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<document id="DE9B7A5272441B3B3069D89FD070AB5D" ID-CLB-Dataset="25313" ID-DOI="10.11646/zootaxa.4679.3.7" ID-GBIF-Dataset="0a75e36a-4254-44be-9a1a-2c2bc6f115fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3471264" ID-ZooBank="1FF46982-622E-4C27-AB8E-8F8A993A419A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570099354004" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03F287B9FFDA1B1AFF10FC750B8B26FE" docLanguage="en" docName="zootaxa.4679.3.7.pdf" docOrigin="Zootaxa 4679 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microscydmus (Microscydmus) kochiensis Jałoszyński 2019, sp. n." docType="treatment" docVersion="6" lastPageNumber="545" masterDocId="FFCBFFC1FFDF1B1CFF87FFF80864257D" masterDocTitle="Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="552" masterPageNumber="539" pageNumber="544" updateTime="1698757590553" updateUser="ExternalLinkService">
<mods:mods id="96A406C2171504A5D3CC0AC15A640705" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2448F492F16D028E0B45E738C1091220">
<mods:title id="021EB55694129D134513D06E68E93A95">Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="97EBBC73FC6230BC11CA82048CE45286" type="personal">
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<mods:roleTerm id="8E6802BCDA93F6260678BE4855652D01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="17C051D102A09E44A8A5212147BBDE62">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="6D6802E9BA2A4597F19A5599ABF766EC">text</mods:typeOfResource>
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<mods:title id="5429443BE8593C554C3B913547490998">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="70E5C868A52160BD270FB8BC012B727E">
<mods:date id="8B46ACC1F9A727092C4E6D92B75F36FD">2019</mods:date>
<mods:detail id="7BB8B21CC0A5B72358C9F3503C8D0317" type="pubDate">
<mods:number id="7D07E3D836BFABA5C0C18795926F8DB5">2019-10-03</mods:number>
</mods:detail>
<mods:detail id="1AC6550502B21013AE0A64B6F7213025" type="volume">
<mods:number id="03A4797390B7190EE486634B731D7CF7">4679</mods:number>
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<mods:number id="654661513A92CCC6F1AF95F568743509">3</mods:number>
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<mods:classification id="E9A9B0483AFBAD601141298544D72FB9">journal article</mods:classification>
<mods:identifier id="3E3F8CBF544F4B63D72CB8A2ECA7F8BA" type="CLB-Dataset">25313</mods:identifier>
<mods:identifier id="451D9ABEB6D3AEDF0E9F4C79A83C4C99" type="DOI">10.11646/zootaxa.4679.3.7</mods:identifier>
<mods:identifier id="7EBB37F70384B1EBD5672ED2C1B24EBB" type="GBIF-Dataset">0a75e36a-4254-44be-9a1a-2c2bc6f115fa</mods:identifier>
<mods:identifier id="4266A1EAF5B4221429B27CA7B41829BF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3A271F4C486E061459A36F4D50686FA7" type="Zenodo-Dep">3471264</mods:identifier>
<mods:identifier id="0E4A52952D9D06A3E8864F5B110F5BF7" type="ZooBank">1FF46982-622E-4C27-AB8E-8F8A993A419A</mods:identifier>
</mods:mods>
<treatment id="03F287B9FFDA1B1AFF10FC750B8B26FE" ID-GBIF-Taxon="159628633" LSID="urn:lsid:plazi:treatment:03F287B9FFDA1B1AFF10FC750B8B26FE" httpUri="http://treatment.plazi.org/id/03F287B9FFDA1B1AFF10FC750B8B26FE" lastPageId="6" lastPageNumber="545" pageId="5" pageNumber="544">
<subSubSection id="C3416524FFDA1B19FF10FC750A8626DA" box="[151,738,909,936]" pageId="5" pageNumber="544" type="nomenclature">
<paragraph id="8BE436AFFFDA1B19FF10FC750A8626DA" blockId="5.[151,738,909,971]" box="[151,738,909,936]" pageId="5" pageNumber="544">
<heading id="D0AC81C3FFDA1B19FF10FC750A8626DA" bold="true" box="[151,738,909,936]" fontSize="11" level="1" pageId="5" pageNumber="544" reason="1">
<emphasis id="B92FEABDFFDA1B19FF10FC750A8626DA" bold="true" box="[151,738,909,936]" pageId="5" pageNumber="544">
<taxonomicName id="4C5B4D2CFFDA1B19FF10FC750AFC26D5" ID-CoL="8Y6FD" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,664,909,936]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="kochiensis" status="sp. nov." subGenus="Microscydmus">
<emphasis id="B92FEABDFFDA1B19FF10FC75092326D5" bold="true" box="[151,327,909,936]" italics="true" pageId="5" pageNumber="544">Microscydmus</emphasis>
(
<emphasis id="B92FEABDFFDA1B19FEDFFC750A6C26D5" bold="true" box="[344,520,909,936]" italics="true" pageId="5" pageNumber="544">Microscydmus</emphasis>
)
<emphasis id="B92FEABDFFDA1B19FD9EFC750AFC26D5" bold="true" box="[537,664,909,936]" italics="true" pageId="5" pageNumber="544">kochiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57C6FFDA1B19FD27FC750A8626DA" box="[672,738,909,935]" pageId="5" pageNumber="544" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDA1B19FF10FC49090926B6" box="[151,365,945,971]" pageId="5" pageNumber="544" type="description">
<paragraph id="8BE436AFFFDA1B19FF10FC49090926B6" blockId="5.[151,738,909,971]" box="[151,365,945,971]" pageId="5" pageNumber="544">
(
<figureCitation id="13602A2AFFDA1B19FF18FC49088C26B6" box="[159,232,945,971]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="5" pageNumber="544">Figs 4</figureCitation>
,
<figureCitation id="13602A2AFFDA1B19FF73FC49097626B6" box="[244,274,945,971]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="5" pageNumber="544">12</figureCitation>
,
<figureCitation id="13602A2AFFDA1B19FE99FC49095926B6" box="[286,317,945,971]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="5" pageNumber="544">20</figureCitation>
,
<figureCitation id="13602A2AFFDA1B19FECFFC49090126B6" box="[328,357,945,971]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="5" pageNumber="544">28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDA1B19FF10FC010C3821DE" pageId="5" pageNumber="544" type="materials_examined">
<paragraph id="8BE436AFFFDA1B19FF10FC010C3821DE" blockId="5.[151,1437,1016,2015]" pageId="5" pageNumber="544">
<emphasis id="B92FEABDFFDA1B19FF10FC01092D216E" bold="true" box="[151,329,1016,1043]" pageId="5" pageNumber="544">Type material.</emphasis>
<materialsCitation id="3B333CF2FFDA1B19FED5FC010A392126" ID-GBIF-Occurrence="2423812457" collectingDate="1999-05-23" collectionCode="JAPAN, EUMJ" collectorName="Pref. &amp; M. Sakai" country="Japan" county="Nishitosa-mura" location="Kochi" municipality="Kuroson" pageId="5" pageNumber="544" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B92FEABDFFDA1B19FED5FC010AB9216E" bold="true" box="[338,733,1016,1043]" pageId="5" pageNumber="544">
<typeStatus id="54E0880DFFDA1B19FED5FC0109A7216E" box="[338,451,1017,1043]" pageId="5" pageNumber="544" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34C763FFFDA1B19FE56FC000A49216E" box="[465,557,1016,1043]" name="Japan" pageId="5" pageNumber="544">JAPAN</collectingCountry>
(Kôchi
<collectorName id="26AE5379FFDA1B19FD17FC000AB4216E" box="[656,720,1016,1043]" pageId="5" pageNumber="544">Pref.</collectorName>
):
</emphasis>
<specimenCount id="9D5DFD26FFDA1B19FD61FC010A99216E" box="[742,765,1017,1043]" pageId="5" pageNumber="544" type="male">♂</specimenCount>
, two labels: &quot;(
<location id="8E846074FFDA1B19FC25FC010B8B216E" LSID="urn:lsid:plazi:treatment:03F287B9FFDA1B1AFF10FC750B8B26FE:8E846074FFDA1B19FC25FC010B8B216E" box="[930,1007,1017,1043]" country="Japan" county="Nishitosa-mura" municipality="Kuroson" name="Kochi" pageId="5" pageNumber="544">Kôchi</location>
:
<collectionCode id="ED4AAE6AFFDA1B19FC7CFC010C30216E" box="[1019,1108,1017,1043]" pageId="5" pageNumber="544">JAPAN</collectionCode>
) /
<collectingMunicipality id="6B80ACD5FFDA1B19FBF2FC010CB1216E" box="[1141,1237,1017,1043]" pageId="5" pageNumber="544">Kuroson</collectingMunicipality>
/
<collectingCounty id="62854E23FFDA1B19FB77FC010DF9216E" box="[1264,1437,1017,1043]" pageId="5" pageNumber="544">Nishitosa-mura</collectingCounty>
/
<date id="FFE5106FFFDA1B19FF20FBE5094A214A" box="[167,302,1053,1079]" pageId="5" pageNumber="544" value="1999-05-23">
<collectingDate id="EFA1E987FFDA1B19FF20FBE5094A214A" box="[167,302,1053,1079]" pageId="5" pageNumber="544" value="1999-05-23">23. V. 1999</collectingDate>
</date>
/
<collectorName id="26AE5379FFDA1B19FEC1FBE509C8214A" box="[326,428,1053,1079]" pageId="5" pageNumber="544">M. Sakai</collectorName>
leg.&quot; [white, printed]; &quot;
<taxonomicName id="4C5B4D2CFFDA1B19FD3BFBE50BF7214A" authorityName="Saulcy &amp; Croissandeau" authorityYear="1893" box="[700,915,1053,1079]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEABDFFDA1B19FD3BFBE50BF7214A" box="[700,915,1053,1079]" italics="true" pageId="5" pageNumber="544">MICROSCYDMUS</emphasis>
</taxonomicName>
(s. str.) /
<emphasis id="B92FEABDFFDA1B19FB84FBE50C1C214A" box="[1027,1144,1053,1079]" italics="true" pageId="5" pageNumber="544">kochiensis</emphasis>
m. / P.
<bibRefCitation id="EFCA4B5EFFDA1B19FB53FBE50DF8214A" author="Jaloszynski, P." box="[1236,1436,1053,1079]" pageId="5" pageNumber="544" pagination="205 - 220" refId="ref9055" refString="Jaloszynski, P. (2019) Taxonomy of the &quot; Afroeudesis group &quot; of glandulariine ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4612 (2), 205 - 220. https: // doi. org / 10.11646 / zootaxa. 4612.2.4" type="journal article" year="2019">Jałoszyński, 2019</bibRefCitation>
/
<typeStatus id="54E0880DFFDA1B19FF20FBB909352126" box="[167,337,1089,1115]" pageId="5" pageNumber="544" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="ED4AAE6AFFDA1B19FD81FBB90A312126" box="[518,597,1089,1115]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="5" pageNumber="544">EUMJ</collectionCode>
)
</materialsCitation>
.
<emphasis id="B92FEABDFFDA1B19FDEEFBB90B2E2126" bold="true" box="[617,842,1088,1115]" pageId="5" pageNumber="544">
<typeStatus id="54E0880DFFDA1B19FDEEFBB90A852126" box="[617,737,1089,1115]" pageId="5" pageNumber="544" type="paratype">Paratypes</typeStatus>
(3 exx.):
</emphasis>
<specimenCount id="9D5DFD26FFDA1B19FCD5FBB90BF72126" box="[850,915,1089,1115]" pageId="5" pageNumber="544" type="female">2 ♀♀</specimenCount>
,
<materialsCitation id="3B333CF2FFDA1B19FC26FBB90AAE21DE" ID-GBIF-Occurrence="2423812463" collectingDate="1997-10-18" collectionCode="JAPAN" collectorName="Tatsuya Kan &amp; M. Sakai" country="India" county="Kuroson" location="Kochi" municipality="Ohdaba" pageId="5" pageNumber="544" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
same data as for holotype, except for
<collectingDate id="EFA1E987FFDA1B19FACAFBB909D52102" isEnumeration="true" pageId="5" pageNumber="544">
<date id="FFE5106FFFDA1B19FACAFBB908AB2102" pageId="5" pageNumber="544" value="1999-06-21">21. VI. 1999</date>
and
<date id="FFE5106FFFDA1B19FE82FB9D09D52102" box="[261,433,1125,1151]" pageId="5" pageNumber="544" value="1999-05-22" valueMax="1999-05-23" valueMin="1999-05-22">22-23. V. 1999</date>
</collectingDate>
, both
<collectorName id="26AE5379FFDA1B19FE73FB9D0AE62102" box="[500,642,1125,1151]" pageId="5" pageNumber="544">Tatsuya Kan</collectorName>
leg.;
<specimenCount id="9D5DFD26FFDA1B19FD47FB9D0A8E2102" box="[704,746,1125,1151]" pageId="5" pageNumber="544" type="female">1 ♀</specimenCount>
, &quot;
<emphasis id="B92FEABDFFDA1B19FC84FB9D0BCA2102" box="[771,942,1125,1151]" italics="true" pageId="5" pageNumber="544">
<location id="8E846074FFDA1B19FC84FB9D0B222102" LSID="urn:lsid:plazi:treatment:03F287B9FFDA1B1AFF10FC750B8B26FE:8E846074FFDA1B19FC84FB9D0B222102" box="[771,838,1125,1151]" country="India" county="Kuroson" municipality="Ohdaba" name="Kochi" pageId="5" pageNumber="544">Kôchi</location>
:
<collectionCode id="ED4AAE6AFFDA1B19FCDBFB9D0BCA2102" box="[860,942,1125,1151]" pageId="5" pageNumber="544">JAPAN</collectionCode>
</emphasis>
/
<collectingMunicipality id="6B80ACD5FFDA1B19FC43FB9D0C7F2102" box="[964,1051,1125,1151]" pageId="5" pageNumber="544">Ohdaba</collectingMunicipality>
/
<collectingCounty id="62854E23FFDA1B19FBB7FB9D0CF42102" box="[1072,1168,1125,1151]" pageId="5" pageNumber="544">Kuroson</collectingCounty>
/
<date id="FFE5106FFFDA1B19FB22FB9D0D462102" box="[1189,1314,1125,1151]" pageId="5" pageNumber="544" value="1997-10-18">
<collectingDate id="EFA1E987FFDA1B19FB22FB9D0D462102" box="[1189,1314,1125,1151]" pageId="5" pageNumber="544" value="1997-10-18">18. x. 1997</collectingDate>
</date>
/
<collectorName id="26AE5379FFDA1B19FABFFB9D0DF82102" box="[1336,1436,1125,1151]" pageId="5" pageNumber="544">M. Sakai</collectorName>
(
<emphasis id="B92FEABDFFDA1B19FF26FB7108B321DE" box="[161,215,1161,1187]" italics="true" pageId="5" pageNumber="544">litter</emphasis>
)&quot; [white, printed]; all
<typeStatus id="54E0880DFFDA1B19FE54FB710A5A21DE" box="[467,574,1161,1187]" pageId="5" pageNumber="544" type="paratype">paratypes</typeStatus>
with yellow
</materialsCitation>
<materialsCitation id="3B333CF2FFDA1B19FD56FB710C3C21DE" ID-GBIF-Occurrence="2423812461" box="[721,1112,1161,1187]" collectionCode="EUMJ" country="India" pageId="5" pageNumber="544" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E0880DFFDA1B19FD56FB710B1021DE" box="[721,884,1161,1187]" pageId="5" pageNumber="544" type="paratype">PARATYPUS</typeStatus>
labels (cPJ,
<collectionCode id="ED4AAE6AFFDA1B19FB85FB710C3421DE" box="[1026,1104,1161,1187]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="5" pageNumber="544">EUMJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDA1B19FF40FB540A87204E" pageId="5" pageNumber="544" type="diagnosis">
<paragraph id="8BE436AFFFDA1B19FF40FB540A87204E" blockId="5.[151,1437,1016,2015]" pageId="5" pageNumber="544">
<emphasis id="B92FEABDFFDA1B19FF40FB54092621BA" bold="true" box="[199,322,1196,1223]" pageId="5" pageNumber="544">Diagnosis.</emphasis>
Body length ~
<quantity id="4CA39B4AFFDA1B19FE7DFB550A3221BA" box="[506,598,1197,1223]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="544" unit="mm" value="0.8">0.8 mm</quantity>
; entire frons and vertex with distinct, deep, well-defined and very dense punctures; bristles on tempora and vertex short but well-visible; protibia not thickened in apical region, with apical comb of four teeth that occupy distomesal margin; aedeagus with conspicuous ovoid vesicular structure in sub-basal region of median lobe; parameres strongly recurved.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFDA1B1AFF40FAC4090E24AE" lastPageId="6" lastPageNumber="545" pageId="5" pageNumber="544" type="description">
<paragraph id="8BE436AFFFDA1B19FF40FAC409DA2006" blockId="5.[151,1437,1016,2015]" pageId="5" pageNumber="544">
<emphasis id="B92FEABDFFDA1B19FF40FAC4093D202A" bold="true" box="[199,345,1340,1367]" pageId="5" pageNumber="544">Description.</emphasis>
Body of male (
<figureCitation id="13602A2AFFDA1B19FD83FAC50A23202A" box="[516,583,1341,1367]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="5" pageNumber="544">Fig. 4</figureCitation>
) slender, distinctly flattened, pigmentation moderately dark brown, vestiture light brown; BL
<quantity id="4CA39B4AFFDA1B19FED4FA9909DE2006" box="[339,442,1377,1403]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="5" pageNumber="544" unit="mm" value="0.81">0.81 mm</quantity>
.
</paragraph>
<paragraph id="8BE436AFFFDA1B19FF40FA7D08BB230A" blockId="5.[151,1437,1016,2015]" pageId="5" pageNumber="544">
Head (
<figureCitation id="13602A2AFFDA1B19FE94FA7D090120E2" box="[275,357,1413,1439]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="5" pageNumber="544">Fig. 12</figureCitation>
) broadest at eyes, HL
<quantity id="4CA39B4AFFDA1B19FDE0FA7C0AAB20E2" box="[615,719,1412,1439]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="544" unit="mm" value="0.15">0.15 mm</quantity>
, HW
<quantity id="4CA39B4AFFDA1B19FC95FA7D0B1E20E2" box="[786,890,1413,1439]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="5" pageNumber="544" unit="mm" value="0.18">0.18 mm</quantity>
; tempora weakly arcuate, about as long as eyes; posterior margin of vertex nearly straight, vertex and frons confluent and convex; supraantennal tubercles weakly elevated but well-defined; eyes large, strongly convex and coarsely faceted. Frons and vertex uniformly and evenly covered with small but distinct, well-marked and conspicuously dense punctures; setae sparse, short and suberect, bristles on tempora and vertex short but well-visible; long posterolateral setae on vertex not found. Antennae slen- der, AnL
<quantity id="4CA39B4AFFDA1B19FE85F9C1090E232E" box="[258,362,1593,1619]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="544" unit="mm" value="0.3">0.30 mm</quantity>
, antennomeres I and II strongly elongate, III–X each distinctly transverse, XI about as long as broad.
</paragraph>
<paragraph id="8BE436AFFFDA1B19FF40F979088D2232" blockId="5.[151,1437,1016,2015]" pageId="5" pageNumber="544">
Pronotum in dorsal view nearly circular, broadest in front of middle, PL
<quantity id="4CA39B4AFFDA1B19FB98F9790CEC23E6" box="[1055,1160,1665,1691]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="544" unit="mm" value="0.21">0.21 mm</quantity>
, PW
<quantity id="4CA39B4AFFDA1B19FB4BF9790D5223E6" box="[1228,1334,1665,1691]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="544" unit="mm" value="0.23">0.23 mm</quantity>
; anterior margin weakly arcuate; anterior corners not marked; sides strongly rounded; posterior pronotal corners distinct, obtuse-angled; pronotal base with a well-defined, short transverse groove deepened at each end, lateral pit near each hind corner present, but barely discernible, very small and obscured by dense punctures. Punctures on disc similar as those on head, conspicuously dense; setae sparse, short, suberect; lateral bristles dense and relatively long, well visible.
</paragraph>
<paragraph id="8BE436AFFFDA1B19FF40F8A10D7622A2" blockId="5.[151,1437,1016,2015]" pageId="5" pageNumber="544">
Elytra together oval, broadest distinctly in front of middle, EL
<quantity id="4CA39B4AFFDA1B19FC0AF8A00B90220E" box="[909,1012,1880,1907]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="544" unit="mm" value="0.45">0.45 mm</quantity>
, EW
<quantity id="4CA39B4AFFDA1B19FBB4F8A10CFE220E" box="[1075,1178,1881,1907]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="544" unit="mm" value="0.3">0.30 mm</quantity>
, EI 1.50; humeral calli distinct; elytral base with large and deep basal impression; narrow adsutural region in anterior half of each elytron indistinctly impressed; apices of elytra separately rounded. Punctures dense but superficial and very small and diffuse, much less distinct than those on head and pronotum; setae short, sparse and nearly recumbent.
</paragraph>
<paragraph id="8BE436AFFFD91B1AFF40FF61088A25AA" blockId="6.[151,1437,153,899]" pageId="6" pageNumber="545">
Protibia (
<figureCitation id="13602A2AFFD91B1AFEB2FF6109EF25CE" box="[309,395,153,179]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="6" pageNumber="545">Fig. 20</figureCitation>
) not thickened in apical region, with apical comb of four teeth that occupy distomesal margin.
</paragraph>
<paragraph id="8BE436AFFFD91B1AFF40FF190970243E" blockId="6.[151,1437,153,899]" pageId="6" pageNumber="545">
Aedeagus (
<figureCitation id="13602A2AFFD91B1AFECDFF1909C42586" box="[330,416,225,251]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="6" pageNumber="545">Fig. 28</figureCitation>
) moderately slender, AeL
<quantity id="4CA39B4AFFD91B1AFD59FF190B2D2586" box="[734,841,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="545" unit="mm" value="0.13">0.13 mm</quantity>
, endophallus with long median tubular component gradually broadened in its proximal portion to form an ovoid vesicle; parameres strongly sinuate, each with one apical seta.
</paragraph>
<paragraph id="8BE436AFFFD91B1AFF40FEB5090E24AE" blockId="6.[151,1437,153,899]" pageId="6" pageNumber="545">
Female. Externally similar to male, also with thickened protibiae, but lacking apical combs; BL
<quantity id="4CA39B4AFFD91B1AFB7DFEB40DF9241A" box="[1274,1437,332,359]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" metricValueMax="8.5" metricValueMin="8.299999999999999" pageId="6" pageNumber="545" unit="mm" value="0.84" valueMax="0.85" valueMin="0.83">0.83–0.85 mm</quantity>
(mean
<quantity id="4CA39B4AFFD91B1AFF61FE89093524F6" box="[230,337,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="6" pageNumber="545" unit="mm" value="0.83">0.83 mm</quantity>
); HL
<quantity id="4CA39B4AFFD91B1AFE11FE88099A24F6" box="[406,510,368,395]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="545" unit="mm" value="0.15">0.15 mm</quantity>
, HW
<quantity id="4CA39B4AFFD91B1AFDC3FE890A8D24F6" box="[580,745,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="6" pageNumber="545" unit="mm" value="0.195" valueMax="0.2" valueMin="0.19">0.19–0.20 mm</quantity>
(mean
<quantity id="4CA39B4AFFD91B1AFCC6FE890BC824F6" box="[833,940,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="545" unit="mm" value="0.2">0.20 mm</quantity>
), AnL
<quantity id="4CA39B4AFFD91B1AFC79FE890CC724F6" box="[1022,1187,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="6" pageNumber="545" unit="mm" value="0.315" valueMax="0.33" valueMin="0.3">0.30–0.33 mm</quantity>
(mean
<quantity id="4CA39B4AFFD91B1AFB7CFE890D0224F6" box="[1275,1382,369,395]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="545" unit="mm" value="0.32">0.32 mm</quantity>
); PL
<quantity id="4CA39B4AFFD91B1AFF10FE6D095D24D2" box="[151,313,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="6" pageNumber="545" unit="mm" value="0.215" valueMax="0.23" valueMin="0.2">0.20–0.23 mm</quantity>
(mean
<quantity id="4CA39B4AFFD91B1AFE0CFE6D099724D2" box="[395,499,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="545" unit="mm" value="0.22">0.22 mm</quantity>
), PW
<quantity id="4CA39B4AFFD91B1AFDB2FE6C0AB324D2" box="[565,727,404,431]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="6" pageNumber="545" unit="mm" value="0.24" valueMax="0.25" valueMin="0.23">0.23–0.25 mm</quantity>
(mean
<quantity id="4CA39B4AFFD91B1AFCAEFE6D0BF424D2" box="[809,912,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="545" unit="mm" value="0.24">0.24 mm</quantity>
); EL
<quantity id="4CA39B4AFFD91B1AFC4BFE6C0C0A24D2" box="[972,1134,404,431]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="6" pageNumber="545" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
(mean
<quantity id="4CA39B4AFFD91B1AFB47FE6D0D4324D2" box="[1216,1319,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="6" pageNumber="545" unit="mm" value="0.47">0.47 mm</quantity>
), EW
<quantity id="4CA39B4AFFD91B1AFAECFE6D08A124AE" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="545" unit="mm" value="0.33">0.33 mm</quantity>
, EI 1.38–1.46.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD91B1AFF40FE240AE6248A" box="[199,642,476,503]" pageId="6" pageNumber="545" type="distribution">
<paragraph id="8BE436AFFFD91B1AFF40FE240AE6248A" blockId="6.[151,1437,153,899]" box="[199,642,476,503]" pageId="6" pageNumber="545">
<emphasis id="B92FEABDFFD91B1AFF40FE240905248A" bold="true" box="[199,353,476,503]" pageId="6" pageNumber="545">Distribution.</emphasis>
<collectingCountry id="F34C763FFFD91B1AFEE0FE2509CE248A" box="[359,426,477,503]" name="Japan" pageId="6" pageNumber="545">Japan</collectingCountry>
, southern Shikoku.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD91B1AFF40FDF90A8F2766" box="[199,747,513,539]" pageId="6" pageNumber="545" type="etymology">
<paragraph id="8BE436AFFFD91B1AFF40FDF90A8F2766" blockId="6.[151,1437,153,899]" box="[199,747,513,539]" pageId="6" pageNumber="545">
<emphasis id="B92FEABDFFD91B1AFF40FDF9092B2766" bold="true" box="[199,335,513,539]" pageId="6" pageNumber="545">Etymology.</emphasis>
Locotypical, after
<collectingCountry id="F34C763FFFD91B1AFDA3FDF90A0C2766" box="[548,616,513,539]" name="India" pageId="6" pageNumber="545">Kôchi</collectingCountry>
Prefecture.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD91B1AFF40FDDD0B8B26FE" pageId="6" pageNumber="545" type="discussion">
<paragraph id="8BE436AFFFD91B1AFF40FDDD0B8B26FE" blockId="6.[151,1437,153,899]" pageId="6" pageNumber="545">
<emphasis id="B92FEABDFFD91B1AFF40FDDD095F2742" bold="true" box="[199,315,549,575]" pageId="6" pageNumber="545">Remarks.</emphasis>
No other Far Eastern species have so distinctly and densely punctate both the head and pronotum. The male protibial comb composed of only four teeth is similarly small and occupies a similar area on the tibial apex as those in some European species (revised and illustrated by
<bibRefCitation id="EFCA4B5EFFD91B1AFC12FD950C3327FA" author="Orousset, J." box="[917,1111,621,647]" pageId="6" pageNumber="545" pagination="187 - 202" refId="ref9248" refString="Orousset, J. (2017) Le genre Microscydmus Saulcy &amp; Croissandeau, 1893 (Coleoptera, Staphylinidae, Scydmaeninae). L'Entomologiste, 73 (3), 187 - 202." type="journal article" year="2017">Orousset (2017))</bibRefCitation>
, but the aedeagi of the latter do not have the large, sub-basal vesicular structure in their endophalli, and their parameres are nearly straight or weakly curved, whereas
<taxonomicName id="4C5B4D2CFFD91B1AFE2CFD4E0A2127B2" authorityName="Jałoszyński" authorityYear="2019" box="[427,581,693,719]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="kochiensis">
<emphasis id="B92FEABDFFD91B1AFE2CFD4E0A2127B2" box="[427,581,693,719]" italics="true" pageId="6" pageNumber="545">M. kochiensis</emphasis>
</taxonomicName>
has conspicuously recurved, sinuate parameres. The shape of parameres and the sub-basal vesicular endophallic structure are similar in
<taxonomicName id="4C5B4D2CFFD91B1AFCA8FD220BAE278E" authorityName="Jałoszyński" authorityYear="2019" box="[815,970,729,755]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="kochiensis">
<emphasis id="B92FEABDFFD91B1AFCA8FD220BAE278E" box="[815,970,729,755]" italics="true" pageId="6" pageNumber="545">M. kochiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5B4D2CFFD91B1AFB85FD220CEE278E" authorityName="Jałoszyński" authorityYear="2019" box="[1026,1162,729,755]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="elytratus">
<emphasis id="B92FEABDFFD91B1AFB85FD220CEE278E" box="[1026,1162,729,755]" italics="true" pageId="6" pageNumber="545">M. elytratus</emphasis>
</taxonomicName>
; both species have been collected in the same area. However, despite similarities in the aedeagi, their body length is conspicuously different (
<taxonomicName id="4C5B4D2CFFD91B1AFF26FCDA09582646" authorityName="Jałoszyński" authorityYear="2019" box="[161,316,801,827]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="kochiensis">
<emphasis id="B92FEABDFFD91B1AFF26FCDA09582646" box="[161,316,801,827]" italics="true" pageId="6" pageNumber="545">M. kochiensis</emphasis>
</taxonomicName>
is much smaller than
<taxonomicName id="4C5B4D2CFFD91B1AFDBFFCDA0AA52646" authorityName="Jałoszyński" authorityYear="2019" box="[568,705,801,827]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="elytratus">
<emphasis id="B92FEABDFFD91B1AFDBFFCDA0AA52646" box="[568,705,801,827]" italics="true" pageId="6" pageNumber="545">M. elytratus</emphasis>
</taxonomicName>
), and male secondary sexual characters differ profoundly. In
<taxonomicName id="4C5B4D2CFFD91B1AFAF9FCDA089D2622" authorityName="Jałoszyński" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="elytratus">
<emphasis id="B92FEABDFFD91B1AFAF9FCDA089D2622" italics="true" pageId="6" pageNumber="545">M. elytratus</emphasis>
</taxonomicName>
the elytra bear a posteromedian patch of modified setae (absent in
<taxonomicName id="4C5B4D2CFFD91B1AFC6BFCBE0CE32622" authorityName="Jałoszyński" authorityYear="2019" box="[1004,1159,837,863]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="kochiensis">
<emphasis id="B92FEABDFFD91B1AFC6BFCBE0CE32622" box="[1004,1159,837,863]" italics="true" pageId="6" pageNumber="545">M. kochiensis</emphasis>
</taxonomicName>
); and the protibial comb in
<taxonomicName id="4C5B4D2CFFD91B1AFF33FC92095F26FE" authorityName="Jałoszyński" authorityYear="2019" box="[180,315,873,899]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="elytratus">
<emphasis id="B92FEABDFFD91B1AFF33FC92095F26FE" box="[180,315,873,899]" italics="true" pageId="6" pageNumber="545">M. elytratus</emphasis>
</taxonomicName>
is composed of six teeth, vs. only four teeth in
<taxonomicName id="4C5B4D2CFFD91B1AFCC9FC920B8D26FE" authorityName="Jałoszyński" authorityYear="2019" box="[846,1001,873,899]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="kochiensis">
<emphasis id="B92FEABDFFD91B1AFCC9FC920B8D26FE" box="[846,1001,873,899]" italics="true" pageId="6" pageNumber="545">M. kochiensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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