231 lines
30 KiB
XML
231 lines
30 KiB
XML
<document id="2D8BA6EA000DD115FB84578C03942D9B" ID-CLB-Dataset="32391" ID-DOI="10.11646/zootaxa.4306.2.6" ID-GBIF-Dataset="647384bd-cf6a-4546-a70d-1a813f4258cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="843798" ID-ZooBank="D45C3578-0245-4CD0-98BE-79DCB35423AD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502868504205" checkinUser="plazi" docAuthor="Schlüter, Nils & Wiese, Frank" docDate="2017" docId="03F28781FF8DE17087E56987C0ABFEE1" docLanguage="en" docName="zootaxa.4306.2.6.pdf.imf" docOrigin="Zootaxa 4306 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vologesia toucasi" docType="treatment" docVersion="8" lastPageNumber="267" masterDocId="FFCBFFF9FF8BE17787726B50C17AFF84" masterDocTitle="Late Cretaceous species of Vologesia (Echinoidea, Cassiduloida) from northern Spain" masterLastPageNumber="270" masterPageNumber="261" pageNumber="267" updateTime="1698466706665" updateUser="ExternalLinkService">
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<mods:title id="5D8EDF390A322F8E83BAE093FE945DDC">Late Cretaceous species of Vologesia (Echinoidea, Cassiduloida) from northern Spain</mods:title>
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<mods:namePart id="F5B617D68BD7F7499390673DEEA7E2DB">Schlüter, Nils</mods:namePart>
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<mods:namePart id="A2A495E6050166400CB8EFDB427B2CE1">Wiese, Frank</mods:namePart>
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<mods:title id="0042AA5E24F43A710757EA1D8C6BDB2F">Zootaxa</mods:title>
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<mods:date id="AE540C691867082F3FC3791BC6511B0A">2017</mods:date>
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<treatment id="03F28781FF8DE17087E56987C0ABFEE1" ID-DOI="http://doi.org/10.5281/zenodo.6029224" ID-GBIF-Taxon="132749353" ID-Zenodo-Dep="6029224" LSID="urn:lsid:plazi:treatment:03F28781FF8DE17087E56987C0ABFEE1" httpUri="http://treatment.plazi.org/id/03F28781FF8DE17087E56987C0ABFEE1" lastPageId="7" lastPageNumber="267" pageId="6" pageNumber="267">
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<subSubSection id="C341651CFF8DE17187E56987C019FC97" pageId="6" pageNumber="267" type="nomenclature">
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<paragraph id="8BE43697FF8DE17187E56987C343FD76" blockId="6.[151,569,727,787]" box="[151,569,727,754]" pageId="6" pageNumber="267">
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<heading id="D0AC81FBFF8DE17187E56987C343FD76" bold="true" box="[151,569,727,754]" fontSize="11" level="1" pageId="6" pageNumber="267" reason="1">
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<taxonomicName id="4C5B4D14FF8DE17187E56987C343FD76" authority="(Lambert, 1911)" baseAuthorityName="Lambert" baseAuthorityYear="1911" box="[151,569,727,754]" class="Echinoidea" family="Echinolampadidae" genus="Vologesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="6" pageNumber="267" phylum="Echinodermata" rank="species" species="toucasi">
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||
<emphasis id="B92FEA85FF8DE17187E56987C343FD76" bold="true" box="[151,569,727,754]" pageId="6" pageNumber="267">
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<emphasis id="B92FEA85FF8DE17187E56987C012FD75" bold="true" box="[151,360,727,753]" italics="true" pageId="6" pageNumber="267">Vologesia toucasi</emphasis>
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(
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<bibRefCitation id="EFCA4B66FF8DE17186056988C34AFD76" author="Lambert" box="[375,560,728,754]" pageId="6" pageNumber="267" refString="Lambert, J. (1911) Etude sur les Echinides cretaces de Rennes les Bains et des Corbieres. Bulletin de la Societ d'Etudes scientifiques de l'Aude, 22, 1 - 120." type="journal article" year="1911">Lambert, 1911</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BE43697FF8DE17187E569AAC019FC97" blockId="6.[151,569,727,787]" box="[151,355,762,787]" pageId="6" pageNumber="267">
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(
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<figureCitation id="13602A12FF8DE17187ED69AAC196FC97" box="[159,236,762,787]" captionStart="FIGURE 5" captionStartId="5.[151,250,1316,1338]" captionTargetBox="[173,1400,294,1274]" captionTargetId="figure@5.[151,1436,265,1294]" captionTargetPageId="5" captionText="FIGURE 5. Drawings of plating style; A – B, D, F, Vologesia rollingstones sp. nov.; A, D, MB. E. 6226, A: oral view, D: detail of peristomial area; B, holotype, MB. E. 6224, apical view; F, MB. E. 6227, apical disc; A – B, D, F: Langre, Cantabria, northern Spain, Sponge Rhythmites (Scaphites hippocrepis III Zone, lower Campanian); C, E, G, Vologesia toucasi (Lambert, 1911); C, MB. E. 11644, C: oral view, E: detail of peristomial area; G, MB. E. 11645, apical disc; C, E, G: Langre, Cantabria, northern Spain, Bed 200, upper Santonian (Placenticeras polyopsis Zone). Scale bars equal 10 mm (A – C) and 2 mm (D – G)." httpUri="https://zenodo.org/record/843808/files/figure.png" pageId="6" pageNumber="267">Figs. 5</figureCitation>
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C, E, G, 7)
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</paragraph>
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</subSubSection>
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<subSubSection id="C341651CFF8DE17187E56812C2BBFCF3" pageId="6" pageNumber="267" type="description">
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<paragraph id="8BE43697FF8DE17187E56812C278FCDC" blockId="6.[151,961,834,887]" box="[151,770,834,856]" pageId="6" pageNumber="267">
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1911
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<taxonomicName id="4C5B4D14FF8DE17187BD6812C377FCDC" authority="Lambert" authorityName="Lambert" authorityYear="1911" box="[207,525,834,856]" class="Echinoidea" family="Clypeolampadidae" genus="Clypeolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="6" pageNumber="267" phylum="Echinodermata" rank="species" species="toucasi">
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<emphasis id="B92FEA85FF8DE17187BD6812C0D5FCDC" box="[207,431,834,856]" italics="true" pageId="6" pageNumber="267">Clypeolampas toucasi</emphasis>
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Lambert
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</taxonomicName>
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, p. 155, pl. 3,
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<figureCitation id="13602A12FF8DE17185EF6812C384FCDC" box="[669,766,834,856]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1277,1299]" captionStartId-1="3.[151,250,1875,1897]" captionStartId-2="4.[151,250,1093,1115]" captionTargetBox-0="[339,1248,711,1252]" captionTargetBox-1="[236,1343,201,1829]" captionTargetBox-2="[151,1436,193,1071]" captionTargetId-0="figure@2.[339,1248,711,1252]" captionTargetId-1="figure@3.[205,1381,193,1853]" captionTargetId-2="figure@4.[151,1436,193,1071]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Simplified map of the vicinity of the village of Langre, showing the origins of Vologesia toucasi (Lambert, 1911) (A) and Vologesia rollingstones sp. nov. (B)." captionText-1="FIGURE 2. Stratigraphy and lithology of the Langre section." captionText-2="FIGURE 3. Vologesia rollingstones sp. nov.; A – C, holotype, MB. E. 6224; B, paratype, MB. E. 6225; in oral (A, D), apical (B, E) and left lateral (C, F) views. Langre, Cantabria, northern Spain, Sponge Rhythmites (Scaphites hippocrepis III Zone, lower Campanian). Both specimens coated with ammonium chloride prior to photography. Scale bar equals 10 mm." httpUri-0="https://zenodo.org/record/843800/files/figure.png" httpUri-1="https://zenodo.org/record/843802/files/figure.png" httpUri-2="https://zenodo.org/record/843804/files/figure.png" pageId="6" pageNumber="267">Figs. 1–3</figureCitation>
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.
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</paragraph>
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<paragraph id="8BE43697FF8DE17187E56831C2BBFCF3" blockId="6.[151,961,834,887]" box="[151,961,865,887]" pageId="6" pageNumber="267">
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2014
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<taxonomicName id="4C5B4D14FF8DE17187A26832C37FFCF3" authority="(Lambert)" authorityName="Lambert" box="[208,517,865,887]" class="Echinoidea" family="Clypeolampadidae" genus="Hungaresia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="6" pageNumber="267" phylum="Echinodermata" rank="species" species="toucasi">
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<emphasis id="B92FEA85FF8DE17187A26832C0ECFCF3" box="[208,406,866,887]" italics="true" pageId="6" pageNumber="267">Hungaresia toucasi</emphasis>
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(Lambert)
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</taxonomicName>
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, Saura & García p. 306, pl. 104,
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<figureCitation id="13602A12FF8DE17184206831C2E9FCF3" box="[850,915,865,887]" captionStart="FIGURE 1" captionStartId="2.[151,250,1277,1299]" captionTargetBox="[339,1248,711,1252]" captionTargetId="figure@2.[339,1248,711,1252]" captionTargetPageId="2" captionText="FIGURE 1. Simplified map of the vicinity of the village of Langre, showing the origins of Vologesia toucasi (Lambert, 1911) (A) and Vologesia rollingstones sp. nov. (B)." httpUri="https://zenodo.org/record/843800/files/figure.png" pageId="6" pageNumber="267">Figs. 1</figureCitation>
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a–c.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C341651CFF8DE17187E568F4C56DFB58" pageId="6" pageNumber="267" type="materials_examined">
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<paragraph id="8BE43697FF8DE17187E568F4C198FBF4" blockId="6.[151,1437,932,2037]" pageId="6" pageNumber="267">
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<emphasis id="B92FEA85FF8DE17187E568F4C1A2FC39" bold="true" box="[151,216,932,957]" pageId="6" pageNumber="267">Type.</emphasis>
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Holotype, by monotypy, is the specimen illustrated by
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<bibRefCitation id="EFCA4B66FF8DE171841468F4C59AFC38" author="Lambert" box="[870,1248,932,957]" pageId="6" pageNumber="267" refString="Lambert, J. (1911) Etude sur les Echinides cretaces de Rennes les Bains et des Corbieres. Bulletin de la Societ d'Etudes scientifiques de l'Aude, 22, 1 - 120." type="journal article" year="1911">
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Lambert (1911, pl. 3,
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<figureCitation id="13602A12FF8DE171831B68F4C5ACFC38" box="[1129,1238,932,957]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1277,1299]" captionStartId-1="3.[151,250,1875,1897]" captionStartId-2="4.[151,250,1093,1115]" captionTargetBox-0="[339,1248,711,1252]" captionTargetBox-1="[236,1343,201,1829]" captionTargetBox-2="[151,1436,193,1071]" captionTargetId-0="figure@2.[339,1248,711,1252]" captionTargetId-1="figure@3.[205,1381,193,1853]" captionTargetId-2="figure@4.[151,1436,193,1071]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Simplified map of the vicinity of the village of Langre, showing the origins of Vologesia toucasi (Lambert, 1911) (A) and Vologesia rollingstones sp. nov. (B)." captionText-1="FIGURE 2. Stratigraphy and lithology of the Langre section." captionText-2="FIGURE 3. Vologesia rollingstones sp. nov.; A – C, holotype, MB. E. 6224; B, paratype, MB. E. 6225; in oral (A, D), apical (B, E) and left lateral (C, F) views. Langre, Cantabria, northern Spain, Sponge Rhythmites (Scaphites hippocrepis III Zone, lower Campanian). Both specimens coated with ammonium chloride prior to photography. Scale bar equals 10 mm." httpUri-0="https://zenodo.org/record/843800/files/figure.png" httpUri-1="https://zenodo.org/record/843802/files/figure.png" httpUri-2="https://zenodo.org/record/843804/files/figure.png" pageId="6" pageNumber="267">Figs. 1–3</figureCitation>
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)
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</bibRefCitation>
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, in the Lambert Collection at the Muséum national d’Histoire naturelle, Paris
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<emphasis id="B92FEA85FF8DE171843D6897C223FC64" bold="true" box="[847,857,967,992]" pageId="6" pageNumber="267">(</emphasis>
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MNHN.F.
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<accessionNumber id="9408AB74FF8DE17184BB6898C566FC64" box="[969,1052,968,992]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J01723" pageId="6" pageNumber="267" type="EnaNcbi">J01723</accessionNumber>
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) (not seen). It is from Sougraigne in the Corbières (Aude, France), having been collected from the “couches
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<taxonomicName id="4C5B4D14FF8DE171835668BDC5E6FB80" box="[1060,1180,1005,1028]" class="Bivalvia" family="Hippuritidae" genus="Hippurites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hippuritida" pageId="6" pageNumber="267" phylum="Mollusca" rank="genus">
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<emphasis id="B92FEA85FF8DE171835668BDC5E6FB80" box="[1060,1180,1005,1028]" italics="true" pageId="6" pageNumber="267">Hippurites</emphasis>
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</taxonomicName>
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... avec
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<taxonomicName id="4C5B4D14FF8DE171827868BCC0A6FBAC" authority="Grossouvrei. Lambert (1911)" authorityName="Grossouvrei. Lambert" authorityYear="1911" class="Cephalopoda" family="Belemnopseidae" genus="Actinocamax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Belemnitida" pageId="6" pageNumber="267" phylum="Mollusca" rank="genus">
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<emphasis id="B92FEA85FF8DE171827868BCC065FBA3" italics="true" pageId="6" pageNumber="267">Actinocamax Grossouvrei</emphasis>
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.
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<bibRefCitation id="EFCA4B66FF8DE171865F6F5FC0A6FBAC" author="Lambert" box="[301,476,1039,1064]" pageId="6" pageNumber="267" refString="Lambert, J. (1911) Etude sur les Echinides cretaces de Rennes les Bains et des Corbieres. Bulletin de la Societ d'Etudes scientifiques de l'Aude, 22, 1 - 120." type="journal article" year="1911">Lambert (1911)</bibRefCitation>
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</taxonomicName>
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considered those strata to be of Campanian age. According to the revised stratigraphy by
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<bibRefCitation id="EFCA4B66FF8DE17187CC6F64C09AFBC8" box="[190,480,1076,1101]" pageId="6" pageNumber="267" refString="Christensen, W. K., Bilotte, M. & Hansotte, M. (1993) Additional Late Cretaceous belemnitellids from the Corbieres, French Pyrenees and ammonite biostratigraphy of the Santonian. Cretaceous Research, 14, 101 - 106. https: // doi. org / 10.1006 / cres. 1993.1008" type="journal article">
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Christensen
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<emphasis id="B92FEA85FF8DE171863F6F65C0F3FBC9" box="[333,393,1076,1101]" italics="true" pageId="6" pageNumber="267">et al.</emphasis>
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(1993)
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</bibRefCitation>
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, these levels are of late Santonian age (
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<taxonomicName id="4C5B4D14FF8DE17184CB6F64C5B2FBC9" box="[953,1224,1076,1101]" class="Cephalopoda" family="Placenticeratidae" genus="Placenticeras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="6" pageNumber="267" phylum="Mollusca" rank="species" species="polyopsis">
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<emphasis id="B92FEA85FF8DE17184CB6F64C5B2FBC9" box="[953,1224,1076,1101]" italics="true" pageId="6" pageNumber="267">Placenticeras polyopsis</emphasis>
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||
</taxonomicName>
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/
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<taxonomicName id="4C5B4D14FF8DE17183A26F65C1ACFBF4" authority="Zone" authorityName="Zone" class="Cephalopoda" family="Collignoniceratidae" genus="Texanites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="6" pageNumber="267" phylum="Mollusca" rank="species" species="gallicus">
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||
<emphasis id="B92FEA85FF8DE17183A26F65C4E6FBC9" box="[1232,1436,1076,1101]" italics="true" pageId="6" pageNumber="267">Texanites gallicus</emphasis>
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Zone
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||
</taxonomicName>
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||
).
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</paragraph>
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<paragraph id="8BE43697FF8DE17187B56F2CC56DFB58" blockId="6.[151,1437,932,2037]" pageId="6" pageNumber="267">
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||
<materialsCitation id="3B333CCAFF8DE17187B56F2CC56DFB58" ID-GBIF-Occurrence="1586063357" collectorName="Material. A" country="Germany" location="upper Santonian" pageId="6" pageNumber="267" specimenCount="1" stateProvince="Berlin">
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||
<collectorName id="26AE5341FF8DE17187B56F2CC02BFB10" box="[199,337,1148,1173]" pageId="6" pageNumber="267">
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||
<emphasis id="B92FEA85FF8DE17187B56F2CC04CFB11" bold="true" box="[199,310,1148,1173]" pageId="6" pageNumber="267">Material.</emphasis>
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||
A
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</collectorName>
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||
single near-complete and four fragmentarily preserved specimens (MB.E.11463-11467) from the top of Bed 200,
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||
<location id="8E84604CFF8DE17186166FF0C35AFB3C" LSID="urn:lsid:plazi:treatment:03F28781FF8DE17087E56987C0ABFEE1:8E84604CFF8DE17186166FF0C35AFB3C" box="[356,544,1183,1208]" country="Germany" name="upper Santonian" pageId="6" pageNumber="267" stateProvince="Berlin">upper Santonian</location>
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||
(
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||
<taxonomicName id="4C5B4D14FF8DE17185446FCFC2EBFB3C" authority="Zone" authorityName="Zone" box="[566,913,1183,1208]" class="Cephalopoda" family="Placenticeratidae" genus="Placenticeras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="6" pageNumber="267" phylum="Mollusca" rank="species" species="polyopsis">
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||
<emphasis id="B92FEA85FF8DE17185446FCFC23DFB3C" box="[566,839,1183,1208]" italics="true" pageId="6" pageNumber="267">Placenticeras polyopsis</emphasis>
|
||
Zone
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||
</taxonomicName>
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||
), deposited in the Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung (
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||
<collectingRegion id="499FF875FF8DE171843B6F94C2E9FB59" box="[841,915,1220,1245]" country="Germany" name="Berlin" pageId="6" pageNumber="267">Berlin</collectingRegion>
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||
,
|
||
<collectingCountry id="F34C7607FF8DE17184EC6F94C576FB58" box="[926,1036,1220,1244]" name="Germany" pageId="6" pageNumber="267">Germany</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C341651CFF8DE17187B56FB7C335FAE9" pageId="6" pageNumber="267" type="diagnosis">
|
||
<paragraph id="8BE43697FF8DE17187B56FB7C335FAE9" blockId="6.[151,1437,932,2037]" pageId="6" pageNumber="267">
|
||
<emphasis id="B92FEA85FF8DE17187B56FB7C03BFA84" bold="true" box="[199,321,1255,1280]" pageId="6" pageNumber="267">Diagnosis.</emphasis>
|
||
Small species with a domed to conical test and a flattened oral surface. The tetrabasal apical disc, with a prominent madreporite, is in a central position. The pentagonal, laterally elongated peristome is positioned close to the anterior margin. The phyllodes are short and slightly sunken, with an inner and an outer series of pore pairs; bourrelets are weakly developed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C341651CFF8DE17187B56E27C532F829" pageId="6" pageNumber="267" type="description">
|
||
<paragraph id="8BE43697FF8DE17187B56E27C359F9C1" blockId="6.[151,1437,932,2037]" pageId="6" pageNumber="267">
|
||
<emphasis id="B92FEA85FF8DE17187B56E27C023FA14" bold="true" box="[199,345,1399,1424]" pageId="6" pageNumber="267">Description.</emphasis>
|
||
<taxonomicName id="4C5B4D14FF8DE17186126E27C35FFA14" baseAuthorityName="Lambert" baseAuthorityYear="1911" box="[352,549,1399,1424]" class="Echinoidea" family="Echinolampadidae" genus="Vologesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="6" pageNumber="267" phylum="Echinodermata" rank="species" species="toucasi">
|
||
<emphasis id="B92FEA85FF8DE17186126E27C35FFA14" box="[352,549,1399,1424]" italics="true" pageId="6" pageNumber="267">Vologesia toucasi</emphasis>
|
||
</taxonomicName>
|
||
is a small species with a maximum test length of
|
||
<emphasis id="B92FEA85FF8DE171832E6E29C512FA14" box="[1116,1128,1401,1424]" italics="true" pageId="6" pageNumber="267">c</emphasis>
|
||
.
|
||
<quantity id="4CA39B72FF8DE17183056E27C5B0FA0B" box="[1143,1226,1399,1424]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="267" unit="mm" value="35.0">35 mm</quantity>
|
||
. The oval tests are domed to conical in profile, with its longest axis along the anterior-posterior direction in cross section. The ambitus is positioned close to the oral surface, in the lower quarter of test height. The apical disc is positioned at 47% of test length to the anterior margin. The apical disc is not completely preserved; it was presumably tetrabasal, with the madreporite formed by the large genital plate 2, occupying the centre of the apical disc, while other genital plates are much reduced in size (
|
||
<figureCitation id="13602A12FF8DE17186CF6D7CC085F9C1" box="[445,511,1580,1605]" captionStart="FIGURE 5" captionStartId="5.[151,250,1316,1338]" captionTargetBox="[173,1400,294,1274]" captionTargetId="figure@5.[151,1436,265,1294]" captionTargetPageId="5" captionText="FIGURE 5. Drawings of plating style; A – B, D, F, Vologesia rollingstones sp. nov.; A, D, MB. E. 6226, A: oral view, D: detail of peristomial area; B, holotype, MB. E. 6224, apical view; F, MB. E. 6227, apical disc; A – B, D, F: Langre, Cantabria, northern Spain, Sponge Rhythmites (Scaphites hippocrepis III Zone, lower Campanian); C, E, G, Vologesia toucasi (Lambert, 1911); C, MB. E. 11644, C: oral view, E: detail of peristomial area; G, MB. E. 11645, apical disc; C, E, G: Langre, Cantabria, northern Spain, Bed 200, upper Santonian (Placenticeras polyopsis Zone). Scale bars equal 10 mm (A – C) and 2 mm (D – G)." httpUri="https://zenodo.org/record/843808/files/figure.png" pageId="6" pageNumber="267">Fig. 5</figureCitation>
|
||
G).
|
||
</paragraph>
|
||
<paragraph id="8BE43697FF8DE17187B56D1FC0A2F934" blockId="6.[151,1437,932,2037]" pageId="6" pageNumber="267">Petaloid ambulacra are present adapically; pore counts are not available due to poor preservation. The straight petals are relatively short, not reaching below the upper third of test height. Conjugate anisopores can be recognised within the petals.</paragraph>
|
||
<paragraph id="8BE43697FF8DE17187B56DECC240F8E1" blockId="6.[151,1437,932,2037]" pageId="6" pageNumber="267">
|
||
The small peristome is laterally elongated and has a pentagonal outline. It is positioned anteriorly, with a distance to the anterior margin of 29% of test length (at a length of
|
||
<quantity id="4CA39B72FF8DE17184C66DB0C565F97C" box="[948,1055,1760,1784]" metricMagnitude="-2" metricUnit="m" metricValue="2.98" pageId="6" pageNumber="267" unit="mm" value="29.8">29.8 mm</quantity>
|
||
). A weakly pronounced floscelle structure can be seen; the bourrelets are only weakly swollen, in combination with only slightly sunken phyllodes. The phyllodes are short, with an outer series of 10–11 pores, including small buccal pores, and an inner series of 3–4 pores (see
|
||
<figureCitation id="13602A12FF8DE171864D6C1CC0F9F8E0" box="[319,387,1868,1893]" captionStart="FIGURE 4" captionStartId="4.[151,250,1641,1663]" captionTargetBox="[340,1247,1226,1620]" captionTargetId="figure@4.[340,1247,1226,1620]" captionTargetPageId="4" captionText="FIGURE 4. Vologesia rollingstones sp. nov.; A, holotype, MB. E. 6224, detail of apical area; B, paratype, MB. E. 6225, detail of peristomial area; both from Langre, Cantabria, northern Spain, Sponge Rhythmites (Scaphites hippocrepis III Zone, lower Campanian). Both specimens coated with ammonium chloride prior to photography. Scale bars equal 5 mm." httpUri="https://zenodo.org/record/843806/files/figure.png" pageId="6" pageNumber="267">Fig. 4</figureCitation>
|
||
C). The oval periproct is inframarginal.
|
||
</paragraph>
|
||
<paragraph id="8BE43697FF8DE17187B56C3FC532F829" blockId="6.[151,1437,932,2037]" pageId="6" pageNumber="267">The tubercules generally are small, largest on the oral surface and decreasing in size in aboral direction. A narrow, granular band without tubercles extends from the peristome to the periproct.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C341651CFF8DE17087B56CE8C0ABFEE1" lastPageId="7" lastPageNumber="268" pageId="6" pageNumber="267" type="discussion">
|
||
<paragraph id="8BE43697FF8DE17087B56CE8C0ABFEE1" blockId="6.[151,1437,932,2037]" lastBlockId="7.[151,1436,151,357]" lastPageId="7" lastPageNumber="268" pageId="6" pageNumber="267">
|
||
<emphasis id="B92FEA85FF8DE17187B56CE8C036F855" bold="true" box="[199,332,1976,2001]" pageId="6" pageNumber="267">Discussion.</emphasis>
|
||
Our record of
|
||
<taxonomicName id="4C5B4D14FF8DE171869D6CE9C321F854" baseAuthorityName="Lambert" baseAuthorityYear="1911" box="[495,603,1977,2000]" class="Echinoidea" family="Echinolampadidae" genus="Vologesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="6" pageNumber="267" phylum="Echinodermata" rank="species" species="toucasi">
|
||
<emphasis id="B92FEA85FF8DE171869D6CE9C321F854" box="[495,603,1977,2000]" italics="true" pageId="6" pageNumber="267">V. toucasi</emphasis>
|
||
</taxonomicName>
|
||
from the upper Santonian of Langre is in accordance with the stratigraphic position of the
|
||
<typeStatus id="54E08835FF8DE171863A6C8DC002F871" box="[328,376,2013,2037]" pageId="6" pageNumber="267">type</typeStatus>
|
||
series from the Corbières, southern
|
||
<collectingCountry id="F34C7607FF8DE17184526C8DC216F870" box="[800,876,2013,2036]" name="France" pageId="6" pageNumber="267">France</collectingCountry>
|
||
and extends the known geographic range of the species to
|
||
<collectingCountry id="F34C7607FF8CE170867E6BC7C035FF34" box="[268,335,151,176]" name="Spain" pageId="7" pageNumber="268">Spain</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C5B4D14FF8CE17086286BC7C35AFF34" baseAuthorityName="Lambert" baseAuthorityYear="1911" box="[346,544,151,176]" class="Echinoidea" family="Echinolampadidae" genus="Vologesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="7" pageNumber="268" phylum="Echinodermata" rank="species" species="toucasi">
|
||
<emphasis id="B92FEA85FF8CE17086286BC7C35AFF34" box="[346,544,151,176]" italics="true" pageId="7" pageNumber="268">Vologesia toucasi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C5B4D14FF8CE17085C76BC9C218FF34" authority="Schlüter & Wiese, 2017" authorityName="Schlüter & Wiese" authorityYear="2017" box="[693,866,151,176]" class="Echinoidea" family="Echinolampadidae" genus="Vologesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="7" pageNumber="268" phylum="Echinodermata" rank="species" species="rollingstones" status="sp. nov.">
|
||
<emphasis id="B92FEA85FF8CE17085C76BC9C218FF34" box="[693,866,151,176]" italics="true" pageId="7" pageNumber="268">V. rollingstones</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92FEA85FF8CE170841B6BC7C2B9FF34" bold="true" box="[873,963,151,176]" pageId="7" pageNumber="268">
|
||
<taxonomicNameLabel id="A21C57FEFF8CE170841B6BC7C2B9FF34" box="[873,963,151,176]" pageId="7" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by its subconical to conical profile and in phyllode structure that generally reveals a floscelle on account of more pronounced bourrelets. This species has a more inflated and circular shape in comparison to
|
||
<taxonomicName id="4C5B4D14FF8CE17085B56BB1C20AFF7C" authorityName="Ali" authorityYear="1989" box="[711,880,223,248]" class="Echinoidea" family="Echinolampadidae" genus="Vologesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="7" pageNumber="268" phylum="Echinodermata" rank="species" species="rawdahensis">
|
||
<emphasis id="B92FEA85FF8CE17085B56BB1C20AFF7C" box="[711,880,223,248]" italics="true" pageId="7" pageNumber="268">V. rawdahensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B4D14FF8CE17084D46BB1C577FF7C" authorityName="Cotteau & Gauthier" authorityYear="1895" box="[934,1037,225,248]" class="Echinoidea" family="Echinolampadidae" genus="Vologesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="7" pageNumber="268" phylum="Echinodermata" rank="species" species="tataosi">
|
||
<emphasis id="B92FEA85FF8CE17084D46BB1C577FF7C" box="[934,1037,225,248]" italics="true" pageId="7" pageNumber="268">V. tataosi</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the apical disc is more central.
|
||
<bibRefCitation id="EFCA4B66FF8CE17087806A54C0D9FE98" author="Lambert" box="[242,419,260,285]" pageId="7" pageNumber="268" refString="Lambert, J. (1919) Echinides fossiles des environs de Santander recueillis par M. L. Mengaud. Annales de la Societ Linneenne de Lyon, Nouvelle Serie, 66, 1 - 32." type="journal article" year="1919">Lambert (1919)</bibRefCitation>
|
||
mentioned
|
||
<taxonomicName id="4C5B4D14FF8CE170855A6A54C2B6FE98" authority="(Des Moulins, 1837)" baseAuthorityName="Des Moulins" baseAuthorityYear="1837" box="[552,972,260,285]" class="Echinoidea" family="Echinolampadidae" genus="Vologesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="7" pageNumber="268" phylum="Echinodermata" rank="species" species="acuta">
|
||
<emphasis id="B92FEA85FF8CE170855A6A54C3A0FE98" box="[552,730,260,285]" italics="true" pageId="7" pageNumber="268">Vologesia acuta</emphasis>
|
||
(Des Moulins, 1837)
|
||
</taxonomicName>
|
||
from the Santander region. This is better referred to
|
||
<taxonomicName id="4C5B4D14FF8CE17086666A77C0CEFEC4" authorityName="Pomel" authorityYear="1869" box="[276,436,295,320]" class="Echinoidea" family="Clypeolampadidae" genus="Clypeolampas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="7" pageNumber="268" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B92FEA85FF8CE17086666A77C0CEFEC4" box="[276,436,295,320]" italics="true" pageId="7" pageNumber="268">Clypeolampas</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFCA4B66FF8CE17086836A77C306FEC4" author="Smith" box="[497,636,295,320]" pageId="7" pageNumber="268" refString="Smith, A. B. (1995) Late Campanian-Maastrichtian echinoids from the United Arab Emirates - Oman border region. Bulletin of the British Museum (Natural History), Geology, 51, 121 - 240." type="journal article" year="1995">Smith, 1995</bibRefCitation>
|
||
) and differs to
|
||
<taxonomicName id="4C5B4D14FF8CE17084586A77C2EDFEC4" authorityName="Cotteau & Gauthier" authorityYear="1895" box="[810,919,295,320]" class="Echinoidea" family="Echinolampadidae" genus="Vologesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cassiduloida" pageId="7" pageNumber="268" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B92FEA85FF8CE17084586A77C2EDFEC4" box="[810,919,295,320]" italics="true" pageId="7" pageNumber="268">Vologesia</emphasis>
|
||
</taxonomicName>
|
||
by a more centrally positioned peristome and better-developed bourrelets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |