181,1764]" captionTargetPageId="9" captionText="FIGURE 6. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs: (A) head; (B) setae on scape and pedicel of antenna; (C) long feathered bristles on dorsal margin of femur; (D) dense, irregular row of long, simple setae on ventral margin of femur; (E) villopore; (F) tarsal claw. Scale bars: A = 500 µm; B, E, F = 100 µm; C = 200 µm; D = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.6450851" httpUri="https://zenodo.org/record/6450851/files/figure.png" pageId="3" pageNumber="354">Fig. 6C</figureCitation>
); ventral edge with dense irregular row of long, simple submarginal setae (
<figureCitation id="218868EFF327833B34964A8AFCA397DD" box="[690,779,475,502]" captionStart="FIGURE 6" captionStartId="9.[151,250,1788,1812]" captionTargetBox="[151,1435,182,1764]" captionTargetId="figure-19@9.[151,1435,181,1764]" captionTargetPageId="9" captionText="FIGURE 6. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs: (A) head; (B) setae on scape and pedicel of antenna; (C) long feathered bristles on dorsal margin of femur; (D) dense, irregular row of long, simple setae on ventral margin of femur; (E) villopore; (F) tarsal claw. Scale bars: A = 500 µm; B, E, F = 100 µm; C = 200 µm; D = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.6450851" httpUri="https://zenodo.org/record/6450851/files/figure.png" pageId="3" pageNumber="354">Fig. 6D</figureCitation>
); femoral villopore present, usually surrounded by rounded flattened setae (
<figureCitation id="218868EFF327833B376A4AAEFE009431" box="[334,424,511,538]" captionStart="FIGURE 6" captionStartId="9.[151,250,1788,1812]" captionTargetBox="[151,1435,182,1764]" captionTargetId="figure-19@9.[151,1435,181,1764]" captionTargetPageId="9" captionText="FIGURE 6. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs: (A) head; (B) setae on scape and pedicel of antenna; (C) long feathered bristles on dorsal margin of femur; (D) dense, irregular row of long, simple setae on ventral margin of femur; (E) villopore; (F) tarsal claw. Scale bars: A = 500 µm; B, E, F = 100 µm; C = 200 µm; D = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.6450851" httpUri="https://zenodo.org/record/6450851/files/figure.png" pageId="3" pageNumber="354">Fig. 6E</figureCitation>
). Tibiae with two rows of long, feathered, bristlelike setae dorsally and shorter, robust, feathered bristlelike setae ventrally. Tarsi with regular row of long, feathered bristlelike setae dorsally and short, robust, feathered bristlelike setae sparsely interspersed with fine, simple setae ventrally. Tarsal claw with row of 7–9 denticles increasing in length towards apex. All tarsal claws lack subapical setae (
<figureCitation id="218868EFF327833B320D493AFB2D94AE" box="[1065,1157,618,645]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[186,1403,191,1901]" captionTargetId="figure-19@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Acentrella fimbriata sp. nov., nymph, thoracic and abdominal structures: (A) small posterolateral projection on metanotum; (B) foreleg; (C) tarsal claw; (D) abdominal tergum V; (E) abdominal gills I–VII; (F) gill V, enlarged view with details; (G) paraproct; (H) cerci and median filament. Scale bars: A, B, D, E, F = 0.2 mm; C, G = 0.02 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450849" httpUri="https://zenodo.org/record/6450849/files/figure.png" pageId="3" pageNumber="354">Figs 5C</figureCitation>
,
<figureCitation id="218868EFF327833B32B4493AFB1994AD" box="[1168,1201,619,646]" captionStart="FIGURE 6" captionStartId="9.[151,250,1788,1812]" captionTargetBox="[151,1435,182,1764]" captionTargetId="figure-19@9.[151,1435,181,1764]" captionTargetPageId="9" captionText="FIGURE 6. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs: (A) head; (B) setae on scape and pedicel of antenna; (C) long feathered bristles on dorsal margin of femur; (D) dense, irregular row of long, simple setae on ventral margin of femur; (E) villopore; (F) tarsal claw. Scale bars: A = 500 µm; B, E, F = 100 µm; C = 200 µm; D = 300 µm." figureDoi="http://doi.org/10.5281/zenodo.6450851" httpUri="https://zenodo.org/record/6450851/files/figure.png" pageId="3" pageNumber="354">6F</figureCitation>
).
</paragraph>
<paragraph id="B90C746AF327833B36E349DFFBB0931A" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="354">
<emphasis id="8BC7A878F327833B36E349DFFE9D9482" bold="true" box="[199,309,654,681]" italics="true" pageId="3" pageNumber="354">Abdomen</emphasis>
(
<figureCitation id="218868EFF327833B376349DEFE0A9482" box="[327,418,655,681]" captionStart="FIGURE 2" captionStartId="5.[151,250,1751,1775]" captionTargetBox="[151,1436,181,1728]" captionTargetId="figure-19@5.[151,1436,181,1728]" captionTargetPageId="5" captionText="FIGURE 2. Acentrella fimbriata sp. nov., nymph, dorsal views: (A) head and pronotum; (B) thorax; (C) abdomen; (D) abdominal terga V–VII; (E) abdominal gills. Scale bars: A = 0.2 mm; B–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6450843" httpUri="https://zenodo.org/record/6450843/files/figure.png" pageId="3" pageNumber="354">Fig. 2C</figureCitation>
). Ventrally somewhat flattened. General coloration of terga yellowish-brown. Abdominal terga I–V light brown, each with transverse red brown stripe on posterior margin; faint transverse stripe on terga IV–V along median line. Terga I–III with indistinct patterns; terga IV–VIII each with large, dark brown, roundish medioanterior sigilla (cf.
<bibRefCitation id="DD22099BF327833B379749AAFD63953E" author="Kluge, N. J. &amp; Novikova, E. A." box="[435,715,763,789]" pageId="3" pageNumber="354" pagination="1 - 56" refId="ref9109" refString="Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera: Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56. https: // doi. org / 10.15298 / rusentj. 20.1.01" type="journal article" year="2011">Kluge &amp; Novikova 2011</bibRefCitation>
: Fig. 99) decreasing in size backwards; anterior margins of terga IV–IX with transverse dark brown stripes; terga IX–X light brown without patterns. Surfaces of terga with scattered fine setae, sparse scale-like setae along margins. Posterior margins (
<figureCitation id="218868EFF327833B35874812FC559576" box="[931,1021,835,861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1751,1775]" captionTargetBox="[151,1436,181,1728]" captionTargetId="figure-19@5.[151,1436,181,1728]" captionTargetPageId="5" captionText="FIGURE 2. Acentrella fimbriata sp. nov., nymph, dorsal views: (A) head and pronotum; (B) thorax; (C) abdomen; (D) abdominal terga V–VII; (E) abdominal gills. Scale bars: A = 0.2 mm; B–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6450843" httpUri="https://zenodo.org/record/6450843/files/figure.png" pageId="3" pageNumber="354">Fig. 2D</figureCitation>
) of terga each with long translucent scales, more abundant posterolaterally; these margins with small, poorly-developed, acute denticles (
<figureCitation id="218868EFF327833B32DF4836FAF095AA" box="[1275,1368,870,897]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[186,1403,191,1901]" captionTargetId="figure-19@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Acentrella fimbriata sp. nov., nymph, thoracic and abdominal structures: (A) small posterolateral projection on metanotum; (B) foreleg; (C) tarsal claw; (D) abdominal tergum V; (E) abdominal gills I–VII; (F) gill V, enlarged view with details; (G) paraproct; (H) cerci and median filament. Scale bars: A, B, D, E, F = 0.2 mm; C, G = 0.02 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450849" httpUri="https://zenodo.org/record/6450849/files/figure.png" pageId="3" pageNumber="354">Figs 5D</figureCitation>
,
<figureCitation id="218868EFF327833B33474836FA3C95AA" box="[1379,1428,871,897]" captionStart="FIGURE 10" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[152,1435,182,1829]" captionTargetId="figure-19@13.[151,1436,181,1831]" captionTargetPageId="13" captionText="FIGURE 10. Acentrella fimbriata sp. nov., scanning electron micrographs, A–D nymph, E–F egg: (A) abdominal terga IV–VI; (B) setae and scales on abdominal tergum V; (C) gill with long setae; (D) median filament; (E) egg, general view; (F) egg, longitudinal tissue of inner layer. Scale bars: A = 400 µm; B, D = 100 µm; C = 0.5 mm; E = 50 µm; F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6450859" httpUri="https://zenodo.org/record/6450859/files/figure.png" pageId="3" pageNumber="354">10A</figureCitation>
) and long-oval translucent scales with increasing number in posterolateral area; truncate, translucent scales sparsely located along posterior margin (
<figureCitation id="218868EFF327833B37DB48FEFD2595E2" box="[511,653,943,969]" captionStart="FIGURE 10" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[152,1435,182,1829]" captionTargetId="figure-19@13.[151,1436,181,1831]" captionTargetPageId="13" captionText="FIGURE 10. Acentrella fimbriata sp. nov., scanning electron micrographs, A–D nymph, E–F egg: (A) abdominal terga IV–VI; (B) setae and scales on abdominal tergum V; (C) gill with long setae; (D) median filament; (E) egg, general view; (F) egg, longitudinal tissue of inner layer. Scale bars: A = 400 µm; B, D = 100 µm; C = 0.5 mm; E = 50 µm; F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6450859" httpUri="https://zenodo.org/record/6450859/files/figure.png" pageId="3" pageNumber="354">Figs 10A–B</figureCitation>
), with scattered long-oval, translucent scales, increased in number on tergum X. Gills (
<figureCitation id="218868EFF327833B37784882FE1095C6" box="[348,440,979,1005]" captionStart="FIGURE 2" captionStartId="5.[151,250,1751,1775]" captionTargetBox="[151,1436,181,1728]" captionTargetId="figure-19@5.[151,1436,181,1728]" captionTargetPageId="5" captionText="FIGURE 2. Acentrella fimbriata sp. nov., nymph, dorsal views: (A) head and pronotum; (B) thorax; (C) abdomen; (D) abdominal terga V–VII; (E) abdominal gills. Scale bars: A = 0.2 mm; B–E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6450843" httpUri="https://zenodo.org/record/6450843/files/figure.png" pageId="3" pageNumber="354">Figs 2E</figureCitation>
,
<figureCitation id="218868EFF327833B37E34883FE4F95C6" box="[455,487,978,1005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[186,1403,191,1901]" captionTargetId="figure-19@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Acentrella fimbriata sp. nov., nymph, thoracic and abdominal structures: (A) small posterolateral projection on metanotum; (B) foreleg; (C) tarsal claw; (D) abdominal tergum V; (E) abdominal gills I–VII; (F) gill V, enlarged view with details; (G) paraproct; (H) cerci and median filament. Scale bars: A, B, D, E, F = 0.2 mm; C, G = 0.02 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450849" httpUri="https://zenodo.org/record/6450849/files/figure.png" pageId="3" pageNumber="354">5E</figureCitation>
) on segments I–VII, simple, with tracheation, though indistinct on segments I and VII; margins smooth with scattered fine setae; coloration white. Gills on segments II–VII with numerous long, fine, simple setae apically (
<figureCitation id="218868EFF327833B37E94F4AFD8D921E" box="[461,549,1050,1077]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[186,1403,191,1901]" captionTargetId="figure-19@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Acentrella fimbriata sp. nov., nymph, thoracic and abdominal structures: (A) small posterolateral projection on metanotum; (B) foreleg; (C) tarsal claw; (D) abdominal tergum V; (E) abdominal gills I–VII; (F) gill V, enlarged view with details; (G) paraproct; (H) cerci and median filament. Scale bars: A, B, D, E, F = 0.2 mm; C, G = 0.02 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450849" httpUri="https://zenodo.org/record/6450849/files/figure.png" pageId="3" pageNumber="354">Figs 5F</figureCitation>
,
<figureCitation id="218868EFF327833B34154F4AFDCC921E" box="[561,612,1051,1077]" captionStart="FIGURE 10" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[152,1435,182,1829]" captionTargetId="figure-19@13.[151,1436,181,1831]" captionTargetPageId="13" captionText="FIGURE 10. Acentrella fimbriata sp. nov., scanning electron micrographs, A–D nymph, E–F egg: (A) abdominal terga IV–VI; (B) setae and scales on abdominal tergum V; (C) gill with long setae; (D) median filament; (E) egg, general view; (F) egg, longitudinal tissue of inner layer. Scale bars: A = 400 µm; B, D = 100 µm; C = 0.5 mm; E = 50 µm; F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6450859" httpUri="https://zenodo.org/record/6450859/files/figure.png" pageId="3" pageNumber="354">10C</figureCitation>
). Sterna (
<figureCitation id="218868EFF327833B34F44F4AFC9D921E" box="[720,821,1051,1077]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="10.[151,250,1910,1934]" captionStartId-1="11.[151,250,1785,1809]" captionStartId-2="12.[151,250,1882,1906]" captionTargetBox-0="[151,1436,562,1885]" captionTargetBox-1="[152,1436,182,1761]" captionTargetBox-2="[152,1436,183,1857]" captionTargetId-0="figure-205@10.[151,1436,561,1886]" captionTargetId-1="figure-19@11.[151,1436,181,1761]" captionTargetId-2="figure-19@12.[151,1436,181,1858]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 7. Acentrella fimbriata sp. nov., nymph, scanning electron micrograph: abdomen, ventral view. Scale bar = 0.5 mm" captionText-1="FIGURE 8. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A) sterna 3–5; (B) sterna 5–7; (C) sterna 7–10. Scale bars: A, B = 400 µm; C = 500 µm" captionText-2="FIGURE 9. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A–C) details of sternum 8; (D–E) details of sternum 9; (F) paraproct. Scale bars: A = 200 µm; B, D, F = 50 µm; C = 30 µm; E = 20 µm." figureDoi-0="http://doi.org/10.5281/zenodo.6450853" figureDoi-1="http://doi.org/10.5281/zenodo.6450855" figureDoi-2="http://doi.org/10.5281/zenodo.6450857" httpUri-0="https://zenodo.org/record/6450853/files/figure.png" httpUri-1="https://zenodo.org/record/6450855/files/figure.png" httpUri-2="https://zenodo.org/record/6450857/files/figure.png" pageId="3" pageNumber="354">Figs 7–9</figureCitation>
) generally yellowish-white; posterior margins without marginal spines; sterna I–VII (
<figureCitation id="218868EFF327833B37D74F6EFDF19272" box="[499,601,1087,1113]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="10.[151,250,1910,1934]" captionStartId-1="11.[151,250,1785,1809]" captionTargetBox-0="[151,1436,562,1885]" captionTargetBox-1="[152,1436,182,1761]" captionTargetId-0="figure-205@10.[151,1436,561,1886]" captionTargetId-1="figure-19@11.[151,1436,181,1761]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 7. Acentrella fimbriata sp. nov., nymph, scanning electron micrograph: abdomen, ventral view. Scale bar = 0.5 mm" captionText-1="FIGURE 8. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A) sterna 3–5; (B) sterna 5–7; (C) sterna 7–10. Scale bars: A, B = 400 µm; C = 500 µm" figureDoi-0="http://doi.org/10.5281/zenodo.6450853" figureDoi-1="http://doi.org/10.5281/zenodo.6450855" httpUri-0="https://zenodo.org/record/6450853/files/figure.png" httpUri-1="https://zenodo.org/record/6450855/files/figure.png" pageId="3" pageNumber="354">Figs 7–8</figureCitation>
) without multidentated scales; sterna VIII–X (
<figureCitation id="218868EFF327833B324F4F6EFB609272" box="[1131,1224,1087,1113]" captionStart="FIGURE 8" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[152,1436,182,1761]" captionTargetId="figure-19@11.[151,1436,181,1761]" captionTargetPageId="11" captionText="FIGURE 8. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A) sterna 3–5; (B) sterna 5–7; (C) sterna 7–10. Scale bars: A, B = 400 µm; C = 500 µm" figureDoi="http://doi.org/10.5281/zenodo.6450855" httpUri="https://zenodo.org/record/6450855/files/figure.png" pageId="3" pageNumber="354">Figs 8C</figureCitation>
,
<figureCitation id="218868EFF327833B32F14F6EFB4C9272" box="[1237,1252,1087,1113]" captionStart="FIGURE 9" captionStartId="12.[151,250,1882,1906]" captionTargetBox="[152,1436,183,1857]" captionTargetId="figure-19@12.[151,1436,181,1858]" captionTargetPageId="12" captionText="FIGURE 9. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A–C) details of sternum 8; (D–E) details of sternum 9; (F) paraproct. Scale bars: A = 200 µm; B, D, F = 50 µm; C = 30 µm; E = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6450857" httpUri="https://zenodo.org/record/6450857/files/figure.png" pageId="3" pageNumber="354">9</figureCitation>
) with multidentated scales; sterna VIII &amp; IX (
<figureCitation id="218868EFF327833B37CA4F32FD9E9256" box="[494,566,1123,1149]" captionStart="FIGURE 7" captionStartId="10.[151,250,1910,1934]" captionTargetBox="[151,1436,562,1885]" captionTargetId="figure-205@10.[151,1436,561,1886]" captionTargetPageId="10" captionText="FIGURE 7. Acentrella fimbriata sp. nov., nymph, scanning electron micrograph: abdomen, ventral view. Scale bar = 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.6450853" httpUri="https://zenodo.org/record/6450853/files/figure.png" pageId="3" pageNumber="354">Figs 7</figureCitation>
,
<figureCitation id="218868EFF327833B34664F32FDCC9256" box="[578,612,1123,1149]" captionStart="FIGURE 8" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[152,1436,182,1761]" captionTargetId="figure-19@11.[151,1436,181,1761]" captionTargetPageId="11" captionText="FIGURE 8. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A) sterna 3–5; (B) sterna 5–7; (C) sterna 7–10. Scale bars: A, B = 400 µm; C = 500 µm" figureDoi="http://doi.org/10.5281/zenodo.6450855" httpUri="https://zenodo.org/record/6450855/files/figure.png" pageId="3" pageNumber="354">8C</figureCitation>
,
<figureCitation id="218868EFF327833B34544F32FDD79256" box="[624,639,1123,1149]" captionStart="FIGURE 9" captionStartId="12.[151,250,1882,1906]" captionTargetBox="[152,1436,183,1857]" captionTargetId="figure-19@12.[151,1436,181,1858]" captionTargetPageId="12" captionText="FIGURE 9. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A–C) details of sternum 8; (D–E) details of sternum 9; (F) paraproct. Scale bars: A = 200 µm; B, D, F = 50 µm; C = 30 µm; E = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6450857" httpUri="https://zenodo.org/record/6450857/files/figure.png" pageId="3" pageNumber="354">9</figureCitation>
) with pair of anterior patches of scales and scales on posterior margins. Paraproct (
<figureCitation id="218868EFF327833B37374FD6FEDA928A" box="[275,370,1158,1185]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[186,1403,191,1901]" captionTargetId="figure-19@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Acentrella fimbriata sp. nov., nymph, thoracic and abdominal structures: (A) small posterolateral projection on metanotum; (B) foreleg; (C) tarsal claw; (D) abdominal tergum V; (E) abdominal gills I–VII; (F) gill V, enlarged view with details; (G) paraproct; (H) cerci and median filament. Scale bars: A, B, D, E, F = 0.2 mm; C, G = 0.02 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450849" httpUri="https://zenodo.org/record/6450849/files/figure.png" pageId="3" pageNumber="354">Figs 5G</figureCitation>
,
<figureCitation id="218868EFF327833B375B4FD6FE0A928A" box="[383,418,1159,1185]" captionStart="FIGURE 8" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[152,1436,182,1761]" captionTargetId="figure-19@11.[151,1436,181,1761]" captionTargetPageId="11" captionText="FIGURE 8. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A) sterna 3–5; (B) sterna 5–7; (C) sterna 7–10. Scale bars: A, B = 400 µm; C = 500 µm" figureDoi="http://doi.org/10.5281/zenodo.6450855" httpUri="https://zenodo.org/record/6450855/files/figure.png" pageId="3" pageNumber="354">8C</figureCitation>
,
<figureCitation id="218868EFF327833B37944FD6FE66928A" box="[432,462,1159,1185]" captionStart="FIGURE 9" captionStartId="12.[151,250,1882,1906]" captionTargetBox="[152,1436,183,1857]" captionTargetId="figure-19@12.[151,1436,181,1858]" captionTargetPageId="12" captionText="FIGURE 9. Acentrella fimbriata sp. nov., nymph, scanning electron micrographs, abdominal sterna: (A–C) details of sternum 8; (D–E) details of sternum 9; (F) paraproct. Scale bars: A = 200 µm; B, D, F = 50 µm; C = 30 µm; E = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6450857" httpUri="https://zenodo.org/record/6450857/files/figure.png" pageId="3" pageNumber="354">9F</figureCitation>
) brown, without posterolateral projection; surface scattered with numerous pores and fine, short setae and small, blunt denticles on inner margin. Cerci yellowish-brown; length approximately 3/4body length. Posterior margins of segments with short fine spines on each segment, inner margins of cerci each with row of swimming bristles on posterior half, increasing in length towards apex. Median caudal filament reduced, composed of approximately 5–7 segments, without swimming setae (
<figureCitation id="218868EFF327833B354A4E46FC64931A" box="[878,972,1302,1329]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[186,1403,191,1901]" captionTargetId="figure-19@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Acentrella fimbriata sp. nov., nymph, thoracic and abdominal structures: (A) small posterolateral projection on metanotum; (B) foreleg; (C) tarsal claw; (D) abdominal tergum V; (E) abdominal gills I–VII; (F) gill V, enlarged view with details; (G) paraproct; (H) cerci and median filament. Scale bars: A, B, D, E, F = 0.2 mm; C, G = 0.02 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6450849" httpUri="https://zenodo.org/record/6450849/files/figure.png" pageId="3" pageNumber="354">Figs 5H</figureCitation>
,
<figureCitation id="218868EFF327833B35F34E46FBA4931A" box="[983,1036,1303,1329]" captionStart="FIGURE 10" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[152,1435,182,1829]" captionTargetId="figure-19@13.[151,1436,181,1831]" captionTargetPageId="13" captionText="FIGURE 10. Acentrella fimbriata sp. nov., scanning electron micrographs, A–D nymph, E–F egg: (A) abdominal terga IV–VI; (B) setae and scales on abdominal tergum V; (C) gill with long setae; (D) median filament; (E) egg, general view; (F) egg, longitudinal tissue of inner layer. Scale bars: A = 400 µm; B, D = 100 µm; C = 0.5 mm; E = 50 µm; F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6450859" httpUri="https://zenodo.org/record/6450859/files/figure.png" pageId="3" pageNumber="354">10D</figureCitation>
).
</paragraph>
<paragraph id="B90C746AF327833B36E34E6AFAE891BE" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="354">
<emphasis id="8BC7A878F327833B36E34E6AFED7937F" bold="true" box="[199,383,1338,1365]" pageId="3" pageNumber="354">Male subimago</emphasis>
(
<figureCitation id="218868EFF327833B37AB4E6AFE4A937E" box="[399,482,1339,1365]" captionStart="FIGURE 11" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[152,1436,182,1841]" captionTargetId="figure-19@14.[151,1436,181,1841]" captionTargetPageId="14" captionText="FIGURE 11. Acentrella fimbriata sp. nov., male subimago: (A) dorsal habitus; (B) head and anterior thorax, lateral view; (C) abdomen, dorsal view; (D) forewing; (E) genitalia. Scale bars: A, D = 1 mm; B–C = 0.5 mm; E = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6450861" httpUri="https://zenodo.org/record/6450861/files/figure.png" pageId="3" pageNumber="354">Fig. 11</figureCitation>
). Body length
<quantity id="7E4BD98FF327833B34AF4E6BFD4D937E" box="[651,741,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="354" unit="mm" value="5.3">5.3 mm</quantity>
; forewing length
<quantity id="7E4BD98FF327833B35884E6BFBAE937E" box="[940,1030,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="354" unit="mm" value="5.2">5.2 mm</quantity>
.
<emphasis id="8BC7A878F327833B32364E6BFBFF937E" bold="true" box="[1042,1111,1338,1365]" italics="true" pageId="3" pageNumber="354">Head.</emphasis>
Hypognathous, dark brownish. Upper portion of compound eye orange. Lower portion of compound eye slightly darker than upper portion. Inner margins of turbinate eyes (
<figureCitation id="218868EFF327833B34244ED2FDCA93B6" box="[512,610,1411,1437]" captionStart="FIGURE 11" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[152,1436,182,1841]" captionTargetId="figure-19@14.[151,1436,181,1841]" captionTargetPageId="14" captionText="FIGURE 11. Acentrella fimbriata sp. nov., male subimago: (A) dorsal habitus; (B) head and anterior thorax, lateral view; (C) abdomen, dorsal view; (D) forewing; (E) genitalia. Scale bars: A, D = 1 mm; B–C = 0.5 mm; E = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6450861" httpUri="https://zenodo.org/record/6450861/files/figure.png" pageId="3" pageNumber="354">Fig. 11B</figureCitation>
) with space between them. Antennae: Coloration of middle and distal segments of flagellum darker brown than scape and pedicel.
<emphasis id="8BC7A878F327833B353E4EF7FCDE93EA" bold="true" box="[794,886,1446,1473]" italics="true" pageId="3" pageNumber="354">Thorax.</emphasis>
Coloration dark brown. Mesothoracic projection moderately pointed (
<figureCitation id="218868EFF327833B37A74E9AFE4393CE" box="[387,491,1483,1509]" captionStart="FIGURE 11" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[152,1436,182,1841]" captionTargetId="figure-19@14.[151,1436,181,1841]" captionTargetPageId="14" captionText="FIGURE 11. Acentrella fimbriata sp. nov., male subimago: (A) dorsal habitus; (B) head and anterior thorax, lateral view; (C) abdomen, dorsal view; (D) forewing; (E) genitalia. Scale bars: A, D = 1 mm; B–C = 0.5 mm; E = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6450861" httpUri="https://zenodo.org/record/6450861/files/figure.png" pageId="3" pageNumber="354">Fig. 11B</figureCitation>
). Legs: Coloration pale brownish to light brown. Leg lengths. Foreleg: femur
<quantity id="7E4BD98FF327833B334F4E9AFF6D9022" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="354" unit="mm" value="0.34">0.34 mm</quantity>
, tibia
<quantity id="7E4BD98FF327833B372F4EBEFEDA9022" box="[267,370,1519,1545]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="354" unit="mm" value="0.37">0.37 mm</quantity>
, tarsi 4 segments (proximal to distal, excluding claw)
<quantity id="7E4BD98FF327833B35F34EBEFB969022" box="[983,1086,1519,1545]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="354" unit="mm" value="0.08">0.08 mm</quantity>
,
<quantity id="7E4BD98FF327833B326E4EBEFB199022" box="[1098,1201,1519,1545]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="354" unit="mm" value="0.06">0.06 mm</quantity>
,
<quantity id="7E4BD98FF327833B32994EBEFA8C9022" box="[1213,1316,1519,1545]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="354" unit="mm" value="0.04">0.04 mm</quantity>
,
<quantity id="7E4BD98FF327833B33144EBEFA309022" box="[1328,1432,1519,1545]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="3" pageNumber="354" unit="mm" value="0.07">0.07 mm</quantity>
; midleg: femur
<quantity id="7E4BD98FF327833B37644D42FE0F9006" box="[320,423,1555,1581]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="354" unit="mm" value="0.33">0.33 mm</quantity>
, tibia
<quantity id="7E4BD98FF327833B37C94D42FDFC9006" box="[493,596,1555,1581]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="354" unit="mm" value="0.3">0.30 mm</quantity>
, tarsi 4 segments (proximal to distal, excluding claw)
<quantity id="7E4BD98FF327833B32994D43FA8C9006" box="[1213,1316,1554,1581]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="354" unit="mm" value="0.05">0.05 mm</quantity>
,
<quantity id="7E4BD98FF327833B33154D42FA309006" box="[1329,1432,1555,1581]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="354" unit="mm" value="0.03">0.03 mm</quantity>
,
<quantity id="7E4BD98FF327833B36B34D66FF56907A" box="[151,254,1591,1617]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="354" unit="mm" value="0.02">0.02 mm</quantity>
,
<quantity id="7E4BD98FF327833B372E4D67FEDA907A" box="[266,370,1590,1617]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="354" unit="mm" value="0.05">0.05 mm</quantity>
; hindleg: femur
<quantity id="7E4BD98FF327833B34084D66FD3B907A" box="[556,659,1591,1617]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="354" unit="mm" value="0.34">0.34 mm</quantity>
, tibia
<quantity id="7E4BD98FF327833B34FC4D66FC97907A" box="[728,831,1591,1617]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="3" pageNumber="354" unit="mm" value="0.36">0.36 mm</quantity>
, tarsi 4 segments (proximal to distal, excluding claw)
<quantity id="7E4BD98FF327833B36B34D0BFF56905E" box="[151,254,1626,1653]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="354" unit="mm" value="0.05">0.05 mm</quantity>
,
<quantity id="7E4BD98FF327833B372F4D0AFED9905E" box="[267,369,1627,1653]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="354" unit="mm" value="0.04">0.04 mm</quantity>
,
<quantity id="7E4BD98FF327833B375A4D0AFE4D905E" box="[382,485,1627,1653]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="354" unit="mm" value="0.02">0.02 mm</quantity>
,
<quantity id="7E4BD98FF327833B37D54D0BFDF1905E" box="[497,601,1626,1653]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="354" unit="mm" value="0.05">0.05 mm</quantity>
. Forewing (
<figureCitation id="218868EFF327833B34C44D0AFCED905E" box="[736,837,1627,1653]" captionStart="FIGURE 11" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[152,1436,182,1841]" captionTargetId="figure-19@14.[151,1436,181,1841]" captionTargetPageId="14" captionText="FIGURE 11. Acentrella fimbriata sp. nov., male subimago: (A) dorsal habitus; (B) head and anterior thorax, lateral view; (C) abdomen, dorsal view; (D) forewing; (E) genitalia. Scale bars: A, D = 1 mm; B–C = 0.5 mm; E = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6450861" httpUri="https://zenodo.org/record/6450861/files/figure.png" pageId="3" pageNumber="354">Fig. 11D</figureCitation>
) translucent, cloudy in costal and subcostal margins; marginal intercalary veins of first few interspaces distinctly different in length; marginal intercalary veins of middle part of wing more or less subequal; marginal intercalary veins of posterior part of wing tiny. Hind wing absent.
<emphasis id="8BC7A878F327833B36B34D97FEAD90CA" bold="true" box="[151,261,1734,1761]" italics="true" pageId="3" pageNumber="354">Abdomen</emphasis>
(
<figureCitation id="218868EFF327833B37304D96FED190CA" box="[276,377,1735,1761]" captionStart="FIGURE 11" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[152,1436,182,1841]" captionTargetId="figure-19@14.[151,1436,181,1841]" captionTargetPageId="14" captionText="FIGURE 11. Acentrella fimbriata sp. nov., male subimago: (A) dorsal habitus; (B) head and anterior thorax, lateral view; (C) abdomen, dorsal view; (D) forewing; (E) genitalia. Scale bars: A, D = 1 mm; B–C = 0.5 mm; E = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6450861" httpUri="https://zenodo.org/record/6450861/files/figure.png" pageId="3" pageNumber="354">Fig. 11C</figureCitation>
). Abdominal tergum I dark brown with red brown line along posterior margin; terga II–III light brown with red brown line along each posterior margin; terga IV–V light brown, each with pair of red brown areas on posterolateral margins; terga VI–X dark brown. Sterna light brown without distinct markings.
<emphasis id="8BC7A878F327833B32D64C5FFAF59102" bold="true" box="[1266,1373,1806,1833]" italics="true" pageId="3" pageNumber="354">Genitalia</emphasis>
(
<figureCitation id="218868EFF327833B334B4C5EFF6E9166" captionStart="FIGURE 11" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[152,1436,182,1841]" captionTargetId="figure-19@14.[151,1436,181,1841]" captionTargetPageId="14" captionText="FIGURE 11. Acentrella fimbriata sp. nov., male subimago: (A) dorsal habitus; (B) head and anterior thorax, lateral view; (C) abdomen, dorsal view; (D) forewing; (E) genitalia. Scale bars: A, D = 1 mm; B–C = 0.5 mm; E = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6450861" httpUri="https://zenodo.org/record/6450861/files/figure.png" pageId="3" pageNumber="354">Fig. 11E</figureCitation>
). Forceps with hair-like setae present on inner margin towards base; basal segment subquadrate, with length subequal to width; segment II with overall curved appearance, subcylindrical basally, strongly constricted medially, and slightly flared; segment III length near twice its width, rounded apically. Caudal filaments translucent.
</paragraph>
<paragraph id="B90C746AF327833B36E34CCEFE309192" blockId="3.[151,1437,151,1977]" box="[199,408,1951,1977]" pageId="3" pageNumber="354">
<emphasis id="8BC7A878F327833B36E34CCEFEB29192" bold="true" box="[199,282,1951,1977]" pageId="3" pageNumber="354">Imago.</emphasis>
Unknown.
</paragraph>
<caption id="EDCC24E2F320833C36B34CC6FB959184" ID-DOI="http://doi.org/10.5281/zenodo.6450837" ID-Zenodo-Dep="6450837" box="[151,1085,1943,1968]" httpUri="https://zenodo.org/record/6450837/files/figure.png" pageId="4" pageNumber="355" startId="4.[151,250,1943,1967]" targetBox="[174,1413,181,1919]" targetPageId="4">
<paragraph id="B90C746AF320833C36B34CC6FB959184" blockId="4.[151,1085,1943,1968]" box="[151,1085,1943,1968]" pageId="4" pageNumber="355">
<emphasis id="8BC7A878F320833C36B34CC6FEBB9184" bold="true" box="[151,275,1943,1967]" pageId="4" pageNumber="355">FIGURE 1.</emphasis>
<taxonomicName id="7EB30FE9F320833C373D4CC9FE4E919B" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[281,486,1944,1968]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="355" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F320833C373D4CC9FE4E919B" box="[281,486,1944,1968]" italics="true" pageId="4" pageNumber="355">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F320833C37C94CC9FD95919B" bold="true" box="[493,573,1944,1968]" pageId="4" pageNumber="355">
<taxonomicNameLabel id="90F41503F320833C37C94CC9FD95919B" box="[493,573,1944,1968]" pageId="4" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female nymph, dorsal habitus. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="EDCC24E2F321833D36B34D86FC049138" ID-DOI="http://doi.org/10.5281/zenodo.6450843" ID-Zenodo-Dep="6450843" httpUri="https://zenodo.org/record/6450843/files/figure.png" pageId="5" pageNumber="356" startId="5.[151,250,1751,1775]" targetBox="[151,1436,181,1728]" targetPageId="5">
<paragraph id="B90C746AF321833D36B34D86FC049138" blockId="5.[151,1436,1751,1812]" pageId="5" pageNumber="356">
<emphasis id="8BC7A878F321833D36B34D86FEBA90C4" bold="true" box="[151,274,1751,1775]" pageId="5" pageNumber="356">FIGURE 2.</emphasis>
<taxonomicName id="7EB30FE9F321833D373C4D89FE4C90DB" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[280,484,1752,1776]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F321833D373C4D89FE4C90DB" box="[280,484,1752,1776]" italics="true" pageId="5" pageNumber="356">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F321833D37CD4D89FD9190DB" bold="true" box="[489,569,1752,1776]" pageId="5" pageNumber="356">
<taxonomicNameLabel id="90F41503F321833D37CD4D89FD9190DB" box="[489,569,1752,1776]" pageId="5" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymph, dorsal views: (
<emphasis id="8BC7A878F321833D350F4D86FCED90C4" bold="true" box="[811,837,1751,1775]" pageId="5" pageNumber="356">A)</emphasis>
head and pronotum;
<emphasis id="8BC7A878F321833D323D4D86FB9390C4" bold="true" box="[1049,1083,1751,1775]" pageId="5" pageNumber="356">(B)</emphasis>
thorax; (
<emphasis id="8BC7A878F321833D32B24D86FB1990C4" bold="true" box="[1174,1201,1751,1775]" pageId="5" pageNumber="356">C)</emphasis>
abdomen;
<emphasis id="8BC7A878F321833D333B4D86FAEB90C4" bold="true" box="[1311,1347,1751,1775]" pageId="5" pageNumber="356">(D)</emphasis>
abdominal terga V–VII;
<emphasis id="8BC7A878F321833D37604DAAFECE9138" bold="true" box="[324,358,1787,1811]" pageId="5" pageNumber="356">(E)</emphasis>
abdominal gills. Scale bars: A = 0.2 mm; B–E = 0.5 mm.
</paragraph>
</caption>
<caption id="EDCC24E2F322833E36B34CC6FC3D91DC" ID-DOI="http://doi.org/10.5281/zenodo.6450845" ID-Zenodo-Dep="6450845" httpUri="https://zenodo.org/record/6450845/files/figure.png" pageId="6" pageNumber="357" startId="6.[151,250,1943,1967]" targetBox="[194,1391,193,1907]" targetPageId="6">
<paragraph id="B90C746AF322833E36B34CC6FC3D91DC" blockId="6.[151,1437,1943,2040]" pageId="6" pageNumber="357">
<emphasis id="8BC7A878F322833E36B34CC6FEBA9184" bold="true" box="[151,274,1943,1967]" pageId="6" pageNumber="357">FIGURE 3.</emphasis>
<taxonomicName id="7EB30FE9F322833E37334CC9FE4B919B" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[279,483,1944,1968]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F322833E37334CC9FE4B919B" box="[279,483,1944,1968]" italics="true" pageId="6" pageNumber="357">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F322833E37CD4CC9FD91919B" bold="true" box="[489,569,1944,1968]" pageId="6" pageNumber="357">
<taxonomicNameLabel id="90F41503F322833E37CD4CC9FD91919B" box="[489,569,1944,1968]" pageId="6" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
nymph, head structures:
<emphasis id="8BC7A878F322833E35114CC6FCF09184" bold="true" box="[821,856,1943,1967]" pageId="6" pageNumber="357">(A)</emphasis>
head capsule, frontal view;
<emphasis id="8BC7A878F322833E32554CC6FB3B9184" bold="true" box="[1137,1171,1943,1967]" pageId="6" pageNumber="357">(B)</emphasis>
antenna;
<emphasis id="8BC7A878F322833E32D74CC6FABE9184" bold="true" box="[1267,1302,1943,1967]" pageId="6" pageNumber="357">(C)</emphasis>
labrum, with different setae emphasized on each side; (
<emphasis id="8BC7A878F322833E341C4CEAFDFA91F8" bold="true" box="[568,594,1979,2003]" pageId="6" pageNumber="357">D)</emphasis>
left (angulate) mandible;
<emphasis id="8BC7A878F322833E35724CEAFCD091F8" bold="true" box="[854,888,1979,2003]" pageId="6" pageNumber="357">(E)</emphasis>
right (planate) mandible;
<emphasis id="8BC7A878F322833E32584CEAFB3491F8" bold="true" box="[1148,1180,1979,2003]" pageId="6" pageNumber="357">(F)</emphasis>
incisors of left mandible;
<emphasis id="8BC7A878F322833E36B34C8EFF1491DC" bold="true" box="[151,188,2015,2039]" pageId="6" pageNumber="357">(G)</emphasis>
incisors of right mandible. Scale bars: A–E = 0.1 mm; F–G = 0.02 mm.
</paragraph>
</caption>
<caption id="EDCC24E2F323833F36B34CC6FF6E91DC" ID-DOI="http://doi.org/10.5281/zenodo.6450847" ID-Zenodo-Dep="6450847" httpUri="https://zenodo.org/record/6450847/files/figure.png" pageId="7" pageNumber="358" startId="7.[151,250,1943,1967]" targetBox="[192,1392,190,1903]" targetPageId="7">
<paragraph id="B90C746AF323833F36B34CC6FF6E91DC" blockId="7.[151,1437,1943,2039]" pageId="7" pageNumber="358">
<emphasis id="8BC7A878F323833F36B34CC6FEBB9184" bold="true" box="[151,275,1943,1967]" pageId="7" pageNumber="358">FIGURE 4.</emphasis>
<taxonomicName id="7EB30FE9F323833F373E4CC9FE4F919B" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[282,487,1944,1968]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F323833F373E4CC9FE4F919B" box="[282,487,1944,1968]" italics="true" pageId="7" pageNumber="358">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F323833F37CB4CC9FD97919B" bold="true" box="[495,575,1944,1968]" pageId="7" pageNumber="358">
<taxonomicNameLabel id="90F41503F323833F37CB4CC9FD97919B" box="[495,575,1944,1968]" pageId="7" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymph, mouthparts:
<emphasis id="8BC7A878F323833F35044CC6FCEC9184" bold="true" box="[800,836,1943,1967]" pageId="7" pageNumber="358">(A)</emphasis>
maxilla;
<emphasis id="8BC7A878F323833F35804CC6FC6E9184" bold="true" box="[932,966,1943,1967]" pageId="7" pageNumber="358">(B)</emphasis>
apex of galae-lacinia; (
<emphasis id="8BC7A878F323833F32924CC6FB799184" bold="true" box="[1206,1233,1943,1967]" pageId="7" pageNumber="358">C)</emphasis>
maxillary palp;
<emphasis id="8BC7A878F323833F335D4CC6FA359184" bold="true" box="[1401,1437,1943,1967]" pageId="7" pageNumber="358">(D)</emphasis>
labium;
<emphasis id="8BC7A878F323833F36CF4CEAFEA591F8" bold="true" box="[235,269,1979,2003]" pageId="7" pageNumber="358">(E)</emphasis>
glossa;
<emphasis id="8BC7A878F323833F37454CEAFE2A91F8" bold="true" box="[353,386,1979,2003]" pageId="7" pageNumber="358">(F)</emphasis>
paraglossa;
<emphasis id="8BC7A878F323833F34254CEAFD8E91F8" bold="true" box="[513,550,1979,2003]" pageId="7" pageNumber="358">(G)</emphasis>
labial palp;
<emphasis id="8BC7A878F323833F34814CEAFD6291F8" bold="true" box="[677,714,1979,2003]" pageId="7" pageNumber="358">(H)</emphasis>
hypopharynx. Scale bars: A, C = 0.2 mm; B, H = 0.05 mm; D–F = 0.1 mm.
</paragraph>
</caption>
<caption id="EDCC24E2F32C833036B34CC6FAC991DC" ID-DOI="http://doi.org/10.5281/zenodo.6450849" ID-Zenodo-Dep="6450849" httpUri="https://zenodo.org/record/6450849/files/figure.png" pageId="8" pageNumber="359" startId="8.[151,250,1943,1967]" targetBox="[186,1403,191,1901]" targetPageId="8">
<paragraph id="B90C746AF32C833036B34CC6FAC991DC" blockId="8.[151,1436,1943,2040]" pageId="8" pageNumber="359">
<emphasis id="8BC7A878F32C833036B34CC6FEBD919B" bold="true" box="[151,277,1943,1968]" pageId="8" pageNumber="359">FIGURE 5.</emphasis>
<taxonomicName id="7EB30FE9F32C833037394CC9FE44919B" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[285,492,1944,1968]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F32C833037394CC9FE44919B" box="[285,492,1944,1968]" italics="true" pageId="8" pageNumber="359">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F32C833037D14CC9FDEF919B" bold="true" box="[501,583,1944,1968]" pageId="8" pageNumber="359">
<taxonomicNameLabel id="90F41503F32C833037D14CC9FDEF919B" box="[501,583,1944,1968]" pageId="8" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymph, thoracic and abdominal structures:
<emphasis id="8BC7A878F32C833032334CC6FB939184" bold="true" box="[1047,1083,1943,1967]" pageId="8" pageNumber="359">(A)</emphasis>
small posterolateral projection on metanotum;
<emphasis id="8BC7A878F32C8330373C4CEAFE9291F8" bold="true" box="[280,314,1979,2003]" pageId="8" pageNumber="359">(B)</emphasis>
foreleg;
<emphasis id="8BC7A878F32C833037BD4CEAFE1491F8" bold="true" box="[409,444,1979,2003]" pageId="8" pageNumber="359">(C)</emphasis>
tarsal claw;
<emphasis id="8BC7A878F32C833034664CEAFDCE91F8" bold="true" box="[578,614,1979,2003]" pageId="8" pageNumber="359">(D)</emphasis>
abdominal tergum V;
<emphasis id="8BC7A878F32C8330356B4CEAFCD991F8" bold="true" box="[847,881,1979,2003]" pageId="8" pageNumber="359">(E)</emphasis>
abdominal gills I–VII;
<emphasis id="8BC7A878F32C8330324C4CEAFB2091F8" bold="true" box="[1128,1160,1979,2003]" pageId="8" pageNumber="359">(F)</emphasis>
gill V, enlarged view with details;
<emphasis id="8BC7A878F32C833036C34C8EFEA491DC" bold="true" box="[231,268,2015,2039]" pageId="8" pageNumber="359">(G)</emphasis>
paraproct;
<emphasis id="8BC7A878F32C8330375B4C8EFE0C91DC" bold="true" box="[383,420,2015,2039]" pageId="8" pageNumber="359">(H)</emphasis>
cerci and median filament. Scale bars: A, B, D, E, F = 0.2 mm; C, G = 0.02 mm; H = 0.1 mm.
</paragraph>
</caption>
<caption id="EDCC24E2F32D833136B34DADFB599176" ID-DOI="http://doi.org/10.5281/zenodo.6450851" ID-Zenodo-Dep="6450851" httpUri="https://zenodo.org/record/6450851/files/figure.png" pageId="9" pageNumber="360" startId="9.[151,250,1788,1812]" targetBox="[151,1435,182,1764]" targetPageId="9">
<paragraph id="B90C746AF32D833136B34DADFB599176" blockId="9.[151,1436,1788,1885]" pageId="9" pageNumber="360">
<emphasis id="8BC7A878F32D833136B34DADFEBB913F" bold="true" box="[151,275,1788,1812]" pageId="9" pageNumber="360">FIGURE 6.</emphasis>
<taxonomicName id="7EB30FE9F32D8331373E4DADFE4F913F" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[282,487,1788,1812]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="360" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F32D8331373E4DADFE4F913F" box="[282,487,1788,1812]" italics="true" pageId="9" pageNumber="360">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F32D833137CA4DACFD97913E" bold="true" box="[494,575,1789,1813]" pageId="9" pageNumber="360">
<taxonomicNameLabel id="90F41503F32D833137CA4DACFD97913E" box="[494,575,1789,1813]" pageId="9" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymph, scanning electron micrographs: (
<emphasis id="8BC7A878F32D833135C94DADFBAF913F" bold="true" box="[1005,1031,1788,1812]" pageId="9" pageNumber="360">A)</emphasis>
head;
<emphasis id="8BC7A878F32D833132684DADFBC6913F" bold="true" box="[1100,1134,1788,1812]" pageId="9" pageNumber="360">(B)</emphasis>
setae on scape and pedicel of antenna;
<emphasis id="8BC7A878F32D833136D74C71FEBE9113" bold="true" box="[243,278,1824,1848]" pageId="9" pageNumber="360">(C)</emphasis>
long feathered bristles on dorsal margin of femur;
<emphasis id="8BC7A878F32D8331353C4C71FC949113" bold="true" box="[792,828,1824,1848]" pageId="9" pageNumber="360">(D)</emphasis>
dense, irregular row of long, simple setae on ventral margin of femur;
<emphasis id="8BC7A878F32D833136D84C15FEB79177" bold="true" box="[252,287,1860,1884]" pageId="9" pageNumber="360">(E)</emphasis>
villopore;
<emphasis id="8BC7A878F32D833137A94C15FE069177" bold="true" box="[397,430,1860,1884]" pageId="9" pageNumber="360">(F)</emphasis>
tarsal claw. Scale bars: A = 500 µm; B, E, F = 100 µm; C = 200 µm; D = 300 µm.
</paragraph>
</caption>
<paragraph id="B90C746AF32D833136E34CDAFD2A91C6" blockId="9.[151,1437,1930,2029]" pageId="9" pageNumber="360">
<emphasis id="8BC7A878F32D833136E34CDAFF5E918E" bold="true" box="[199,246,1931,1957]" pageId="9" pageNumber="360">Egg</emphasis>
(dissected from last nymphal instar). Oval shaped. Length about 103–115 µm, width about 70–75µm (
<figureCitation id="218868EFF32D8331334B4CDAFF6091E2" captionStart="FIGURE 10" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[152,1435,182,1829]" captionTargetId="figure-19@13.[151,1436,181,1831]" captionTargetPageId="13" captionText="FIGURE 10. Acentrella fimbriata sp. nov., scanning electron micrographs, A–D nymph, E–F egg: (A) abdominal terga IV–VI; (B) setae and scales on abdominal tergum V; (C) gill with long setae; (D) median filament; (E) egg, general view; (F) egg, longitudinal tissue of inner layer. Scale bars: A = 400 µm; B, D = 100 µm; C = 0.5 mm; E = 50 µm; F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6450859" httpUri="https://zenodo.org/record/6450859/files/figure.png" pageId="9" pageNumber="360">Fig. 10E</figureCitation>
). Extrachorion with transverse striation completely surrounding egg. Chorionic sculpturing with longitudinal striation on surface of entire egg (
<figureCitation id="218868EFF32D833134364C82FDDD91C6" box="[530,629,2003,2029]" captionStart="FIGURE 10" captionStartId="13.[151,250,1855,1879]" captionTargetBox="[152,1435,182,1829]" captionTargetId="figure-19@13.[151,1436,181,1831]" captionTargetPageId="13" captionText="FIGURE 10. Acentrella fimbriata sp. nov., scanning electron micrographs, A–D nymph, E–F egg: (A) abdominal terga IV–VI; (B) setae and scales on abdominal tergum V; (C) gill with long setae; (D) median filament; (E) egg, general view; (F) egg, longitudinal tissue of inner layer. Scale bars: A = 400 µm; B, D = 100 µm; C = 0.5 mm; E = 50 µm; F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6450859" httpUri="https://zenodo.org/record/6450859/files/figure.png" pageId="9" pageNumber="360">Fig. 10F</figureCitation>
).
</paragraph>
<paragraph id="B90C746AF32E833236E34BC6FC9997FA" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="361">
<emphasis id="8BC7A878F32E833236E34BC6FE1B969B" bold="true" box="[199,435,150,177]" pageId="10" pageNumber="361">Nymphal diagnosis.</emphasis>
The new species,
<taxonomicName id="7EB30FE9F32E833234A04BC6FCCF969A" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[644,871,151,177]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F32E833234A04BC6FCCF969A" box="[644,871,151,177]" italics="true" pageId="10" pageNumber="361">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F32E8332354A4BC6FC61969A" bold="true" box="[878,969,151,177]" pageId="10" pageNumber="361">
<taxonomicNameLabel id="90F41503F32E8332354A4BC6FC61969A" box="[878,969,151,177]" pageId="10" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from all other congeners by a combination of morphological characters (
<tableCitation id="F43141D1F32E8332348B4BEAFCAC96FE" box="[687,772,187,213]" captionStart="TABLE 1" captionStartId="23.[151,239,735,761]" captionTargetPageId="23" captionText="TABLE 1. Comparative morphological characters of Acentrella fimbriata sp. nov., Acentrella lapponica, and Acentrella (Liebebiella) vera." httpUri="http://table.plazi.org/id/EDCC24E2F333832F36B3498EF9C294D2" pageId="10" pageNumber="361" tableUuid="EDCC24E2F333832F36B3498EF9C294D2">Table 1</tableCitation>
): (i) body covered with long dense setae; (ii) the proximal segment of the maxillary palp approximately two times longer than the distal segment, and the distal segment with a nipple-like tip; (iii) margins of femora with irregular rows of dense setae; (iv) metatibia with two rows of feathered bristlelike setae on dorsal margin; (v) the absence of preapical setae on the tarsus; (vi) subapical setae of tarsal claw vestigial or absent; (vii) abdominal terga scattered with numerous long, translucent scales, at least some of which appear truncated; and the posterior margins of abdominal terga generally smooth, but sparsely with tiny, blunt denticles; (viii) margins of gills II–VII with densely covered with long, hairlike setae; and (ix) median filament reduced in length, being comprised of approximately 5–7 segments.
</paragraph>
</subSubSection>
<subSubSection id="F1A927E1F32E833236E34A8AFB9D9432" pageId="10" pageNumber="361" type="etymology">
<paragraph id="B90C746AF32E833236E34A8AFB9D9432" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="361">
<emphasis id="8BC7A878F32E833236E34A8AFEE097DE" bold="true" box="[199,328,475,501]" pageId="10" pageNumber="361">Etymology</emphasis>
. The specific epithet is from the Latin word for fringed, an allusion to the setose margins of various body parts. The common name of this species is the fringed small minnow mayfly.
</paragraph>
</subSubSection>
<caption id="EDCC24E2F32E833236B34C27FA3891A6" ID-DOI="http://doi.org/10.5281/zenodo.6450853" ID-Zenodo-Dep="6450853" box="[151,1424,1910,1934]" httpUri="https://zenodo.org/record/6450853/files/figure.png" pageId="10" pageNumber="361" startId="10.[151,250,1910,1934]" targetBox="[151,1436,562,1885]" targetPageId="10">
<paragraph id="B90C746AF32E833236B34C27FA3891A6" blockId="10.[151,1424,1910,1934]" box="[151,1424,1910,1934]" pageId="10" pageNumber="361">
<emphasis id="8BC7A878F32E833236B34C27FEBB91A5" bold="true" box="[151,275,1910,1934]" pageId="10" pageNumber="361">FIGURE 7.</emphasis>
<taxonomicName id="7EB30FE9F32E8332373D4C27FE4E91A5" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[281,486,1910,1934]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F32E8332373D4C27FE4E91A5" box="[281,486,1910,1934]" italics="true" pageId="10" pageNumber="361">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F32E833237C94C27FD9591A5" bold="true" box="[493,573,1910,1934]" pageId="10" pageNumber="361">
<taxonomicNameLabel id="90F41503F32E833237C94C27FD9591A5" box="[493,573,1910,1934]" pageId="10" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymph, scanning electron micrograph: abdomen, ventral view. Scale bar = 0.5 mm
</paragraph>
</caption>
<caption id="EDCC24E2F32F833336B34DA8FCA3911D" ID-DOI="http://doi.org/10.5281/zenodo.6450855" ID-Zenodo-Dep="6450855" httpUri="https://zenodo.org/record/6450855/files/figure.png" pageId="11" pageNumber="362" startId="11.[151,250,1785,1809]" targetBox="[152,1436,182,1761]" targetPageId="11">
<paragraph id="B90C746AF32F833336B34DA8FCA3911D" blockId="11.[151,1436,1785,1846]" pageId="11" pageNumber="362">
<emphasis id="8BC7A878F32F833336B34DA8FEBB913A" bold="true" box="[151,275,1785,1809]" pageId="11" pageNumber="362">FIGURE 8.</emphasis>
<taxonomicName id="7EB30FE9F32F8333373D4DA8FE4D913A" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[281,485,1785,1809]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F32F8333373D4DA8FE4D913A" box="[281,485,1785,1809]" italics="true" pageId="11" pageNumber="362">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F32F833337C84DA8FD94913A" bold="true" box="[492,572,1785,1809]" pageId="11" pageNumber="362">
<taxonomicNameLabel id="90F41503F32F833337C84DA8FD94913A" box="[492,572,1785,1809]" pageId="11" pageNumber="362" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymph, scanning electron micrographs, abdominal sterna:
<emphasis id="8BC7A878F32F833332BD4DA8FB15913A" bold="true" box="[1177,1213,1785,1809]" pageId="11" pageNumber="362">(A)</emphasis>
sterna 3–5;
<emphasis id="8BC7A878F32F8333331C4DA8FAF2913A" bold="true" box="[1336,1370,1785,1809]" pageId="11" pageNumber="362">(B)</emphasis>
sterna 5–7;
<emphasis id="8BC7A878F32F833336EF4C4CFF46911E" bold="true" box="[203,238,1821,1845]" pageId="11" pageNumber="362">(C)</emphasis>
sterna 7–10. Scale bars: A, B = 400 µm; C = 500 µm
</paragraph>
</caption>
<caption id="EDCC24E2F328833436B34C0BFAA891BC" ID-DOI="http://doi.org/10.5281/zenodo.6450857" ID-Zenodo-Dep="6450857" httpUri="https://zenodo.org/record/6450857/files/figure.png" pageId="12" pageNumber="363" startId="12.[151,250,1882,1906]" targetBox="[152,1436,183,1857]" targetPageId="12">
<paragraph id="B90C746AF328833436B34C0BFAA891BC" blockId="12.[151,1437,1882,1943]" pageId="12" pageNumber="363">
<emphasis id="8BC7A878F328833436B34C0BFEBB9159" bold="true" box="[151,275,1882,1906]" pageId="12" pageNumber="363">FIGURE 9.</emphasis>
<taxonomicName id="7EB30FE9F3288334373E4C0BFE4F9159" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[282,487,1882,1906]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F3288334373E4C0BFE4F9159" box="[282,487,1882,1906]" italics="true" pageId="12" pageNumber="363">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F328833437C94C0AFD969158" bold="true" box="[493,574,1883,1907]" pageId="12" pageNumber="363">
<taxonomicNameLabel id="90F41503F328833437C94C0AFD969158" box="[493,574,1883,1907]" pageId="12" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, nymph, scanning electron micrographs, abdominal sterna:
<emphasis id="8BC7A878F328833432BA4C0BFB489159" bold="true" box="[1182,1248,1882,1906]" pageId="12" pageNumber="363">(A–C)</emphasis>
details of sternum 8;
<emphasis id="8BC7A878F328833436954C2FFF5A91BD" bold="true" box="[177,242,1918,1942]" pageId="12" pageNumber="363">(D–E)</emphasis>
details of sternum 9;
<emphasis id="8BC7A878F328833437EA4C2FFE4791BD" bold="true" box="[462,495,1918,1942]" pageId="12" pageNumber="363">(F)</emphasis>
paraproct. Scale bars: A = 200 µm; B, D, F = 50 µm; C = 30 µm; E = 20 µm.
</paragraph>
</caption>
<caption id="EDCC24E2F329833536B34C6EFB48918B" ID-DOI="http://doi.org/10.5281/zenodo.6450859" ID-Zenodo-Dep="6450859" httpUri="https://zenodo.org/record/6450859/files/figure.png" pageId="13" pageNumber="364" startId="13.[151,250,1855,1879]" targetBox="[152,1435,182,1829]" targetPageId="13">
<paragraph id="B90C746AF329833536B34C6EFB48918B" blockId="13.[151,1436,1855,1952]" pageId="13" pageNumber="364">
<emphasis id="8BC7A878F329833536B34C6EFEB7917C" bold="true" box="[151,287,1855,1879]" pageId="13" pageNumber="364">FIGURE 10.</emphasis>
<taxonomicName id="7EB30FE9F329833537014C6EFE5A917C" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[293,498,1855,1879]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F329833537014C6EFE5A917C" box="[293,498,1855,1879]" italics="true" pageId="13" pageNumber="364">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F329833537DC4C6EFDE0917C" bold="true" box="[504,584,1855,1879]" pageId="13" pageNumber="364">
<taxonomicNameLabel id="90F41503F329833537DC4C6EFDE0917C" box="[504,584,1855,1879]" pageId="13" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, scanning electron micrographs, A–D nymph, E–F egg:
<emphasis id="8BC7A878F329833532A54C6EFB0D917C" bold="true" box="[1153,1189,1855,1879]" pageId="13" pageNumber="364">(A)</emphasis>
abdominal terga IV–VI;
<emphasis id="8BC7A878F329833536B34C32FF119150" bold="true" box="[151,185,1891,1915]" pageId="13" pageNumber="364">(B)</emphasis>
setae and scales on abdominal tergum V;
<emphasis id="8BC7A878F329833534454C32FD2C9150" bold="true" box="[609,644,1891,1915]" pageId="13" pageNumber="364">(C)</emphasis>
gill with long setae;
<emphasis id="8BC7A878F3298335357C4C32FCD49150" bold="true" box="[856,892,1891,1915]" pageId="13" pageNumber="364">(D)</emphasis>
median filament;
<emphasis id="8BC7A878F329833532104C32FBFE9150" bold="true" box="[1076,1110,1891,1915]" pageId="13" pageNumber="364">(E)</emphasis>
egg, general view;
<emphasis id="8BC7A878F329833533384C32FA949150" bold="true" box="[1308,1340,1891,1915]" pageId="13" pageNumber="364">(F)</emphasis>
egg, longitudinal tissue of inner layer. Scale bars: A = 400 µm; B, D = 100 µm; C = 0.5 mm; E = 50 µm; F = 10 µm.
</paragraph>
</caption>
<caption id="EDCC24E2F32A833636B34C18FB7191AE" ID-DOI="http://doi.org/10.5281/zenodo.6450861" ID-Zenodo-Dep="6450861" httpUri="https://zenodo.org/record/6450861/files/figure.png" pageId="14" pageNumber="365" startId="14.[151,250,1865,1889]" targetBox="[152,1436,182,1841]" targetPageId="14">
<paragraph id="B90C746AF32A833636B34C18FB7191AE" blockId="14.[151,1436,1865,1926]" pageId="14" pageNumber="365">
<emphasis id="8BC7A878F32A833636B34C18FEB7914A" bold="true" box="[151,287,1865,1889]" pageId="14" pageNumber="365">FIGURE 11.</emphasis>
<taxonomicName id="7EB30FE9F32A833637014C18FE5B914A" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[293,499,1865,1889]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="365" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F32A833637014C18FE5B914A" box="[293,499,1865,1889]" italics="true" pageId="14" pageNumber="365">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F32A833637DE4C1BFDE39149" bold="true" box="[506,587,1866,1890]" pageId="14" pageNumber="365">
<taxonomicNameLabel id="90F41503F32A833637DE4C1BFDE39149" box="[506,587,1866,1890]" pageId="14" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male subimago:
<emphasis id="8BC7A878F32A833635254C18FC8C914A" bold="true" box="[769,804,1865,1889]" pageId="14" pageNumber="365">(A)</emphasis>
dorsal habitus;
<emphasis id="8BC7A878F32A833635E24C18FC40914A" bold="true" box="[966,1000,1865,1889]" pageId="14" pageNumber="365">(B)</emphasis>
head and anterior thorax, lateral view;
<emphasis id="8BC7A878F32A8336335D4C18FA34914A" bold="true" box="[1401,1436,1865,1889]" pageId="14" pageNumber="365">(C)</emphasis>
abdomen, dorsal view;
<emphasis id="8BC7A878F32A833637A04C3CFE0F91AE" bold="true" box="[388,423,1901,1925]" pageId="14" pageNumber="365">(D)</emphasis>
forewing;
<emphasis id="8BC7A878F32A833634324C3CFD9091AE" bold="true" box="[534,568,1901,1925]" pageId="14" pageNumber="365">(E)</emphasis>
genitalia. Scale bars: A, D = 1 mm; B–C = 0.5 mm; E = 0.05 mm.
</paragraph>
</caption>
<caption id="EDCC24E2F32B833736B34EA5FBB19026" ID-DOI="http://doi.org/10.5281/zenodo.6450863" ID-Zenodo-Dep="6450863" box="[151,1049,1524,1549]" httpUri="https://zenodo.org/record/6450863/files/figure.png" pageId="15" pageNumber="366" startId="15.[151,250,1524,1548]" targetBox="[152,1436,181,1501]" targetPageId="15">
<paragraph id="B90C746AF32B833736B34EA5FBB19026" blockId="15.[151,1049,1524,1549]" box="[151,1049,1524,1549]" pageId="15" pageNumber="366">
<emphasis id="8BC7A878F32B833736B34EA5FE889026" bold="true" box="[151,288,1524,1549]" pageId="15" pageNumber="366">FIGURE 12.</emphasis>
Sampling site:
<emphasis id="8BC7A878F32B8337379A4EA5FE4A9027" bold="true" box="[446,482,1524,1548]" pageId="15" pageNumber="366">(A)</emphasis>
Khun Korn stream, general view;
<emphasis id="8BC7A878F32B833735644EA5FC299027" bold="true" box="[832,897,1524,1548]" pageId="15" pageNumber="366">(B–C)</emphasis>
microhabitats.
</paragraph>
</caption>
<subSubSection id="F1A927E1F32B833736E34D6AFD359122" pageId="15" pageNumber="366" type="discussion">
<paragraph id="B90C746AF32B833736E34D6AFD359122" blockId="15.[151,1437,1595,2018]" pageId="15" pageNumber="366">
<emphasis id="8BC7A878F32B833736E34D6AFE93907E" bold="true" box="[199,315,1595,1621]" pageId="15" pageNumber="366">Remarks.</emphasis>
<taxonomicName id="7EB30FE9F32B833737674D6AFD8E907E" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[323,550,1595,1621]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F32B833737674D6AFD8E907E" box="[323,550,1595,1621]" italics="true" pageId="15" pageNumber="366">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F32B8337340B4D6AFD21907E" bold="true" box="[559,649,1595,1621]" pageId="15" pageNumber="366">
<taxonomicNameLabel id="90F41503F32B8337340B4D6AFD21907E" box="[559,649,1595,1621]" pageId="15" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a curious maxillary palp morphology. In at least preserved specimens, the distinctive terminal segments of the maxillary palps are held against the body in a way that nearly mimics the placement and orientation of the procoxal osmobranchiae that are found on some other related baetids, such as
<taxonomicName id="7EB30FE9F32B8337369C4DF9FD3B90EA" authority="Muller-Liebenau, 1984" authorityName="Muller-Liebenau" authorityYear="1984" box="[184,659,1703,1729]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="8BC7A878F32B8337369C4DF9FE2A90EA" box="[184,386,1703,1729]" italics="true" pageId="15" pageNumber="366">Baetis sabahensis</emphasis>
<bibRefCitation id="DD22099BF32B833737AF4DF6FD3B90EA" author="Muller-Liebenau, I." box="[395,659,1703,1729]" pageId="15" pageNumber="366" pagination="85 - 99" refId="ref10073" refString="Muller-Liebenau, I. (1984) Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa, V., Soldan, T. &amp; Tonner, M. (Eds.), Proceedings of the Fourth International Conference on Ephemeroptera. Institute of Entomology, Czechoslovak Academy of Sciences, Ceske Budejovice, pp. 85 - 99." type="book chapter" year="1984">Müller-Liebenau, 1984</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EB30FE9F32B833734844DF6FB8590E9" authority="Kennedy, 1985" authorityName="Kennedy" authorityYear="1985" box="[672,1069,1703,1730]" class="Insecta" family="Baetidae" genus="Barbaetis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="benfieldi">
<emphasis id="8BC7A878F32B833734844DF6FCD090EA" box="[672,888,1703,1729]" italics="true" pageId="15" pageNumber="366">Barbaetis benfieldi</emphasis>
Kennedy, 1985
</taxonomicName>
(in
<bibRefCitation id="DD22099BF32B8337327A4DF6FA8E90E9" author="Waltz, R. D. &amp; McCafferty, W. P. &amp; Kennedy, J. H." box="[1118,1318,1703,1730]" pageId="15" pageNumber="366" pagination="161 - 165" refId="ref10827" refString="Waltz, R. D., McCafferty, W. P. &amp; Kennedy, J. H. (1985) Barbaetis: a new genus of eastern Nearctic mayflies (Ephemeroptera: Baetidae). Great Lakes Entomologist, 18 (4), 161 - 165." type="journal article" year="1985">
Waltz
<emphasis id="8BC7A878F32B833732834DF9FB4A90EA" box="[1191,1250,1703,1729]" italics="true" pageId="15" pageNumber="366">et al.</emphasis>
1985
</bibRefCitation>
), and four species of the North American genus
<taxonomicName id="7EB30FE9F32B833734644D9AFC6C90CD" authority="McDunnough, 1925" authorityName="McDunnough" authorityYear="1925" box="[576,964,1739,1766]" class="Insecta" family="Baetidae" genus="Heterocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="8BC7A878F32B833734644D9AFD7F90CE" box="[576,727,1739,1765]" italics="true" pageId="15" pageNumber="366">Heterocloeon</emphasis>
<bibRefCitation id="DD22099BF32B833734FB4D9AFC6C90CD" author="McDunnough, J." box="[735,964,1739,1766]" pageId="15" pageNumber="366" pagination="168 - 176" refId="ref9714" refString="McDunnough, J. (1925) New Canadian Ephemeridae with notes, III. Canadian Entomologist, 57, 168 - 176 + 185 - 192. https: // doi. org / 10.4039 / Ent 57185 - 8" type="journal article" year="1925">McDunnough, 1925</bibRefCitation>
</taxonomicName>
. This may suggest some analogous or additional functions of these different structures.
</paragraph>
</subSubSection>
<subSubSection id="F1A927E1F32B832836E34C42FED5976A" lastPageId="16" lastPageNumber="367" pageId="15" pageNumber="366" type="biology_ecology">
<paragraph id="B90C746AF32B833736E34C42FD2D91CA" blockId="15.[151,1437,1595,2018]" pageId="15" pageNumber="366">
<emphasis id="8BC7A878F32B833736E34C42FDA89106" bold="true" box="[199,512,1811,1837]" pageId="15" pageNumber="366">Ecology and Distribution.</emphasis>
The nymphs of
<taxonomicName id="7EB30FE9F32B833734E64C42FC009106" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[706,936,1811,1837]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F32B833734E64C42FC009106" box="[706,936,1811,1837]" italics="true" pageId="15" pageNumber="366">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F32B833735964C42FBA69106" bold="true" box="[946,1038,1811,1837]" pageId="15" pageNumber="366">
<taxonomicNameLabel id="90F41503F32B833735964C42FBA69106" box="[946,1038,1811,1837]" pageId="15" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
always were collected from slowflowing streams with a substrate mainly consisting of cobbles and pebbles covered by sediment. They were found under the cobble and covered by much sediment (
<figureCitation id="218868EFF32B833734E54C0AFCBC915E" box="[705,788,1883,1909]" captionStart="FIGURE 12" captionStartId="15.[151,250,1524,1548]" captionTargetBox="[152,1436,181,1501]" captionTargetId="figure-19@15.[151,1436,181,1501]" captionTargetPageId="15" captionText="FIGURE 12. Sampling site: (A) Khun Korn stream, general view; (B–C) microhabitats." figureDoi="http://doi.org/10.5281/zenodo.6450863" httpUri="https://zenodo.org/record/6450863/files/figure.png" pageId="15" pageNumber="366">Fig. 12</figureCitation>
). The microhabitats where we found them were usually in the upper stream, with relatively cool water temperature. Water temperature was 25ºC at time of sampling; dissolved oxygen was
<quantity id="7E4BD98FF32B833737074CF2FE179196" box="[291,447,1955,1982]" metricMagnitude="-6" metricUnit="kg" metricValue="7.609999999999999" metricValueMax="7.64" metricValueMin="7.58" pageId="15" pageNumber="366" unit="mg" value="7.609999999999999" valueMax="7.64" valueMin="7.58">7.58–7.64 mg</quantity>
/L; pH was 8.1; the electric conductivity of the water was 155–158 µS/cm, and total dissolved solid in the water was 205–209 ppm.
</paragraph>
<paragraph id="B90C746AF334832836E34BC6FC0E96D1" blockId="16.[151,1437,151,322]" pageId="16" pageNumber="367">
The microhabitat of our new species in slow flow areas laden with sediment is unusual among
<taxonomicName id="7EB30FE9F334832832D74BC6FF6296FE" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8BC7A878F334832832D74BC6FACF969A" box="[1267,1383,151,177]" italics="true" pageId="16" pageNumber="367">Acentrella</emphasis>
species.
</taxonomicName>
Most species are found on the tops and sides of objects in riffles and rapids, in areas of the stream that are relatively free of sediments (
<bibRefCitation id="DD22099BF334832837F34B8EFD7496D2" author="McCafferty, W. P. &amp; Wigle, M. J. &amp; Waltz, R. D." box="[471,732,223,249]" pageId="16" pageNumber="367" pagination="301 - 308" refId="ref9616" refString="McCafferty, W. P., Wigle, M. J. &amp; Waltz, R. D. (1994) Systematics and Biology of Acentrella turbida (McDunnough) (Ephemeroptera: Baetidae). Pan-Pacific Entomologist, 70 (4), 301 - 308." type="journal article" year="1994">
McCafferty
<emphasis id="8BC7A878F334832834464B8EFD3296D2" box="[610,666,223,249]" italics="true" pageId="16" pageNumber="367">et al.</emphasis>
1994
</bibRefCitation>
; LMJ, pers. obs.).
</paragraph>
<paragraph id="B90C746AF334832836E34A52FED5976A" blockId="16.[151,1437,151,322]" pageId="16" pageNumber="367">
<taxonomicName id="7EB30FE9F334832836E34A52FE059736" authority="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus, 2022" authorityName="Tungpairojwong &amp; Phlai-Ngam &amp; Jacobus" authorityYear="2022" box="[199,429,259,285]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="fimbriata" status="sp. nov.">
<emphasis id="8BC7A878F334832836E34A52FE059736" box="[199,429,259,285]" italics="true" pageId="16" pageNumber="367">Acentrella fimbriata</emphasis>
</taxonomicName>
<emphasis id="8BC7A878F334832837934A52FDBC9736" bold="true" box="[439,532,259,285]" pageId="16" pageNumber="367">
<taxonomicNameLabel id="90F41503F334832837934A52FDBC9736" box="[439,532,259,285]" pageId="16" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has been found at only three sites in northern
<collectingCountry id="C1A434FAF3348328321C4A52FB349736" box="[1080,1180,259,285]" name="Thailand" pageId="16" pageNumber="367">Thailand</collectingCountry>
(
<collectingRegion id="7B77BA88F334832832894A52FA999736" box="[1197,1329,259,285]" country="Thailand" name="Chiang Rai" pageId="16" pageNumber="367">Chiang Rai</collectingRegion>
and
<collectingRegion id="7B77BA88F3348328334A4A52FA359736" box="[1390,1437,259,285]" country="Thailand" name="Nan" pageId="16" pageNumber="367">Nan</collectingRegion>
provinces) (
<figureCitation id="218868EFF334832837394A76FED8976A" box="[285,368,295,321]" captionStart="FIGURE 18" captionStartId="22.[151,250,1528,1552]" captionTargetBox="[151,1436,181,1505]" captionTargetId="figure-19@22.[151,1436,181,1505]" captionTargetPageId="22" captionText="FIGURE 18. Distribution map of Acentrella fimbriata sp. nov. and Acentrella (Liebebiella) vera in Thailand. Red squares: Acentrella fimbriata sp. nov. Blue circles: Acentrella (Liebebiella) vera." figureDoi="http://doi.org/10.5281/zenodo.6450875" httpUri="https://zenodo.org/record/6450875/files/figure.png" pageId="16" pageNumber="367">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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