304 lines
37 KiB
XML
304 lines
37 KiB
XML
<document id="B5E2AFD8428FD369AAD95424057CFD19" ID-CLB-Dataset="42408" ID-DOI="10.5281/zenodo.235127" ID-GBIF-Dataset="16b434af-bed4-43fd-bdfc-0f29bcaec7bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="235127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460931586988" checkinUser="plazi" docAuthor="Magalhães, Célio, Wehrtmann, Ingo S., Lara, Luis Rólier & Mantelatto, Fernando L." docDate="2015" docId="710D87BB291E1601D6BDFD17FE9BFECC" docLanguage="en" docName="zt03905p344.pdf" docOrigin="Zootaxa 3905 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptychophallus costaricensis Villalobos 1974" docType="treatment" docVersion="14" lastPageNumber="309" masterDocId="8D34FFC329161608D62AFFA3FFD1FFD0" masterDocTitle="Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)" masterLastPageNumber="344" masterPageNumber="301" pageNumber="309" updateTime="1698582371579" updateUser="ExternalLinkService">
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<mods:title id="87CF8D950D0594082B16A5B3916A4401">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<mods:namePart id="CC956B5B08C91C5758A8B086B01775E9">Magalhães, Célio</mods:namePart>
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<mods:namePart id="EF0CFA6EB249ED87A9B1D75C9B0ACE87">Wehrtmann, Ingo S.</mods:namePart>
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<mods:namePart id="D9B05027EB03625963C453584F2F625D">Lara, Luis Rólier</mods:namePart>
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<mods:namePart id="57DD466A02D25CD850090DAD1D7CA5C0">Mantelatto, Fernando L.</mods:namePart>
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<mods:date id="E270882C7981C71C69CF0B5B3FAECB82">2015</mods:date>
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<treatment id="710D87BB291E1601D6BDFD17FE9BFECC" ID-DOI="http://doi.org/10.5281/zenodo.5491512" ID-GBIF-Taxon="119590156" ID-Zenodo-Dep="5491512" LSID="urn:lsid:plazi:treatment:710D87BB291E1601D6BDFD17FE9BFECC" httpUri="http://treatment.plazi.org/id/710D87BB291E1601D6BDFD17FE9BFECC" lastPageId="9" lastPageNumber="309" pageId="8" pageNumber="309">
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<subSubSection id="B1BE6526291E1600D6BDFD17FE89FD3F" pageId="8" pageNumber="309" type="nomenclature">
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<paragraph id="F91B36AD291E1600D6BDFD17FD63FD1E" blockId="8.[151,690,692,751]" box="[151,690,692,718]" pageId="8" pageNumber="309">
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<heading id="A25381C1291E1600D6BDFD17FD63FD1E" bold="true" box="[151,690,692,718]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
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<taxonomicName id="3EA44D2E291E1600D6BDFD17FD63FD1E" ID-CoL="84H8R" authority="Villalobos, 1974" authorityName="Villalobos" authorityYear="1974" box="[151,690,692,718]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="costaricensis">
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<emphasis id="CBD0EABF291E1600D6BDFD17FD63FD1E" bold="true" box="[151,690,692,718]" pageId="8" pageNumber="309">
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<emphasis id="CBD0EABF291E1600D6BDFD17FE34FD1E" bold="true" box="[151,485,692,718]" italics="true" pageId="8" pageNumber="309">Ptychophallus costaricensis</emphasis>
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<bibRefCitation id="9D354B5C291E1600D7C6FD17FD63FD1E" author="Villalobos" box="[492,690,692,718]" pageId="8" pageNumber="309" refString="Villalobos, C. R. (1974) Ptychophallus costaricensis, a new freshwater crab from Costa Rica. Revista de Biologia Tropical, 21 (2), 197 - 203." type="journal article" year="1974">Villalobos, 1974</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="F91B36AD291E1600D6BDFD75FE89FD3F" blockId="8.[151,690,692,751]" box="[151,344,726,751]" pageId="8" pageNumber="309">
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(
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||
<figureCitation id="619F2A28291E1600D6B5FD75FEF7FD3F" box="[159,294,726,751]" captionStart="FIGURES 10 – 21" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[159,1427,197,1724]" captionTargetId="figure@7.[157,1429,193,1725]" captionTargetPageId="7" captionText="FIGURES 10 – 21. Ptychophallus colombianus (Rathbun 1896): 10, left first gonopod, INPA 1845, caudo-mesial view; 11, same data, latero-cephalic view of distal part; 12, outline of left first gonopod, INPA 1654, caudo-mesial view; 13, same data, INPA 1659; 14, same data, NHMW 4001. Ptychophallus costaricensis Villalobos, 1974: 15, right first gonopod, paratype, UCR-MZ 719, latero-cephalic view of distal part; 16, same data, caudo-mesial view. Ptychophallus goldmanni Pretzmann, 1965: 17, left first gonopod, holotype, USNM 54044, caudo-mesial view; 18, same data, cephalic-apical view of distal part; 19, same data, latero-cephalic view of distal part. Ptychophallus kuna Campos & Lemaitre, 1999: 20, left first gonopod, holotype, USNM 184340, latero-cephalic view of distal part; 21, same data, caudo-mesial view (dr, disto-caudal ridge; lp, lateral process; me, mesial process; mp, marginal process). Scale bars = 1 mm." httpUri="https://zenodo.org/record/235129/files/figure.png" pageId="8" pageNumber="309">Figs. 16, 17</figureCitation>
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,
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<figureCitation id="619F2A28291E1600D718FD74FE9EFD3F" box="[306,335,727,751]" captionStart="FIGURE 80" captionStartId="34.[151,250,1534,1556]" captionTargetBox="[163,1432,524,1513]" captionTargetId="figure@34.[150,1436,519,1513]" captionTargetPageId="34" captionText="FIGURE 80. Distribution map of Ptychophallus costaricensis Villalobos, 1974, and P. exilipes (Rathbun, 1898)." httpUri="https://zenodo.org/record/235138/files/figure.png" pageId="8" pageNumber="309">80</figureCitation>
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)
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</paragraph>
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<subSubSection id="B1BE6526291E1600D6BDFCBDFCF6FCA2" pageId="8" pageNumber="309" type="reference_group">
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<paragraph id="F91B36AD291E1600D6BDFCBDFCD8FC83" blockId="8.[151,1436,798,882]" pageId="8" pageNumber="309">
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<treatmentCitationGroup id="D9B41183291E1600D6BDFCBDFCD8FC83" pageId="8" pageNumber="309">
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<taxonomicName id="3EA44D2E291E1600D6BDFCBDFD60FCE4" ID-CoL="84H8R" authority="Villalobos, 1974: 197" authorityName="Villalobos" authorityPageNumber="197" authorityYear="1974" box="[151,689,798,821]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="costaricensis">
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<emphasis id="CBD0EABF291E1600D6BDFCBDFE6BFCE5" box="[151,442,798,821]" italics="true" pageId="8" pageNumber="309">Ptychophallus costaricensis</emphasis>
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<treatmentCitation id="780510BC291E1600D7ECFCBDFD60FCE4" author="Villalobos" box="[454,689,798,820]" page="197" pageId="8" pageNumber="309" year="1974">
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<bibRefCitation id="9D354B5C291E1600D7ECFCBDFDA9FCE4" author="Villalobos" box="[454,632,798,820]" pageId="8" pageNumber="309" refString="Villalobos, C. R. (1974) Ptychophallus costaricensis, a new freshwater crab from Costa Rica. Revista de Biologia Tropical, 21 (2), 197 - 203." type="journal article" year="1974">Villalobos, 1974</bibRefCitation>
|
||
: 197
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||
</treatmentCitation>
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||
</taxonomicName>
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||
, figs. 1–4.—
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||
<treatmentCitation id="780510BC291E1600D516FCBDFBF5FCE4" author="Rodriguez" box="[828,1060,798,820]" page="184" pageId="8" pageNumber="309" year="1992">
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<bibRefCitation id="9D354B5C291E1600D516FCBDFC3EFCE4" author="Rodriguez" box="[828,1007,798,820]" pageId="8" pageNumber="309" refString="Rodriguez, G. (1992) The freshwater crabs of America. Family Trichodactylidae and supplement to the family Pseudothelphusidae. ORSTOM, Paris, 189 pp. [Faune Tropicale 31]" type="book" year="1992">Rodríguez 1992</bibRefCitation>
|
||
: 184
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||
</treatmentCitation>
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||
.—
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||
<treatmentCitation id="780510BC291E1600D268FCBDFA4DFCE4" author="Campos" box="[1090,1436,798,820]" page="560" pageId="8" pageNumber="309" year="1999">
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||
<bibRefCitation id="9D354B5C291E1600D268FCBDFAB4FCE4" author="Campos" box="[1090,1381,798,820]" pageId="8" pageNumber="309" refString="Campos, M. R. & Lemaitre, R. (1999) Two new freshwater crabs of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) from Panama, with notes on the distribution of the genus. Proceedings of the Biological Society of Washington, 112 (3), 553 - 561." type="journal article" year="1999">Campos & Lemaitre 1999</bibRefCitation>
|
||
: 560
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||
</treatmentCitation>
|
||
(map).—
|
||
<treatmentCitation id="780510BC291E1600D70AFC9EFD60FC83" author="Villalobos" box="[288,689,829,851]" page="297" pageId="8" pageNumber="309" year="2008">
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||
<bibRefCitation id="9D354B5C291E1600D70AFC9EFD50FC83" author="Villalobos" box="[288,641,829,851]" pageId="8" pageNumber="309" refString="Villalobos Hiriart, J. L. & Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. & Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart & Álvarez 2008</bibRefCitation>
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||
: 297
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||
</treatmentCitation>
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||
(in list).
|
||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="F91B36AD291E1600D6BDFCFFFCF6FCA2" blockId="8.[151,1436,798,882]" box="[151,807,860,882]" pageId="8" pageNumber="309">
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||
<treatmentCitationGroup id="D9B41183291E1600D6BDFCFFFCF6FCA2" box="[151,807,860,882]" pageId="8" pageNumber="309">
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<taxonomicName id="3EA44D2E291E1600D6BDFCFFFE6EFCA2" box="[151,447,860,882]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="coastaricensis">
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||
<emphasis id="CBD0EABF291E1600D6BDFCFFFE6EFCA2" box="[151,447,860,882]" italics="true" pageId="8" pageNumber="309">Ptychophallus coastaricensis</emphasis>
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||
</taxonomicName>
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||
[sic]—
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||
<treatmentCitation id="780510BC291E1600D420FCFFFD01FCA2" author="Ng" box="[522,720,860,882]" page="176" pageId="8" pageNumber="309" year="2008">
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<bibRefCitation id="9D354B5C291E1600D420FCFFFD71FCA2" author="Ng" box="[522,672,860,882]" pageId="8" pageNumber="309" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
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||
Ng
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||
<emphasis id="CBD0EABF291E1600D41BFCFEFDB5FCA2" box="[561,612,860,882]" italics="true" pageId="8" pageNumber="309">et al.</emphasis>
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||
2008
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||
</bibRefCitation>
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||
: 176
|
||
</treatmentCitation>
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||
(in list).
|
||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="B1BE6526291E1600D6BDFC3DFE0FFA7F" pageId="8" pageNumber="309" type="materials_examined">
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<paragraph id="F91B36AD291E1600D6BDFC3DFE0FFA7F" blockId="8.[151,1437,926,2031]" pageId="8" pageNumber="309">
|
||
<emphasis id="CBD0EABF291E1600D6BDFC3DFE51FC67" bold="true" box="[151,384,926,951]" pageId="8" pageNumber="309">Material examined</emphasis>
|
||
.
|
||
<collectingCountry id="81B3763D291E1600D7BEFC3CFD93FC67" box="[404,578,927,951]" name="Costa Rica" pageId="8" pageNumber="309">COSTA RICA</collectingCountry>
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||
. Cartago Province, Atlantic drainage:
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||
<specimenCount id="EFA2FD24291E1600D224FC3CFBB3FC67" box="[1038,1122,926,951]" pageId="8" pageNumber="309" type="male">1 male</specimenCount>
|
||
,
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||
<collectionCode id="9FB5AE68291E1600D25EFC3CFB14FC67" LSID="urn:lsid:biocol.org:col:34871" box="[1140,1221,927,951]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="309">USNM</collectionCode>
|
||
1089241, Tapantí National Park, unnamed tributaries, approximately
|
||
<quantity id="3E5C9B48291E1600D4FEFC60FCDDFC0B" box="[724,780,962,987]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="309" unit="km" value="9.0">9 km</quantity>
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||
(road) NW tunnel,
|
||
<quantity id="3E5C9B48291E1600D5CDFC60FBECFC0A" box="[999,1085,963,987]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="309" unit="m" value="1400.0">1400 m</quantity>
|
||
,
|
||
<geoCoordinate id="9C90506A291E1600D263FC60FB48FC0B" box="[1097,1177,963,987]" direction="north" orientation="latitude" pageId="8" pageNumber="309" precision="555" value="9.72">9.72°N</geoCoordinate>
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<geoCoordinate id="9C90506A291E1600D28AFC60FAD4FC0B" box="[1184,1285,963,987]" direction="west" orientation="longitude" pageId="8" pageNumber="309" precision="555" value="-83.78">83.78°W</geoCoordinate>
|
||
,
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||
<date id="8D1A106D291E1600D325FC61FA49FC0B" box="[1295,1432,962,987]" pageId="8" pageNumber="309" value="1988-06-08" valueMax="1988-06-09" valueMin="1988-06-08">8–9.vi.1988</date>
|
||
, C.M. & O.S. Flint Jr. leg.—San José Province, Pacific drainage:
|
||
<specimenCount id="EFA2FD24291E1600D5AFFC44FC04FC2F" box="[901,981,998,1023]" pageId="8" pageNumber="309" type="male">1 male</specimenCount>
|
||
,
|
||
<typeStatus id="261F880F291E1600D5C9FC44FB97FC2F" box="[995,1094,999,1023]" pageId="8" pageNumber="309" type="paratype">paratype</typeStatus>
|
||
, UCR-MZ 707, Dota Canton, Santa María, Finca El Cedral,
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||
<date id="8D1A106D291E1600D7C5FBA9FDB6FBF3" box="[495,615,1034,1059]" pageId="8" pageNumber="309" value="1972-04-23">23.iv.1972</date>
|
||
, R. Saenz, leg.;
|
||
<specimenCount id="EFA2FD24291E1600D50BFBA9FCA9FBF3" box="[801,888,1034,1059]" pageId="8" pageNumber="309" type="male">6 males</specimenCount>
|
||
(cw 17.0, cl 11.6 – cw 25.8, cl 16.0),
|
||
<typeStatus id="261F880F291E1600D300FBA8FA49FBF3" box="[1322,1432,1035,1059]" pageId="8" pageNumber="309" type="paratype">paratypes</typeStatus>
|
||
, UCR-MZ 721, Dota Canton, creek approximately 5.0 km of house at El Cedral Farm, road to Naranjo,
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||
<date id="8D1A106D291E1600D33FFB8DFAA6FB97" box="[1301,1399,1070,1095]" pageId="8" pageNumber="309" value="1972-05-05">5.v.1972</date>
|
||
, R. Saenz, V. Ramírez & C. Villalobos leg.;
|
||
<specimenCount id="EFA2FD24291E1600D471FBF0FD60FBBB" box="[603,689,1106,1131]" pageId="8" pageNumber="309" type="male">2 males</specimenCount>
|
||
(cw 23.1, cl 14.0; cw 33.5, cl 20.3),
|
||
<specimenCount id="EFA2FD24291E1600D266FBF0FB17FBBB" box="[1100,1222,1106,1131]" pageId="8" pageNumber="309" type="female">10 females</specimenCount>
|
||
(cw 18.6, cl 11.8 – cw 25.2, cl 15.2),
|
||
<typeStatus id="261F880F291E1600D742FBD4FE07FB5F" box="[360,470,1143,1167]" pageId="8" pageNumber="309" type="paratype">paratypes</typeStatus>
|
||
, UCR-MZ 720, San Lucas, creek aprox.
|
||
<quantity id="3E5C9B48291E1600D587FBD5FC2DFB5F" box="[941,1020,1142,1167]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="309" unit="km" value="5.5">5.5 km</quantity>
|
||
of house at El Cedral Farm, road to Naranjo,
|
||
<date id="8D1A106D291E1600D729FB38FEB4FB63" box="[259,357,1179,1203]" pageId="8" pageNumber="309" value="1972-05-07">7.v.1972</date>
|
||
, R. Saez, V. Ramírez & C. Villalobos leg.;
|
||
<specimenCount id="EFA2FD24291E1600D542FB39FC13FB63" box="[872,962,1178,1203]" pageId="8" pageNumber="309" type="male">6 males</specimenCount>
|
||
(cw 15.5, cl 10.1 – cw 29.1, cl 18.0),
|
||
<specimenCount id="EFA2FD24291E1600D3A4FB38FF20FB07" pageId="8" pageNumber="309" type="female">4 females</specimenCount>
|
||
,
|
||
<typeStatus id="261F880F291E1600D6D6FB1CFEBBFB07" box="[252,362,1215,1239]" pageId="8" pageNumber="309" type="paratype">paratypes</typeStatus>
|
||
, UCR-MZ 722, Naranjo, Naranjo River, Guillermo Solís’ farm,
|
||
<date id="8D1A106D291E1600D214FB1CFB71FB07" box="[1086,1184,1215,1239]" pageId="8" pageNumber="309" value="1972-05-07">7.v.1972</date>
|
||
, R. Saenz, V. Ramírez & C. Villalobos leg.;
|
||
<specimenCount id="EFA2FD24291E1600D7A7FB40FE0BFB2B" box="[397,474,1250,1275]" pageId="8" pageNumber="309" type="male">1 male</specimenCount>
|
||
(cw 33.8, cl 21.4),
|
||
<specimenCount id="EFA2FD24291E1600D497FB40FCF1FB2B" box="[701,800,1250,1275]" pageId="8" pageNumber="309" type="female">1 female</specimenCount>
|
||
(cw 28.2, cl 17.5),
|
||
<typeStatus id="261F880F291E1600D229FB40FBA0FB2B" box="[1027,1137,1251,1275]" pageId="8" pageNumber="309" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<collectionCode id="9FB5AE68291E1600D254FB40FB6CFB2B" box="[1150,1213,1251,1275]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="8" pageNumber="309">INPA</collectionCode>
|
||
1844, same data;
|
||
<specimenCount id="EFA2FD24291E1600D3A4FB40FF1CFACF" pageId="8" pageNumber="309" type="male">1 male</specimenCount>
|
||
(cw 34.8, cl 20.3),
|
||
<specimenCount id="EFA2FD24291E1600D780FAA4FDDAFACF" box="[426,523,1286,1311]" pageId="8" pageNumber="309" type="female">1 female</specimenCount>
|
||
(cw 26.5, cl 16.8),
|
||
<typeStatus id="261F880F291E1600D4C2FAA4FC87FACF" box="[744,854,1287,1311]" pageId="8" pageNumber="309" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<collectionCode id="9FB5AE68291E1600D548FAA4FC65FACE" box="[866,948,1287,1310]" httpUri="http://grbio.org/cool/1h73-pg9e" name="Tianjin Museum of Natural History" pageId="8" pageNumber="309">TMNH</collectionCode>
|
||
6408, San Isidro de El General, Los Tubos Creek,
|
||
<date id="8D1A106D291E1600D6C0FA89FE85FA93" box="[234,340,1322,1347]" pageId="8" pageNumber="309" value="1972-04-07">7.iv.1972</date>
|
||
, C. Villalobos leg.;
|
||
<specimenCount id="EFA2FD24291E1600D410FA88FD43FA93" box="[570,658,1322,1347]" pageId="8" pageNumber="309" type="male">2 males</specimenCount>
|
||
plus 1 G1,
|
||
<typeStatus id="261F880F291E1600D536FA88FC5BFA93" box="[796,906,1323,1347]" pageId="8" pageNumber="309" type="paratype">paratypes</typeStatus>
|
||
, UCR-MZ 719, San Isidro de El General, Los Tubos Creek,
|
||
<quantity id="3E5C9B48291E1600D71EFAECFEBFFAB7" box="[308,366,1358,1383]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="309" unit="km" value="1.0">1 km</quantity>
|
||
W Boquete, Adan Bonilla’s farm,
|
||
<date id="8D1A106D291E1600D4DDFAEDFCB0FAB7" box="[759,865,1358,1383]" pageId="8" pageNumber="309" value="1972-04-07">7.iv.1972</date>
|
||
, V. Juárez leg.;
|
||
<specimenCount id="EFA2FD24291E1600D23EFAECFBBCFAB7" box="[1044,1133,1358,1383]" pageId="8" pageNumber="309" type="male">4 males</specimenCount>
|
||
,
|
||
<specimenCount id="EFA2FD24291E1600D250FAECFB38FAB7" box="[1146,1257,1358,1383]" pageId="8" pageNumber="309" type="female">2 females</specimenCount>
|
||
, UCR-MZ 742- 0 1, San Isidro del General, Los Tubos Creek,
|
||
<quantity id="3E5C9B48291E1600D4B4FAD0FD09FA5B" box="[670,728,1394,1419]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="309" unit="km" value="1.0">1 km</quantity>
|
||
W Boquete, Adan Bonilla’s farm,
|
||
<date id="8D1A106D291E1600D248FAD1FB0AFA5B" box="[1122,1243,1394,1419]" pageId="8" pageNumber="309" value="1972-04-22">22.iv.1972</date>
|
||
, V. Juárez leg.;
|
||
<specimenCount id="EFA2FD24291E1600D3A4FAD0FF01FA7F" pageId="8" pageNumber="309" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="9FB5AE68291E1600D6F1FA34FECBFA7F" box="[219,282,1431,1455]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="8" pageNumber="309">INPA</collectionCode>
|
||
1858, same data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE6526291E1600D6EDFA19FB9FF91F" pageId="8" pageNumber="309" type="description">
|
||
<paragraph id="F91B36AD291E1600D6EDFA19FB9FF91F" blockId="8.[151,1437,926,2031]" pageId="8" pageNumber="309">
|
||
<emphasis id="CBD0EABF291E1600D6EDFA19FE3CFA03" bold="true" box="[199,493,1466,1491]" pageId="8" pageNumber="309">Description of gonopod</emphasis>
|
||
. Straight in mesiocaudal view, apex bent approximately 90° in laterocephalic direction. Marginal suture on mesial side, straight; marginal process rounded, produced beyond distal border of apex. Mesial process small, subtriangular, slightly downturned, with 2 short spines on laterocephalic side, proximal spine slightly sharper than distal spine. Lateral process long, strongly bilobed, approximately 70% of stem length (from proximal opening to caudal border of apex), usually as wide as lateral end of apex, with deep notch in the middle; lateral border with deep median incision, both lobes subequal in size. Distocaudal ridge short, narrow well separated from distal border of lateral process by distinct depression. Apex oblong, narrow, with narrow notch on lateral end. Field of apical spines well developed, facing towards cephalolateral side.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE6526291E1600D6EDF979FABAF88F" pageId="8" pageNumber="309" type="materials_examined">
|
||
<paragraph id="F91B36AD291E1600D6EDF979FABAF88F" blockId="8.[151,1437,926,2031]" pageId="8" pageNumber="309">
|
||
<emphasis id="CBD0EABF291E1600D6EDF979FE2EF923" bold="true" box="[199,511,1754,1779]" pageId="8" pageNumber="309">
|
||
<typeStatus id="261F880F291E1600D6EDF979FEE5F923" box="[199,308,1754,1779]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="261F880F291E1600D75AF979FE72F923" box="[368,419,1754,1779]" pageId="8" pageNumber="309">type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
. Male, cw 44.4, cl 27.1, UCR-MZ 392.
|
||
<collectingCountry id="81B3763D291E1600D5EFF978FB92F923" box="[965,1091,1754,1779]" name="Costa Rica" pageId="8" pageNumber="309">Costa Rica</collectingCountry>
|
||
, San José Province, San Isidro del General (Pacific drainage). The
|
||
<typeStatus id="261F880F291E1600D405F95DFD40F8C7" box="[559,657,1790,1815]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
|
||
could not be found in the UCR-MZ holdings. It is a male originally separated from lot CVS-079-72 (number in C. Villalobos field notes) (
|
||
<bibRefCitation id="9D354B5C291E1600D5CBF881FB74F8EB" author="Villalobos" box="[993,1189,1826,1851]" pageId="8" pageNumber="309" refString="Villalobos, C. R. (1974) Ptychophallus costaricensis, a new freshwater crab from Costa Rica. Revista de Biologia Tropical, 21 (2), 197 - 203." type="journal article" year="1974">Villalobos, 1974</bibRefCitation>
|
||
), which received the catalogue number UCR-MZ 719. The G1 of one male
|
||
<typeStatus id="261F880F291E1600D4D3F8E4FC88F88F" box="[761,857,1863,1887]" pageId="8" pageNumber="309" type="paratype">paratype</typeStatus>
|
||
from this same lot is illustrated in
|
||
<figureCitation id="619F2A28291E1600D2F5F8E5FAB6F88F" box="[1247,1383,1862,1887]" captionStart="FIGURES 10 – 21" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[159,1427,197,1724]" captionTargetId="figure@7.[157,1429,193,1725]" captionTargetPageId="7" captionText="FIGURES 10 – 21. Ptychophallus colombianus (Rathbun 1896): 10, left first gonopod, INPA 1845, caudo-mesial view; 11, same data, latero-cephalic view of distal part; 12, outline of left first gonopod, INPA 1654, caudo-mesial view; 13, same data, INPA 1659; 14, same data, NHMW 4001. Ptychophallus costaricensis Villalobos, 1974: 15, right first gonopod, paratype, UCR-MZ 719, latero-cephalic view of distal part; 16, same data, caudo-mesial view. Ptychophallus goldmanni Pretzmann, 1965: 17, left first gonopod, holotype, USNM 54044, caudo-mesial view; 18, same data, cephalic-apical view of distal part; 19, same data, latero-cephalic view of distal part. Ptychophallus kuna Campos & Lemaitre, 1999: 20, left first gonopod, holotype, USNM 184340, latero-cephalic view of distal part; 21, same data, caudo-mesial view (dr, disto-caudal ridge; lp, lateral process; me, mesial process; mp, marginal process). Scale bars = 1 mm." httpUri="https://zenodo.org/record/235129/files/figure.png" pageId="8" pageNumber="309">Figs. 15, 16</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE6526291E1600D6EDF8C9FC8DF877" pageId="8" pageNumber="309" type="distribution">
|
||
<paragraph id="F91B36AD291E1600D6EDF8C9FC8DF877" blockId="8.[151,1437,926,2031]" pageId="8" pageNumber="309">
|
||
<emphasis id="CBD0EABF291E1600D6EDF8C9FE88F853" bold="true" box="[199,345,1898,1923]" pageId="8" pageNumber="309">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="3EA44D2E291E1600D746F8C9FD7AF853" box="[364,683,1898,1923]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="costaricensis">
|
||
<emphasis id="CBD0EABF291E1600D746F8C9FD7AF853" box="[364,683,1898,1923]" italics="true" pageId="8" pageNumber="309">Ptychophallus costaricensis</emphasis>
|
||
</taxonomicName>
|
||
is distributed on both Atlantic and Pacific drainages of central southern
|
||
<collectingCountry id="81B3763D291E1600D6D4F82CFEAAF877" box="[254,379,1934,1959]" name="Costa Rica" pageId="8" pageNumber="309">Costa Rica</collectingCountry>
|
||
, Cartago and San José Provinces (
|
||
<figureCitation id="619F2A28291E1600D4D1F82DFC81F877" box="[763,848,1934,1959]" captionStart="FIGURE 80" captionStartId="34.[151,250,1534,1556]" captionTargetBox="[163,1432,524,1513]" captionTargetId="figure@34.[150,1436,519,1513]" captionTargetPageId="34" captionText="FIGURE 80. Distribution map of Ptychophallus costaricensis Villalobos, 1974, and P. exilipes (Rathbun, 1898)." httpUri="https://zenodo.org/record/235138/files/figure.png" pageId="8" pageNumber="309">Fig. 80</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE6526291E1601D6EDF811FE9BFECC" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309" type="discussion">
|
||
<paragraph id="F91B36AD291E1601D6EDF811FE9BFECC" blockId="8.[151,1437,926,2031]" lastBlockId="9.[151,1436,151,285]" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
|
||
<emphasis id="CBD0EABF291E1600D6EDF811FEE5F81B" bold="true" box="[199,308,1970,1995]" pageId="8" pageNumber="309">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="3EA44D2E291E1600D769F811FDAFF81B" box="[323,638,1970,1995]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="costaricensis">
|
||
<emphasis id="CBD0EABF291E1600D769F811FDAFF81B" box="[323,638,1970,1995]" italics="true" pageId="8" pageNumber="309">Ptychophallus costaricensis</emphasis>
|
||
</taxonomicName>
|
||
is very close to
|
||
<taxonomicName id="3EA44D2E291E1600D516F817FC20F81B" box="[828,1009,1970,1995]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="micracanthus">
|
||
<emphasis id="CBD0EABF291E1600D516F817FC20F81B" box="[828,1009,1970,1995]" italics="true" pageId="8" pageNumber="309">P. micracanthus</emphasis>
|
||
</taxonomicName>
|
||
. They differ by the number of spines on the laterocephalic side of the mesial process (two in
|
||
<taxonomicName id="3EA44D2E291E1600D52BF874FC7FF83E" box="[769,942,2007,2030]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="costaricensis">
|
||
<emphasis id="CBD0EABF291E1600D52BF874FC7FF83E" box="[769,942,2007,2030]" italics="true" pageId="8" pageNumber="309">P. costaricensis</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="619F2A28291E1600D596F875FBC0F83F" box="[956,1041,2006,2031]" captionStart="FIGURES 10 – 21" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[159,1427,197,1724]" captionTargetId="figure@7.[157,1429,193,1725]" captionTargetPageId="7" captionText="FIGURES 10 – 21. Ptychophallus colombianus (Rathbun 1896): 10, left first gonopod, INPA 1845, caudo-mesial view; 11, same data, latero-cephalic view of distal part; 12, outline of left first gonopod, INPA 1654, caudo-mesial view; 13, same data, INPA 1659; 14, same data, NHMW 4001. Ptychophallus costaricensis Villalobos, 1974: 15, right first gonopod, paratype, UCR-MZ 719, latero-cephalic view of distal part; 16, same data, caudo-mesial view. Ptychophallus goldmanni Pretzmann, 1965: 17, left first gonopod, holotype, USNM 54044, caudo-mesial view; 18, same data, cephalic-apical view of distal part; 19, same data, latero-cephalic view of distal part. Ptychophallus kuna Campos & Lemaitre, 1999: 20, left first gonopod, holotype, USNM 184340, latero-cephalic view of distal part; 21, same data, caudo-mesial view (dr, disto-caudal ridge; lp, lateral process; me, mesial process; mp, marginal process). Scale bars = 1 mm." httpUri="https://zenodo.org/record/235129/files/figure.png" pageId="8" pageNumber="309">Fig. 15</figureCitation>
|
||
]; one in
|
||
<taxonomicName id="3EA44D2E291E1600D25AF874FAF5F83F" box="[1136,1316,2006,2031]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="micracanthus">
|
||
<emphasis id="CBD0EABF291E1600D25AF874FAF5F83F" box="[1136,1316,2006,2031]" italics="true" pageId="8" pageNumber="309">P. micracanthus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="619F2A28291E1600D31BF875FA56F83F" box="[1329,1415,2006,2031]" captionStart="FIGURES 33 – 43" captionStartId="15.[151,264,1763,1786]" captionTargetBox="[178,1412,197,1737]" captionTargetId="figure@15.[174,1412,193,1742]" captionTargetPageId="15" captionText="FIGURES 33 – 43. Ptychophallus micracanthus Rodríguez, 1994: 33, left first gonopod, holotype, USNM 240106, caudomesial view; 34, same data, latero-cephalic view of distal part. Ptychophallus montanus (Rathbun, 1898): 35, left first gonopod, lectotype, USNM 19486, caudo-mesial view; 36, same data, latero-cephalic view; 37, outline of left first gonopod, INPA 1843, caudo-mesial view; 38, same data, INPA 1847, caudo-mesial view. Ptychophallus tristani (Rathbun, 1896): 39, left first gonopod (partially damaged on proximal part), holotype, USNM 19047, caudo-mesial view; 40, same data, latero-cephalic view of distal part; 41, left first gonopod, holotype of Ptychophallus (Ptychophallus) campylus Pretzmann, 1968, NHMW 21849, caudo-mesial view; 42, outline of right gonopod, INPA 1836, caudo-mesial view; 43, outline of left first gonopod, INPA 1851, caudo-mesial view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235131/files/figure.png" pageId="8" pageNumber="309">Fig. 34</figureCitation>
|
||
]), and the development of the marginal process (more developed, reaching beyond the caudal border of apex in
|
||
<taxonomicName id="3EA44D2E291F1601D3ADFF3AFEF8FF04" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="costaricensis">
|
||
<emphasis id="CBD0EABF291F1601D3ADFF3AFEF8FF04" italics="true" pageId="9" pageNumber="310">P. costaricensis</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="619F2A28291F1601D710FF1FFE42FF05" box="[314,403,188,213]" captionStart="FIGURES 10 – 21" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[159,1427,197,1724]" captionTargetId="figure@7.[157,1429,193,1725]" captionTargetPageId="7" captionText="FIGURES 10 – 21. Ptychophallus colombianus (Rathbun 1896): 10, left first gonopod, INPA 1845, caudo-mesial view; 11, same data, latero-cephalic view of distal part; 12, outline of left first gonopod, INPA 1654, caudo-mesial view; 13, same data, INPA 1659; 14, same data, NHMW 4001. Ptychophallus costaricensis Villalobos, 1974: 15, right first gonopod, paratype, UCR-MZ 719, latero-cephalic view of distal part; 16, same data, caudo-mesial view. Ptychophallus goldmanni Pretzmann, 1965: 17, left first gonopod, holotype, USNM 54044, caudo-mesial view; 18, same data, cephalic-apical view of distal part; 19, same data, latero-cephalic view of distal part. Ptychophallus kuna Campos & Lemaitre, 1999: 20, left first gonopod, holotype, USNM 184340, latero-cephalic view of distal part; 21, same data, caudo-mesial view (dr, disto-caudal ridge; lp, lateral process; me, mesial process; mp, marginal process). Scale bars = 1 mm." httpUri="https://zenodo.org/record/235129/files/figure.png" pageId="9" pageNumber="310">Fig. 16</figureCitation>
|
||
]; less developed, reaching about the caudal border of apex in
|
||
<taxonomicName id="3EA44D2E291F1601D24CFF1EFACCFF05" box="[1126,1309,188,213]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="micracanthus">
|
||
<emphasis id="CBD0EABF291F1601D24CFF1EFACCFF05" box="[1126,1309,188,213]" italics="true" pageId="9" pageNumber="310">P. micracanthus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="619F2A28291F1601D304FF1FFA56FF04" box="[1326,1415,188,213]" captionStart="FIGURES 33 – 43" captionStartId="15.[151,264,1763,1786]" captionTargetBox="[178,1412,197,1737]" captionTargetId="figure@15.[174,1412,193,1742]" captionTargetPageId="15" captionText="FIGURES 33 – 43. Ptychophallus micracanthus Rodríguez, 1994: 33, left first gonopod, holotype, USNM 240106, caudomesial view; 34, same data, latero-cephalic view of distal part. Ptychophallus montanus (Rathbun, 1898): 35, left first gonopod, lectotype, USNM 19486, caudo-mesial view; 36, same data, latero-cephalic view; 37, outline of left first gonopod, INPA 1843, caudo-mesial view; 38, same data, INPA 1847, caudo-mesial view. Ptychophallus tristani (Rathbun, 1896): 39, left first gonopod (partially damaged on proximal part), holotype, USNM 19047, caudo-mesial view; 40, same data, latero-cephalic view of distal part; 41, left first gonopod, holotype of Ptychophallus (Ptychophallus) campylus Pretzmann, 1968, NHMW 21849, caudo-mesial view; 42, outline of right gonopod, INPA 1836, caudo-mesial view; 43, outline of left first gonopod, INPA 1851, caudo-mesial view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235131/files/figure.png" pageId="9" pageNumber="310">Fig. 33</figureCitation>
|
||
]).
|
||
<taxonomicName id="3EA44D2E291F1601D6BDFF7CFE00FF28" box="[151,465,223,248]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="costaricensis">
|
||
<emphasis id="CBD0EABF291F1601D6BDFF7CFE00FF28" box="[151,465,223,248]" italics="true" pageId="9" pageNumber="310">Ptychophallus costaricensis</emphasis>
|
||
</taxonomicName>
|
||
occurs only in central-southern
|
||
<collectingCountry id="81B3763D291F1601D56AFF43FC6FFF28" box="[832,958,223,248]" name="Costa Rica" pageId="9" pageNumber="310">Costa Rica</collectingCountry>
|
||
, whereas
|
||
<taxonomicName id="3EA44D2E291F1601D204FF42FB35FF28" box="[1070,1252,223,248]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="micracanthus">
|
||
<emphasis id="CBD0EABF291F1601D204FF42FB35FF28" box="[1070,1252,223,248]" italics="true" pageId="9" pageNumber="310">P. micracanthus</emphasis>
|
||
</taxonomicName>
|
||
is distributed in central
|
||
<collectingCountry id="81B3763D291F1601D6C0FEA6FE97FECC" box="[234,326,261,284]" name="Panama" pageId="9" pageNumber="310">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |