212 lines
17 KiB
XML
212 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1458" ID-GBIF-Dataset="c8fded56-3ddb-4e26-8863-ba8d55862689" ID-PMC="PMC3260749" ID-Pensoft-Pub="1313-2970-130-57" ID-PubMed="22259266" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-57" ModsDocOrigin="ZooKeys 130" ModsDocTitle="Rasnitsynala sigambrorum gen. et sp. n., a small odonatopterid (“Eomeganisoptera”, “Erasipteridae”) from the early Late Carboniferous of Hagen-Vorhalle (Germany)" checkinTime="1451249895056" checkinUser="pensoft" docAuthor="Zessin, Wolfgang, Brauckmann, Carsten & Groening, Elke" docDate="2011" docId="3D3250E4394E3D8A325E8DF9A233F104" docLanguage="en" docName="ZooKeys 130: 57-66" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1458" docTitle="Rasnitsynala sigambrorum Zessin, Brauckmann & Groening, 2011, sp. n." docType="treatment" docVersion="4" lastPageNumber="60" masterDocId="EC02FF9D2D103463FF94FFB86874FFA2" masterDocTitle="Rasnitsynala sigambrorum gen. et sp. n., a small odonatopterid (" Eomeganisoptera ", " Erasipteridae ") from the early Late Carboniferous of Hagen-Vorhalle (Germany)" masterLastPageNumber="66" masterPageNumber="57" pageNumber="59" updateTime="1668152291818" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Rasnitsynala sigambrorum gen. et sp. n., a small odonatopterid (" Eomeganisoptera ", " Erasipteridae ") from the early Late Carboniferous of Hagen-Vorhalle (Germany)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Zessin, Wolfgang</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Brauckmann, Carsten</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Groening, Elke</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2011</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>130</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>57</mods:start>
|
||
<mods:end>66</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.130.1458</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.130.1458</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-130-57</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="152031064" LSID="urn:lsid:zoobank.org:act:2DA218E6-D010-45C2-9763-5C78DC7F0758" httpUri="http://treatment.plazi.org/id/3D3250E4394E3D8A325E8DF9A233F104" lastPageId="4" lastPageNumber="60" pageId="2" pageNumber="59">
|
||
<subSubSection pageId="2" pageNumber="59" type="nomenclature">
|
||
<paragraph pageId="2" pageNumber="59">
|
||
<taxonomicName LSID="urn:lsid:zoobank.org:act:2DA218E6-D010-45C2-9763-5C78DC7F0758" class="Insecta" family="Erasipteridae" genus="Rasnitsynala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rasnitsynala sigambrorum" order="Geroptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="sigambrorum">Rasnitsynala sigambrorum</taxonomicName>
|
||
<taxonomicNameLabel pageId="2" pageNumber="59">sp. n.</taxonomicNameLabel>
|
||
Figs 2-3
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="59" type="nomenclature-citation">
|
||
<paragraph pageId="2" pageNumber="59">
|
||
<bibRefCitation author="Zessin, W" journalOrPublisher="Virgo, Mitteilungsblatt des Entomologischen Vereins Mecklenburg" pageId="4" pageNumber="61" pagination="5 - 32" title="Ueberblick ueber die palaeozoischen Libellen (Insecta, Odonatoptera)." volume="11" year="2008 a">Zessin 2008a</bibRefCitation>
|
||
: 5 - kleine, neue unbeschriebene Art von Vorhalle (= small, yet undescribed new species from Vorhalle)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="59" type="holotype">
|
||
<paragraph pageId="2" pageNumber="59">Holotype.</paragraph>
|
||
<paragraph pageId="2" pageNumber="59">
|
||
Former Sippel collection no. 182 A and B, now collection of the LWL-Museum
|
||
<normalizedToken originalValue="für">fuer</normalizedToken>
|
||
Naturkunde,
|
||
<normalizedToken originalValue="Westfälisches">Westfaelisches</normalizedToken>
|
||
Landesmuseum mit Planetarium in
|
||
<normalizedToken originalValue="Münster">Muenster</normalizedToken>
|
||
(Germany), positive imprint: no. WMf.N P27781, negative imprint: no. WMf.N N3182B.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="60" type="type locality">
|
||
<paragraph pageId="3" pageNumber="60">
|
||
<pageBreakToken pageId="3" pageNumber="60" start="start">Type</pageBreakToken>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph pageId="3" pageNumber="60">
|
||
Abandoned brickyard quarry in Hagen-Vorhalle, North Rhine-Westphalia, Germany (topographic map 1: 25,000 sheet no. 4610 Hagen/Westfalen;
|
||
<geoCoordinate direction="north" orientation="latitude" precision="9" value="51.381332">51°22.88'N</geoCoordinate>
|
||
;
|
||
<geoCoordinate direction="east" orientation="longitude" precision="9" value="7.4461665">007°26.77'E</geoCoordinate>
|
||
, ~115 m a.s.l.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="60" type="stratum typicum">
|
||
<paragraph pageId="3" pageNumber="60">Stratum typicum.</paragraph>
|
||
<paragraph pageId="3" pageNumber="60">Early Late Carboniferous (Early Pennsylvanian: late Namurian B, late Marsdenian), Ziegelschiefer Formation.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="60" type="diagnosis">
|
||
<paragraph pageId="3" pageNumber="60">Diagnosis.</paragraph>
|
||
<paragraph pageId="3" pageNumber="60">
|
||
Thus far the smallest species of the "
|
||
<taxonomicName family="Erasipteridae" lsidName="" pageId="3" pageNumber="60" rank="family">Erasipteridae</taxonomicName>
|
||
" with the following main characters: Mesothoracic wing: (i) Length: 27 mm; (ii) anterior margin very slightly convex; (iii)
|
||
<normalizedToken originalValue="ScP–">ScP-</normalizedToken>
|
||
very long, nearly reaching apex; (iv) subcostal area (Sc area) distally very narrow between
|
||
<normalizedToken originalValue="ScP–">ScP-</normalizedToken>
|
||
and
|
||
<normalizedToken originalValue="“PC/CA+/CP–/ScA+”">"PC/CA+/CP-/ScA+"</normalizedToken>
|
||
; (v) AA1+ short, forming an angle of about 45° with Cu and CuA+; (vi) basal parts of CuA+ and
|
||
<normalizedToken originalValue="CuP–">CuP-</normalizedToken>
|
||
rather long; (vii) basal intercubital area (ICu area) a relatively high triangle with its point downwards; (viii) kink between CuA+ and
|
||
<normalizedToken originalValue="CuP–">CuP-</normalizedToken>
|
||
very short. Metathoracic wing: (i) Length: 22 mm; (ii) anterior margin very slightly convex. Body: (i) Preserved length (without head and posterior 3 abdominal segments): 24.5 mm, estimated total length: 32 mm; (ii) abdominal segments narrow and long.
|
||
</paragraph>
|
||
<caption pageId="3" pageNumber="60">
|
||
<paragraph pageId="3" pageNumber="60">
|
||
Figure 2.
|
||
<taxonomicName class="Insecta" family="Erasipteridae" genus="Rasnitsynala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rasnitsynala sigambrorum" order="Geroptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="sigambrorum">Rasnitsynala sigambrorum</taxonomicName>
|
||
gen. et sp. n., holotype, former Sippel collection no. 182 A, now collection of the LWL-Museum
|
||
<normalizedToken originalValue="für">fuer</normalizedToken>
|
||
Naturkunde,
|
||
<normalizedToken originalValue="Westfälisches">Westfaelisches</normalizedToken>
|
||
Landesmuseum mit Planetarium in
|
||
<normalizedToken originalValue="Münster">Muenster</normalizedToken>
|
||
(Germany), positive imprint WMf.N P27781, nearly complete specimen; Early Late Carboniferous (Early Pennsylvanian: Namurian B, Marsdenian); abandoned brickyard quarry, Hagen-Vorhalle (Germany). Preserved length (= without head and posterior three segments): 24,5 mm, length of mesothoracic wing: 27 mm. Photograph by LWL- Museum
|
||
<normalizedToken originalValue="für">fuer</normalizedToken>
|
||
Naturkunde, Berenika Oblonczyk.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="3" pageNumber="60">
|
||
<paragraph pageId="3" pageNumber="60">
|
||
Figure 3. Drawing of
|
||
<taxonomicName class="Insecta" family="Erasipteridae" genus="Rasnitsynala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rasnitsynala sigambrorum" order="Geroptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="sigambrorum">Rasnitsynala sigambrorum</taxonomicName>
|
||
gen. et sp. n., holotype, former Sippel collection no. 182 A, now collection of the LWL-Museum
|
||
<normalizedToken originalValue="für">fuer</normalizedToken>
|
||
Naturkunde,
|
||
<normalizedToken originalValue="Westfälisches">Westfaelisches</normalizedToken>
|
||
Landesmuseum mit Planetarium in
|
||
<normalizedToken originalValue="Münster">Muenster</normalizedToken>
|
||
(Germany), positive imprint WMf.N P27781, nearly complete specimen; Early Late Carboniferous (Early Pennsylvanian: Namurian B, Marsdenian); abandoned brickyard quarry, Hagen-Vorhalle (Germany). Body structure schematized. Preserved length (= without head and posterior three segments): 24.5 mm, length of mesothoracic wing: 27 mm. Drawing by Wolfgang Zessin, Jasnitz (Germany).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="60" type="preservation">
|
||
<paragraph pageId="3" pageNumber="60">Preservation.</paragraph>
|
||
<paragraph pageId="3" pageNumber="60">
|
||
The rather well preserved fossil lies in dorsal view. The head with antennae, legs, the posterior three abdominal segments and the left wings are lacking as well as the posterior regions of the right wings. Nevertheless, the preserved parts of the right wings clearly exhibit the main diagnostic features. The original corrugation is nearly completely flattened by tectonic processes but can be reconstructed by comparison with the characters of the ground plan (
|
||
<bibRefCitation author="Kukalova-Peck, J" journalOrPublisher="Palaeodiversity" pageId="4" pageNumber="61" pagination="169 - 198" title="Carboniferous protodonatoid dragonfly nymphs and the synapomorphies of Odonatoptera and Ephemeroptera (Insecta: Palaeoptera)." volume="2" year="2009">
|
||
<normalizedToken originalValue="Kukalová-Peck">Kukalova-Peck</normalizedToken>
|
||
2009
|
||
</bibRefCitation>
|
||
). The body is compressed, too, and a part of the thoracic segments seems to have been split off. Possibly the right mesothoracic wing was basally disconnected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="60" type="measurements (in mm)">
|
||
<paragraph pageId="3" pageNumber="60">Measurements (in mm).</paragraph>
|
||
<paragraph pageId="3" pageNumber="60">(i) Length of mesothoracic wing, 27; (ii) length of metathoracic wing, 22; (iii) preserved length of body (without posterior three abdominal segments), 24.5, estimated total length, 32.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="60" type="description">
|
||
<paragraph pageId="3" pageNumber="60">Description.</paragraph>
|
||
<paragraph pageId="3" pageNumber="60">
|
||
Mesothoracic wing: Anterior margin very slightly convex;
|
||
<normalizedToken originalValue="ScP–">ScP-</normalizedToken>
|
||
very long, nearly reaching apex; subcostal area (Sc area) distally very narrow between
|
||
<normalizedToken originalValue="ScP–">ScP-</normalizedToken>
|
||
and complex PC/CA+/
|
||
<normalizedToken originalValue="CP–">CP-</normalizedToken>
|
||
/ScA+; AA1+ short, forming an angle of about 45° with Cu and CuA+; basal parts of CuA+ and
|
||
<normalizedToken originalValue="CuP–">CuP-</normalizedToken>
|
||
rather long; basal intercubital area (ICu area) resembling a relatively high triangle with its point downwards; kink between CuA+ and
|
||
<normalizedToken originalValue="CuP–">CuP-</normalizedToken>
|
||
very short. Metathoracic wing: Anterior margin very slightly convex, nearly straight. Body: Abdominal segments slender and long.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="60" type="etymology">
|
||
<paragraph pageId="3" pageNumber="60">Etymology.</paragraph>
|
||
<paragraph pageId="3" pageNumber="60">After the Latin name of the Germanic tribe Sigambri (or Sugambri) who inhabited the type region of the species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60" type="discussion">
|
||
<paragraph pageId="3" pageNumber="60">Discussion.</paragraph>
|
||
<paragraph lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
|
||
With a wing-span of about 55mm,
|
||
<taxonomicName class="Insecta" family="Erasipteridae" genus="Rasnitsynala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rasnitsynala sigambrorum" order="Geroptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="sigambrorum">Rasnitsynala sigambrorum</taxonomicName>
|
||
gen. et sp. n. represents the smallest species known to date of the presumably paraphyletic "
|
||
<taxonomicName family="Erasipteridae" lsidName="" pageId="3" pageNumber="60" rank="family">Erasipteridae</taxonomicName>
|
||
". It shows some characters already known from the very plesiomorphic
|
||
<taxonomicName genus="Geroptera" lsidName="Geroptera" pageId="3" pageNumber="60" rank="genus">Geroptera</taxonomicName>
|
||
:
|
||
<taxonomicName family="Eugeropteridae" lsidName="" pageId="3" pageNumber="60" rank="family">Eugeropteridae</taxonomicName>
|
||
(metathoracic wing of
|
||
<taxonomicName class="Insecta" family="Eugeropteridae" genus="Eugeropteron" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eugeropteron lunatum" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="lunatum">Eugeropteron lunatum</taxonomicName>
|
||
Riek in Riek &
|
||
<normalizedToken originalValue="Kukalová-Peck">Kukalova-Peck</normalizedToken>
|
||
, 1984) or
|
||
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Meganisoptera" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Meganisoptera" order="Malvales" pageId="3" pageNumber="60" phylum="Tracheophyta" rank="genus">Meganisoptera</taxonomicName>
|
||
:
|
||
<taxonomicName family="Aulertupidae" lsidName="" pageId="3" pageNumber="60" rank="family">Aulertupidae</taxonomicName>
|
||
(
|
||
<taxonomicName class="Insecta" family="Aulertupidae" genus="Aulertupus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aulertupus tembrocki" order="Meganisoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="tembrocki">Aulertupus tembrocki</taxonomicName>
|
||
), e.g. (i) the morphology of the intercubital area (ICu area) and (ii) the course of AA1+. The abdominal morphology is unknown in most Palaeozoic
|
||
<taxonomicName genus="Odonatoptera" lsidName="Odonatoptera" pageId="3" pageNumber="60" rank="genus">Odonatoptera</taxonomicName>
|
||
and cannot be
|
||
<pageBreakToken pageId="4" pageNumber="61" start="start">compered</pageBreakToken>
|
||
. In
|
||
<taxonomicName class="Insecta" family="Erasipteridae" genus="Rasnitsynala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rasnitsynala sigambrorum" order="Geroptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="sigambrorum">Rasnitsynala sigambrorum</taxonomicName>
|
||
gen. et sp. n. it is extremely slender and longer than the wings. It is even relatively longer and narrower than in the contemporaneous
|
||
<taxonomicName class="Insecta" genus="Namurotypus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Namurotypus sippeli" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="sippeli">Namurotypus sippeli</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" genus="Erasipteroides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Erasipteroides valentini" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="valentini">Erasipteroides valentini</taxonomicName>
|
||
but seems to be similar in
|
||
<taxonomicName class="Insecta" genus="Zessinella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zessinella siope" order="Megasecoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="siope">Zessinella siope</taxonomicName>
|
||
, again of the same age and locality. This character is common in several Recent
|
||
<taxonomicName genus="Zygoptera" higherTaxonomySource="GBIF" kingdom="Protozoa" lsidName="Zygoptera" pageId="4" pageNumber="61" rank="genus">Zygoptera</taxonomicName>
|
||
. It demonstrates that the basic ground plan of
|
||
<taxonomicName genus="Odonatoptera" lsidName="Odonatoptera" pageId="4" pageNumber="61" rank="genus">Odonatoptera</taxonomicName>
|
||
was already established in the very early phase of their evolution, presumably long before the Namurian B (about 319 m.y. ago).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |