133 lines
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133 lines
14 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.349.6164" ID-GBIF-Dataset="cd871e7e-7e75-4f2e-8c8b-7af9cb4803d4" ID-PMC="PMC3837407" ID-Pensoft-Pub="1313-2970-349-81" ID-PubMed="24294080" ID-ZBK="45895E0B79E74794A0155CD74FEDDF22" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-349-81" ModsDocOrigin="ZooKeys 349" ModsDocTitle="Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)" checkinTime="1451246687136" checkinUser="pensoft" docAuthor="Tian, Mingyi & Deuve, Thierry" docDate="2013" docId="09451E5D69D99D43A05E24662A7BD11A" docLanguage="en" docName="ZooKeys 349: 81-100" docOrigin="ZooKeys 349" docSource="http://dx.doi.org/10.3897/zookeys.349.6164" docTitle="Orthogonius carinatus Tian & Deuve, sp. n." docType="treatment" docUuid="55C5C2C1-598B-4F4C-98D9-50B26CA66935" docUuidSource="ZooBank" docVersion="5" lastPageNumber="93" masterDocId="B203D41F1D41C83E8603FFE5D1499035" masterDocTitle="Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)" masterLastPageNumber="100" masterPageNumber="81" pageNumber="89" updateTime="1668156835216" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tian, Mingyi</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Deuve, Thierry</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>349</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>81</mods:start>
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<mods:end>100</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.349.6164</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.349.6164</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-349-81</mods:identifier>
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<mods:identifier type="ZBK">45895E0B79E74794A0155CD74FEDDF22</mods:identifier>
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<mods:identifier type="ZooBank">45895E0B79E74794A0155CD74FEDDF22</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152049513" LSID="urn:lsid:zoobank.org:act:55C5C2C1-598B-4F4C-98D9-50B26CA66935" httpUri="http://treatment.plazi.org/id/09451E5D69D99D43A05E24662A7BD11A" lastPageId="12" lastPageNumber="93" pageId="8" pageNumber="89">
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<subSubSection pageId="8" pageNumber="89" type="nomenclature">
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<paragraph pageId="8" pageNumber="89">
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<taxonomicName LSID="http://zoobank.org/55C5C2C1-598B-4F4C-98D9-50B26CA66935" authority="Tian & Deuve" class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius carinatus" order="Coleoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="carinatus">Orthogonius carinatus Tian & Deuve</taxonomicName>
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<taxonomicNameLabel pageId="8" pageNumber="89">sp. n.</taxonomicNameLabel>
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Figs 1, 8-9, 18-19
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="89" type="diagnosis">
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<paragraph pageId="8" pageNumber="89">Diagnosis.</paragraph>
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<paragraph pageId="8" pageNumber="89">Medium to large sized, body elongate, whole dorsal surface extraordinarily and densely punctate; head as long as wide, eyes moderate, labrum roundly protruding at front; pronotum narrow, lateral expanded margins distinctly reflexed throughout; elytra carinate on basal 1/3 of interval 7 in both sexes, and at base of intervals 4-6 in female, but normal in male.</paragraph>
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<paragraph pageId="8" pageNumber="89">Length: 14.0-16 mm; width: 5.5-6.0 mm. Body slender and elongate. Habitus as in Figs 8-9.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="91" pageId="8" pageNumber="89" type="description">
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<paragraph pageId="8" pageNumber="89">Description.</paragraph>
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<paragraph pageId="8" pageNumber="89">Black, antennomeres 1, 3-4, legs (except tarsomeres) and ventral surface dark brown, mandibles, labrum, palps, other antennomeres and tarsomeres reddish brown.</paragraph>
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<paragraph pageId="8" pageNumber="89">Macrosculpture: Head extraordinarily and densely punctate, pronotum distinctly punctate in male, but faintly punctate or impunctate in female; elytra with tiny, sparse and faint punctures in both sexes; underside smooth and glabrous.</paragraph>
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<paragraph pageId="8" pageNumber="89">Microsculpture: Engraved meshes densely isodiametric or slightly transverse on elytra, faint or not clear on head and pronotum.</paragraph>
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<paragraph lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90">
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<pageBreakToken pageId="9" pageNumber="90" start="start">Head</pageBreakToken>
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stout, as long as wide,eyes moderate, prominent, frons and vertex moderately convex, frontal impressions not well defined, shallow; clypeus bisetose, base area rugose; labrum broadly protruding at front, sexsetose; mandibles rather short; ligula broad, bisetose at apex; maxillary palps moderate, subcylindrical, maxillary palpomere 3 slightly longer than 4, both glabrous except for palpomere 3 with two short setae at apex; labial palpomeres stout, palpomere 3 somewhat expanded, palpomere 2 cylindrical, as long as palpomere 3, bisetose on inner margin, and with two or three additional setae at subapex and apex; palpiger asetose, mentum and submentum each with one
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<pageBreakToken pageId="10" pageNumber="91" start="start">pair</pageBreakToken>
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of setae; mentum without median tooth. Antennae long and slender, extending beyond basal 1/3 of elytra; pubescent from basal 1/3 of antennomere 4, antennomere 3 slightly shorter than 4.
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</paragraph>
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<paragraph pageId="10" pageNumber="91">Pronotum transverse, PW/PL=1.55-1.60, disc moderately convex; sides evenly rounded, widest at middle, both basal and fore margins beaded, lateral expanded margins well defined, wide and obviously reflexed throughout; fore and hind angles rounded; both fore and hind transversal impressions faint, basal foveae large and wide, connected to expanded margins; median line clear.</paragraph>
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<paragraph pageId="10" pageNumber="91">Elytra elongate, EL/EW=1.72-1.78, moderately convex, base completely bordered; sides nearly parallel at middle, striae deep, intervals convex, subequal in width in middle; apex broadly truncate; carinate on bases of intervals 6-8, basal 1/3 of interval 7 in both sexes, and at base of intervals 4-6 in female (carination shortest on interval 4, and longest on 6); interval 3 with only the basal and middle setiferous pores, the subapical one absent.</paragraph>
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<paragraph pageId="10" pageNumber="91">Legs slender; middle and hind coxae glabrous in median portion; fore tibia slightly dilated at apex, outer angle obtuse, apex slightly and obliquely truncate, outer margin slightly subserrate; middle and hind tibiae hardly dilated at apex, slender and strongly curved in middle portion in male, normal in female; hind tarsomeres 1 and 3 much longer than 2 and 4 respectively, length ratio of tarsomeres 1-4 as 3.9: 2.6: 1.8: 1.0; tarsomere 4 deeply emarginate at apex, lobes asymmetric, longer lobe as long as 1/3 of the joint; all tarsal claws strongly pectinate.</paragraph>
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<paragraph pageId="10" pageNumber="91">Prosternal process faintly bordered at apex. Abdominal ventrite VII distinctly emarginate at apical margin in male.</paragraph>
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<paragraph pageId="10" pageNumber="91">
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Male genitalia (Figs 18-19). Rather slender, strongly sinuate ventrally, distinctly arcuate at apical 1/3, slightly dilated in middle portion, dorsal opening large, subapex nearly paralleled towards apex, broadly obtused at apex; in dorsal view, apical lamella almost as long as wide, not
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<normalizedToken originalValue="parallel–sided">parallel-sided</normalizedToken>
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, apex broadly rounded.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="91" type="remarks">
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<paragraph pageId="10" pageNumber="91">Remarks.</paragraph>
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<paragraph pageId="10" pageNumber="91">
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Similar to
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<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius lancangjiang" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="lancangjiang">Orthogonius lancangjiang</taxonomicName>
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, but intervals of elytra carinate; the median lobe of aedeagus distinctly arcuate and the apical lamella somewhat stouter. It is easily distinguished from the next species,
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<taxonomicName class="Insecta" family="Carabidae" genus="Orthogonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthogonius macrophthalmus" order="Coleoptera" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="macrophthalmus">Orthogonius macrophthalmus</taxonomicName>
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sp. n., by its small and less prominent eyes, narrower pronotum, and elytra carinate on base of intervals 4-6 in female.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="12" lastPageNumber="93" pageId="10" pageNumber="91" type="material examined">
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<paragraph pageId="10" pageNumber="91">Material examined.</paragraph>
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<paragraph pageId="10" pageNumber="91">
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Holotype: male, labeled:
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<normalizedToken originalValue="“Laos–NE">"Laos-NE</normalizedToken>
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: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.12 N/ 104.01 E, 1500-1900 m, 17.V.-3.VI. 2007, M. Brancucci Leg.", and "NHMB Basel Expeditionto Laos, 2007", in NHMB.
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</paragraph>
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<paragraph lastPageId="12" lastPageNumber="93" pageId="10" pageNumber="91">
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Paratypes. 1 male and 3 female, idem; 1 male 3 females,
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<normalizedToken originalValue="“Laos–NE">"Laos-NE</normalizedToken>
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: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.1309N/ 103.5954-104.0003E, 1480-1550 m, 9-16. VI. 2009, David Hauck Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 1 male and 5 females,
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<normalizedToken originalValue="“Laos–NE">"Laos-NE</normalizedToken>
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: Xieng Khouang Province, 30 km NE Phonsaven, Phou Sane Mt., 19.37-8N/ 103.20-21E, 1420 m, 30 km NE Phonsaven, Ban Na Lam to Phou Sane Mt., 1300-1700 m, 10.-30.V. 2009, M. Geiser Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 1 male and 2 females,
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<normalizedToken originalValue="“Laos–NE">"Laos-NE</normalizedToken>
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: Houa Phan Province, Ban Saluei to Phou
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<pageBreakToken pageId="11" pageNumber="92" start="start">Pane</pageBreakToken>
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Mt., 20.13N/ 104.00E, 1350-1500 m, 1-16. VI. 2009, M. Brancucci Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 3 females,
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<normalizedToken originalValue="“Laos–NE">"Laos-NE</normalizedToken>
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: Xieng Khouang Province, 30 km NE Phonsaven: Ban Na Lam to Phou Sane Mt., 19.37-38N/ 103.20E, 1300-1500 m, 10.-30.V. 2009, M. Brancucci Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 2 female,
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<normalizedToken originalValue="“Laos–NE">"Laos-NE</normalizedToken>
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: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.12-13N/ 103.595-104.01E, 1340-1870 m, 10. V.-16. VI. 2009, M. Brancucci and local coll. Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M.
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<pageBreakToken pageId="12" pageNumber="93" start="start">Geiser</pageBreakToken>
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, Z. Kraus, D. Hauck, V. Kuban"; 1 male 6 female,
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<normalizedToken originalValue="“Laos–NE">"Laos-NE</normalizedToken>
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: Xieng Khouang Province, Phonsaven (30 km NE), Phou Sane Mt., 19.37-38N/ 103.20E, 1400-1500 m, 10.-30.V. 2009, Z. Kraus Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 1 females,
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<normalizedToken originalValue="“Laos–N">"Laos-N</normalizedToken>
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: (Oudom Xai), 17 km, NEE of Oudom Xai, 20.45N/ 102.09E, 1-9. V. 2002, V. Kuban leg."; 7 females,
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<normalizedToken originalValue="“Laos–NE">"Laos-NE</normalizedToken>
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: Xieng Khouang Province, Phonsaven (30 km NE), Phou Sane Mt., 19.37-38N/ 103.20-21E, 1400-1700 m, 10.-30.V. 2009, D. Hauck Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 1 female,
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<normalizedToken originalValue="“Laos–NE">"Laos-NE</normalizedToken>
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: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.11-13N/ 103.59-104.01E, 1300-1900 m, 9-17.VI. 2009, Michael Geiser Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban", all in NHMB except two males and ten females in MNHN and SCAU, respectively. 4 female, "Lao, Phongsaly Prov., 21°41-2'N /102°6-8'E, 28. v.-20. vi. 2003, Phongsaly env., 1500 m,
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<normalizedToken originalValue="Pacholátko">Pacholatko</normalizedToken>
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leg.", and "Collection Naturhistorisches Museum Basel", all in NHMB except one in MNHN; 1 female, ibid, except
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<normalizedToken originalValue="“Pacholátko">"Pacholatko</normalizedToken>
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leg" replaced by "Brancucci leg.", in NHMB.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="93" type="etymology">
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<paragraph pageId="12" pageNumber="93">Etymology.</paragraph>
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<paragraph pageId="12" pageNumber="93">Referring to the character of elytral carination.</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="93" type="distribution">
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<paragraph pageId="12" pageNumber="93">Distribution.</paragraph>
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<paragraph pageId="12" pageNumber="93">Northern Laos (Fig. 1).</paragraph>
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</subSubSection>
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</treatment>
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</document> |