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<document id="6899B77A1219E5F286CF9DE5207DA90A" ID-DOI="10.5281/zenodo.200788" ID-GBIF-Dataset="fd436f04-a6b6-47eb-a746-d54e2192e0e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="200788" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460338949804" checkinUser="plazi" docAuthor="Miller, Kelly B. &amp; Garcia, Mauricio" docDate="2011" docId="EE7587C2FFB5FFD0FF437E66FE533F8E" docLanguage="en" docName="zt02996p056.pdf" docOrigin="Zootaxa 2996" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Spanglerodessus shorti Miller and Garcia, new species" docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="124CFFBAFFB7FFD4FFD4796CFFBB3E09" masterDocTitle="Spanglerodessus shorti and Incomptodessus camachoi, new genera and species of Bidessini from Guyana and Venezuela (Coleoptera: Dytiscidae: Hydroporinae)" masterLastPageNumber="56" masterPageNumber="49" pageNumber="51" updateTime="1698260536294" updateUser="plazi">
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<mods:title id="E3EB3173A529EBF8CA1ABDA0DBACF9EC">Spanglerodessus shorti and Incomptodessus camachoi, new genera and species of Bidessini from Guyana and Venezuela (Coleoptera: Dytiscidae: Hydroporinae)</mods:title>
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<mods:name id="F0314F6064C0D3CE4BA82BB688E71E6D" type="personal">
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<mods:namePart id="09DBBE2C22D192094F639A4E3B8FFBDB">Miller, Kelly B.</mods:namePart>
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<mods:namePart id="FC15C44937DF29233C4D73156F888333">Garcia, Mauricio</mods:namePart>
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<mods:date id="CB1A9DDF86CA5C8397610BF0A8F9D7D0">2011</mods:date>
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<treatment id="EE7587C2FFB5FFD0FF437E66FE533F8E" ID-DOI="http://doi.org/10.5281/zenodo.6183245" ID-GBIF-Taxon="119405909" ID-Zenodo-Dep="6183245" LSID="urn:lsid:plazi:treatment:EE7587C2FFB5FFD0FF437E66FE533F8E" httpUri="http://treatment.plazi.org/id/EE7587C2FFB5FFD0FF437E66FE533F8E" lastPageId="4" lastPageNumber="52" pageId="2" pageNumber="51">
<subSubSection id="2EC6655FFFB5FFD6FF437E66FEEB394C" pageId="2" pageNumber="51" type="nomenclature">
<paragraph id="666336D4FFB5FFD6FF437E66FC8E392D" blockId="2.[151,821,1802,1861]" box="[151,821,1802,1828]" pageId="2" pageNumber="51">
<heading id="3D2B81B8FFB5FFD6FF437E66FC8E392D" bold="true" box="[151,821,1802,1828]" fontSize="11" level="1" pageId="2" pageNumber="51" reason="1">
<emphasis id="54A8EAC6FFB5FFD6FF437E66FC8E392D" bold="true" box="[151,821,1802,1828]" pageId="2" pageNumber="51">
<taxonomicName id="A1DC4D57FFB5FFD6FF437E66FD22392D" ID-CoL="8LXXS" authority="Miller and Garcia" authorityName="Miller and Garcia" box="[151,665,1802,1828]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="shorti" status="sp. nov.">
<emphasis id="54A8EAC6FFB5FFD6FF437E66FE15392D" bold="true" box="[151,430,1802,1828]" italics="true" pageId="2" pageNumber="51">Spanglerodessus shorti</emphasis>
Miller and Garcia
</taxonomicName>
,
<taxonomicNameLabel id="4F9B57BDFFB5FFD6FD727E66FC8E392D" box="[678,821,1802,1828]" pageId="2" pageNumber="51" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="666336D4FFB5FFD6FF437E41FEEB394C" blockId="2.[151,821,1802,1861]" box="[151,336,1837,1861]" pageId="2" pageNumber="51">
(
<figureCitation id="FEE72A51FFB5FFD6FF4B7E41FF5B394C" box="[159,224,1837,1861]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="2" pageNumber="51">Figs 1</figureCitation>
6,10,11)
</paragraph>
</subSubSection>
<subSubSection id="2EC6655FFFB5FFD6FF437E19FAFF3984" box="[151,1348,1908,1934]" pageId="2" pageNumber="51" type="materials_examined">
<paragraph id="666336D4FFB5FFD6FF437E19FAFF3984" blockId="2.[151,1436,1908,2005]" box="[151,1348,1908,1934]" pageId="2" pageNumber="51">
<emphasis id="54A8EAC6FFB5FFD6FF437E19FE8E3987" bold="true" box="[151,309,1909,1934]" pageId="2" pageNumber="51">
<typeStatus id="B9678876FFB5FFD6FF437E19FF693984" box="[151,210,1909,1933]" pageId="2" pageNumber="51">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="1ECB7644FFB5FFD6FEE87E19FE093984" box="[316,434,1909,1933]" name="Venezuela" pageId="2" pageNumber="51">Venezuela</collectingCountry>
, Bolivar State, rocky stream along La Escalera, 6°1ʹ38.0&quot;N 61°23ʹ41.1&quot;W,
<specimenCount id="70DAFD5DFFB5FFD6FB297E19FAFF3984" box="[1277,1348,1909,1933]" pageId="2" pageNumber="51" type="generic">
<quantity id="A1249B31FFB5FFD6FB297E19FAFB3984" box="[1277,1344,1909,1933]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="2" pageNumber="51" unit="m" value="690.0">690m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="2EC6655FFFB5FFD7FF117EF4FCF93EDD" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51" type="diagnosis">
<paragraph id="666336D4FFB5FFD7FF117EF4FCF93EDD" blockId="2.[151,1436,1908,2005]" lastBlockId="3.[151,1437,151,1039]" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">
<emphasis id="54A8EAC6FFB5FFD6FF117EF4FE8439B8" bold="true" box="[197,319,1944,1969]" pageId="2" pageNumber="51">Diagnosis.</emphasis>
<taxonomicName id="A1DC4D57FFB5FFD6FE927EF5FDF139B8" box="[326,586,1945,1969]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="shorti">
<emphasis id="54A8EAC6FFB5FFD6FE927EF5FDF139B8" box="[326,586,1945,1969]" italics="true" pageId="2" pageNumber="51">Spanglerodessus shorti</emphasis>
</taxonomicName>
is the only member of this genus and is characterized by the diagnostic features of
<taxonomicName id="A1DC4D57FFB5FFD6FF277ED1FE1539DC" box="[243,430,1981,2005]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="54A8EAC6FFB5FFD6FF277ED1FE1539DC" box="[243,430,1981,2005]" italics="true" pageId="2" pageNumber="51">Spanglerodessus</emphasis>
</taxonomicName>
. Likely species level diagnostic features include the overall dark brown dorsal coloration with transverse medial yellow-brown region on pronotum (
<figureCitation id="FEE72A51FFB4FFD7FCE279FBFCC33EA6" box="[822,888,151,175]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 1</figureCitation>
) and the shape of the male genitalia (
<figureCitation id="FEE72A51FFB4FFD7FAFC79FBFA343EA6" box="[1320,1423,151,175]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Figs 46</figureCitation>
). The median lobe is very broad and apically trilobate (
<figureCitation id="FEE72A51FFB4FFD7FD3B79D0FC8F3EDD" box="[751,820,188,212]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2EC6655FFFB4FFD7FF1179B3FBE23D55" pageId="3" pageNumber="52" type="description">
<paragraph id="666336D4FFB4FFD7FF1179B3FF503F6D" blockId="3.[151,1437,151,1039]" pageId="3" pageNumber="52">
<emphasis id="54A8EAC6FFB4FFD7FF1179B3FEED3EF1" bold="true" box="[197,342,223,248]" pageId="3" pageNumber="52">Description.</emphasis>
<emphasis id="54A8EAC6FFB4FFD7FE8B798DFDBA3EF1" box="[351,513,225,248]" italics="true" pageId="3" pageNumber="52">Measurements</emphasis>
. TL =
<quantity id="A1249B31FFB4FFD7FD8179B3FD5B3EFE" box="[597,736,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="3" pageNumber="52" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.51.7 mm</quantity>
, GW = 0.91.0 mm, PW =
<quantity id="A1249B31FFB4FFD7FBF779B3FB143EFE" box="[1059,1199,223,247]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="3" pageNumber="52" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9 mm</quantity>
, HW =
<quantity id="A1249B31FFB4FFD7FAD879B3FA233EFE" box="[1292,1432,223,247]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="52" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
, EW =
<quantity id="A1249B31FFB4FFD7FF347868FED13F15" box="[224,362,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="52" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.30.4 mm</quantity>
, TL/GW = 1.51.6. Body broad, robust, globular (
<figureCitation id="FEE72A51FFB4FFD7FC747868FC5A3F15" box="[928,993,260,284]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 1</figureCitation>
); lateral outline discontinuous between pronotum and elytron; lateral margins of pronotum broadly and evenly curved; lateral margins of elytron broadly curved.
</paragraph>
<paragraph id="666336D4FFB4FFD7FF13781CFAE33FA5" blockId="3.[151,1437,151,1039]" pageId="3" pageNumber="52">
<emphasis id="54A8EAC6FFB4FFD7FF13781CFEF93F81" box="[199,322,368,392]" italics="true" pageId="3" pageNumber="52">Coloration</emphasis>
. Head dark brown, pronotum dark brown along posterior half and anterior margin, medial and lateral region yellow-brown; elytron dark brown. Ventral surfaces and appendages red-brown to yellow-brown.
</paragraph>
<paragraph id="666336D4FFB4FFD7FF1378D4FC8F3CE6" blockId="3.[151,1437,151,1039]" pageId="3" pageNumber="52">
<emphasis id="54A8EAC6FFB4FFD7FF1378D4FE6B3FD9" box="[199,464,440,464]" italics="true" pageId="3" pageNumber="52">Sculpture and structure</emphasis>
. Head with very fine, inconspicuous, irregular punctation, surface between punctures with microsculpture in the form of small, isodiametric cells; eyes medium in size (
<figureCitation id="FEE72A51FFB4FFD7FB9278B0FB323FFD" box="[1094,1161,476,500]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 1</figureCitation>
, HW/EW = 1.541.61). Pronotal surface similar to that of head; with posterior angles obtuse; lateral bead broad, of even width throughout (
<figureCitation id="FEE72A51FFB4FFD7FF4B7B48FF593C35" box="[159,226,548,572]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 1</figureCitation>
); pronotal striae prominent, extending about 1/3 distance across pronotum (
<figureCitation id="FEE72A51FFB4FFD7FBEB7B48FB393C35" box="[1087,1154,548,572]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 1</figureCitation>
). Elytron with anterolateral angle obtuse, not strongly extended anteriorly; surface similar to pronotum. Prosternal process broad, lateral margins subparallel, apex of process broadly rounded, extending to metasternal fork (
<figureCitation id="FEE72A51FFB4FFD7FBB97B00FB083C8D" box="[1133,1203,620,644]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 2</figureCitation>
); metacoxal process with lateral lobe minute but distinct; metaventrite finely, sparsely punctate, metacoxa impunctate, both metaventrite and metacoxae with distinct microsculpture. Pro- and mesotarsi moderately broad in male, slightly narrower in female. Metatrochanter large relative to metafemur (
<figureCitation id="FEE72A51FFB4FFD7FD367BBBFC9D3CE6" box="[738,806,727,751]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="666336D4FFB4FFD7FF137B90FBE23D55" blockId="3.[151,1437,151,1039]" pageId="3" pageNumber="52">
<emphasis id="54A8EAC6FFB4FFD7FF137B90FED63D1D" box="[199,365,764,788]" italics="true" pageId="3" pageNumber="52">Male genitalia</emphasis>
. Median lobe in ventral aspect broad, expanded apically with apical margin weakly trilobate (
<figureCitation id="FEE72A51FFB4FFD7FF4B7A73FF533D3E" box="[159,232,799,823]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 4</figureCitation>
); in lateral aspect slender, straight to ventrally-curved apex (
<figureCitation id="FEE72A51FFB4FFD7FC667A73FC413D3E" box="[946,1018,799,823]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 5</figureCitation>
). Lateral lobe in lateral and ventral aspects narrow, broadly curved throughout length, apically narrowly rounded (
<figureCitation id="FEE72A51FFB4FFD7FBD27A28FBF73D55" box="[1030,1100,836,860]" captionStart="FIGURES 1 9" captionStartId="2.[151,264,1607,1630]" captionTargetBox="[334,1257,211,1561]" captionTargetId="figure@2.[313,1273,193,1586]" captionTargetPageId="2" captionText="FIGURES 1 9. Figs 1 6. Spanglerodessus shorti. 1, dorsal habitus, bar = 1.0 mm. 2, prosternal process. 3, right metathoracic leg, ventral aspect. 4 6, male genitalia, 4, median lobe and right lateral lobe, dorsal aspect, 5, median lobe, left lateral aspect, 6, left lateral lobe, left lateral aspect. Figs 7 9. Incomptodessus camachoi. 7, dorsal habitus, bar = 1.0 mm. 8, male median lobe and right lateral lobe, right lateral aspect. 9, male median lobe, ventral aspect." httpUri="https://zenodo.org/record/200789/files/figure.png" pageId="3" pageNumber="52">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2EC6655FFFB4FFD7FF117A04FF723DAD" pageId="3" pageNumber="52" type="etymology">
<paragraph id="666336D4FFB4FFD7FF117A04FF723DAD" blockId="3.[151,1437,151,1039]" pageId="3" pageNumber="52">
<emphasis id="54A8EAC6FFB4FFD7FF117A04FEF73D89" bold="true" box="[197,332,872,896]" pageId="3" pageNumber="52">Etymology.</emphasis>
This species is named in honor of our good friend and colleague A.E.Z. Short (University of Kansas).
</paragraph>
</subSubSection>
<subSubSection id="2EC6655FFFB4FFD7FF117AC3FE953A06" pageId="3" pageNumber="52" type="distribution">
<paragraph id="666336D4FFB4FFD7FF117AC3FE953A06" blockId="3.[151,1437,151,1039]" pageId="3" pageNumber="52">
<emphasis id="54A8EAC6FFB4FFD7FF117AC3FE553DC1" bold="true" box="[197,494,943,968]" pageId="3" pageNumber="52">Distribution and habitat.</emphasis>
Label data indicate the
<collectingCountry id="1ECB7644FFB4FFD7FD2D7AC3FCEB3DCE" box="[761,848,943,967]" name="Guyana" pageId="3" pageNumber="52">Guyana</collectingCountry>
specimens were collected in pools in the stream bed at the base of a large waterfall. Other specimens were collected along a rocky stream, and the species is hygropetric (
<figureCitation id="FEE72A51FFB4FFD7FF4B7A9BFE993A06" box="[159,290,1015,1039]" captionStart="FIGURES 11 14" captionStartId="5.[151,264,1741,1764]" captionTargetBox="[217,1369,409,1719]" captionTargetId="figure@5.[217,1369,409,1720]" captionTargetPageId="5" captionText="FIGURES 11 14. Habitats. 11, Spanglerodessus shorti, AS- 08 - 058. 12 14, Incomptodessus comachoi; 12, VZ 09 - 0114 - 01 C; 13, VZ 09 - 0112 - 01 B; 14, VZ 09 - 0115 - 01 D. Codes correspond to collection locality codes listed in “ Material examined ” section for each species." httpUri="https://zenodo.org/record/200791/files/figure.png" pageId="3" pageNumber="52">Figs 1214</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="32A3665CFFB4FFD7FF437E36FAB93978" box="[151,1282,1882,1905]" httpUri="https://zenodo.org/record/200790/files/figure.png" pageId="3" pageNumber="52" targetBox="[159,1425,1102,1851]" targetPageId="3">
<paragraph id="666336D4FFB4FFD7FF437E36FAB93978" blockId="3.[151,1282,1882,1905]" box="[151,1282,1882,1905]" pageId="3" pageNumber="52">
<emphasis id="54A8EAC6FFB4FFD7FF437E36FEA43979" bold="true" box="[151,287,1882,1905]" pageId="3" pageNumber="52">FIGURE 10.</emphasis>
Known distribution of
<taxonomicName id="A1DC4D57FFB4FFD7FDD97E37FD433978" box="[525,760,1883,1905]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="shorti">
<emphasis id="54A8EAC6FFB4FFD7FDD97E37FD433978" box="[525,760,1883,1905]" italics="true" pageId="3" pageNumber="52">Spanglerodessus shorti</emphasis>
</taxonomicName>
(triangles) and
<taxonomicName id="A1DC4D57FFB4FFD7FC437E37FB1A3978" box="[919,1185,1883,1905]" class="Insecta" family="Dytiscidae" genus="Incomptodessus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="camachoi">
<emphasis id="54A8EAC6FFB4FFD7FC437E37FB1A3978" box="[919,1185,1883,1905]" italics="true" pageId="3" pageNumber="52">Incomptodessus camachoi</emphasis>
</taxonomicName>
(circles).
</paragraph>
</caption>
<subSubSection id="2EC6655FFFB4FFD0FF117EF1FE533F8E" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52" type="materials_examined">
<paragraph id="666336D4FFB4FFD0FF117EF1FE533F8E" blockId="3.[151,1436,1948,2010]" lastBlockId="4.[151,1437,151,391]" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52">
<emphasis id="54A8EAC6FFB4FFD7FF117EF1FE0739BF" bold="true" box="[197,444,1949,1974]" pageId="3" pageNumber="52">Material examined.</emphasis>
<typeStatus id="B9678876FFB4FFD7FE047EF2FDD339BF" box="[464,616,1950,1974]" pageId="3" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
: ♂ in
<collectionCode id="00CDAE11FFB4FFD7FD077EF2FCA039BF" LSID="urn:lsid:biocol.org:col:33834" box="[723,795,1950,1974]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="3" pageNumber="52">MIZA</collectionCode>
labeled, “
<collectingCountry id="1ECB7644FFB4FFD7FC717EF2FBE939BF" box="[933,1106,1950,1974]" name="Venezuela" pageId="3" pageNumber="52">VENEZUELA</collectingCountry>
: Bolivar State 6°1ʹ38.0&quot;N 61°23ʹ41.1&quot;W,
<quantity id="A1249B31FFB4FFD7FE917EAEFE3439D3" box="[325,399,1986,2010]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="3" pageNumber="52" unit="m" value="690.0">690 m</quantity>
Along La Escalera;
<date id="12621014FFB4FFD7FD547EAEFD4439D3" box="[640,767,1986,2010]" pageId="3" pageNumber="52" value="2008-07-31">31.vii.2008</date>
leg. A.Short, M.Garcia, L.Joly AS-08-059; rocky stream/ [barcode] SM0828289 KUNHM-ENT/
<typeStatus id="B9678876FFB3FFD0FD8F79FBFD483EA6" box="[603,755,151,175]" pageId="4" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
:
<taxonomicName id="A1DC4D57FFB3FFD0FCD479F4FA9E3EA6" authority="Miller and Garcia, 2011" authorityName="Miller and Garcia" authorityYear="2011" box="[768,1317,151,176]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="shorti">
<emphasis id="54A8EAC6FFB3FFD0FCD479F4FBBC3EB9" box="[768,1031,152,176]" italics="true" pageId="4" pageNumber="53">Spanglerodessus shorti</emphasis>
Miller and Garcia, 2011
</taxonomicName>
[red label with black line border].” 37
<typeStatus id="B9678876FFB3FFD0FDD079D1FDCA3EDC" box="[516,625,189,213]" pageId="4" pageNumber="53" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="00CDAE11FFB3FFD0FD5D79D0FD6C3EDD" box="[649,727,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="53">SEMC</collectionCode>
specimen numbers SM0828281-SM0828284, SM0828286- SM0828287, SM0828290, SM0828292, SM0827367-SM0827386, SM0827388-SM
<date id="12621014FFB3FFD0FB8879B3FB083EFE" box="[1116,1203,223,247]" pageId="4" pageNumber="53" value="39-08-27">082739</date>
); 9 labeled same as
<typeStatus id="B9678876FFB3FFD0FF437868FF463F15" box="[151,253,260,284]" pageId="4" pageNumber="53" type="holotype">holotype</typeStatus>
; 24 labeled same as
<typeStatus id="B9678876FFB3FFD0FE307868FDFD3F15" box="[484,582,260,284]" pageId="4" pageNumber="53" type="holotype">holotype</typeStatus>
except “…6°2ʹ10.5&quot;N 61°23ʹ57.8&quot;W,
<quantity id="A1249B31FFB3FFD0FC2C7868FB833F15" box="[1016,1080,260,284]" metricMagnitude="2" metricUnit="m" metricValue="6.3" pageId="4" pageNumber="53" unit="m" value="630.0">630m</quantity>
AS-08-058”; 4 labeled, “GUY- ANA Lethem (25kmSE), KumuFalls 3°15.9ʹN 59°43.6ʹW
<date id="12621014FFB3FFD0FCF8784BFC273F36" box="[812,924,295,319]" pageId="4" pageNumber="53" value="1994-04-04">4Apr1994</date>
,PJSpangler/ Collection #14 side pool at base of falls.” Each
<typeStatus id="B9678876FFB3FFD0FE927821FE1D3F6C" box="[326,422,333,357]" pageId="4" pageNumber="53" type="paratype">paratype</typeStatus>
with label reading, “
<typeStatus id="B9678876FFB3FFD0FD4C7820FC903F6D" box="[664,811,332,356]" pageId="4" pageNumber="53" type="paratype">PARATYPE</typeStatus>
:
<taxonomicName id="A1DC4D57FFB3FFD0FCE37820FAE23F6D" authority="Miller and Garcia, 2011" authorityName="Miller and Garcia" authorityYear="2011" box="[823,1369,332,356]" class="Insecta" family="Dytiscidae" genus="Spanglerodessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="shorti">
<emphasis id="54A8EAC6FFB3FFD0FCE37820FB863F6D" box="[823,1085,332,356]" italics="true" pageId="4" pageNumber="53">Spanglerodessus shorti</emphasis>
Miller and Garcia, 2011
</taxonomicName>
[blue label with black line border].”
</paragraph>
</subSubSection>
</treatment>
</document>