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<document id="08E4DC7881A7E6711383E2FA68AFA292" ID-DOI="10.5281/zenodo.294189" ID-GBIF-Dataset="b2775d25-e230-4f10-9430-f0d6dfd5a28e" ID-ISSN="1175-5326" ID-Zenodo-Dep="294189" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460245078205" checkinUser="plazi" docAuthor="O’Loughlin, Mark &amp; Whitfield, Emily" docDate="2010" docId="03F287FA1970BD75FF12FEA3FA2CFD29" docLanguage="en" docName="zt02528p068.pdf" docOrigin="Zootaxa 2528" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Psolus lockhartae O’Loughlin &amp; Whitfield, 2010, sp. nov." docType="treatment" docVersion="6" lastPageNumber="68" masterDocId="FFCBFF821975BD72FF85FF8EFFBFFFCB" masterDocTitle="New species of Psolus Oken from Antarctica (Echinodermata: Holothuroidea: Psolidae)" masterLastPageNumber="68" masterPageNumber="61" pageNumber="66" updateTime="1698241549117" updateUser="plazi">
<mods:mods id="46B51050AEE2CB8A7EF63658FF2328B6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="66D4D6C4126DD634AA7ACFD4DC7ABDA1">
<mods:title id="455E1B2A0CA066A84CED33C9E829E7F2">New species of Psolus Oken from Antarctica (Echinodermata: Holothuroidea: Psolidae)</mods:title>
</mods:titleInfo>
<mods:name id="53A366F33D11A938B7447A4B32708BA4" type="personal">
<mods:role id="7C6971006A924800AE2C55DB10DF2104">
<mods:roleTerm id="144051178E73CD9AC8BBC35533360127">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="332CF2CED106A175ED20219A333258BA">O’Loughlin, Mark</mods:namePart>
</mods:name>
<mods:name id="664BBB7C587D38B1E9970B9D28FFE81A" type="personal">
<mods:role id="21507703069BD0A4F50F33CF7D65D5F7">
<mods:roleTerm id="B8EB8787A4D5CA681849AC178A028018">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F876046289D389151D5D79C296FD63E3">Whitfield, Emily</mods:namePart>
</mods:name>
<mods:typeOfResource id="9685ECE276838687A8EB112A6D27EEC8">text</mods:typeOfResource>
<mods:relatedItem id="E532D3B244112CD58D6558FA026744C1" type="host">
<mods:titleInfo id="1C3E6FE9FF8DE4D73E3E214DBDE4D167">
<mods:title id="845D512B3B0AB46785E18F097051C66D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9C548C57AF7EB92D46C2D839644D8071">
<mods:date id="09B296A6C5AFC7291CEDD253D9B62975">2010</mods:date>
<mods:detail id="C84CAD3F3FEE74582AE13F91D660D674" type="volume">
<mods:number id="94CC17990985E510399C7CE2759ECBE0">2528</mods:number>
</mods:detail>
<mods:extent id="FFF34B4123F3BEF9992A2E15C7788B03" unit="page">
<mods:start id="F16977CBC579C2CDDC1ACA37DF91C88E">61</mods:start>
<mods:end id="A22F4A08FF8BAF329DA47658EB0272CD">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4010C5D6AF1F395AFF6E712203C37C48">journal article</mods:classification>
<mods:identifier id="7CADEE0D0BFB1A4A71356DFAFF5A35DA" type="DOI">10.5281/zenodo.294189</mods:identifier>
<mods:identifier id="43FE926CE76D69E5A1C8A29CA5ECA16C" type="GBIF-Dataset">b2775d25-e230-4f10-9430-f0d6dfd5a28e</mods:identifier>
<mods:identifier id="B75B03958E1FC3252E1BF66495E64CED" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="53719D594BE3F71EC062B6AC97DCF33D" type="Zenodo-Dep">294189</mods:identifier>
</mods:mods>
<treatment id="03F287FA1970BD75FF12FEA3FA2CFD29" ID-DOI="http://doi.org/10.5281/zenodo.6196621" ID-GBIF-Taxon="119390316" ID-Zenodo-Dep="6196621" LSID="urn:lsid:plazi:treatment:03F287FA1970BD75FF12FEA3FA2CFD29" httpUri="http://treatment.plazi.org/id/03F287FA1970BD75FF12FEA3FA2CFD29" lastPageId="7" lastPageNumber="68" pageId="5" pageNumber="66">
<subSubSection id="C34165671970BD77FF12FEA3FEF0FEA1" pageId="5" pageNumber="66" type="nomenclature">
<paragraph id="8BE436EC1970BD77FF12FEA3FE6CFE8C" blockId="5.[151,467,301,362]" box="[151,467,301,327]" pageId="5" pageNumber="66">
<heading id="D0AC81801970BD77FF12FEA3FE6CFE8C" bold="true" box="[151,467,301,327]" fontSize="11" level="1" pageId="5" pageNumber="66" reason="1">
<emphasis id="B92FEAFE1970BD77FF12FEA3FE6CFE8C" bold="true" box="[151,467,301,327]" pageId="5" pageNumber="66">
<taxonomicName id="4C5B4D6F1970BD77FF12FEA3FED1FE8C" ID-CoL="4PL9G" box="[151,366,301,327]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="5" pageNumber="66" phylum="Echinodermata" rank="species" species="lockhartae" status="sp. nov.">
<emphasis id="B92FEAFE1970BD77FF12FEA3FED1FE8C" bold="true" box="[151,366,301,327]" italics="true" pageId="5" pageNumber="66">Psolus lockhartae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57851970BD77FEF0FEA3FE6CFE8C" box="[373,467,301,327]" pageId="5" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE436EC1970BD77FF12FEDEFEF0FEA1" blockId="5.[151,467,301,362]" box="[151,335,336,362]" pageId="5" pageNumber="66">
<figureCitation id="13602A691970BD77FF12FEDEFEBAFEA1" box="[151,261,336,362]" captionStart="FIGURE 4" captionStartId="5.[151,255,1904,1928]" captionTargetBox="[156,1436,427,1874]" captionTargetId="figure@5.[151,1436,418,1882]" captionTargetPageId="5" captionText="FIGURE 4. Psolus lockhartae sp. nov. holotype (a – c, NMV F 168861): a, dorsal view; b, lateral view; c, ventral view; paratype (d – f, NMV F 168860): d, tentacle ossicles; e, ossicles from sole; f, ossicles from tube foot." httpUri="https://zenodo.org/record/294193/files/figure.png" pageId="5" pageNumber="66">Figures 4</figureCitation>
a–f, 5.
</paragraph>
</subSubSection>
<caption id="DF2466641970BD77FF12F8FEFB05F86C" httpUri="https://zenodo.org/record/294193/files/figure.png" pageId="5" pageNumber="66" targetBox="[156,1436,427,1874]" targetPageId="5">
<paragraph id="8BE436EC1970BD77FF12F8FEFB05F86C" blockId="5.[151,1436,1904,1959]" pageId="5" pageNumber="66">
<emphasis id="B92FEAFE1970BD77FF12F8FEFEA5F843" bold="true" box="[151,282,1904,1928]" pageId="5" pageNumber="66">FIGURE 4.</emphasis>
<taxonomicName id="4C5B4D6F1970BD77FEA7F8FEFE5CF84C" box="[290,483,1904,1927]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="5" pageNumber="66" phylum="Echinodermata" rank="species" species="lockhartae" status="sp. nov.">
<emphasis id="B92FEAFE1970BD77FEA7F8FEFE5CF84C" box="[290,483,1904,1927]" italics="true" pageId="5" pageNumber="66">Psolus lockhartae</emphasis>
</taxonomicName>
<emphasis id="B92FEAFE1970BD77FE6EF8FEFDFEF843" bold="true" box="[491,577,1904,1928]" pageId="5" pageNumber="66">
<taxonomicNameLabel id="A21C57851970BD77FE6EF8FEFDFEF843" box="[491,577,1904,1928]" pageId="5" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype (a–c, NMV F168861): a, dorsal view; b, lateral view; c, ventral view; paratype (d–f, NMV F168860): d, tentacle ossicles; e, ossicles from sole; f, ossicles from tube foot.
</paragraph>
</caption>
<caption id="DF2466641973BD74FF12FC53FC6FFBDF" httpUri="https://zenodo.org/record/294194/files/figure.png" pageId="6" pageNumber="67" targetBox="[151,1435,181,966]" targetPageId="6">
<paragraph id="8BE436EC1973BD74FF12FC53FC6FFBDF" blockId="6.[151,1435,989,1045]" pageId="6" pageNumber="67">
<emphasis id="B92FEAFE1973BD74FF12FC53FEABFC3E" bold="true" box="[151,276,989,1013]" pageId="6" pageNumber="67">FIGURE 5</emphasis>
.
<taxonomicName id="4C5B4D6F1973BD74FEA7FC50FD6FFC3F" authority="Mortensen, 1910" authorityName="Mortensen" authorityYear="1910" box="[290,720,989,1013]" class="Echinoidea" family="Ctenocidaridae" genus="Ctenocidaris" kingdom="Animalia" order="Cidaroida" pageId="6" pageNumber="67" phylum="Echinodermata" rank="species" species="speciosa">
<emphasis id="B92FEAFE1973BD74FEA7FC50FDADFC3E" box="[290,530,990,1013]" italics="true" pageId="6" pageNumber="67">Ctenocidaris speciosa</emphasis>
Mortensen, 1910
</taxonomicName>
spine with specimens of
<taxonomicName id="4C5B4D6F1973BD74FC61FC50FB19FC3E" box="[996,1190,990,1013]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="67" phylum="Echinodermata" rank="species" species="lockhartae" status="sp. nov.">
<emphasis id="B92FEAFE1973BD74FC61FC50FB19FC3E" box="[996,1190,990,1013]" italics="true" pageId="6" pageNumber="67">Psolus lockhartae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57851973BD74FB2BFC53FAB9FC3E" box="[1198,1286,989,1013]" pageId="6" pageNumber="67" rank="species">
<emphasis id="B92FEAFE1973BD74FB2BFC53FAB9FC3E" bold="true" box="[1198,1286,989,1013]" pageId="6" pageNumber="67">sp. nov.</emphasis>
</taxonomicNameLabel>
above (NMV F168857) and
<taxonomicName id="4C5B4D6F1973BD74FEB7FC70FD7BFBDF" authority="Gutt, 1990" authorityName="Gutt" authorityYear="1990" box="[306,708,1021,1045]" class="Holothuroidea" family="Psolidae" genus="Echinopsolus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="67" phylum="Echinodermata" rank="species" species="acanthocola">
<emphasis id="B92FEAFE1973BD74FEB7FC70FDF2FBDE" box="[306,589,1022,1045]" italics="true" pageId="6" pageNumber="67">Echinopsolus acanthocola</emphasis>
Gutt, 1990
</taxonomicName>
below (NMV F168858).
</paragraph>
</caption>
<subSubSection id="C34165671973BD74FF12FBCDFC6EFAA9" pageId="6" pageNumber="67" type="materials_examined">
<paragraph id="8BE436EC1973BD74FF12FBCDFC15FB49" blockId="6.[151,1437,1091,2013]" pageId="6" pageNumber="67">
<emphasis id="B92FEAFE1973BD74FF12FBCDFE31FB96" bold="true" box="[151,398,1091,1117]" pageId="6" pageNumber="67">Material examined.</emphasis>
<typeStatus id="54E0884E1973BD74FE13FBCDFDB7FB96" box="[406,520,1091,1117]" pageId="6" pageNumber="67" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F34C767C1973BD74FD96FBCDFD2EFB96" box="[531,657,1091,1117]" name="Antarctica" pageId="6" pageNumber="67">Antarctica</collectingCountry>
,
<collectingCountry id="F34C767C1973BD74FD1BFBCDFD57FB96" box="[670,744,1091,1117]" name="United Kingdom" pageId="6" pageNumber="67">Scotia</collectingCountry>
Sea, South
<collectingCountry id="F34C767C1973BD74FCFDFBCDFC6EFB96" box="[888,977,1091,1117]" name="United Kingdom" pageId="6" pageNumber="67">Orkney</collectingCountry>
Is,
<geoCoordinate id="EE6F502B1973BD74FC78FBCDFBC2FB96" box="[1021,1149,1091,1117]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="9" value="-60.6035">60º36.21'S</geoCoordinate>
<geoCoordinate id="EE6F502B1973BD74FB00FBCDFAAFFB96" box="[1157,1296,1091,1117]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="9" value="-44.327835">44º19.67'W</geoCoordinate>
,
<quantity id="4CA39B091973BD74FA99FBCDFAD6FB96" box="[1308,1385,1091,1117]" metricMagnitude="2" metricUnit="m" metricValue="2.11" pageId="6" pageNumber="67" unit="m" value="211.0">211 m</quantity>
,
<collectingCountry id="F34C767C1973BD74FAF3FBCDFA23FB96" box="[1398,1436,1091,1117]" name="United States of America" pageId="6" pageNumber="67">US</collectingCountry>
<collectionCode id="ED4AAE291973BD74FF12FBE6FF53FB49" box="[151,236,1128,1154]" pageId="6" pageNumber="67">AMLR</collectionCode>
2009 stn 19–13, S. Lockhart,
<date id="FFE5102C1973BD74FDC8FBE6FD5DFB49" box="[589,738,1128,1154]" pageId="6" pageNumber="67" value="2009-02-13">13 Feb 2009</date>
,
<collectionCode id="ED4AAE291973BD74FD6BFBE6FC8DFB49" LSID="urn:lsid:biocol.org:col:34978" box="[750,818,1128,1154]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" pageNumber="67">NMV</collectionCode>
F168861.
</paragraph>
<paragraph id="8BE436EC1973BD74FF40FB00FCEAFB06" blockId="6.[151,1437,1091,2013]" pageId="6" pageNumber="67">
<typeStatus id="54E0884E1973BD74FF40FB00FEFFFB63" box="[197,320,1166,1192]" pageId="6" pageNumber="67" type="paratype">Paratypes</typeStatus>
. South
<collectingCountry id="F34C767C1973BD74FE19FB00FE48FB63" box="[412,503,1166,1192]" name="United Kingdom" pageId="6" pageNumber="67">Orkney</collectingCountry>
Is,
<geoCoordinate id="EE6F502B1973BD74FDA2FB00FD15FB63" box="[551,682,1166,1192]" direction="south" orientation="latitude" pageId="6" pageNumber="67" precision="9" value="-60.442">60º26.52'S</geoCoordinate>
<geoCoordinate id="EE6F502B1973BD74FD36FB00FCFDFB63" box="[691,834,1166,1192]" direction="west" orientation="longitude" pageId="6" pageNumber="67" precision="9" value="-45.2985">45º17.91'W</geoCoordinate>
,
<quantity id="4CA39B091973BD74FCCAFB00FC1FFB63" box="[847,928,1166,1192]" metricMagnitude="2" metricUnit="m" metricValue="4.97" pageId="6" pageNumber="67" unit="m" value="497.0">497 m</quantity>
,
<collectingCountry id="F34C767C1973BD74FC28FB00FC6BFB63" box="[941,980,1166,1192]" name="United States of America" pageId="6" pageNumber="67">US</collectingCountry>
<collectionCode id="ED4AAE291973BD74FC58FB00FB8BFB63" box="[989,1076,1166,1192]" pageId="6" pageNumber="67">AMLR</collectionCode>
2009 stn 12–7,
<date id="FFE5102C1973BD74FB7EFB00FA27FB63" box="[1275,1432,1166,1192]" pageId="6" pageNumber="67" value="2009-02-11">11 Feb 2009</date>
,
<collectionCode id="ED4AAE291973BD74FF12FB3DFF63FB06" LSID="urn:lsid:biocol.org:col:34978" box="[151,220,1203,1229]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" pageNumber="67">NMV</collectionCode>
F168857 (1);
<collectionCode id="ED4AAE291973BD74FE06FB3DFE77FB06" LSID="urn:lsid:biocol.org:col:34978" box="[387,456,1203,1229]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" pageNumber="67">NMV</collectionCode>
F168859 (5);
<collectionCode id="ED4AAE291973BD74FDF5FB3DFD0AFB06" LSID="urn:lsid:biocol.org:col:34978" box="[624,693,1203,1229]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" pageNumber="67">NMV</collectionCode>
F168860 (3).
</paragraph>
<paragraph id="8BE436EC1973BD74FF40FB56FC6EFAA9" blockId="6.[151,1437,1091,2013]" pageId="6" pageNumber="67">
<emphasis id="B92FEAFE1973BD74FF40FB56FE2DFB39" bold="true" box="[197,402,1240,1266]" pageId="6" pageNumber="67">Other material.</emphasis>
South
<collectingCountry id="F34C767C1973BD74FE6EFB56FDF9FB39" box="[491,582,1240,1266]" name="United Kingdom" pageId="6" pageNumber="67">Orkney</collectingCountry>
Is,
<quantity id="4CA39B091973BD74FDF0FB56FCBBFB39" box="[629,772,1240,1266]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.02" metricValueMin="2.98" pageId="6" pageNumber="67" unit="m" value="300.0" valueMax="302.0" valueMin="298.0">298–302 m</quantity>
,
<collectionCode id="ED4AAE291973BD74FC94FB56FCD7FB39" LSID="urn:lsid:biocol.org:col:34871" box="[785,872,1240,1266]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="67">USNM</collectionCode>
<accessionNumber id="9408AB0F1973BD74FCF4FB56FC70FB39" box="[881,975,1240,1266]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E43073" pageId="6" pageNumber="67" type="EnaNcbi">E43073</accessionNumber>
(3);
<quantity id="4CA39B091973BD74FB8BFB56FB23FB39" box="[1038,1180,1240,1266]" metricMagnitude="2" metricUnit="m" metricValue="5.955" metricValueMax="5.9799999999999995" metricValueMin="5.93" pageId="6" pageNumber="67" unit="m" value="595.5" valueMax="598.0" valueMin="593.0">593–598 m</quantity>
,
<collectionCode id="ED4AAE291973BD74FB2CFB56FABFFB39" LSID="urn:lsid:biocol.org:col:34871" box="[1193,1280,1240,1266]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="67">USNM</collectionCode>
<accessionNumber id="9408AB0F1973BD74FA8CFB56FAD7FB39" box="[1289,1384,1240,1266]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E43074" pageId="6" pageNumber="67" type="EnaNcbi">E43074</accessionNumber>
(2);
<quantity id="4CA39B091973BD74FF12FB70FE84FAD3" box="[151,315,1278,1304]" metricMagnitude="3" metricUnit="m" metricValue="2.626" metricValueMax="2.897" metricValueMin="2.355" pageId="6" pageNumber="67" unit="m" value="2626.0" valueMax="2897.0" valueMin="2355.0">2355–2897 m</quantity>
,
<collectionCode id="ED4AAE291973BD74FECCFB70FE21FAD3" LSID="urn:lsid:biocol.org:col:34871" box="[329,414,1278,1304]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="67">USNM</collectionCode>
<accessionNumber id="9408AB0F1973BD74FE23FB70FDBEFAD3" box="[422,513,1278,1304]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E43076" pageId="6" pageNumber="67" type="EnaNcbi">E43076</accessionNumber>
(63);
<quantity id="4CA39B091973BD74FDCFFB70FD6DFAD3" box="[586,722,1278,1304]" metricMagnitude="2" metricUnit="m" metricValue="6.1899999999999995" metricValueMax="6.709999999999999" metricValueMin="5.67" pageId="6" pageNumber="67" unit="m" value="619.0" valueMax="671.0" valueMin="567.0">567–671 m</quantity>
,
<collectionCode id="ED4AAE291973BD74FD5AFB70FC8BFAD3" LSID="urn:lsid:biocol.org:col:34871" box="[735,820,1278,1304]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="67">USNM</collectionCode>
<accessionNumber id="9408AB0F1973BD74FCB9FB70FC29FAD3" box="[828,918,1278,1304]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E43080" pageId="6" pageNumber="67" type="EnaNcbi">E43080</accessionNumber>
(3);
<quantity id="4CA39B091973BD74FC54FB70FBE7FAD3" box="[977,1112,1278,1304]" metricMagnitude="2" metricUnit="m" metricValue="8.08" metricValueMax="8.57" metricValueMin="7.59" pageId="6" pageNumber="67" unit="m" value="808.0" valueMax="857.0" valueMin="759.0">759–857 m</quantity>
,
<collectionCode id="ED4AAE291973BD74FBE3FB70FB04FAD3" LSID="urn:lsid:biocol.org:col:34871" box="[1126,1211,1278,1304]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="67">USNM</collectionCode>
<accessionNumber id="9408AB0F1973BD74FB47FB70FAA2FAD3" box="[1218,1309,1278,1304]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E43081" pageId="6" pageNumber="67" type="EnaNcbi">E43081</accessionNumber>
(7); South
<collectingCountry id="F34C767C1973BD74FF12FAADFEBCFAF6" box="[151,259,1315,1341]" name="United Kingdom" pageId="6" pageNumber="67">Shetland</collectingCountry>
Is,
<quantity id="4CA39B091973BD74FEB7FAADFE3BFAF6" box="[306,388,1315,1341]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="67" unit="m" value="300.0">300 m</quantity>
,
<collectionCode id="ED4AAE291973BD74FE14FAADFE57FAF6" LSID="urn:lsid:biocol.org:col:34871" box="[401,488,1315,1341]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="67">USNM</collectionCode>
<accessionNumber id="9408AB0F1973BD74FE74FAADFDEEFAF6" box="[497,593,1315,1341]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E43075" pageId="6" pageNumber="67" type="EnaNcbi">E43075</accessionNumber>
(3);
<quantity id="4CA39B091973BD74FD0BFAADFCA2FAF6" box="[654,797,1315,1341]" metricMagnitude="2" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="6" pageNumber="67" unit="m" value="230.0" valueMax="240.0" valueMin="220.0">220–240 m</quantity>
,
<collectionCode id="ED4AAE291973BD74FCAFFAADFC3EFAF6" LSID="urn:lsid:biocol.org:col:34871" box="[810,897,1315,1341]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="67">USNM</collectionCode>
<accessionNumber id="9408AB0F1973BD74FC0FFAADFC56FAF6" box="[906,1001,1315,1341]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E43078" pageId="6" pageNumber="67" type="EnaNcbi">E43078</accessionNumber>
(1);
<collectingCountry id="F34C767C1973BD74FBADFAADFB61FAF6" box="[1064,1246,1315,1341]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="67">South Georgia</collectingCountry>
,
<quantity id="4CA39B091973BD74FB6EFAADFA27FAF6" box="[1259,1432,1315,1341]" metricMagnitude="3" metricUnit="m" metricValue="2.078" metricValueMax="2.188" metricValueMin="1.968" pageId="6" pageNumber="67" unit="m" value="2078.0" valueMax="2188.0" valueMin="1968.0">1968–2188 m</quantity>
,
<collectionCode id="ED4AAE291973BD74FF12FAC6FF53FAA9" LSID="urn:lsid:biocol.org:col:34871" box="[151,236,1352,1378]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="67">USNM</collectionCode>
<accessionNumber id="9408AB0F1973BD74FF76FAC6FEF2FAA9" box="[243,333,1352,1378]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E43079" pageId="6" pageNumber="67" type="EnaNcbi">E43079</accessionNumber>
(3); Burdwood Bank,
<quantity id="4CA39B091973BD74FDD3FAC6FD61FAA9" box="[598,734,1352,1378]" metricMagnitude="2" metricUnit="m" metricValue="3.4799999999999995" metricValueMax="3.5700000000000003" metricValueMin="3.3899999999999997" pageId="6" pageNumber="67" unit="m" value="348.0" valueMax="357.0" valueMin="339.0">339–357 m</quantity>
,
<collectionCode id="ED4AAE291973BD74FD6CFAC6FC81FAA9" LSID="urn:lsid:biocol.org:col:34871" box="[745,830,1352,1378]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="67">USNM</collectionCode>
<accessionNumber id="9408AB0F1973BD74FCC3FAC6FC1FFAA9" box="[838,928,1352,1378]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E43077" pageId="6" pageNumber="67" type="EnaNcbi">E43077</accessionNumber>
(4).
</paragraph>
</subSubSection>
<subSubSection id="C34165671973BD74FF43FAE3FDADF816" pageId="6" pageNumber="67" type="diagnosis">
<paragraph id="8BE436EC1973BD74FF43FAE3FE3CF979" blockId="6.[151,1437,1091,2013]" pageId="6" pageNumber="67">
<emphasis id="B92FEAFE1973BD74FF43FAE3FEF9FA4C" bold="true" box="[198,326,1389,1415]" pageId="6" pageNumber="67">Diagnosis.</emphasis>
<taxonomicName id="4C5B4D6F1973BD74FED5FAE0FE21FA4C" box="[336,414,1390,1415]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="67" phylum="Echinodermata" rank="genus">
<emphasis id="B92FEAFE1973BD74FED5FAE0FE21FA4C" box="[336,414,1390,1415]" italics="true" pageId="6" pageNumber="67">Psolus</emphasis>
</taxonomicName>
species up to
<quantity id="4CA39B091973BD74FDD4FAE0FD18FA43" box="[593,679,1390,1416]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="67" unit="mm" value="20.0">20 mm</quantity>
long,
<quantity id="4CA39B091973BD74FD72FAE0FCF2FA43" box="[759,845,1390,1416]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="67" unit="mm" value="10.0">10 mm</quantity>
high,
<quantity id="4CA39B091973BD74FC18FAE0FC5BFA43" box="[925,996,1390,1416]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="67" unit="mm" value="8.0">8 mm</quantity>
wide; form sub-cylindrical, smooth, rounded oral and anal ends, lacking oral and anal cone; scales up to
<quantity id="4CA39B091973BD74FC31FA1DFC48FA66" box="[948,1015,1427,1453]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="67" unit="mm" value="3.0">3 mm</quantity>
wide, imbricating, macroscopically smooth, microscopically finely beaded, mosaic arrangement of large and small, rounded and angular scales; lacking 5 discrete oral valves, oral scales numerous, digitiform or narrow pointed or triangular pointed; tentacles 10, 8 large, 2 small ventrally; calcareous ring with digitiform radial and interradial anterior projections, radial projections slightly swollen distally, concave radial and interradial posterior indentations; thin calcareous sole, single to zig-zag to double series of large inner peripheral tube feet, lacking outer peripheral series of smaller tube feet, rare mid-ventral tube feet present; single ventral polian vesicle; gonad tubules unbranched.
</paragraph>
<paragraph id="8BE436EC1973BD74FF43F930FC17F873" blockId="6.[151,1437,1091,2013]" pageId="6" pageNumber="67">
Dorsal and lateral multilayered ossicles only. Sole with crosses, thick, finely knobbed or bluntly-spinous, shallow concave or cupped, small and large, commonly 88–112
<emphasis id="B92FEAFE1973BD74FC30F96DFC7BF937" box="[949,964,1763,1788]" italics="true" pageId="6" pageNumber="67">µ</emphasis>
m long, rarely 168
<emphasis id="B92FEAFE1973BD74FB33F96DFB7AF937" box="[1206,1221,1763,1788]" italics="true" pageId="6" pageNumber="67">µ</emphasis>
m long, upturned branched arms of crosses may join to form cups with up to 8 perforations, marginal projections sometimes joined to create anastomosing branches over cups or transverse bridges across cups. Largest tentacle ossicles irregularly oval or triangular or narrow perforated plates, up to 480
<emphasis id="B92FEAFE1973BD74FC72F8DDFBB9F8A7" box="[1015,1030,1875,1900]" italics="true" pageId="6" pageNumber="67">µ</emphasis>
m long, surface smooth, bluntly denticulate margin. Largest tube feet support ossicles up to 560
<emphasis id="B92FEAFE1973BD74FC2CF8F7FC07F859" box="[937,952,1913,1938]" italics="true" pageId="6" pageNumber="67">µ</emphasis>
m long, thick smooth perforated plates, irregularly oval or triangular or narrow, bluntly denticulate margin.
</paragraph>
<paragraph id="8BE436EC1973BD74FF43F84DFDADF816" blockId="6.[151,1437,1091,2013]" box="[198,530,1987,2013]" pageId="6" pageNumber="67">
<emphasis id="B92FEAFE1973BD74FF43F84DFE04F816" bold="true" box="[198,443,1987,2013]" pageId="6" pageNumber="67">Colour (preserved).</emphasis>
White.
</paragraph>
</subSubSection>
<subSubSection id="C34165671972BD75FF43FF19FE5AFF37" pageId="7" pageNumber="68" type="description">
<paragraph id="8BE436EC1972BD75FF43FF19FE5AFF37" blockId="7.[151,1437,151,738]" pageId="7" pageNumber="68">
<emphasis id="B92FEAFE1972BD75FF43FF19FEAAFF7A" bold="true" box="[198,277,151,177]" pageId="7" pageNumber="68">Habit.</emphasis>
Lives with the lateral margins of the sole wrapped around
<taxonomicName id="4C5B4D6F1972BD75FC5EFF16FBF2FF79" box="[987,1101,152,178]" class="Echinoidea" family="Cidaridae" kingdom="Animalia" order="Cidaroida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="family">Cidaridae</taxonomicName>
echinoid spines of different species, with
<typeStatus id="54E0884E1972BD75FEB2FF33FED5FF1C" box="[311,362,189,215]" pageId="7" pageNumber="68">type</typeStatus>
specimens found on spines of
<taxonomicName id="4C5B4D6F1972BD75FD53FF33FB15FF1C" authority="Mortensen, 1910" authorityName="Mortensen" authorityYear="1910" box="[726,1194,189,215]" class="Echinoidea" family="Ctenocidaridae" genus="Ctenocidaris" kingdom="Animalia" order="Cidaroida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="species" species="speciosa">
<emphasis id="B92FEAFE1972BD75FD53FF33FC64FF1D" box="[726,987,189,214]" italics="true" pageId="7" pageNumber="68">Ctenocidaris speciosa</emphasis>
<bibRefCitation id="EFCA4B1D1972BD75FC66FF33FB15FF1C" author="Mortensen" box="[995,1194,189,215]" pageId="7" pageNumber="68" refString="Mortensen, T. (1910) The Echinoidea of the Swedish south polar expedition. Schwedische Sudpolar-Expedition 1901 - 1903, 6, 1 - 114." type="journal article" year="1910">Mortensen, 1910</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C5B4D6F1972BD75FB61FF33FE5FFF37" authority="Mortensen, 1909" authorityName="Mortensen" authorityYear="1909" class="Echinoidea" family="Ctenocidaridae" genus="Rhynchocidaris" kingdom="Animalia" order="Cidaroida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="species" species="triplopora">
<emphasis id="B92FEAFE1972BD75FB61FF33FEAFFF30" italics="true" pageId="7" pageNumber="68">Rhynchocidaris triplopora</emphasis>
<bibRefCitation id="EFCA4B1D1972BD75FE93FF6CFE5FFF37" author="Mortensen" box="[278,480,226,252]" pageId="7" pageNumber="68" refString="Mortensen, T. (1909) Die Echinoiden der Deutchen Sudpolar-Expedition 1901 - 1903. Deutsche Sudpolar-Expedition, 11, 1 - 113." type="journal article" year="1909">Mortensen, 1909</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34165671972BD75FF43FE89FD2EFE8C" pageId="7" pageNumber="68" type="distribution">
<paragraph id="8BE436EC1972BD75FF43FE89FD2EFE8C" blockId="7.[151,1437,151,738]" pageId="7" pageNumber="68">
<emphasis id="B92FEAFE1972BD75FF43FE89FEE0FEEA" bold="true" box="[198,351,263,289]" pageId="7" pageNumber="68">Distribution</emphasis>
. South Atlantic Ocean, Burdwood Bank;
<collectingCountry id="F34C767C1972BD75FCCDFE86FC0AFEE9" box="[840,949,264,290]" name="Antarctica" pageId="7" pageNumber="68">Antarctic</collectingCountry>
Ocean,
<collectingCountry id="F34C767C1972BD75FB93FE86FBE0FEE9" box="[1046,1119,264,290]" name="United Kingdom" pageId="7" pageNumber="68">Scotia</collectingCountry>
Sea,
<collectingCountry id="F34C767C1972BD75FB1AFE86FAF3FEE9" box="[1183,1356,264,290]" name="South Georgia and the South Sandwich Islands" pageId="7" pageNumber="68">South Georgia</collectingCountry>
, South
<collectingCountry id="F34C767C1972BD75FF12FEA3FF41FE8C" box="[151,254,301,327]" name="United Kingdom" pageId="7" pageNumber="68">Shetland</collectingCountry>
Is, South
<collectingCountry id="F34C767C1972BD75FEF1FEA3FE73FE8C" box="[372,460,301,327]" name="United Kingdom" pageId="7" pageNumber="68">Orkney</collectingCountry>
Is,
<quantity id="4CA39B091972BD75FE72FEA3FD33FE8C" box="[503,652,301,327]" metricMagnitude="3" metricUnit="m" metricValue="1.554" metricValueMax="2.897" metricValueMin="0.211" pageId="7" pageNumber="68" unit="m" value="1554.0" valueMax="2897.0" valueMin="211.0">211–2897 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34165671972BD75FF43FEDCFDA2FE7C" pageId="7" pageNumber="68" type="etymology">
<paragraph id="8BE436EC1972BD75FF43FEDCFDA2FE7C" blockId="7.[151,1437,151,738]" pageId="7" pageNumber="68">
<emphasis id="B92FEAFE1972BD75FF43FEDCFEECFEA7" bold="true" box="[198,339,338,364]" pageId="7" pageNumber="68">Etymology.</emphasis>
Named for Susie Lockhart (
<collectingCountry id="F34C767C1972BD75FD31FEDCFD64FEA7" box="[692,731,338,364]" name="United States of America" pageId="7" pageNumber="68">US</collectingCountry>
National Oceanic and Atmospheric Administration), with appreciation of her generous collaborative assistance in creating the opportunity for us to study
<collectingCountry id="F34C767C1972BD75FA9DFEF6FA81FE59" box="[1304,1342,376,402]" name="United States of America" pageId="7" pageNumber="68">US</collectingCountry>
AMLR
<collectingCountry id="F34C767C1972BD75FF12FE13FEBBFE7C" box="[151,260,413,439]" name="Antarctica" pageId="7" pageNumber="68">Antarctic</collectingCountry>
holothuroid specimens.
</paragraph>
</subSubSection>
<subSubSection id="C34165671972BD75FF43FE4CFA2CFD29" pageId="7" pageNumber="68" type="discussion">
<paragraph id="8BE436EC1972BD75FF43FE4CFA2CFD29" blockId="7.[151,1437,151,738]" pageId="7" pageNumber="68">
<emphasis id="B92FEAFE1972BD75FF43FE4CFE80FE17" bold="true" box="[198,319,450,476]" pageId="7" pageNumber="68">Remarks.</emphasis>
<taxonomicName id="4C5B4D6F1972BD75FECDFE4CFDA3FE10" box="[328,540,450,475]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="species" species="lockhartae" status="sp. nov.">
<emphasis id="B92FEAFE1972BD75FECDFE4CFDA3FE10" box="[328,540,450,475]" italics="true" pageId="7" pageNumber="68">Psolus lockhartae</emphasis>
</taxonomicName>
<emphasis id="B92FEAFE1972BD75FDA0FE4CFD3AFE17" bold="true" box="[549,645,450,476]" pageId="7" pageNumber="68">
<taxonomicNameLabel id="A21C57851972BD75FDA0FE4CFD3AFE17" box="[549,645,450,476]" pageId="7" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only known species of
<taxonomicName id="4C5B4D6F1972BD75FC6EFE4CFB86FE10" box="[1003,1081,450,475]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="genus">
<emphasis id="B92FEAFE1972BD75FC6EFE4CFB86FE10" box="[1003,1081,450,475]" italics="true" pageId="7" pageNumber="68">Psolus</emphasis>
</taxonomicName>
that lives on echinoid spines. The psolids
<taxonomicName id="4C5B4D6F1972BD75FEA0FE66FD60FDC9" authority="Gutt, 1990" authorityName="Gutt" authorityYear="1990" box="[293,735,488,514]" class="Holothuroidea" family="Psolidae" genus="Echinopsolus" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="species" species="acanthocola">
<emphasis id="B92FEAFE1972BD75FEA0FE66FDE5FDCA" box="[293,602,488,513]" italics="true" pageId="7" pageNumber="68">Echinopsolus acanthocola</emphasis>
<bibRefCitation id="EFCA4B1D1972BD75FDE4FE66FD60FDC9" author="Gutt" box="[609,735,488,514]" pageId="7" pageNumber="68" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt, 1990</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C5B4D6F1972BD75FC93FE66FB7BFDC9" authority="Massin, 1992" authorityName="Massin" authorityYear="1992" box="[790,1220,488,514]" class="Holothuroidea" family="Psolidae" genus="Echinopsolus" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="species" species="parvipes">
<emphasis id="B92FEAFE1972BD75FC93FE66FB9FFDCA" box="[790,1056,488,513]" italics="true" pageId="7" pageNumber="68">Echinopsolus parvipes</emphasis>
<bibRefCitation id="EFCA4B1D1972BD75FBA3FE66FB7BFDC9" author="Massin" box="[1062,1220,488,514]" pageId="7" pageNumber="68" refString="Massin, C. (1992) Three new species of Dendrochirotida (Holothuroidea, Echinodermata) from the Weddell Sea (Antarctica). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie, 62, 179 - 191." type="journal article" year="1992">Massin, 1992</bibRefCitation>
</taxonomicName>
have a similar life habit. In this study one specimen of
<taxonomicName id="4C5B4D6F1972BD75FDC8FD83FC9BFDED" box="[589,804,525,550]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="species" species="lockhartae" status="sp. nov.">
<emphasis id="B92FEAFE1972BD75FDC8FD83FC9BFDED" box="[589,804,525,550]" italics="true" pageId="7" pageNumber="68">Psolus lockhartae</emphasis>
</taxonomicName>
<emphasis id="B92FEAFE1972BD75FCA9FD83FC33FDEC" bold="true" box="[812,908,525,551]" pageId="7" pageNumber="68">
<taxonomicNameLabel id="A21C57851972BD75FCA9FD83FC33FDEC" box="[812,908,525,551]" pageId="7" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(NMV F168857) was found attached to the same spine from the echinoid
<taxonomicName id="4C5B4D6F1972BD75FD89FDBCFCA1FD80" box="[524,798,562,587]" class="Echinoidea" family="Ctenocidaridae" genus="Ctenocidaris" kingdom="Animalia" order="Cidaroida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="species" species="speciosa">
<emphasis id="B92FEAFE1972BD75FD89FDBCFCA1FD80" box="[524,798,562,587]" italics="true" pageId="7" pageNumber="68">Ctenocidaris speciosa</emphasis>
</taxonomicName>
as a specimen of
<taxonomicName id="4C5B4D6F1972BD75FC7BFDBCFAFBFD80" box="[1022,1348,562,587]" class="Holothuroidea" family="Psolidae" genus="Echinopsolus" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="species" species="acanthocola">
<emphasis id="B92FEAFE1972BD75FC7BFDBCFAFBFD80" box="[1022,1348,562,587]" italics="true" pageId="7" pageNumber="68">Echinopsolus acanthocola</emphasis>
</taxonomicName>
(NMV F168858). In addition to a unique habit, the morphological characters that distinguish this species from all other
<collectingCountry id="F34C767C1972BD75FF5AFDF3FEEDFD5C" box="[223,338,637,663]" name="Antarctica" pageId="7" pageNumber="68">Antarctic</collectingCountry>
<taxonomicName id="4C5B4D6F1972BD75FEDFFDF3FE13FD5D" box="[346,428,637,662]" class="Holothuroidea" family="Psolidae" genus="Psolus" kingdom="Animalia" order="Dendrochirotida" pageId="7" pageNumber="68" phylum="Echinodermata" rank="genus">
<emphasis id="B92FEAFE1972BD75FEDFFDF3FE13FD5D" box="[346,428,637,662]" italics="true" pageId="7" pageNumber="68">Psolus</emphasis>
</taxonomicName>
species are a combination of: smooth sub-cylindrical body form; absence of oral valves; absence of an outer peripheral series of smaller tube feet on the sole; distinctive thick knobbed and bluntly spinous concave crosses and cups in the sole; lacking ossicles other than scales dorsally and laterally.
</paragraph>
</subSubSection>
</treatment>
</document>
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