283 lines
33 KiB
XML
283 lines
33 KiB
XML
<document ID-CLB-Dataset="24300" ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="3DAA2F51B5B205102782C5F093C90C0B" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole fisaka Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="2BA075C8-7EA3-444A-B35F-CE9BA0DC0798" docUuidSource="ZooBank" docVersion="7" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1732649601836" updateUser="ExternalLinkService">
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<mods:titleInfo id="397C225599DFECF7462DC8010B9E60D1">
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<mods:title id="BBAD0AD0A54F6EF20D1DBD92C9A47AC3">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:namePart id="7E13E0F6FC4BEB5844147F6C4CE89D60">Salata, Sebastian</mods:namePart>
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<mods:name id="EFF6C05A76E472E4D8584FCB5C66CD85" type="personal">
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<mods:namePart id="2FBC8D34C684C7013B94A8BD51AEA756">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="847DF2911F6A54CFEEF79B3A6B44C1D4">text</mods:typeOfResource>
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<mods:title id="06087D069E87A110C748D1F7BA9D4D7B">ZooKeys</mods:title>
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<mods:part id="A56A249F775C53AFBBBC8D67CB5D37D6">
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<mods:date id="15E38820149A975A0DA3D69E8A334EF6">2020</mods:date>
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<mods:detail id="BC66F60C0CC0E4CC9016431101BD618E" type="volume">
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<mods:number id="6F7D05F4AE1E6F314CC3E568ED5BD0CF">905</mods:number>
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<mods:start id="12DDE981502AD5707FE3DD74C6EFE2DD">1</mods:start>
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<mods:end id="150BDDC94E3A77B2F858FC5EE46965EB">235</mods:end>
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<mods:url id="B5B44637AA9E2BCBF6B53FAA503E658A">http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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<mods:classification id="DAA6604A6C356E144E7AA69B5FD79545">journal article</mods:classification>
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<mods:identifier id="A7D850E6B6F3B59810D8F75BC4EF93FB" type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier id="6C3048D0FD17980CF70E5B450B545DA8" type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<treatment id="3DAA2F51B5B205102782C5F093C90C0B" ID-GBIF-Taxon="161777008" LSID="urn:lsid:zoobank.org:act:2BA075C8-7EA3-444A-B35F-CE9BA0DC0798" httpUri="http://treatment.plazi.org/id/3DAA2F51B5B205102782C5F093C90C0B" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="33515FC8FEE2373D92AA2FF4B52D7726" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="9216567B5DC0D31B8ABDF7A1D1BF4ADF" pageId="0" pageNumber="1">
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<taxonomicName id="63DAAFBBBB5492186E1C6CB9AD0CED66" ID-CoL="CML5Q" LSID="http://zoobank.org/2BA075C8-7EA3-444A-B35F-CE9BA0DC0798" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka" status="sp. nov.">Pheidole fisaka</taxonomicName>
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<taxonomicNameLabel id="C7E1B156FFE6F8A2EE5890CAC2DD0337" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="F7A7A185AA7F28728B07C87546E21A65" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">Figs 79A-F</figureCitation>
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<figureCitation id="02D33B241308BB4E4C83A041C8F65178" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84R</figureCitation>
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<figureCitation id="C076000203B1E5365682D825BCE3EE2A" captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86R</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1536337C24BC49DD9CB2F1B16CB4C210" pageId="0" pageNumber="1" type="type material">
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<paragraph id="92ED8C4513A492531A52A12EA93DDBD9" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="7D639D0192C098F6BE7DB58A3907C905" pageId="0" pageNumber="1">
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<emphasis id="AA4E6576F2B56E68925019579B20DC72" bold="true" pageId="0" pageNumber="1">
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<emphasis id="A2D9491BE16F577C5258CE66218A5F5E" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Toamasina; Ankerana;
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<geoCoordinate id="5660A5C72996EB627F734032134F23A3" degrees="18.40062" direction="south" orientation="latitude" precision="1" value="-18.40062">-18.40062</geoCoordinate>
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,
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<geoCoordinate id="02DA677F48D157F43C1D968B96D3AF90" degrees="48.81311" direction="east" orientation="longitude" precision="1" value="48.81311">48.81311</geoCoordinate>
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; alt. 865 m; 17 Jan 2012; B.L. Fisher et al. leg.; CASENT0274930 (CASC).
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<emphasis id="A5B60C849EA51677BC2A3D3F54A56FB3" bold="true" pageId="0" pageNumber="1">
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<emphasis id="B46387360410138D6B463E7301168473" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
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.
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</emphasis>
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Madagascar. •1w.; same data as for holotype; BLF27851, CASENT0923180 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="41F6765108077A8D31E5435A235BD199" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="902C0275B4CB74BC8BE560C08F05325F" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="3F748E6315AF0415DF200EDEAFB830DD" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="514B9E347F20387DB9743B5A2F5535F6" bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
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: •1w., 1s.; Ankerana;
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<geoCoordinate id="DF1D4C98FBA3900C584826BCC24270F8" degrees="18.4061" direction="south" orientation="latitude" precision="5" value="-18.4061">-18.4061</geoCoordinate>
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,
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<geoCoordinate id="D0A03C6264349AE22B3620EAC53FF4D8" degrees="48.82029" direction="east" orientation="longitude" precision="1" value="48.82029">48.82029</geoCoordinate>
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; alt. 725 m; 16 Jan 2012; B.L. Fisher et al. leg.; CASENT0273490, CASENT0273492 (CASC). •2w., 1m.; Ankerana;
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<geoCoordinate id="0BF826DC96A08990263E544C41213780" degrees="18.40829" direction="south" orientation="latitude" precision="1" value="-18.40829">-18.40829</geoCoordinate>
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,
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<geoCoordinate id="3496FF6FB9BBBABD571ED55498F32D7C" degrees="48.82107" direction="east" orientation="longitude" precision="1" value="48.82107">48.82107</geoCoordinate>
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; alt. 750 m; 21 Jan 2012; B.L. Fisher et al. leg.; CASENT0275257, CASENT0275258 (CASC). •3w., 1q.; Ankerana;
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<geoCoordinate id="72995E31D45251F91481225A99779F88" degrees="18.4104" direction="south" orientation="latitude" precision="5" value="-18.4104">-18.4104</geoCoordinate>
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,
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<geoCoordinate id="6802CB872D76A1D2C127DF68E7D5CE33" degrees="48.8189" direction="east" orientation="longitude" precision="5" value="48.8189">48.8189</geoCoordinate>
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; alt. 855 m; 27 Jan 2012; B.L. Fisher et al. leg.; CASENT0274031, CASENT0274032, CASENT0274033 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="D9CDB3B7757D5977089A4968877F7B52" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="A53CEA9FAEA2A1EFD199E914E031AA0B" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="E7A1939A96243461E3CE4ECC145ED224" pageId="0" pageNumber="1">
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<emphasis id="6C584DB142F6DACCE6468ECBF31C1D5C" bold="true" pageId="0" pageNumber="1">
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<emphasis id="CC18675BC7920253C085BE67106C4FEB" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Small species: HL: 1.01-1.0; HW: 0.99-0.98, WL: 0.8-0.76; head in full-face view sub-rectangular, anterior of eyes straight, posterior of eyes convex; sides of the head with moderately dense, short, erect pilosity; antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, foveolate with thick, longitudinal to irregular, and long rugae; inner hypostomal teeth distinct, moderately high, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth thinner and narrower than inner hypostomal teeth, dentate; gaster finely shagreened; body yellowish brown.
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<emphasis id="9A1D56A206361D9A1F8850D328CB374E" bold="true" pageId="0" pageNumber="1">
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<emphasis id="1F65C2E074BC296D7EDBA9F164CAEE22" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Head foveolate, genae smooth; promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; propodeal spines minute; mesosoma foveolate, lateral sides of pronotum and propodeum with fading sculpture, katepisternum smooth.
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</paragraph>
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</subSubSection>
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<subSubSection id="4F0C22CFAE7A0F7FBFF42A769F387256" pageId="0" pageNumber="1" type="description">
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<paragraph id="2F1EDCD25CBCA01CAE415800DE8F9C5A" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="4974EA9FB0CA41266AA4CACFC8BAFF20" pageId="0" pageNumber="1">
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<emphasis id="D8603CB464983DC99D3D3A3FF65F9A0F" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="4B537D3EF976D79878D25841B1468F35" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 2): HL: 1.01-1.0; HW: 0.99-0.98; SL: 0.44-0.45; EL: 0.13-0.13; WL: 0.8-0.76; PSL: 0.17-0.14; MTL: 0.440.44; PNW: 0.58-0.54; PTW: 0.16-0.14; PPW: 0.39-0.35; CI: 98.5-97.7; SI: 44.5-46.2; PSLI: 17.1-13.6; PPI: 40.6-40.5; PNI: 58.0-55.0; MTI: 44.6-45.3.
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<emphasis id="6FB05D6CB677D7210F232B62D7934160" bold="true" pageId="0" pageNumber="1">
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<emphasis id="027E4B3CBA5D5F2531FC3E2753D056F3" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view longer than wide, anterior of eyes straight, posterior of eyes convex; slightly widening posteriorly (Fig.
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<figureCitation id="AFAB49474F3832EB20E049CAD8171F8E" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces relatively flat; dorsal face finely depressed posteriorly, forming shallow, transverse depression between frons and occipital lobes; inner hypostomal teeth visible. Sides of the head with moderately dense, short, erect pilosity; whole head with dense, short, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, foveolate with thick, longitudinal to irregular, and long rugae. Occipital lobes shiny, with thick, irregular rugae, interspaces with distinct, irregular rugulae; frons with moderately dense, thick and longitudinal rugae, interspaces smooth to rugo-foveolate in the posterior part; genae shiny, with dense and thin, irregular rugoreticulation; malar area with thin, longitudinal, dense rugoreticulation. Centre of clypeus shiny and smooth, lateral sides with longitudinal rugulae; median notch present, wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae present. Scape, when laid back, reaching the midlength of head; pilosity suberect (Fig.
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<figureCitation id="8C11A528901575708F89F9A850D5C3E2" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79B, D</figureCitation>
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). Inner hypostomal teeth distinct, moderately high, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth thinner and narrower than inner hypostomal teeth, dentate (Fig.
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<figureCitation id="E77374685B65247025B903DBD1E37859" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84R</figureCitation>
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).
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<emphasis id="EC1B131F7EF2377974F11AD59708A452" bold="true" pageId="0" pageNumber="1">
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<emphasis id="E1DE5AB00494588285212B090F14912B" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum low and arched, posterior mesonotum steep, without tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines short, triangular, with rounded apex and wide base; humeral area laterally well produced (Fig.
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<figureCitation id="1297EA4B88F9A0D0856BEEE4CF1FBD53" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D</figureCitation>
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). Surface shiny, with fine and moderately dense foveolae and additional rugoreticulation on the lateral sides of propodeum; dorsal surface of promesonotum with fading foveolae; katepisternum smooth. Pilosity moderately dense and long, erect (Fig.
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<figureCitation id="3E74A61C902BA20AE27E229CB9B26118" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
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).
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<emphasis id="46A1855ADDB248CF002188B39BCB0BFE" bold="true" pageId="0" pageNumber="1">
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<emphasis id="3522E291F07DD33218047F1F001782B6" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny and with fine foveolae; peduncle short, with indistinct horizontal lobes on its basal part; node relatively high, triangular, with rounded apex, in rear view node slightly dorsoventrally concave; pilosity moderately sparse and erect (Fig.
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<figureCitation id="10F3C4E5E4DE2AEB4BE31C73FECC2F1A" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
|
|
).
|
|
<emphasis id="FDBC042229597020CD8BAB8B5EF784C2" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="FEAABCD2A511671B1B61E3897431B607" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Shiny, finely shagreened; in dorsal view sides with moderately long, acute, and triangular projections; pilosity long, moderately long, and erect (Fig.
|
|
<figureCitation id="544244C428F8C848B4666924C20EF69E" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
|
|
).
|
|
<emphasis id="D544B39F4735275352E8FDA694B4520A" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="D2E11E7EF649434CB14B3F096C72CDA3" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Shiny and finely shagreened; pilosity dense, moderately long, and erect (Fig.
|
|
<figureCitation id="10ECD8ADB7EA171118A80413504226E6" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
|
|
).
|
|
<emphasis id="1AC1670355AAEE797D2D2E74D98C0047" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="C6085675A7CE19FCB871442A11A0B7CA" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Yellow; frons, occipital lobes; dorsal surface of mesosoma and gaster yellowish brown to brown (Fig.
|
|
<figureCitation id="0495D1CB44FF97CC862C40BA07AE7410" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79D, F</figureCitation>
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).
|
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</paragraph>
|
|
<caption id="51E95D96F0320399197AC3F2DD539B67" doi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1" start="Figure 79" startId="F79">
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|
<paragraph id="3F07A5B86BA82E9E4406D60E5C3A2D97" pageId="0" pageNumber="1">
|
|
<emphasis id="EF75469F7D76AA2C18E4048EB50EF371" bold="true" pageId="0" pageNumber="1">Figure 79.</emphasis>
|
|
<taxonomicName id="955844D29F633A64B496BB41E7C8E6B2" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka">
|
|
<emphasis id="983D7267FF1009BA1F4F4B559A026038" italics="true" pageId="0" pageNumber="1">Pheidole fisaka</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., full-face view (
|
|
<emphasis id="A880D495DF7D5249183E516942928107" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), profile (
|
|
<emphasis id="41CA4E5A0C0428317E6A3B8656555488" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), and dorsal view (
|
|
<emphasis id="61ABD88060841AFBEE816B7A99A17DA1" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
) of paratype minor worker (CASENT0923180) and full-face view (
|
|
<emphasis id="A08B8C8A2CE7C5D2FDFD19EE1677069D" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
), profile (
|
|
<emphasis id="6CD7B1B1419F69FC839AEB26715597F9" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
), and dorsal view (
|
|
<emphasis id="3BACF2934D66B67CC58581D658400C3A" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
) of holotype major worker (CASENT0274930).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="455F161764F16612366CC29B6114BDE9" pageId="0" pageNumber="1">
|
|
<emphasis id="6301E02C8CEEE15555A4CD5F13C19EEC" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
|
Measurements (
|
|
<emphasis id="7226EA5108FD4FE3DAD0512E2B13F9F6" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 2): HL: 0.43-0.43; HW: 0.4-0.39; SL: 0.38-0.37; EL: 0.09-0.08; WL: 0.48-0.48; PSL: 0.07-0.06; MTL: 0.28-0.29; PNW: 0.25-0.25; PTW: 0.06-0.06; PPW: 0.09-0.09; CI: 92.9-91.8; SI: 95.0-93.1; PSLI: 15.2-14.5; PPI: 68.5-63.4; PNI: 62.3-63.9; MTI: 70.5-73.0.
|
|
<emphasis id="3C7C9710247A28DF31C82652ED2377D5" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="4114BED1244160386FA109ACDAACA3BC" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
|
.
|
|
</emphasis>
|
|
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
|
|
<figureCitation id="E56257BB12C8C36B5618EBABDAB3B2E7" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79A</figureCitation>
|
|
). Pilosity moderately dense, short, suberect to erect. Head foveolate; genae smooth. Clypeus foveolate; median longitudinal carina absent; absence of two lateral longitudinal carinae. Scape, when laid back, reaching the posterior head margin; pilosity erect (Fig.
|
|
<figureCitation id="844625782239F5D0AEC984577667C52F" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79A, C</figureCitation>
|
|
).
|
|
<emphasis id="89C171217267C18E43E4C54CAF182D6A" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="4A09E4A422C31FC90059519C769E91AF" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
|
.
|
|
</emphasis>
|
|
In lateral view, promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; promesonotal groove absent; metanotal groove indistinct; propodeal spines minute, triangular, with acute apex (Fig.
|
|
<figureCitation id="8B32E85ABAB36FB74D9F5AF2D8ABDCDD" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C</figureCitation>
|
|
). Sculpture foveolate; lateral sides of pronotum and propodeum with fading sculpture; katepisternum smooth. Pilosity moderately sparse, moderately long, and erect (Fig.
|
|
<figureCitation id="C89C4CB61CEAD199999C227971C60BF4" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
<emphasis id="0FD59D3A6CAC7FD30B50E352E6F4DB29" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="F6AE350A86114D9FD1371B9FAF5B37C5" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Peduncle very short and thin; with few short, erect setae (Fig.
|
|
<figureCitation id="929910E7ECB8280A209B0074A7492527" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
<emphasis id="89A9655C5E839349F144683A9279748F" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="4DBF335C8A43A389BD379E3AE990487C" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Short, low and convex; with moderately short, erect setae (Fig.
|
|
<figureCitation id="52B870B0C46EDDC81077617B0D0ADDC6" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
<emphasis id="98AEDCE4F64AC26580FCC81E3E2C7DC5" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="339C469E6E338060BB4ADFD18CA7CE20" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
With sparse, erect pilosity (Fig.
|
|
<figureCitation id="91FED884A9C37593AC69AB273E05EC1F" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
<emphasis id="2436001EAD2BAD4576218AD01210E29F" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="AECEA460FCED83EEC6B2390E2DBA12F8" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Yellow (Fig.
|
|
<figureCitation id="01E6ACC1D9C5F40D617F3CD864521A75" captionStart="Figure 79" captionStartId="F79" captionText="Figure 79. Pheidole fisaka sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923180) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0274930)." figureDoi="10.3897/zookeys.905.39592.figure79" httpUri="https://binary.pensoft.net/fig/373126" pageId="0" pageNumber="1">79C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="085F4D0EF082D559225221970CF4973D" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="219F25614A0984BF8939CF63908CF0B7" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="80EC08E1E9D100EDE921172362DEF0D8" pageId="0" pageNumber="1">Malagasy for flat, in reference for shape of the head in major workers.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="DDC9294433868465A7ABE512275A8CE5" pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph id="18CF7825E2B60B4298EA936E58C2B585" pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph id="98E54AB5199A004465F58666E8E0674A" pageId="0" pageNumber="1">The species was collected between 725-865 m in elevation, in rainforest. Nests were located in rotten logs.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0CE2AF51CE7981C5198E9187F3C97DF5" pageId="0" pageNumber="1" type="comments">
|
|
<paragraph id="EDEE700BA9264C9C623DA634D07FFE28" pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph id="972B702C3496602AD5A3F1FB7EDFA24A" pageId="0" pageNumber="1">
|
|
<taxonomicName id="6AC436E4425D6CEC7E987E3375B94AEA" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka">
|
|
<emphasis id="0788EE2F22A3DC0184EDE2C4B12C78C4" italics="true" pageId="0" pageNumber="1">Pheidole fisaka</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is most similar to
|
|
<taxonomicName id="2633ED04398935E890A3227F5BB5B6FB" lsidName="P. binara" pageId="0" pageNumber="1" rank="species" species="binara">
|
|
<emphasis id="23A9AA69607C25FB034A9C6C45D77E1B" italics="true" pageId="0" pageNumber="1">P. binara</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis id="D619DC280877D4C9EBFD0697C30AFF28" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="A8ACBE3AD4405149975008ACDE54E9AF" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName id="07C02687A671F18F254FD44CA154224C" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka">
|
|
<emphasis id="96448D30B15CF457AB3374FAF87BBFCD" italics="true" pageId="0" pageNumber="1">Pheidole fisaka</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName id="34F860D751197228FDF4EFC997BCCD5A" lsidName="P. binara" pageId="0" pageNumber="1" rank="species" species="binara">
|
|
<emphasis id="1EC3987D9BDB50B60340747A886DFE1C" italics="true" pageId="0" pageNumber="1">P. binara</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in smooth surface between rugae on frons and katepisternum and fading foveolae on propodeal dorsum.
|
|
<emphasis id="3CA1F01E18497DE5B01DD15104ADFC71" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="F642F0AEDE5718C8196BA45D33297A7D" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName id="8E09D3D197E2BE613E1FA6BDDA18365C" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fisaka" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fisaka">
|
|
<emphasis id="634CF5D764E5E9E89D2F8201F7BBF1CD" italics="true" pageId="0" pageNumber="1">Pheidole fisaka</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName id="E22E053952F038C772B166C51BBB6C12" lsidName="P. binara" pageId="0" pageNumber="1" rank="species" species="binara">
|
|
<emphasis id="6576C72B4647ACCC69E8F49740EB1648" italics="true" pageId="0" pageNumber="1">P. binara</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in shorter and more convex promesonotum, and lateral sides of pronotum and propodeum with fading foveolae and smooth katepisternum.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |