275 lines
37 KiB
XML
275 lines
37 KiB
XML
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<mods:title id="B45939FD191131AA44809C00BA9205A5">Re-description of Grammapsychops lebedevi Martynova, 1954 (Neuroptera: Psychopsidae) with notes on the Late Cretaceous psychopsoids</mods:title>
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<taxonomicName id="4C624285FFBA84187387028529D0C896" ID-CoL="3H5XZ" authority="Martynova, 1954" authorityName="Martynova" authorityYear="1954" box="[151,693,151,178]" class="Insecta" family="Psychopsidae" genus="Grammapsychops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="lebedevi">
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<emphasis id="B916E514FFBA84187387028529D0C896" bold="true" box="[151,693,151,178]" pageId="2" pageNumber="583">
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<emphasis id="B916E514FFBA8418738702852ABDC895" bold="true" box="[151,472,151,177]" italics="true" pageId="2" pageNumber="583">Grammapsychops lebedevi</emphasis>
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<bibRefCitation id="EFF344F7FFBA841872F0028A29D0C896" author="Martynova, O. M." box="[480,693,152,178]" pageId="2" pageNumber="583" pagination="1167 - 1169" refId="ref7737" refString="Martynova, O. M. (1954) Neuroptera from the Cretaceous deposits of Siberia. Doklady Akademii Nauk SSSR, 94, 1167 - 1169. [in Russian]" type="journal article" year="1954">Martynova, 1954</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3786A8DFFBA8418738702A82B9CC8F7" box="[151,249,186,211]" pageId="2" pageNumber="583" type="description">
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<paragraph id="8BDD3906FFBA8418738702A82B9CC8F7" blockId="2.[151,693,151,211]" box="[151,249,186,211]" pageId="2" pageNumber="583">
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<figureCitation id="13592583FFBA8418738702A82B9CC8F7" box="[151,249,186,211]" captionStart-0="FIGURES 1" captionStart-1="FIGURES 2" captionStart-2="FIGURES 3" captionStartId-0="2.[151,264,1504,1526]" captionStartId-1="3.[151,264,1939,1961]" captionStartId-2="4.[151,264,1587,1609]" captionTargetBox-0="[171,1425,425,1488]" captionTargetBox-1="[173,1417,517,1926]" captionTargetBox-2="[152,1418,908,1558]" captionTargetId-0="figure@2.[171,1426,425,1488]" captionTargetId-1="figure@3.[173,1417,517,1926]" captionTargetId-2="figure@4.[151,1436,895,1578]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1. Grammapsychops lebedevi Martynova, 1954, holotype PIN 846/1-2. A, specimen as preserved; B, original field label; C, PIN label. Scale bar is 2 mm." captionText-1="FIGURES 2. Grammapsychops lebedevi Martynova, 1954, holotype PIN 846/1-2. A, proximal part of the forewing; B, a portion of RP and its branches (some setal sockets on veins are shown by red arrows). Scale bars are 2 mm (A), 0.4 mm (B)." captionText-2="FIGURES 3. Forewing venation of Grammapsychops lebedevi Martynova, 1954, holotype PIN 846/1-2. Scale bar is 2 mm." httpUri-0="https://zenodo.org/record/2610785/files/figure.png" httpUri-1="https://zenodo.org/record/2610787/files/figure.png" httpUri-2="https://zenodo.org/record/2610789/files/figure.png" pageId="2" pageNumber="583">Figs 1–3</figureCitation>
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</paragraph>
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</subSubSection>
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<paragraph id="8BDD3906FFBA8418738703102EF9C913" blockId="2.[151,1437,258,372]" pageId="2" pageNumber="583">
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<treatmentCitationGroup id="AB721E28FFBA8418738703102EF9C913" pageId="2" pageNumber="583">
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<taxonomicName id="4C624285FFBA84187387031029C9C93C" ID-CoL="3H5XZ" authority="Martynova 1954: 1167" authorityName="Martynova" authorityPageNumber="1167" authorityYear="1954" box="[151,684,258,280]" class="Insecta" family="Psychopsidae" genus="Grammapsychops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="lebedevi">
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<emphasis id="B916E514FFBA8418738703102ACBC93C" box="[151,430,258,280]" italics="true" pageId="2" pageNumber="583">Grammapsychops lebedevi</emphasis>
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<treatmentCitation id="0AC31F17FFBA841872AA031129C9C93C" author="Martynova, O. M." box="[442,684,258,280]" page="1167" pageId="2" pageNumber="583" year="1954">
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<bibRefCitation id="EFF344F7FFBA841872AA0311290EC93C" author="Martynova, O. M." box="[442,619,258,280]" pageId="2" pageNumber="583" pagination="1167 - 1169" refId="ref7737" refString="Martynova, O. M. (1954) Neuroptera from the Cretaceous deposits of Siberia. Doklady Akademii Nauk SSSR, 94, 1167 - 1169. [in Russian]" type="journal article" year="1954">Martynova 1954</bibRefCitation>
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: 1167
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</treatmentCitation>
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</taxonomicName>
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–1169,
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<figureCitation id="13592583FFBA841871EC0310285BC93C" box="[764,830,258,280]" captionStart="FIGURES 1" captionStartId="2.[151,264,1504,1526]" captionTargetBox="[171,1425,425,1488]" captionTargetId="figure@2.[171,1426,425,1488]" captionTargetPageId="2" captionText="FIGURES 1. Grammapsychops lebedevi Martynova, 1954, holotype PIN 846/1-2. A, specimen as preserved; B, original field label; C, PIN label. Scale bar is 2 mm." httpUri="https://zenodo.org/record/2610785/files/figure.png" pageId="2" pageNumber="583">Fig. 1</figureCitation>
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;
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<treatmentCitation id="0AC31F17FFBA8418704003102F52C93C" author="Rohdendorf, B. B." box="[848,1079,258,280]" page="40" pageId="2" pageNumber="583" year="1957">
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<bibRefCitation id="EFF344F7FFBA8418704003102F6EC93C" author="Rohdendorf, B. B." box="[848,1035,258,280]" pageId="2" pageNumber="583" pagination="1 - 102" refId="ref8144" refString="Rohdendorf, B. B. (1957) Paleoentomological investigations in the USSR. Trudy Paleontologicheskogo Instituta, 66, 1 - 102. [in Russian]" type="journal article" year="1957">Rohdendorf 1957</bibRefCitation>
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: 40
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</treatmentCitation>
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, 88, Fig. 18 [
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<taxonomicName id="4C624285FFBA841877C103102E31C93C" baseAuthorityName="Handlirsch" baseAuthorityYear="1906" box="[1233,1364,258,280]" class="Insecta" family="Psychopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="family">Psychopidae</taxonomicName>
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(sic)];
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<treatmentCitation id="0AC31F17FFBA841873D703332AFDC913" author="Carpenter, F. M." box="[199,408,289,311]" page="351" pageId="2" pageNumber="583" year="1992">
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<bibRefCitation id="EFF344F7FFBA841873D703332A03C913" author="Carpenter, F. M." box="[199,358,289,311]" pageId="2" pageNumber="583" pagination="279 - 655" refId="ref5999" refString="Carpenter, F. M. (1992) Superclass Hexapoda. In: Moore, R. C. & Kaesler, R. L. (Eds.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Fol. 4. Geological Society of America, Boulder, CO and University of Kansas, Lawrence, KS, pp. iii + 279 - 655." type="book chapter" year="1992">Carpenter 1992</bibRefCitation>
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||
: 351
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||
</treatmentCitation>
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||
, Fig. 195,1;
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<treatmentCitation id="0AC31F17FFBA841871050333284CC913" author="Engel, M. S. & Grimaldi, D. A." box="[533,809,289,311]" page="13" pageId="2" pageNumber="583" year="2008">
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<bibRefCitation id="EFF344F7FFBA8418710503332862C913" author="Engel, M. S. & Grimaldi, D. A." box="[533,775,289,311]" pageId="2" pageNumber="583" pagination="1 - 86" refId="ref6144" refString="Engel, M. S. & Grimaldi, D. A. (2008) Diverse Neuropterida in Cretaceous amber, with particular reference to the paleofauna of Myanmar (Insecta). Nova Supplementa Entomologica, 20, 1 - 86." type="journal article" year="2008">Engel & Grimaldi 2008</bibRefCitation>
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: 13
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</treatmentCitation>
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||
(Tab. 6) [
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<taxonomicName id="4C624285FFBA8418709903332F73C913" ID-CoL="626V6" authorityName="Handlirsch" authorityYear="1906" box="[905,1046,289,311]" class="Insecta" family="Psychopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="family">Psychopsidae</taxonomicName>
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s. str.];
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<treatmentCitation id="0AC31F17FFBA8418777503302E5CC913" author="Peng, Y. Y. & Makarkin, V. N. & Wang, X. D. & Ren, D." box="[1125,1337,289,311]" page="120" pageId="2" pageNumber="583" year="2011">
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||
<bibRefCitation id="EFF344F7FFBA8418777503302E6EC913" author="Peng, Y. Y. & Makarkin, V. N. & Wang, X. D. & Ren, D." box="[1125,1291,289,311]" pageId="2" pageNumber="583" pagination="217 - 228" refId="ref8023" refString="Peng, Y. Y., Makarkin, V. N., Wang, X. D. & Ren, D. (2011) A new fossil silky lacewing genus (Neuroptera: Psychopsidae) from the Early Cretaceous Yixian Formation of China. ZooKeys, 130, 217 - 228. https: // doi. org / 10.3897 / zookeys. 130.1576" type="journal article" year="2011">
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Peng
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<emphasis id="B916E514FFBA8418778D03302FACC913" box="[1181,1225,289,311]" italics="true" pageId="2" pageNumber="583">et al</emphasis>
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. 2011
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</bibRefCitation>
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: 120
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</treatmentCitation>
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(Tabl. 1).
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</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8BDD3906FFBA8418738703522AEDC950" blockId="2.[151,1437,258,372]" pageId="2" pageNumber="583">
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<treatmentCitationGroup id="AB721E28FFBA8418738703522AEDC950" pageId="2" pageNumber="583">
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<emphasis id="B916E514FFBA8418738703522A2BC972" box="[151,334,320,342]" italics="true" pageId="2" pageNumber="583">Grammopsychops</emphasis>
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<taxonomicName id="4C624285FFBA84187245035229FCC972" ID-CoL="3H5XZ" authority=": Martynova 1962" authorityName="Martynova" authorityYear="1962" box="[341,665,320,342]" class="Insecta" family="Psychopsidae" genus="Grammapsychops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="lebedevi">
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[sic]
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<emphasis id="B916E514FFBA8418729903522AB8C972" box="[393,477,320,342]" italics="true" pageId="2" pageNumber="583">lebedevi</emphasis>
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:
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<bibRefCitation id="EFF344F7FFBA841872FC035329FCC972" author="Martynova, O. M." box="[492,665,320,342]" pageId="2" pageNumber="583" pagination="383 - 404" refId="ref7769" refString="Martynova, O. M. (1962) Superorder Neuropteroidea. In: Rohdendorf, B. B. (Vol. Ed.), Fundamentals of Paleontology. Arthropoda-Tracheata and Chelicerata. Academy of Science of the USSR, Moscow, pp. 269 - 282. [in Russian; English translation: (1991). In: Davis, D. R. (Ed.), Fundamentals of Paleontology. Fol. 9. Arthropoda, Tracheata, Chelicerata. Smithsonian Institution Libraries and the National Science Foundation, Washington, D. C., pp. 383 - 404]" type="book chapter" year="1962">Martynova 1962</bibRefCitation>
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</taxonomicName>
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||
: Fig. 864;
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<treatmentCitation id="0AC31F17FFBA841870160352289FC972" author="Kolosnitsyna, G. R." box="[774,1018,320,342]" page="149" pageId="2" pageNumber="583" year="1964">
|
||
<bibRefCitation id="EFF344F7FFBA84187016035228ACC972" author="Kolosnitsyna, G. R." box="[774,969,320,342]" pageId="2" pageNumber="583" pagination="114 - 150" refId="ref6832" refString="Kolosnitsyna, G. R. (1964) New fossil insects from the Jurassic of the Irkutsk Basin]. In: Martinson, G. G. (Ed.), Stratigraphy and Paleontology of the Mesozoic and Cenozoic Deposits of Eastern Siberia and the Soviet Far East. Trudy Limnologicheskogo Instituta, 4 (24), pp. 114 - 150. [in Russian]" type="journal article" year="1964">Kolosnitsyna 1964</bibRefCitation>
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: 149
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||
</treatmentCitation>
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[Psychopsydidae (sic)];
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<treatmentCitation id="0AC31F17FFBA841877E703522B85C950" author="Zherikhin, V. V." page="80" pageId="2" pageNumber="583" year="1978">
|
||
<bibRefCitation id="EFF344F7FFBA841877E703522EFDC972" author="Zherikhin, V. V." box="[1271,1432,320,342]" pageId="2" pageNumber="583" pagination="1 - 200" refId="ref8576" refString="Zherikhin, V. V. (1978) Development and changes of Cretaceous and Cenozoic faunistic complexes (tracheates and chelicerates). Trudy Paleontologicheskogo Instituta, 165, 1 - 200. [in Russian]" type="journal article" year="1978">Zherikhin 1978</bibRefCitation>
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: 80
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</treatmentCitation>
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||
[
|
||
<taxonomicName id="4C624285FFBA841873FC034C2A1BC950" ID-CoL="626V6" authorityName="Handlirsch" authorityYear="1906" box="[236,382,350,372]" class="Insecta" family="Psychopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="family">Psychopsidae</taxonomicName>
|
||
].
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||
</treatmentCitationGroup>
|
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</paragraph>
|
||
<caption id="DF1D698EFFBA8418738707F22979CE30" httpUri="https://zenodo.org/record/2610785/files/figure.png" pageId="2" pageNumber="583" startId="2.[151,264,1504,1526]" targetBox="[171,1425,425,1488]" targetPageId="2">
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<paragraph id="8BDD3906FFBA8418738707F22979CE30" blockId="2.[151,1435,1504,1556]" pageId="2" pageNumber="583">
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||
<emphasis id="B916E514FFBA8418738707F22A44CDD2" bold="true" box="[151,289,1504,1526]" pageId="2" pageNumber="583">FIGURES 1.</emphasis>
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||
<taxonomicName id="4C624285FFBA8418723807F22997CDD2" ID-CoL="3H5XZ" authority="Martynova, 1954" authorityName="Martynova" authorityYear="1954" box="[296,754,1504,1526]" class="Insecta" family="Psychopsidae" genus="Grammapsychops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="lebedevi">
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||
<emphasis id="B916E514FFBA8418723807F2295CCDD2" box="[296,569,1504,1526]" italics="true" pageId="2" pageNumber="583">Grammapsychops lebedevi</emphasis>
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Martynova, 1954
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||
</taxonomicName>
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, holotype PIN 846/1-2. A, specimen as preserved; B, original field label; C, PIN label. Scale bar is 2 mm.
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||
</paragraph>
|
||
</caption>
|
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<paragraph id="8BDD3906FFBA84197387042E2F1BC9F4" blockId="2.[151,1437,1596,2016]" lastBlockId="3.[151,1437,151,464]" lastPageId="3" lastPageNumber="584" pageId="2" pageNumber="583">
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<emphasis id="B916E514FFBA84187387042E2962CE71" bold="true" box="[151,519,1596,1621]" pageId="2" pageNumber="583">
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Re-description of the
|
||
<materialsCitation id="3B0A335BFFBA841872B1042E2962CE71" ID-GBIF-Occurrence="2028522480" box="[417,519,1596,1621]" collectionCode="RA, RP, MP" pageId="2" pageNumber="583" specimenCode="RP6, RP1, RP4, RP5, RP11, RP12, RP13" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54D987A4FFBA841872B1042E2962CE71" box="[417,519,1596,1621]" pageId="2" pageNumber="583" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
</emphasis>
|
||
. Hind wing ca.
|
||
<quantity id="4C9A94E3FFBA841871D6042E287DCE70" box="[710,792,1596,1620]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="583" unit="mm" value="41.0">41 mm</quantity>
|
||
long; maximum width (measured perpendicular to RA)
|
||
<quantity id="4C9A94E3FFBA84187387044D2B99CE53" box="[151,252,1631,1656]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="2" pageNumber="583" unit="mm" value="18.5">18.5 mm</quantity>
|
||
as preserved. Distal nygma absent; basal nygma not detected. Costal space strongly dilated along entire preserved length, slightly narrowed towards apex. Subcostal veinlets closely spaces, nearly all once to twice forked, becoming more oblique towards apex. Costal crossveins numerous, mainly arranged in gradate series orientated obliquely to anterior wing margin. Subcostal space narrow along entire length (its width in
|
||
<figureCitation id="13592583FFBA8418761904DE2E2FCEC0" box="[1289,1354,1740,1765]" captionStart="FIGURES 3" captionStartId="4.[151,264,1587,1609]" captionTargetBox="[152,1418,908,1558]" captionTargetId="figure@4.[151,1436,895,1578]" captionTargetPageId="4" captionText="FIGURES 3. Forewing venation of Grammapsychops lebedevi Martynova, 1954, holotype PIN 846/1-2. Scale bar is 2 mm." httpUri="https://zenodo.org/record/2610789/files/figure.png" pageId="2" pageNumber="583">Fig. 3</figureCitation>
|
||
is seen slightly narrower than in the reality as RA is convex and the space is at the angle); crossveins not preserved. ScP apically not fused with RA, entering margin well before wing apex. R broad; RA strongly convex, entering margin just before wing apex, with three long veinlets. RA (between RA and RP) space very narrow, slightly broadened distad; few distal crossveins detected. RP originates close to wing base, with 13 pectinate branches; broad proximally (up to origin of
|
||
<specimenCode id="DBC4917DFFBA841872DB056D2965CFBC" box="[459,512,1919,1944]" collectionCode="RP" pageId="2" pageNumber="583">RP6</specimenCode>
|
||
).
|
||
<specimenCode id="DBC4917DFFBA84187103056D2925CFBC" box="[531,576,1919,1944]" collectionCode="RP" pageId="2" pageNumber="583">RP1</specimenCode>
|
||
–
|
||
<specimenCode id="DBC4917DFFBA8418715F056D29E4CFBC" box="[591,641,1919,1944]" collectionCode="RP" pageId="2" pageNumber="583">RP6</specimenCode>
|
||
broad proximally;
|
||
<specimenCode id="DBC4917DFFBA84187049059228E2CFBC" box="[857,903,1920,1944]" collectionCode="RP" pageId="2" pageNumber="583">RP1</specimenCode>
|
||
–
|
||
<specimenCode id="DBC4917DFFBA84187085059228ADCFBC" box="[917,968,1920,1944]" collectionCode="RP" pageId="2" pageNumber="583">RP4</specimenCode>
|
||
probably shallowly branched (their distal parts not preserved);
|
||
<specimenCode id="DBC4917DFFBA8418729605B62AD3CF99" box="[390,438,1956,1981]" collectionCode="RP" pageId="2" pageNumber="583">RP5</specimenCode>
|
||
–
|
||
<specimenCode id="DBC4917DFFBA841872D605B62967CF99" box="[454,514,1956,1981]" collectionCode="RP" pageId="2" pageNumber="583">RP11</specimenCode>
|
||
deeply dichotomously branched;
|
||
<specimenCode id="DBC4917DFFBA8418709105B628A4CF98" box="[897,961,1956,1980]" collectionCode="RP" pageId="2" pageNumber="583">RP12</specimenCode>
|
||
,
|
||
<specimenCode id="DBC4917DFFBA841870DD05B62F6ECF98" box="[973,1035,1956,1980]" collectionCode="RP" pageId="2" pageNumber="583">RP13</specimenCode>
|
||
dichotomously branched, but more distally. Proximal branches of RP basally with setal sockets arranged mainly in two rows (
|
||
<figureCitation id="13592583FFBA8418778405D52F89CFC4" box="[1172,1260,1991,2016]" captionStart="FIGURES 2" captionStartId="3.[151,264,1939,1961]" captionTargetBox="[173,1417,517,1926]" captionTargetId="figure@3.[173,1417,517,1926]" captionTargetPageId="3" captionText="FIGURES 2. Grammapsychops lebedevi Martynova, 1954, holotype PIN 846/1-2. A, proximal part of the forewing; B, a portion of RP and its branches (some setal sockets on veins are shown by red arrows). Scale bars are 2 mm (A), 0.4 mm (B)." httpUri="https://zenodo.org/record/2610787/files/figure.png" pageId="2" pageNumber="583">Fig. 2B</figureCitation>
|
||
); distal parts of RP branches appear thin as located in deep depressions, and setal sockets not detected. M forked slightly distad origin of
|
||
<specimenCode id="DBC4917DFFBB841973EF02AE2A57C8F0" box="[255,306,188,212]" collectionCode="RP" pageId="3" pageNumber="584">RP1</specimenCode>
|
||
. MA, MP parallel, not branched for preserved proximal half (not preserved distally). Cu divided into CuA and CuP close to base of wing. CuA strongly concave, pectinately branched with nine preserved, very oblique branches (two of these deeply forked), their terminal parts not preserved. CuP with two anteriorly directed, parallel branches; their terminal parts not preserved. Anterior trace of AA1 fragmentarily preserved, close and parallel to posterior trace of CuP. Crossveins in radial to cubital spaces numerous, arranged mainly in many gradate series; distinct outer gradate series not detected; basal crossveins between R/RP and M/MA irregularly spaced. Distinct long convex fold between each vein and veinlet in their distal parts. Trichosors along costal and apical margins indistinct. Wing pattern not preserved, there are only small patches of dark pigmentation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1D698EFFBB8419738705812EE7CFEC" httpUri="https://zenodo.org/record/2610787/files/figure.png" pageId="3" pageNumber="584" startId="3.[151,264,1939,1961]" targetBox="[173,1417,517,1926]" targetPageId="3">
|
||
<paragraph id="8BDD3906FFBB8419738705812EE7CFEC" blockId="3.[151,1436,1939,1992]" pageId="3" pageNumber="584">
|
||
<emphasis id="B916E514FFBB8419738705812A41CF8D" bold="true" box="[151,292,1939,1961]" pageId="3" pageNumber="584">FIGURES 2.</emphasis>
|
||
<taxonomicName id="4C624285FFBB8419723E05812864CF8D" authority="Martynova, 1954" authorityName="Martynova" authorityYear="1954" box="[302,769,1939,1961]" class="Insecta" family="Psychopsidae" genus="Grammapsychops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="lebedevi">
|
||
<emphasis id="B916E514FFBB8419723E05812927CF8D" box="[302,578,1939,1961]" italics="true" pageId="3" pageNumber="584">Grammapsychops lebedevi</emphasis>
|
||
Martynova, 1954
|
||
</taxonomicName>
|
||
, holotype PIN 846/1-2. A, proximal part of the forewing; B, a portion of RP and its branches (some setal sockets on veins are shown by red arrows). Scale bars are 2 mm (A), 0.4 mm (B).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3786A8DFFBC841E73D702852FA4C939" pageId="4" pageNumber="585" type="materials_examined">
|
||
<paragraph id="8BDD3906FFBC841E73D702852EF8C894" blockId="4.[151,1437,151,861]" box="[199,1437,151,176]" pageId="4" pageNumber="585">
|
||
<materialsCitation id="3B0A335BFFBC841E73D702852EF8C894" ID-GBIF-Occurrence="2028522491" box="[199,1437,151,176]" collectionCode="PIN" collectorName="Ivan V. Lebedev & An" pageId="4" pageNumber="585" specimenCode="PIN 846" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B916E514FFBC841E73D702852A5EC894" bold="true" box="[199,315,151,176]" pageId="4" pageNumber="585">
|
||
<typeStatus id="54D987A4FFBC841E73D702852A53C894" box="[199,310,151,176]" pageId="4" pageNumber="585" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Specimen
|
||
<specimenCode id="DBC4917DFFBC841E72A6028A297DC894" box="[438,536,151,176]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="4" pageNumber="585">PIN 846</specimenCode>
|
||
/1-2 (part only), collected by
|
||
<collectorName id="26975CD0FFBC841E704A028A2F75C894" box="[858,1040,151,176]" pageId="4" pageNumber="585">Ivan V. Lebedev</collectorName>
|
||
in 1951.
|
||
<collectorName id="26975CD0FFBC841E7769028A2FFEC894" box="[1145,1179,152,176]" pageId="4" pageNumber="585">An</collectorName>
|
||
incomplete hind wing.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BDD3906FFBC841E73D702AE2FA4C939" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="585">
|
||
<emphasis id="B916E514FFBC841E73D702AE2965C8F1" bold="true" box="[199,512,188,213]" pageId="4" pageNumber="585">Type locality and horizon.</emphasis>
|
||
<materialsCitation id="3B0A335BFFBC841E711802AE2FD8C939" ID-GBIF-Occurrence="2028522479" country="Russia" county="Region" latitude="57.97137" location="Kem' River" longLatPrecision="1" longitude="92.230034" municipality="District of Pirovskoe" pageId="4" pageNumber="585" specimenCount="1" stateProvince="Krasnoyarsk">
|
||
<collectingCountry id="F3757996FFBC841E711802AE2932C8F1" box="[520,599,188,213]" name="Russia" pageId="4" pageNumber="585">Russia</collectingCountry>
|
||
:
|
||
<collectingRegion id="49A6F7E4FFBC841E717202AE298AC8F1" box="[610,751,188,213]" country="Russia" name="Krasnoyarsk" pageId="4" pageNumber="585">Krasnoyarsk</collectingRegion>
|
||
<collectingCounty id="62BC418AFFBC841E71E702AE2829C8F1" box="[759,844,188,213]" pageId="4" pageNumber="585">Region</collectingCounty>
|
||
:
|
||
<collectingMunicipality id="6BB9A37CFFBC841E704802AE2F22C8F1" box="[856,1095,188,213]" pageId="4" pageNumber="585">District of Pirovskoe</collectingMunicipality>
|
||
: right bank of the
|
||
<location id="8EBD6FDDFFBC841E760602AE2EF9C8F1" LSID="urn:lsid:plazi:treatment:03CB8810FFBA841E73870285295FCB78:8EBD6FDDFFBC841E760602AE2EF9C8F1" box="[1302,1436,188,213]" country="Russia" county="Region" latitude="57.97137" longLatPrecision="1" longitude="92.230034" municipality="District of Pirovskoe" name="Kem' River" pageId="4" pageNumber="585" stateProvince="Krasnoyarsk">Kem’ River</location>
|
||
(left tributary of
|
||
<location id="8EBD6FDDFFBC841E727C02CD2973C8DC" LSID="urn:lsid:plazi:treatment:03CB8810FFBA841E73870285295FCB78:8EBD6FDDFFBC841E727C02CD2973C8DC" box="[364,534,223,248]" country="Russia" county="Region" latitude="57.97137" longLatPrecision="1" longitude="92.230034" municipality="District of Pirovskoe" name="Yenisey River" pageId="4" pageNumber="585" stateProvince="Krasnoyarsk">Yenisey River</location>
|
||
), ca.
|
||
<quantity id="4C9A94E3FFBC841E717202CD29C1C8DC" box="[610,676,223,248]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="585" unit="km" value="5.0">5 km</quantity>
|
||
upstream from the mouth of the
|
||
<location id="8EBD6FDDFFBC841E774102CD2F88C8DC" LSID="urn:lsid:plazi:treatment:03CB8810FFBA841E73870285295FCB78:8EBD6FDDFFBC841E774102CD2F88C8DC" box="[1105,1261,223,248]" country="Russia" county="Region" latitude="57.97137" longLatPrecision="1" longitude="92.230034" municipality="District of Pirovskoe" name="Belaya River" pageId="4" pageNumber="585" stateProvince="Krasnoyarsk">Belaya River</location>
|
||
[
|
||
<geoCoordinate id="EE565FC1FFBC841E761302CD2EF9C8DC" box="[1283,1436,223,248]" direction="north" orientation="latitude" pageId="4" pageNumber="585" precision="1" value="57.97137">57.971372°N</geoCoordinate>
|
||
<geoCoordinate id="EE565FC1FFBC841E738703162A54C939" box="[151,305,260,285]" direction="east" orientation="longitude" pageId="4" pageNumber="585" precision="1" value="92.230034">92.230036°E</geoCoordinate>
|
||
]; exposure 0 92, horizon 3. Late Cretaceous: Cenomanian (Simonovo Formation)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3786A8DFFBC841E73D70335295FCB78" pageId="4" pageNumber="585" type="discussion">
|
||
<paragraph id="8BDD3906FFBC841E73D703352A61CA3C" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="585">
|
||
<emphasis id="B916E514FFBC841E73D703352A5EC964" bold="true" box="[199,315,295,320]" pageId="4" pageNumber="585">Remarks.</emphasis>
|
||
<bibRefCitation id="EFF344F7FFBC841E7255033A2973C964" author="Martynova, O. M." box="[325,534,295,320]" pageId="4" pageNumber="585" pagination="1167 - 1169" refId="ref7737" refString="Martynova, O. M. (1954) Neuroptera from the Cretaceous deposits of Siberia. Doklady Akademii Nauk SSSR, 94, 1167 - 1169. [in Russian]" type="journal article" year="1954">Martynova (1954)</bibRefCitation>
|
||
interpreted this wing as the left forewing, and she believed that its posterior margin is preserved (actually, it is not preserved, possibly inwardly folded). Based on this assumption, she designated veins accordingly and incorrectly, i.e., her M is actually five proximal branches of RP; Cu (CuA and CuP) is M (MA and MP); A1 is Cu (CuA and CuP); and A2 is AA1. The venation of the
|
||
<typeStatus id="54D987A4FFBC841E776503862FB2C989" box="[1141,1239,404,429]" pageId="4" pageNumber="585" type="holotype">holotype</typeStatus>
|
||
shows that this is a hind wing because its CuA is clearly concave. Also, M is forked very proximally and the anterior trace of AA1 is proximally parallel to the posterior trace of CuP, which are more characteristic of the hind wing than of the forewing.
|
||
</paragraph>
|
||
<paragraph id="8BDD3906FFBC841E73D700372EF3CA8C" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="585">
|
||
Martynova believed that the margins of the broad basal part of the vein R, then RP, are two parallel thin veins, i.e., R, then RP, and M in her interpretation (see
|
||
<bibRefCitation id="EFF344F7FFBC841E71D4005A28E6CA44" author="Martynova, O. M." box="[708,899,583,608]" pageId="4" pageNumber="585" pagination="1167 - 1169" refId="ref7737" refString="Martynova, O. M. (1954) Neuroptera from the Cretaceous deposits of Siberia. Doklady Akademii Nauk SSSR, 94, 1167 - 1169. [in Russian]" type="journal article" year="1954">Martynova 1954</bibRefCitation>
|
||
:
|
||
<figureCitation id="13592583FFBC841E709F005528B4CA44" box="[911,977,583,608]" captionStart="FIGURES 1" captionStartId="2.[151,264,1504,1526]" captionTargetBox="[171,1425,425,1488]" captionTargetId="figure@2.[171,1426,425,1488]" captionTargetPageId="2" captionText="FIGURES 1. Grammapsychops lebedevi Martynova, 1954, holotype PIN 846/1-2. A, specimen as preserved; B, original field label; C, PIN label. Scale bar is 2 mm." httpUri="https://zenodo.org/record/2610785/files/figure.png" pageId="4" pageNumber="585">Fig. 1</figureCitation>
|
||
). So, she considered ‘M’ running to the wing base as a separate vein and deeply forked into five branches (my RP1 to RP5). However, it is clearly visible that the basal parts of R and RP are broad and no break of RP between the origins of RP1 and RP6 exists (
|
||
<figureCitation id="13592583FFBC841E7621009D2EEDCA8C" box="[1329,1416,655,680]" captionStart="FIGURES 2" captionStartId="3.[151,264,1939,1961]" captionTargetBox="[173,1417,517,1926]" captionTargetId="figure@3.[173,1417,517,1926]" captionTargetPageId="3" captionText="FIGURES 2. Grammapsychops lebedevi Martynova, 1954, holotype PIN 846/1-2. A, proximal part of the forewing; B, a portion of RP and its branches (some setal sockets on veins are shown by red arrows). Scale bars are 2 mm (A), 0.4 mm (B)." httpUri="https://zenodo.org/record/2610787/files/figure.png" pageId="4" pageNumber="585">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDD3906FFBC841E73D700A6295FCB78" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="585">
|
||
The costal space of this wing is very broad assuming that it might be a forewing. However, the costal space of the hind wing of
|
||
<taxonomicName id="4C624285FFBC841E727100CA282DCAD4" authority="Peng et al., 2010" authorityName="Peng" authorityYear="2010" box="[353,840,728,752]" class="Insecta" family="Psychopsidae" genus="Cretapsychops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="decipiens">
|
||
<emphasis id="B916E514FFBC841E727100CA291FCAD4" box="[353,634,728,752]" italics="true" pageId="4" pageNumber="585">Cretapsychops decipiens</emphasis>
|
||
<bibRefCitation id="EFF344F7FFBC841E719300CA282DCAD4" author="Peng, Y. Y. & Makarkin, V. N. & Yang, Q. & Ren, D." box="[643,840,728,752]" pageId="4" pageNumber="585" pagination="59 - 67" refId="ref8091" refString="Peng, Y. Y., Makarkin, V. N., Yang, Q. & Ren, D. (2010) A new silky lacewing (Neuroptera: Psychopsidae) from the Middle Jurassic of Inner Mongolia, China. Zootaxa, 2663, 59 - 67." type="journal article" year="2010">
|
||
Peng
|
||
<emphasis id="B916E514FFBC841E71D400CB299DCAD4" box="[708,760,728,752]" italics="true" pageId="4" pageNumber="585">et al</emphasis>
|
||
., 2010
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
is similarly configured to that of
|
||
<taxonomicName id="4C624285FFBC841E77C200CA2B91CB30" authorityName="Martynova" authorityYear="1954" class="Insecta" family="Psychopsidae" genus="Grammapsychops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="lebedevi">
|
||
<emphasis id="B916E514FFBC841E77C200CA2B91CB30" italics="true" pageId="4" pageNumber="585">Grammapsychops lebedevi</emphasis>
|
||
</taxonomicName>
|
||
: it is very broad, wider than that of the forewing, and proximally more dilated than distally (see
|
||
<bibRefCitation id="EFF344F7FFBC841E763600EF2BB1CB1C" author="Peng, Y. Y. & Makarkin, V. N. & Yang, Q. & Ren, D." pageId="4" pageNumber="585" pagination="59 - 67" refId="ref8091" refString="Peng, Y. Y., Makarkin, V. N., Yang, Q. & Ren, D. (2010) A new silky lacewing (Neuroptera: Psychopsidae) from the Middle Jurassic of Inner Mongolia, China. Zootaxa, 2663, 59 - 67." type="journal article" year="2010">
|
||
Peng
|
||
<emphasis id="B916E514FFBC841E767400EF2EF0CB30" box="[1380,1429,764,788]" italics="true" pageId="4" pageNumber="585">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
:
|
||
<figureCitation id="13592583FFBC841E73CE010D2A44CB1C" box="[222,289,799,824]" captionStart="FIGURES 3" captionStartId="4.[151,264,1587,1609]" captionTargetBox="[152,1418,908,1558]" captionTargetId="figure@4.[151,1436,895,1578]" captionTargetPageId="4" captionText="FIGURES 3. Forewing venation of Grammapsychops lebedevi Martynova, 1954, holotype PIN 846/1-2. Scale bar is 2 mm." httpUri="https://zenodo.org/record/2610789/files/figure.png" pageId="4" pageNumber="585">Fig. 3</figureCitation>
|
||
). Also, the costal spaces of the hind wing of
|
||
<taxonomicName id="4C624285FFBC841E700901322F5ACB1C" authorityName="Martins-Neto" authorityYear="1997" box="[793,1087,800,824]" class="Insecta" family="Psychopsidae" genus="Pulchroptilonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="espatifata">
|
||
<emphasis id="B916E514FFBC841E700901322F5ACB1C" box="[793,1087,800,824]" italics="true" pageId="4" pageNumber="585">Pulchroptilonia espatifata</emphasis>
|
||
</taxonomicName>
|
||
are configured rather similarly (see
|
||
<bibRefCitation id="EFF344F7FFBC841E73DB01562AC0CB78" author="Martins-Neto, R. G." box="[203,421,836,861]" pageId="4" pageNumber="585" pagination="68 - 83" refId="ref7609" refString="Martins-Neto, R. G. (1997) Neuropteros (Insecta, Planipennia) da Formacao Santana (Cretaceo Inferior), Bacia do Araripe, nordeste do Brasil. X - Descricao de novos taxons (Chrysopidae, Babinskaiidae, Myrmeleontidae, Ascalaphidae e Psychopsidae). Revista Universidade de Guarulhos (Ciencias de Exatas e Technologicas), 2 (4), 68 - 83." type="journal article" year="1997">Martins-Neto 1997</bibRefCitation>
|
||
: Figs 8A, B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |