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<document id="AC783B7502054548D017A4FABE99311E" ID-DOI="10.5281/zenodo.10667642" ID-ISSN="1407-8619" ID-Zenodo-Dep="10667642" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1708024310785" checkinUser="felipe" docAuthor="Vitali, Francesco" docDate="2018" docId="304F87FF0719A34CCE96F9CB38EDFCEF" docLanguage="en" docName="BalticJColeopterol.18.1.33-38.pdf" docOrigin="Baltic Journal of Coleopterology 18 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.1:BalticJColeopterol.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeopterol.2012-2021.journal_article" docStyleVersion="1" docTitle="Procleomenes gouverneuri Vitali 2018, sp. n." docType="treatment" docVersion="1" lastPageNumber="37" masterDocId="CC76FF870718A348CE75FF893903FFB8" masterDocTitle="Procleomenes gouverneuri sp. n. (Coleoptera: Cerambycidae) from Baltic amber: the first fossil member of the tribe Sestyrini Lacordaire, 1869" masterLastPageNumber="38" masterPageNumber="33" pageNumber="34" updateTime="1711545112766" updateUser="valdenar" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="49EC4DBC5A0E1C45BD68F28E9E73EF02">Procleomenes gouverneuri sp. n. (Coleoptera: Cerambycidae) from Baltic amber: the first fossil member of the tribe Sestyrini Lacordaire, 1869</mods:title>
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<mods:namePart id="D8BE291B03E27AC3F6C2FB6CF62EA26F">Vitali, Francesco</mods:namePart>
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<mods:title id="BF30F6114E796AD8538805E49224B319">Baltic Journal of Coleopterology</mods:title>
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<mods:part id="49FFA3F3B61087FE30C92DC5AF22C5D0">
<mods:date id="77611A314CD614B77D9359455F57AC9C">2018</mods:date>
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<mods:number id="1502B485194AD315C539D9DAD20E02EB">2018-09-10</mods:number>
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<subSubSection id="F0FC65620719A349CE96F9CB3B5FF9E2" box="[227,604,1602,1626]" pageId="1" pageNumber="34" type="nomenclature">
<paragraph id="B85936E90719A349CE96F9CB3B5FF9E2" blockId="1.[227,692,1506,1658]" box="[227,604,1602,1626]" pageId="1" pageNumber="34">
<heading id="E31181850719A349CE96F9CB3B5FF9E2" bold="true" box="[227,604,1602,1626]" fontSize="10" level="2" pageId="1" pageNumber="34" reason="4">
<emphasis id="8A92EAFB0719A349CE96F9CB3B5FF9E2" bold="true" box="[227,604,1602,1626]" pageId="1" pageNumber="34">
<taxonomicName id="7FE64D6A0719A349CE96F9CB3B1BF9E2" authority="Vitali, 2018" authorityName="Vitali" authorityYear="2018" box="[227,536,1602,1626]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="gouverneuri" status="sp. nov.">
<emphasis id="8A92EAFB0719A349CE96F9CB3B1BF9E2" bold="true" box="[227,536,1602,1626]" italics="true" pageId="1" pageNumber="34">Procleomenes gouverneuri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157800719A349CC6AF9CB3B5FF9E2" box="[543,604,1602,1626]" pageId="1" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="F0FC65620719A349CE96F9EB385AF9C2" box="[227,345,1634,1658]" pageId="1" pageNumber="34" type="description">
<paragraph id="B85936E90719A349CE96F9EB385AF9C2" blockId="1.[227,692,1506,1658]" box="[227,345,1634,1658]" pageId="1" pageNumber="34">
(
<figureCitation id="20DD2A6C0719A349CE9EF9EB3853F9C2" box="[235,336,1634,1658]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[303,343,895,919]" captionStartId-1="2.[303,343,1688,1712]" captionStartId-2="3.[227,271,1692,1716]" captionStartId-3="3.[737,778,1420,1444]" captionTargetBox-0="[300,1361,182,855]" captionTargetBox-1="[300,1361,981,1647]" captionTargetBox-2="[227,680,904,1655]" captionTargetBox-3="[718,1319,196,1380]" captionTargetId-0="figure-91@2.[303,1355,515,520]" captionTargetId-1="figure-241@2.[303,1361,1310,1315]" captionTargetId-2="figure-222@3.[227,610,1278,1284]" captionTargetId-3="figure-541@3.[710,1319,786,794]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-3="3" captionText-0="Fig. 1. Procleomenes gouverneuri sp. n., dorsal view" captionText-1="Fig. 2. Procleomenes gouverneuri sp. n., ventral view" captionText-2="Fig. 3. Procleomenes gouverneuri sp. n., reconstruction" captionText-3="Fig. 4. Procleomenes gouverneuri sp. n., colour reconstruction" pageId="1" pageNumber="34">Figs. 1-4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F0FC65620719A349CE96F92B3A91FEEF" pageId="1" pageNumber="34" type="materials_examined">
<paragraph id="B85936E90719A349CE96F92B3A91FEEF" blockId="1.[227,736,1698,1754]" lastBlockId="1.[774,1284,223,343]" pageId="1" pageNumber="34">
<emphasis id="8A92EAFB0719A349CE96F92B3857F902" bold="true" box="[227,340,1698,1722]" pageId="1" pageNumber="34">
<typeStatus id="675D884B0719A349CE96F92B384CF902" box="[227,335,1698,1722]" pageId="1" pageNumber="34" type="holotype">Holotype</typeStatus>
.
</emphasis>
Baltic amber, ex coll. J. Damzen, authors coll. FS71BS40. The death position with opened wings suggests that the specimen was attracted by the flowing resin and drowned, swimming inside for a certain time.
</paragraph>
</subSubSection>
<subSubSection id="F0FC65620719A349CD73FEF63C06FCCF" pageId="1" pageNumber="34" type="diagnosis">
<paragraph id="B85936E90719A349CD73FEF63D0FFE2F" blockId="1.[774,1285,383,888]" box="[774,1036,383,407]" pageId="1" pageNumber="34">
<emphasis id="8A92EAFB0719A349CD73FEF63D0FFE2F" bold="true" box="[774,1036,383,407]" pageId="1" pageNumber="34">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="B85936E90719A349CD73FE163C06FCCF" blockId="1.[774,1285,383,888]" pageId="1" pageNumber="34">
<taxonomicName id="7FE64D6A0719A349CD73FE163D2DFE0F" authority="Vitali, 2018" authorityName="Vitali" authorityYear="2018" box="[774,1070,415,439]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="gouverneuri" status="sp. nov.">
<emphasis id="8A92EAFB0719A349CD73FE163D2DFE0F" box="[774,1070,415,439]" italics="true" pageId="1" pageNumber="34">Procleomenes gouverneuri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157800719A349CA4DFE293D75FE0F" box="[1080,1142,416,439]" pageId="1" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
differs from all known extant species in the elytra long and with a peculiar pattern. Concerning the pattern, a scutellar spot can be found only in the Bornean
<taxonomicName id="7FE64D6A0719A349CD73FDA93D39FD8F" authority="Niisato, 1986" authorityName="Niisato" authorityYear="1986" box="[774,1082,543,567]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="tenuiformis">
<emphasis id="8A92EAFB0719A349CD73FDA93A9CFD8F" box="[774,927,543,567]" italics="true" pageId="1" pageNumber="34">P. tenuiformis</emphasis>
Niisato, 1986
</taxonomicName>
and
<taxonomicName id="7FE64D6A0719A349CA01FDA93A9DFDEF" authority="Niisato, 1986" authorityName="Niisato" authorityYear="1986" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="8A92EAFB0719A349CA01FDA93C06FD8F" box="[1140,1285,543,567]" italics="true" pageId="1" pageNumber="34">P. longicollis</emphasis>
Niisato, 1986
</taxonomicName>
, but none of these species show a yellow elytral apex (Niisato, 1986). Moreover, all species with three elytral bands (basal, premedian and postmedian) always show a premedian yellow band and a black apex (both absent in
<taxonomicName id="7FE64D6A0719A349CD0FFD563DABFD4F" authority="Vitali, 2018" authorityName="Vitali" authorityYear="2018" box="[890,1192,735,759]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="gouverneuri" status="sp. nov.">
<emphasis id="8A92EAFB0719A349CD0FFD563DABFD4F" box="[890,1192,735,759]" italics="true" pageId="1" pageNumber="34">Procleomenes gouverneuri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A157800719A349CAC6FD693DF6FD4F" box="[1203,1269,735,759]" pageId="1" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
). The elytra cover the whole abdomen, whereas they leave uncovered one or two urites in modern species. However, the variability of such characters suggests to not introduce new names.
</paragraph>
</subSubSection>
<subSubSection id="F0FC65620719A34CCD73FC163BFDFD2F" lastPageId="4" lastPageNumber="37" pageId="1" pageNumber="34" type="description">
<paragraph id="B85936E90719A349CD73FC163A96FC0F" blockId="1.[774,1285,927,1239]" box="[774,917,927,951]" pageId="1" pageNumber="34">
<emphasis id="8A92EAFB0719A349CD73FC163A96FC0F" bold="true" box="[774,917,927,951]" pageId="1" pageNumber="34">Description</emphasis>
</paragraph>
<paragraph id="B85936E90719A349CD73FC363D39FB6F" blockId="1.[774,1285,927,1239]" pageId="1" pageNumber="34">
Female (for general habitus and relatively short antennae), body length ~
<quantity id="7F1E9B0C0719A349CA7AFC563D4CFC4F" box="[1039,1103,991,1015]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="34" unit="mm" value="8.0">8 mm</quantity>
. Body elongated, black; antennae yellow with scape and apex of each segment black; legs black with femoral and tibial base more or less yellow; elytra black with a yellow pattern disposed as follows: a cordiform large scutellar spot, a pre-apical spot not reaching both marginal and sutural margin, and a transverse apical band.
</paragraph>
<paragraph id="B85936E90719A349CD73FB763C06F96F" blockId="1.[774,1285,1279,1751]" pageId="1" pageNumber="34">Head large; forehead oblique, short; antennal tubercles close and not elevated; inter-antennal furrow reaching the base of the neck; eyes large, finely facetted, strongly reniform, very prominent laterally (as large as pronotum including lateral teeth), separated above by about one-third of the greatest width of head; neck with some raised setae. Palpi small; last maxillar and labial palpomere securiform, obliquely truncate at apex. Antennae elevensegmented, barely shorter than body; antennomeres IV-X toothed externally, with some apical setae; antennomeres II-IV with median setae as well; scape weekly clavate, straight; pedicle elongate, one-fourth as long as</paragraph>
<caption id="EC996661071AA34ACF5AFCF63A7AFC2F" box="[303,889,895,919]" pageId="2" pageNumber="35" startId="2.[303,343,895,919]" targetBox="[300,1361,182,855]" targetPageId="2" targetType="figure">
<paragraph id="B85936E9071AA34ACF5AFCF63A7AFC2F" blockId="2.[303,889,895,919]" box="[303,889,895,919]" pageId="2" pageNumber="35">
Fig. 1.
<taxonomicName id="7FE64D6A071AA34ACF0AFCF63BA6FC2F" authority="Vitali, 2018" authorityName="Vitali" authorityYear="2018" box="[383,677,895,919]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="gouverneuri" status="sp. nov.">
<emphasis id="8A92EAFB071AA34ACF0AFCF63BA6FC2F" box="[383,677,895,919]" italics="true" pageId="2" pageNumber="35">Procleomenes gouverneuri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A15780071AA34ACCD8FC093BEFFC2F" box="[685,748,896,919]" pageId="2" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
, dorsal view
</paragraph>
</caption>
<caption id="EC996661071AA34ACF5AF9113A7CF908" box="[303,895,1688,1713]" pageId="2" pageNumber="35" startId="2.[303,343,1688,1712]" targetBox="[300,1361,981,1647]" targetPageId="2" targetType="figure">
<paragraph id="B85936E9071AA34ACF5AF9113A7CF908" blockId="2.[303,895,1688,1713]" box="[303,895,1688,1713]" pageId="2" pageNumber="35">
Fig. 2.
<taxonomicName id="7FE64D6A071AA34ACF0AF9113BA7F909" authority="Vitali, 2018" authorityName="Vitali" authorityYear="2018" box="[383,676,1688,1713]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="gouverneuri" status="sp. nov.">
<emphasis id="8A92EAFB071AA34ACF0AF9113BA7F909" box="[383,676,1688,1713]" italics="true" pageId="2" pageNumber="35">Procleomenes gouverneuri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A15780071AA34ACCD9F9103BE9F908" box="[684,746,1689,1712]" pageId="2" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
, ventral view
</paragraph>
</caption>
<paragraph id="B85936E9071BA34BCCA4FF183A1BFF1C" blockId="3.[721,792,145,164]" box="[721,792,145,164]" pageId="3" pageNumber="36">
<emphasis id="8A92EAFB071BA34BCCA4FF183A1BFF1C" box="[721,792,145,164]" italics="true" pageId="3" pageNumber="36">Vitali F.</emphasis>
</paragraph>
<paragraph id="B85936E9071BA34BCE96FF4938ADFE0F" blockId="3.[227,689,191,439]" pageId="3" pageNumber="36">
scape; antennomere III less than two-thirds as long as scape; antennomere
<collectionCode id="DEF7AE2C071BA34BCC4EFF693B5BFF4F" box="[571,600,224,247]" pageId="3" pageNumber="36">IV</collectionCode>
,
<collectionCode id="DEF7AE2C071BA34BCC16FF693B7CFF4F" box="[611,639,224,247]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="36">VI</collectionCode>
and
<collectionCode id="DEF7AE2C071BA34BCE96FE89380AFEAF" box="[227,265,256,279]" pageId="3" pageNumber="36">VII</collectionCode>
equal, about one-third as long as III; antennomere
<collectionCode id="DEF7AE2C071BA34BCFE9FEA938F2FE8F" box="[412,497,288,311]" pageId="3" pageNumber="36">VIII-X</collectionCode>
progressively shortened; antennomere XI as long as IX (antennomere proportions according to the formula: 1.6: 0.4: 1.0: 1.3: 1.4: 1.3: 1.3: 1.2: 1.1: 0.9: 1.1).
</paragraph>
<paragraph id="B85936E9071BA34BCE96FE56388BFCEF" blockId="3.[227,689,479,855]" pageId="3" pageNumber="36">Prothorax cylindrical, elongate, 2.3 times as long as wide, slightly larger at apex than at base; apex with a large apical collar, then slightly enlarged; sides armed with a large blunt conical tubercle at the middle and covered by some raised setae on the apical half; base one-half as wide as elytral base, biarcuate and acutely produced posteriorly in the middle, flattened along the posterior margin; disc covered with extremely fine dense punctures. Scutellum minute, subrectangular.</paragraph>
<caption id="EC996661071BA34BCC94FA053A81FA7C" pageId="3" pageNumber="36" startId="3.[737,778,1420,1444]" targetBox="[718,1319,196,1380]" targetPageId="3" targetType="figure">
<paragraph id="B85936E9071BA34BCC94FA053A81FA7C" blockId="3.[737,1286,1420,1476]" pageId="3" pageNumber="36">
Fig. 4.
<taxonomicName id="7FE64D6A071BA34BCD4DFA053D67FA1C" authority="Vitali, 2018" authorityName="Vitali" authorityYear="2018" box="[824,1124,1420,1444]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="gouverneuri" status="sp. nov.">
<emphasis id="8A92EAFB071BA34BCD4DFA053D67FA1C" box="[824,1124,1420,1444]" italics="true" pageId="3" pageNumber="36">Procleomenes gouverneuri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A15780071BA34BCA1AFA043DB2FA1C" box="[1135,1201,1421,1444]" pageId="3" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
, colour reconstruction
</paragraph>
</caption>
<paragraph id="B85936E9071BA34BCD79F9853AC0F97C" blockId="3.[780,1289,1548,1732]" pageId="3" pageNumber="36">Elytra elongate, slightly dehiscent posteriorly, 3.4 times as long as wide at humeri; base anteriorly concave; humeri squared; sides constricted in the middle; apex concavely truncated; disc flat, covered with extremely fine dense punctures.</paragraph>
<caption id="EC996661071BA34BCE96F9153887F96C" pageId="3" pageNumber="36" startId="3.[227,271,1692,1716]" targetBox="[227,680,904,1655]" targetPageId="3" targetType="figure">
<paragraph id="B85936E9071BA34BCE96F9153887F96C" blockId="3.[227,736,1692,1748]" pageId="3" pageNumber="36">
Fig. 3.
<taxonomicName id="7FE64D6A071BA34BCF34F9153B85F90C" authority="Vitali, 2018" authorityName="Vitali" authorityYear="2018" box="[321,646,1692,1716]" class="Insecta" family="Cerambycidae" genus="Procleomenes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="gouverneuri" status="sp. nov.">
<emphasis id="8A92EAFB071BA34BCF34F9153B85F90C" box="[321,646,1692,1716]" italics="true" pageId="3" pageNumber="36">Procleomenes gouverneuri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91A15780071BA34BCCE6F9143BD8F90C" box="[659,731,1693,1716]" pageId="3" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
, reconstruction
</paragraph>
</caption>
<paragraph id="B85936E9071CA34CCF5AFF36388CFE0F" blockId="4.[303,813,191,439]" pageId="4" pageNumber="37">Ventral side covered with pubescence long and raised on head and prosternum, long and recumbent on the metasterum and fine and recumbent on the abdomen; procoxae globose, procoxal cavities rounded, posteriorly closed; urosternite visible I-V progressively shortened; urosternite VIII (genital segment) posteriorly rounded.</paragraph>
<paragraph id="B85936E9071CA34CCF5AFE693BFDFD2F" blockId="4.[303,813,479,663]" pageId="4" pageNumber="37">Legs relatively long, femora pedunculate, covered with long setae; tibiae evidently shorter than femora, feebly curved, covered with some sub-erect short setae; tarsi short; metatarsi one-half as long as metatibiae; metotarsomere I as long as metatarsomeres II and III together.</paragraph>
</subSubSection>
<subSubSection id="F0FC6562071CA34CCF5AFD3638EDFCEF" pageId="4" pageNumber="37" type="etymology">
<paragraph id="B85936E9071CA34CCF5AFD3638B2FD6F" blockId="4.[303,813,703,855]" box="[303,433,703,727]" pageId="4" pageNumber="37">
<emphasis id="8A92EAFB071CA34CCF5AFD3638B2FD6F" bold="true" box="[303,433,703,727]" pageId="4" pageNumber="37">Etymology</emphasis>
</paragraph>
<paragraph id="B85936E9071CA34CCF5AFD5638EDFCEF" blockId="4.[303,813,703,855]" pageId="4" pageNumber="37">This new species is dedicated to Xavier Gouverneur, enthusiastic explorer and excellent entomologist of the South-eastern Asian Cerambycofauna.</paragraph>
</subSubSection>
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