191 lines
26 KiB
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191 lines
26 KiB
XML
<document id="252ECEA4A199B9A72500B48C9D42003F" ID-DOI="10.5281/zenodo.200788" ID-GBIF-Dataset="fd436f04-a6b6-47eb-a746-d54e2192e0e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="200788" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460338949804" checkinUser="plazi" docAuthor="Miller, Kelly B. & Garcia, Mauricio" docDate="2011" docId="EE7587C2FFB6FFD5FF437BE2FD4339D0" docLanguage="en" docName="zt02996p056.pdf" docOrigin="Zootaxa 2996" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Spanglerodessus Miller and Garcia, new genus" docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="124CFFBAFFB7FFD4FFD4796CFFBB3E09" masterDocTitle="Spanglerodessus shorti and Incomptodessus camachoi, new genera and species of Bidessini from Guyana and Venezuela (Coleoptera: Dytiscidae: Hydroporinae)" masterLastPageNumber="56" masterPageNumber="49" pageNumber="50" updateTime="1698260536294" updateUser="plazi">
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<mods:titleInfo id="6D93C1212769A63F7B4F62B7E209FEE6">
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<mods:title id="79E98D167E92FF1805C74A3AA293DE10">Spanglerodessus shorti and Incomptodessus camachoi, new genera and species of Bidessini from Guyana and Venezuela (Coleoptera: Dytiscidae: Hydroporinae)</mods:title>
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<mods:name id="E1A74C7765DD97FEAE511CFBA0C33FD1" type="personal">
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<mods:namePart id="CED5D68D92EEBFC4EACA5E4767794256">Miller, Kelly B.</mods:namePart>
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<mods:name id="9C346650F3448E395CD06DB850D3C99D" type="personal">
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<mods:roleTerm id="E460E0E5AE5D99A76BCE19DF89D72DCF">Author</mods:roleTerm>
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<mods:namePart id="31FAE5B46127942DEBEBDA87535D30F1">Garcia, Mauricio</mods:namePart>
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<mods:typeOfResource id="531A574E11E3FCC613AECC84D8F13C13">text</mods:typeOfResource>
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<mods:titleInfo id="3341366B8F8DA9E70B8D9F31C5A07C71">
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<mods:title id="B99D20B91C435417E12472B08E20244F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C0D376A429E9106C0B5159F0FC61B7BB">
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<mods:date id="8691215F10F42962EA772AD1EDB48B14">2011</mods:date>
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<mods:detail id="9F0E22BA2FF1E13391058522EC427622" type="volume">
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<mods:number id="F183C4088A5E74074A291C44D2551DF2">2996</mods:number>
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<mods:classification id="E304B1565B51A7A17E257CC34490CCE5">journal article</mods:classification>
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<mods:identifier id="76B49075E20546CCBD68D2A99F5EE6A8" type="DOI">10.5281/zenodo.200788</mods:identifier>
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<mods:identifier id="1F0DAF3B1F29FD0F589E1F7C035BA260" type="GBIF-Dataset">fd436f04-a6b6-47eb-a746-d54e2192e0e3</mods:identifier>
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<mods:identifier id="81CC895FD3AA6112D6962A704982010A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="322B2FB51C605D0E406F58E2B7AC1D49" type="Zenodo-Dep">200788</mods:identifier>
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<treatment id="EE7587C2FFB6FFD5FF437BE2FD4339D0" ID-DOI="http://doi.org/10.5281/zenodo.6183243" ID-GBIF-Taxon="119405907" ID-Zenodo-Dep="6183243" LSID="urn:lsid:plazi:treatment:EE7587C2FFB6FFD5FF437BE2FD4339D0" httpUri="http://treatment.plazi.org/id/EE7587C2FFB6FFD5FF437BE2FD4339D0" lastPageNumber="50" pageId="1" pageNumber="50">
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<subSubSection id="2EC6655FFFB6FFD5FF437BE2FEEC3CC0" pageId="1" pageNumber="50" type="nomenclature">
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<paragraph id="666336D4FFB6FFD5FF437BE2FD613CA1" blockId="1.[151,730,654,713]" box="[151,730,654,680]" pageId="1" pageNumber="50">
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<heading id="3D2B81B8FFB6FFD5FF437BE2FD613CA1" bold="true" box="[151,730,654,680]" fontSize="11" level="1" pageId="1" pageNumber="50" reason="1">
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<emphasis id="54A8EAC6FFB6FFD5FF437BE2FD613CA1" bold="true" box="[151,730,654,680]" pageId="1" pageNumber="50">
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<taxonomicName id="A1DC4D57FFB6FFD5FF437BE2FDF73CA1" ID-CoL="8LSYV" authority="Miller and Garcia" authorityName="Miller and Garcia" box="[151,588,654,680]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="54A8EAC6FFB6FFD5FF437BE2FEDA3CA1" bold="true" box="[151,353,654,680]" italics="true" pageId="1" pageNumber="50">Spanglerodessus</emphasis>
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Miller and Garcia
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</taxonomicName>
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,
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<taxonomicNameLabel id="4F9B57BDFFB6FFD5FD8C7BE2FD613CA1" box="[600,730,654,680]" pageId="1" pageNumber="50" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="666336D4FFB6FFD5FF437BDDFEEC3CC0" blockId="1.[151,730,654,713]" box="[151,343,689,713]" pageId="1" pageNumber="50">
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(
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<figureCitation id="FEE72A51FFB6FFD5FF4B7BDDFF533CC0" box="[159,232,689,713]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Figs 1 – 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 – 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 – 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="1" pageNumber="50">Figs. 1</figureCitation>
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–6,10,11)
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</paragraph>
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</subSubSection>
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<subSubSection id="2EC6655FFFB6FFD5FF437B96FB0A3D18" box="[151,1201,761,786]" pageId="1" pageNumber="50" type="materials_examined">
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<paragraph id="666336D4FFB6FFD5FF437B96FB0A3D18" blockId="1.[151,1437,761,2009]" box="[151,1201,761,786]" pageId="1" pageNumber="50">
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<emphasis id="54A8EAC6FFB6FFD5FF437B96FE893D1B" bold="true" box="[151,306,761,786]" pageId="1" pageNumber="50">
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<typeStatus id="B9678876FFB6FFD5FF437B96FF693D1B" box="[151,210,762,786]" pageId="1" pageNumber="50">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="A1DC4D57FFB6FFD5FEED7B95FCB53D18" authority="Miller and Garcia" authorityName="Miller and Garcia" box="[313,782,761,785]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="shorti" status="sp. nov.">
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<emphasis id="54A8EAC6FFB6FFD5FEED7B95FD863D18" box="[313,573,761,785]" italics="true" pageId="1" pageNumber="50">Spanglerodessus shorti</emphasis>
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Miller and Garcia
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</taxonomicName>
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,
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<taxonomicNameLabel id="4F9B57BDFFB6FFD5FCCC7B96FC193D18" box="[792,930,761,785]" pageId="1" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="2EC6655FFFB6FFD5FF117A71FE413A04" pageId="1" pageNumber="50" type="diagnosis">
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<paragraph id="666336D4FFB6FFD5FF117A71FE413A04" blockId="1.[151,1437,761,2009]" pageId="1" pageNumber="50">
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<emphasis id="54A8EAC6FFB6FFD5FF117A71FDBE3D3F" bold="true" box="[197,517,797,822]" pageId="1" pageNumber="50">Diagnosis and description.</emphasis>
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This genus differs from others in the tribe by the combination of: 1) transverse occipital line absent (
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<figureCitation id="FEE72A51FFB6FFD5FE527A2DFE7C3D50" box="[390,455,833,857]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Figs 1 – 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 – 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 – 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="1" pageNumber="50">Fig. 1</figureCitation>
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), 2) basal pronotal striae present (
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<figureCitation id="FEE72A51FFB6FFD5FC937A2DFC333D50" box="[839,904,833,857]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Figs 1 – 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 – 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 – 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="1" pageNumber="50">Fig. 1</figureCitation>
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), 3) basal elytral stria absent (
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<figureCitation id="FEE72A51FFB6FFD5FB087A2DFAA53D50" box="[1244,1310,833,857]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Figs 1 – 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 – 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 – 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="1" pageNumber="50">Fig. 1</figureCitation>
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), 4) elytral sutural stria faintly present in some specimens, 5) anterior clypeal margin unmodified, 6) elytron without longitudinal carinae (
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<figureCitation id="FEE72A51FFB6FFD5FEF47AE5FEDA3DA8" box="[288,353,905,929]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Figs 1 – 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 – 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 – 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="1" pageNumber="50">Fig. 1</figureCitation>
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), 7) epipleuron without transverse carina at humeral angle, 8) lateral lobes of aedeagus two-segmented (
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<figureCitation id="FEE72A51FFB6FFD5FF2D7AC1FEDA3DCC" box="[249,353,941,965]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Figs 1 – 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 – 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 – 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="1" pageNumber="50">Figs 4, 6</figureCitation>
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), 9) habitus robust, short, lateral margins of pronotum and elytron conspicuously rounded (
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<figureCitation id="FEE72A51FFB6FFD5FABA7AC1FF1F3DE0" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Figs 1 – 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 – 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 – 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="1" pageNumber="50">Fig. 1</figureCitation>
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), and 10) lateral bead on pronotum broad (
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<figureCitation id="FEE72A51FFB6FFD5FD477ABDFD6D3DE0" box="[659,726,977,1001]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Figs 1 – 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 – 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 – 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="1" pageNumber="50">Fig. 1</figureCitation>
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). Individuals lack natatory setae on the legs, which are robust and spinous, in general (
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<figureCitation id="FEE72A51FFB6FFD5FE7C7A99FE563A04" box="[424,493,1013,1037]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Figs 1 – 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 – 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 – 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="1" pageNumber="50">Fig. 3</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="2EC6655FFFB6FFD5FF117D76FB413A5C" pageId="1" pageNumber="50" type="etymology">
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<paragraph id="666336D4FFB6FFD5FF117D76FB413A5C" blockId="1.[151,1437,761,2009]" pageId="1" pageNumber="50">
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<emphasis id="54A8EAC6FFB6FFD5FF117D76FEF73A3B" bold="true" box="[197,332,1050,1074]" pageId="1" pageNumber="50">Etymology.</emphasis>
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The genus name is formed from a combination of “
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<emphasis id="54A8EAC6FFB6FFD5FC717D75FBBC3A38" box="[933,1031,1049,1073]" italics="true" pageId="1" pageNumber="50">spangler</emphasis>
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” in honor of the great water beetle worker and friend, Paul J. Spangler, and “
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<emphasis id="54A8EAC6FFB6FFD5FDBF7D51FD0F3A5C" box="[619,692,1085,1109]" italics="true" pageId="1" pageNumber="50">dessus</emphasis>
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,” a common root in other genus names in this tribe.
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</paragraph>
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</subSubSection>
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<subSubSection id="2EC6655FFFB6FFD5FF117D0DFDDC3AC8" pageId="1" pageNumber="50" type="distribution">
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<paragraph id="666336D4FFB6FFD5FF117D0DFDDC3AC8" blockId="1.[151,1437,761,2009]" pageId="1" pageNumber="50">
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<emphasis id="54A8EAC6FFB6FFD5FF117D0DFE4E3A73" bold="true" box="[197,501,1121,1146]" pageId="1" pageNumber="50">Distribution and habitat.</emphasis>
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<taxonomicName id="A1DC4D57FFB6FFD5FE2B7D0DFD013A70" box="[511,698,1121,1145]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="54A8EAC6FFB6FFD5FE2B7D0DFD013A70" box="[511,698,1121,1145]" italics="true" pageId="1" pageNumber="50">Spanglerodessus</emphasis>
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</taxonomicName>
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is known from the following newly described species from the Kumu Falls in
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<collectingCountry id="1ECB7644FFB6FFD5FE947DE9FE2C3A94" box="[320,407,1157,1181]" name="Guyana" pageId="1" pageNumber="50">Guyana</collectingCountry>
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and nearby localities in
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<collectingCountry id="1ECB7644FFB6FFD5FD7A7DE9FC9A3A94" box="[686,801,1157,1181]" name="Venezuela" pageId="1" pageNumber="50">Venezuela</collectingCountry>
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(
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<figureCitation id="FEE72A51FFB6FFD5FCE47DE9FC393A94" box="[816,898,1157,1181]" captionStart="FIGURES 11 – 14" captionStartId="5.[151,264,1741,1764]" captionTargetBox="[217,1369,409,1719]" captionTargetId="figure@5.[217,1369,409,1720]" captionTargetPageId="5" captionText="FIGURES 11 – 14. Habitats. 11, Spanglerodessus shorti, AS- 08 - 058. 12 – 14, Incomptodessus comachoi; 12, VZ 09 - 0114 - 01 C; 13, VZ 09 - 0112 - 01 B; 14, VZ 09 - 0115 - 01 D. Codes correspond to collection locality codes listed in “ Material examined ” section for each species." httpUri="https://zenodo.org/record/200791/files/figure.png" pageId="1" pageNumber="50">Fig. 11</figureCitation>
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). Label data indicated that specimens were collected in pools at the base of Kumu Falls.
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</paragraph>
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</subSubSection>
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<subSubSection id="2EC6655FFFB6FFD5FF117DA1FD4339D0" pageId="1" pageNumber="50" type="discussion">
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<paragraph id="666336D4FFB6FFD5FF117DA1FD783B7C" blockId="1.[151,1437,761,2009]" pageId="1" pageNumber="50">
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<emphasis id="54A8EAC6FFB6FFD5FF117DA1FEF13AEF" bold="true" box="[197,330,1229,1254]" pageId="1" pageNumber="50">Discussion.</emphasis>
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In addition to the species described here, four specimens of what appears to be another new species in this genus are in the USNM. These specimens are represented by only females and, so, are not described here. The specimens were found at the same site as
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<taxonomicName id="A1DC4D57FFB6FFD5FD217C79FC403B24" box="[757,1019,1301,1325]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="shorti">
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<emphasis id="54A8EAC6FFB6FFD5FD217C79FC403B24" box="[757,1019,1301,1325]" italics="true" pageId="1" pageNumber="50">Spanglerodessus shorti</emphasis>
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</taxonomicName>
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||
. Whereas that species was found in “side pools at base of falls,” label data indicate the specimens of the other species were collected from “wet boulders,” possibly a distinctly different microhabitat.
|
||
</paragraph>
|
||
<paragraph id="666336D4FFB6FFD5FF117CEDFD4339D0" blockId="1.[151,1437,761,2009]" pageId="1" pageNumber="50">
|
||
This genus keys to
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FE747CEDFD213B90" authority="Regimbart" authorityName="Regimbart" box="[416,666,1409,1433]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="54A8EAC6FFB6FFD5FE747CEDFDA73B90" box="[416,540,1409,1433]" italics="true" pageId="1" pageNumber="50">Bidessodes</emphasis>
|
||
Régimbart
|
||
</taxonomicName>
|
||
in Biström’s (1988) key to the Bidessini genera of the world.
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FA8F7CEEFEA13BB4" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="54A8EAC6FFB6FFD5FA8F7CEEFEA13BB4" italics="true" pageId="1" pageNumber="50">Spanglerodessus</emphasis>
|
||
</taxonomicName>
|
||
specimens are considerably more robust than members of
|
||
<emphasis id="54A8EAC6FFB6FFD5FC687CC9FB843BB4" box="[956,1087,1445,1469]" italics="true" pageId="1" pageNumber="50">
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FC687CC9FB803BB4" box="[956,1083,1445,1469]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">Bidessodes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
which are elongate. Additionally,
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FF017CA5FEE93BE8" box="[213,338,1481,1505]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="54A8EAC6FFB6FFD5FF017CA5FEE93BE8" box="[213,338,1481,1505]" italics="true" pageId="1" pageNumber="50">Bidessodes</emphasis>
|
||
</taxonomicName>
|
||
species are pale in coloration, typically with longitudinal yellow fasciae and conspicuous iridescence on the elytra, and have the lateral bead on the pronotum very narrow. Despite these numerous differences, it seems likely that
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FE1F7F7DFDF33820" box="[459,584,1553,1577]" class="Insecta" family="Dytiscidae" genus="Bidessodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="54A8EAC6FFB6FFD5FE1F7F7DFDF33820" box="[459,584,1553,1577]" italics="true" pageId="1" pageNumber="50">Bidessodes</emphasis>
|
||
</taxonomicName>
|
||
may be closely related to
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FCAB7F7DFB813820" box="[895,1082,1553,1577]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="54A8EAC6FFB6FFD5FCAB7F7DFB813820" box="[895,1082,1553,1577]" italics="true" pageId="1" pageNumber="50">Spanglerodessus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FB9F7F7DFABD3820" box="[1099,1286,1553,1577]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="54A8EAC6FFB6FFD5FB9F7F7DFABD3820" box="[1099,1286,1553,1577]" italics="true" pageId="1" pageNumber="50">Spanglerodessus</emphasis>
|
||
</taxonomicName>
|
||
may also be closely related to the recently described
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FDBE7F59FC5B3844" authority="Miller and Spangler" authorityName="Miller and Spangler" box="[618,992,1589,1613]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="54A8EAC6FFB6FFD5FDBE7F59FD4B3844" box="[618,752,1589,1613]" italics="true" pageId="1" pageNumber="50">Fontidessus</emphasis>
|
||
Miller and Spangler
|
||
</taxonomicName>
|
||
, but
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FBC87F59FB6C3844" box="[1052,1239,1589,1613]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="54A8EAC6FFB6FFD5FBC87F59FB6C3844" box="[1052,1239,1589,1613]" italics="true" pageId="1" pageNumber="50">Spanglerodessus</emphasis>
|
||
</taxonomicName>
|
||
are considerably more robust and lack natatory setae on the legs. Nevertheless, given the habitat and morphology of
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FB2F7F35FF00389C" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="54A8EAC6FFB6FFD5FB2F7F35FF00389C" italics="true" pageId="1" pageNumber="50">Spanglerodessus</emphasis>
|
||
</taxonomicName>
|
||
, which are quite similar to
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FE237F11FDC6389C" box="[503,637,1661,1685]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="54A8EAC6FFB6FFD5FE237F11FDC6389C" box="[503,637,1661,1685]" italics="true" pageId="1" pageNumber="50">Fontidessus</emphasis>
|
||
</taxonomicName>
|
||
, it appears
|
||
<taxonomicName id="A1DC4D57FFB6FFD5FCD67F11FC06389C" box="[770,957,1661,1685]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="49" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="54A8EAC6FFB6FFD5FCD67F11FC06389C" box="[770,957,1661,1685]" italics="true" pageId="1" pageNumber="50">Spanglerodessus</emphasis>
|
||
</taxonomicName>
|
||
may be closely related to that genus. It is always a difficult challenge to decide whether to create a new genus or expand the definition of an existing genus in such circumstances. This problem is particularly acute in Bidessini where the many genera are defined by a few characters which occur in combinations clearly implying considerable homoplasy.
|
||
<bibRefCitation id="024D4B25FFB6FFD5FBED7F85FAD73908" author="Miller" box="[1081,1388,1769,1793]" pageId="1" pageNumber="50" refString="Miller, K. B. and Spangler, P. J. (2008) Fontidessus Miller and Spangler, a new genus of Bidessini from Venezuela (Coleoptera: Dytiscidae: Hydroporinae) with three new species. Zootaxa 1827, 45 - 52." type="journal article" year="2008">Miller and Spangler (2008)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="024D4B25FFB6FFD5FF437E61FE17392C" author="Balke" box="[151,428,1805,1829]" pageId="1" pageNumber="50" refString="Balke, M. and Ribera, I. (2004) Jumping across Wallace's line: Allodessus Guignot and Limbodessus Guignot revisited (Coleoptera: Dytiscidae, Bidessini) based on molecular-phylogenetic and morphological data. Australian Journal of Entomology 43, 114 - 128." type="journal article" year="2004">Balke and Ribera (2004)</bibRefCitation>
|
||
(among others) pointed out some of the serious problems with the distribution of the few characters (particularly the transverse occipital line) and their degree of homoplasy in the tribe. Nothing short of a comprehensive phylogenetic analysis of Bidessini with broad taxon sampling and extensive molecular data will likely help clarify how general these features are distributed and in what combination. In this case, we believe that, for now, a new genus is warranted in the absence of a comprehensive cladistic analysis of the tribe to help determine character distributions in a phylogenetic context.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |