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<mods:title id="1A249D8322D57847B2B2CC80F008114B">A taxonomic revision of the species of Licea subg. Licea (Myxomycetes)</mods:title>
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<mods:namePart id="86FC741F3BA669603E3E2C885E37A4A9">Basanta, Diana Wrigley De</mods:namePart>
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<mods:affiliation id="F0E2D4F090749C2C5267CB11BD3F92C7">Real Jardín Botánico, CSIC, Plaza de Murillo 2, 28014 Madrid (Spain). &amp; dwbasanta @ gmail. com, https: // orcid. org / 0000 - 0002 - 7700 - 8399</mods:affiliation>
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<mods:namePart id="8D21A0DA2EBBC896E9858766C0DD78F5">Mier, Carlos De</mods:namePart>
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<mods:affiliation id="4C2290E50D3CC2D265B5B0C4531FF2CA">Real Jardín Botánico, CSIC, Plaza de Murillo 2, 28014 Madrid (Spain). &amp; cmier @ rjb. csic. es, https: // orcid. org / 0009 - 0008 - 1431 - 4410</mods:affiliation>
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<mods:affiliation id="DFB7D4DF215C0529BF42E5EF8A2AC0ED">Real Jardín Botánico, CSIC, Plaza de Murillo 2, 28014 Madrid (Spain). &amp; lado @ rjb. csic. es, https: // orcid. org / 0000 - 0002 - 6135 - 2873</mods:affiliation>
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<emphasis id="B96AF8089B0FFFC7FF1606AFFF67FCB2" bold="true" box="[136,171,763,789]" pageId="23" pageNumber="118">20.</emphasis>
<taxonomicName id="4C1E5F999B0FFFC7FF2C06AFFCC8FCB1" authority="(Meyl.) Ing, Trans. Brit. Mycol. Soc." baseAuthorityName="Meyl." box="[178,772,763,790]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="species" species="pygmaea">
<treatmentCitation id="0ABF020B9B0FFFC7FF2C06AFFEACFCB1" author="Ing" box="[178,352,763,790]" issue="3" journal="Trans. Brit. Mycol. Soc." page="443" pageId="23" pageNumber="118" score="5.464730587576831" volume="78" year="1982">
<emphasis id="B96AF8089B0FFFC7FF2C06AFFEACFCB1" bold="true" box="[178,352,763,790]" italics="true" pageId="23" pageNumber="118">Licea pygmaea</emphasis>
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(Meyl.) Ing, Trans. Brit. Mycol. Soc.
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78(3): 443 (1982) 
<figureCitation id="1325389F9B0FFFC7FC4706AFFBAAFCB1" box="[985,1126,763,790]" captionStart="FIGURE 17" captionStartId="24.[136,229,1392,1414]" captionTargetBox="[151,1436,187,1364]" captionTargetId="figure-22@24.[151,1436,187,1364]" captionTargetPageId="24" captionText="FIGURE 17. Licea pygmaea.A–L (Myxo 055, LAU, Lectotype).A. Herbarium box. B–E. Sessile sporocarps. F–G. Peridial platelets and spore by TL. H. Whole open sporocarp by SEM. I–J. Peridial platelet margins by SEM. K–L. Spores by SEM. Scale bars: B = 0.1 mm, C–E, H = 50 µm, F–G = 10 µm, I–L = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276083" httpUri="https://zenodo.org/record/10276083/files/figure.png" pageId="23" pageNumber="118">Figs. 17A–L</figureCitation>
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<paragraph id="8BA1241A9B0FFFC7FF160770FCA1FC9F" blockId="23.[136,1126,763,825]" box="[136,877,803,825]" pageId="23" pageNumber="118">
≡ 
<treatmentCitation id="0ABF020B9B0FFFC7FF3C0770FCA1FC9F" author="Meyl." box="[162,877,803,825]" journal="Bull. Soc. Vaud. Sci. Nat." page="89" pageId="23" pageNumber="118" score="29.60634419892665" volume="58" year="1933">
<taxonomicName id="4C1E5F999B0FFFC7FF3C0770FD83FC9F" authority="Meyl., Bull. Soc." authorityName="Meyl., Bull. Soc." box="[162,591,803,825]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="variety" species="pusilla" variety="pygmaea">
<emphasis id="B96AF8089B0FFFC7FF3C0770FED2FC9F" box="[162,286,803,825]" italics="true" pageId="23" pageNumber="118">Licea pusilla</emphasis>
var. 
<emphasis id="B96AF8089B0FFFC7FED00770FE69FC9E" box="[334,421,804,825]" italics="true" pageId="23" pageNumber="118">pygmaea</emphasis>
Meyl., Bull. Soc.
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. Sci. Nat. 58: 89 (1933)
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</paragraph>
<paragraph id="8BA1241A9B0FFFC7FF160734FBF1FBEA" blockId="23.[136,1452,864,1736]" pageId="23" pageNumber="118">
Sporophores sporocarpic, scattered to gregarious, sessile. Sporocarps subglobose to pulvinate, angular, 
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diam., brown. Peridium double, outer layer horny, inner layer membranous; dehiscence into platelets along shiny ridges, platelet margins with prominent outgrowths. Spores free, dark olive to dull dark brown in mass, greenish yellow by TL, globose, 12–13 µm diam., minutely evenly warted. By SEM the inner peridium inner surface is warted with different sized warts and some bare patches (
<figureCitation id="1325389F9B0FFFC7FD5A07B9FC84FBA0" box="[708,840,1005,1031]" captionStart="FIGURE 17" captionStartId="24.[136,229,1392,1414]" captionTargetBox="[151,1436,187,1364]" captionTargetId="figure-22@24.[151,1436,187,1364]" captionTargetPageId="24" captionText="FIGURE 17. Licea pygmaea.A–L (Myxo 055, LAU, Lectotype).A. Herbarium box. B–E. Sessile sporocarps. F–G. Peridial platelets and spore by TL. H. Whole open sporocarp by SEM. I–J. Peridial platelet margins by SEM. K–L. Spores by SEM. Scale bars: B = 0.1 mm, C–E, H = 50 µm, F–G = 10 µm, I–L = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276083" httpUri="https://zenodo.org/record/10276083/files/figure.png" pageId="23" pageNumber="118">Figs. 17I–J</figureCitation>
); the platelet margins have different sized outgrowths mostly in a single row but sometimes overlapping, fused and sausage-shaped (
<figureCitation id="1325389F9B0FFFC7FB950044FB42FB8D" box="[1035,1166,1040,1066]" captionStart="FIGURE 17" captionStartId="24.[136,229,1392,1414]" captionTargetBox="[151,1436,187,1364]" captionTargetId="figure-22@24.[151,1436,187,1364]" captionTargetPageId="24" captionText="FIGURE 17. Licea pygmaea.A–L (Myxo 055, LAU, Lectotype).A. Herbarium box. B–E. Sessile sporocarps. F–G. Peridial platelets and spore by TL. H. Whole open sporocarp by SEM. I–J. Peridial platelet margins by SEM. K–L. Spores by SEM. Scale bars: B = 0.1 mm, C–E, H = 50 µm, F–G = 10 µm, I–L = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276083" httpUri="https://zenodo.org/record/10276083/files/figure.png" pageId="23" pageNumber="118">Figs. 17I–J</figureCitation>
); the epispore is covered with flattened bacula evenly dispersed, sometimes two or more fused (
<figureCitation id="1325389F9B0FFFC7FC000067FBFDFBEA" box="[926,1073,1075,1101]" captionStart="FIGURE 17" captionStartId="24.[136,229,1392,1414]" captionTargetBox="[151,1436,187,1364]" captionTargetId="figure-22@24.[151,1436,187,1364]" captionTargetPageId="24" captionText="FIGURE 17. Licea pygmaea.A–L (Myxo 055, LAU, Lectotype).A. Herbarium box. B–E. Sessile sporocarps. F–G. Peridial platelets and spore by TL. H. Whole open sporocarp by SEM. I–J. Peridial platelet margins by SEM. K–L. Spores by SEM. Scale bars: B = 0.1 mm, C–E, H = 50 µm, F–G = 10 µm, I–L = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276083" httpUri="https://zenodo.org/record/10276083/files/figure.png" pageId="23" pageNumber="118">Figs. 17K–L</figureCitation>
).
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<emphasis id="B96AF8089B0FFFC7FF230002FE5DFBD7" box="[189,401,1110,1136]" italics="true" pageId="23" pageNumber="118">Material examined</emphasis>
: 
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. 
<collectingCountry id="F309648A9B0FFFC7FDB70003FD38FBD6" box="[553,756,1111,1137]" name="Switzerland" pageId="23" pageNumber="118">SWITZERLAND</collectingCountry>
. 
<collectingRegion id="49DAEAF89B0FFFC7FD630002FCF6FBD7" box="[765,826,1110,1136]" country="Switzerland" name="Vaud" pageId="23" pageNumber="118">Vaud</collectingRegion>
, nr. Ste Croix, La Gittaz, 
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, 
<date id="FFA002DA9B0FFFC7FB2B0003FAE2FBD6" box="[1205,1326,1111,1137]" pageId="23" pageNumber="118" value="1930-10">OCT 1930</date>
. Myxo 055 (
<collectionCode id="ED0FBCDF9B0FFFC7FF0F002EFF19FB33" box="[145,213,1146,1172]" country="Switzerland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15274" lsid="urn:lsid:biocol.org:col:15274" name="Musée et Jardins Botaniques Cantonaux" pageId="23" pageNumber="118" type="Herbarium">LAU</collectionCode>
)!. (Designated by 
<bibCitation id="5E342C769B0FFFC7FE03002EFCA5FB33" author="Kowlaski" box="[413,873,1145,1172]" journal="Mycologia" page="448" pageId="23" pageNumber="118" score="9.729594737742866" volume="67" year="1975">
Kowlaski, 
<bibCitation id="5E342C769B0FFFC7FD88002EFCA5FB33" box="[534,873,1145,1172]" journal="Mycologia" page="448" pageId="23" pageNumber="118" score="9.729594737742866" volume="67" year="1975">Mycologia 67: 448-494. 1975</bibCitation>
</bibCitation>
).
</paragraph>
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<emphasis id="B96AF8089B0FFFC7FF2300C9FED0FB10" box="[189,284,1181,1207]" italics="true" pageId="23" pageNumber="118">Habitat:</emphasis>
on bark of living and dead trees, trunks, stumps and debris of angiosperms and gymnosperms.
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<emphasis id="B96AF8089B0FFFC7FF230094FE83FB7D" box="[189,335,1216,1242]" italics="true" pageId="23" pageNumber="118">Distribution:</emphasis>
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, 
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, 
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, 
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, 
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, 
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, 
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, 
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, 
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, 
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, 
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, 
<collectingCountry id="F309648A9B0FFFC7FDE400B7FD71FB5A" box="[634,701,1251,1277]" name="Spain" pageId="23" pageNumber="118">Spain</collectingCountry>
, 
<collectingCountry id="F309648A9B0FFFC7FD5500B7FC98FB5A" box="[715,852,1251,1277]" name="Switzerland" pageId="23" pageNumber="118">Switzerland</collectingCountry>
, 
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, 
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, 
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, 
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, 
<collectingCountry id="F309648A9B0FFFC7FD4B0153FCF2FA86" box="[725,830,1287,1313]" name="Paraguay" pageId="23" pageNumber="118">Paraguay</collectingCountry>
, 
<collectingCountry id="F309648A9B0FFFC7FCD60152FC7EFA87" box="[840,946,1286,1312]" name="Australia" pageId="23" pageNumber="118">Australia</collectingCountry>
, 
<collectingCountry id="F309648A9B0FFFC7FC220153FB9AFA87" box="[956,1110,1286,1313]" name="New Zealand" pageId="23" pageNumber="118">New Zealand</collectingCountry>
.
</paragraph>
<paragraph id="8BA1241A9B0FFFC7FF23017EFD81FA53" blockId="23.[136,1452,864,1736]" pageId="23" pageNumber="118">
<emphasis id="B96AF8089B0FFFC7FF23017EFECCFAE4" box="[189,256,1322,1347]" italics="true" pageId="23" pageNumber="118">Icon.:</emphasis>
<bibRefCitation id="EF8F59EB9B0FFFC7FE97017EFD58FAE3" author="Nannenga-Bremekamp, N. E." box="[265,660,1321,1348]" pageId="23" pageNumber="118" pagination="131 - 147" refId="ref21519" refString="Nannenga-Bremekamp, N. E. (1965) Notes on Myxomycetes IX. The genus Licea in the Netherlands. Acta Botanica Neerlandica 14: 131 - 147." type="journal article" year="1965">Nannenga-Bremekamp (1965: 140</bibRefCitation>
, 
<figureCitation id="1325389F9B0FFFC7FD3D017EFCEBFAE3" box="[675,807,1322,1348]" captionStart="FIGURE 7" captionStartId="11.[136,229,1357,1378]" captionTargetBox="[151,1436,187,1329]" captionTargetId="figure-19@11.[151,1436,187,1329]" captionTargetPageId="11" captionText="FIGURE 7. Licea castaneoides. A–I (NENB 16135, ex Y.Y. 6407, BR5020070191601, Holotype). A. Herbarium box. B–C. Sessile sporocarps. D. Peridial platelet and spores by TL. E. Spores by TL. F. Dehisced sporocarp by SEM. G. Platelet by SEM showing double peridium with no outgrowths on the edge. H–I. Spores by SEM. Scale bars: B–C = 0.1 mm, D–F = 10 µm, G–I = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276061" httpUri="https://zenodo.org/record/10276061/files/figure.png" pageId="23" pageNumber="118">Figs. 7B–C</figureCitation>
, as 
<taxonomicName id="4C1E5F999B0FFFC7FCC8017EFBAAFAE4" box="[854,1126,1322,1348]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="variety" species="pusilla" variety="pygmaea">
<emphasis id="B96AF8089B0FFFC7FCC8017EFC0EFAE3" box="[854,962,1322,1348]" italics="true" pageId="23" pageNumber="118">L. pusilla</emphasis>
var. 
<emphasis id="B96AF8089B0FFFC7FC60017EFBAAFAE4" box="[1022,1126,1322,1347]" italics="true" pageId="23" pageNumber="118">pygmaea</emphasis>
</taxonomicName>
<emphasis id="B96AF8089B0FFFC7FBF8017EFBA1FAE4" box="[1126,1133,1322,1347]" italics="true" pageId="23" pageNumber="118">;</emphasis>
1975: 66, as 
<taxonomicName id="4C1E5F999B0FFFC7FA92017EFF21FAC0" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="variety" species="pusilla" variety="pygmaea">
<emphasis id="B96AF8089B0FFFC7FA92017EFAB4FAE3" box="[1292,1400,1322,1348]" italics="true" pageId="23" pageNumber="118">L. pusilla</emphasis>
var. 
<emphasis id="B96AF8089B0FFFC7FF16011AFF21FAC0" box="[136,237,1358,1383]" italics="true" pageId="23" pageNumber="118">pygmaea</emphasis>
</taxonomicName>
; 1991: 45, as 
<taxonomicName id="4C1E5F999B0FFFC7FE11011AFD54FAC0" box="[399,664,1357,1383]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="variety" species="pusilla" variety="pygmaea">
<emphasis id="B96AF8089B0FFFC7FE11011AFE35FAC0" box="[399,505,1357,1383]" italics="true" pageId="23" pageNumber="118">L. pusilla</emphasis>
var. 
<emphasis id="B96AF8089B0FFFC7FDAD011AFD54FAC0" box="[563,664,1358,1383]" italics="true" pageId="23" pageNumber="118">pygmaea</emphasis>
</taxonomicName>
), Gilert (1996: 517, 
<figureCitation id="1325389F9B0FFFC7FC1D0119FC23FAC0" box="[899,1007,1357,1383]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[136,229,999,1021]" captionStartId-1="4.[136,229,1880,1902]" captionTargetBox-0="[151,1436,188,973]" captionTargetBox-1="[151,1436,674,1852]" captionTargetId-0="figure-19@3.[151,1436,188,973]" captionTargetId-1="figure-250@4.[151,1436,674,1852]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Licea alpina. A–F (AH 28531, Holotype).A. Herbarium boxes and slides. B–D. Stalked and subsessile sporocarps. E. Spores and peridial platelets by TL. F. Spores by TL. Scale bars: B–D = 0.1 mm, E–F = 10 µm." captionText-1="FIGURE 2. Licea ampulliformis. A–H, K–M (dwb 1988, MA-Fungi 98835, Holotype), I–J (dwb 2535).A. Herbarium box. B–D. Sessile sporocarps. E. Spores and peridial platelets by TL showing clear margins to the platelets (arrows). F–H. Spores by TL. I. detail of platelet margin showing clear area by SEM (arrow). J. Inner surface of a platelet showing warted surface. K–L Detail of open sporocarp showing platelets and spore by SEM. M. Spores by SEM. Scale bars: B–D = 0.1 mm, E = 20 µm, F–H, L = 10 µm, I–J, M = 5 µm, K = 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.10276049" figureDoi-1="http://doi.org/10.5281/zenodo.10276051" httpUri-0="https://zenodo.org/record/10276049/files/figure.png" httpUri-1="https://zenodo.org/record/10276051/files/figure.png" pageId="23" pageNumber="118">Figs. 1–2</figureCitation>
, 
<figureCitation id="1325389F9B0FFFC7FC620119FBE5FACF" box="[1020,1065,1357,1384]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="8.[136,234,1628,1650]" captionStartId-1="9.[136,229,1876,1898]" captionTargetBox-0="[151,1436,458,1600]" captionTargetBox-1="[151,1436,278,1848]" captionTargetId-0="figure-127@8.[151,1436,458,1600]" captionTargetId-1="figure-52@9.[151,1436,278,1848]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 5. Licea capacia. A–I (12486.Oltra, AH 42711, Holotype).A. Herbarium box. B–D. Sessile sporocarps. E. Peridial platelets by TL. F. Angular spores by TL. G. Peridial platelet margins by SEM showing double peridium (arrow) and no outgrowths. H–I. Spores by SEM. Scale bars: B–D = 0.1 mm, E–F = 10 µm, G–I = 5 µm." captionText-1="FIGURE 6. Licea castanea. A (BM 3025, BM001089212, Lectotype); B–G (MA-Fungi 16013); H, J (MA-Fungi 29564); I (MA-Fungi 15932); K–S (Holotype of L. bryocorticola, HK 201113-07a, M-0313079). A. Herbarium box. B–D. Sessile sporocarps. E. Spores and peridial platelets by TL showing outgrowths on the edges. F–G. Spores by TL. H. Spore by SEM. I. Peridial platelets by SEM, showing outgrowths along the margins. J. Spore by SEM. K. Herbarium specimen. L–N sessile sporocarps. O–Q. Peridial platelets by TL showing outgrowths on the edges. R–S. Spores by TL. Scale bars: B–D, L–N = 0.1 mm, E–G, P–S = 10 µm, H–J = 5 µm, O = 20 µm." figureDoi-0="http://doi.org/10.5281/zenodo.10276057" figureDoi-1="http://doi.org/10.5281/zenodo.10276059" httpUri-0="https://zenodo.org/record/10276057/files/figure.png" httpUri-1="https://zenodo.org/record/10276059/files/figure.png" pageId="23" pageNumber="118">5–6</figureCitation>
; 518, 
<figureCitation id="1325389F9B0FFFC7FBF00119FB16FAC0" box="[1134,1242,1357,1383]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="11.[136,229,1357,1378]" captionStartId-1="12.[136,229,1357,1378]" captionTargetBox-0="[151,1436,187,1329]" captionTargetBox-1="[151,1436,187,1329]" captionTargetId-0="figure-19@11.[151,1436,187,1329]" captionTargetId-1="figure-22@12.[151,1436,187,1329]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 7. Licea castaneoides. A–I (NENB 16135, ex Y.Y. 6407, BR5020070191601, Holotype). A. Herbarium box. B–C. Sessile sporocarps. D. Peridial platelet and spores by TL. E. Spores by TL. F. Dehisced sporocarp by SEM. G. Platelet by SEM showing double peridium with no outgrowths on the edge. H–I. Spores by SEM. Scale bars: B–C = 0.1 mm, D–F = 10 µm, G–I = 5 µm." captionText-1="FIGURE 8. Licea chelonoides. A–J (NENB 5620, BR5020057361348, Isotype). A–B. Herbarium box. C–D. Sessile sporocarps. E. Peridial platelet and spores by TL. F. Spores by TL. G. Dehisced sporocarp base by SEM. H. Platelet outer surface by SEM. Whole platelet showing well-defined edges. I. Edge of a platelet by SEM showing double peridium and no outgrowths on the edge. J. Spore by SEM. Scale bars: C–D = 0.1 mm, E–F, H = 10 µm, G = 50 µm, I–J = 5 µm." figureDoi-0="http://doi.org/10.5281/zenodo.10276061" figureDoi-1="http://doi.org/10.5281/zenodo.10276063" httpUri-0="https://zenodo.org/record/10276061/files/figure.png" httpUri-1="https://zenodo.org/record/10276063/files/figure.png" pageId="23" pageNumber="118">Figs. 7–8</figureCitation>
, 
<figureCitation id="1325389F9B0FFFC7FB780119FAE2FAC0" box="[1254,1326,1357,1383]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="16.[136,229,1716,1738]" captionStartId-1="17.[136,229,1772,1794]" captionTargetBox-0="[151,1436,903,1688]" captionTargetBox-1="[151,1436,602,1744]" captionTargetId-0="figure-298@16.[151,1436,903,1688]" captionTargetId-1="figure-164@17.[151,1436,602,1744]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 11. Licea mariae. A–H (MYX 11149, Holotype).A. Herbarium fragment. B–C. Sessile sporocarps. D. Peridial platelets by TL. E–F. Peridial platelets with row of conical outgrowths (arrows). G–H. Spores by TL. Scale bars: B–C = 0.1 mm, D–H = 10 µm." captionText-1="FIGURE 12. Licea mínima.A–B, E. G, I–K (MA-Fungi 23775, Neotype); C–D, F, H (MA-Fungi 17214).A. Herbarium box. B–D. Sessile sporocarps. E–F. Peridial platelets by TL. G–H. Spores by TL. I–J. Detail of inner peridium surface and platelet edges by SEM. K. Spore by SEM. Scale bars: B–D = 0.1 mm, E, I = 20 µm, F–H, J = 10 µm, K = 5 µm." figureDoi-0="http://doi.org/10.5281/zenodo.10276069" figureDoi-1="http://doi.org/10.5281/zenodo.10276071" httpUri-0="https://zenodo.org/record/10276069/files/figure.png" httpUri-1="https://zenodo.org/record/10276071/files/figure.png" pageId="23" pageNumber="118">11–12</figureCitation>
; 520, 
<figureCitation id="1325389F9B0FFFC7FAEA0119FF1CFA2D" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="16.[136,229,1716,1738]" captionStartId-1="17.[136,229,1772,1794]" captionTargetBox-0="[151,1436,903,1688]" captionTargetBox-1="[151,1436,602,1744]" captionTargetId-0="figure-298@16.[151,1436,903,1688]" captionTargetId-1="figure-164@17.[151,1436,602,1744]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 11. Licea mariae. A–H (MYX 11149, Holotype).A. Herbarium fragment. B–C. Sessile sporocarps. D. Peridial platelets by TL. E–F. Peridial platelets with row of conical outgrowths (arrows). G–H. Spores by TL. Scale bars: B–C = 0.1 mm, D–H = 10 µm." captionText-1="FIGURE 12. Licea mínima.A–B, E. G, I–K (MA-Fungi 23775, Neotype); C–D, F, H (MA-Fungi 17214).A. Herbarium box. B–D. Sessile sporocarps. E–F. Peridial platelets by TL. G–H. Spores by TL. I–J. Detail of inner peridium surface and platelet edges by SEM. K. Spore by SEM. Scale bars: B–D = 0.1 mm, E, I = 20 µm, F–H, J = 10 µm, K = 5 µm." figureDoi-0="http://doi.org/10.5281/zenodo.10276069" figureDoi-1="http://doi.org/10.5281/zenodo.10276071" httpUri-0="https://zenodo.org/record/10276069/files/figure.png" httpUri-1="https://zenodo.org/record/10276071/files/figure.png" pageId="23" pageNumber="118">Figs. 11–12</figureCitation>
, 
<figureCitation id="1325389F9B0FFFC7FF420124FF36FA2D" box="[220,250,1392,1418]" captionStart="FIGURE 14" captionStartId="20.[136,229,1504,1526]" captionTargetBox="[151,1436,192,1477]" captionTargetId="figure-22@20.[151,1436,192,1477]" captionTargetPageId="20" captionText="FIGURE 14. Licea nannengae. A–I (246 Pando, MA-Fungi 16056, Holotype). A. Herbarium box with holotype indicated with arrow. B–C. Sessile sporocarps. D–E. Peridial platelets by TL. F. spore by TL. G. Dehisced sporocarp by SEM. H. Detail of inner peridium surface at platelet edges by SEM. I. Spore by SEM. Scale bars: B–C = 0.1 mm, D–F, H = 10 µm, G = 50 µm, I = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276077" httpUri="https://zenodo.org/record/10276077/files/figure.png" pageId="23" pageNumber="118">14</figureCitation>
, all the images as 
<taxonomicName id="4C1E5F999B0FFFC7FE530125FDFBFA2D" box="[461,567,1392,1418]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="species" species="pusilla">
<emphasis id="B96AF8089B0FFFC7FE530125FDFBFA2D" box="[461,567,1392,1418]" italics="true" pageId="23" pageNumber="118">L. pusilla</emphasis>
</taxonomicName>
), 
<bibRefCitation id="EF8F59EB9B0FFFC7FDCE0124FCB7FA2D" author="Lado, C. &amp; Pando, F." box="[592,891,1392,1418]" pageId="23" pageNumber="118" pagination="1 - 323" refId="ref20332" refString="Lado, C. &amp; Pando, F. (1997) Myxomycetes, I. Ceratiomyxales, Echinosteliales, Licelaes, Trichiales. Flora Mycologica Iberica 2: 1 - 323." type="journal article" year="1997">Lado &amp; Pando (1997: 130</bibRefCitation>
, Figs. 37a–b), Yamamoto (2002a: 3. 
<figureCitation id="1325389F9B0FFFC7FABE0124FA6AFA2D" box="[1312,1446,1392,1418]" captionStart="FIGURE 1" captionStartId="3.[136,229,999,1021]" captionTargetBox="[151,1436,188,973]" captionTargetId="figure-19@3.[151,1436,188,973]" captionTargetPageId="3" captionText="FIGURE 1. Licea alpina. A–F (AH 28531, Holotype).A. Herbarium boxes and slides. B–D. Stalked and subsessile sporocarps. E. Spores and peridial platelets by TL. F. Spores by TL. Scale bars: B–D = 0.1 mm, E–F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10276049" httpUri="https://zenodo.org/record/10276049/files/figure.png" pageId="23" pageNumber="118">Figs. 1A–D</figureCitation>
; 2004: 23, 
<figureCitation id="1325389F9B0FFFC7FF6601C7FEB0FA0A" box="[248,380,1427,1453]" captionStart="FIGURE 3" captionStartId="5.[136,229,1807,1829]" captionTargetBox="[151,1436,566,1780]" captionTargetId="figure-149@5.[151,1436,566,1780]" captionTargetPageId="5" captionText="FIGURE 3. Licea aurea. A–G. (dwb 3815, MA-Fungi 91750, Holotype), H–J. (dwb 3533). A. Herbarium box. B–D. Sporocarps. E. Whole sporocarp and spores by TL. F. Fragment of the inner surface of the peridium and spores. G. Spores by TL with thick walls and thinner germination area. H. Whole open sporocarp showing spores by SEM. I. Group of spores by SEM. J. Spore by SEM. Scale bars: B–D = 0.1 mm, E–G, I = 10 µm, H = 50 µm, J = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276053" httpUri="https://zenodo.org/record/10276053/files/figure.png" pageId="23" pageNumber="118">Figs. 3A–C</figureCitation>
; 2006: 35, 
<figureCitation id="1325389F9B0FFFC7FE6601C7FD47FA09" box="[504,651,1427,1454]" captionStart="FIGURE 15" captionStartId="21.[136,229,1406,1428]" captionTargetBox="[151,1436,530,1379]" captionTargetId="figure-149@21.[151,1436,530,1379]" captionTargetPageId="21" captionText="FIGURE 15. Licea nigromarginata.A–D (NENB 13886, BR5020071551626, Isotype).A.Isotype slide.B–D. Spores and peridial platelets by TL. Scale bars: B–C = 50 µm, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10276079" httpUri="https://zenodo.org/record/10276079/files/figure.png" pageId="23" pageNumber="118">Figs. 15A–D</figureCitation>
; 2021: 148, Figs. A–D), Poulain 
<emphasis id="B96AF8089B0FFFC7FC6501C0FBFFFA0A" box="[1019,1075,1427,1453]" italics="true" pageId="23" pageNumber="118">et al.</emphasis>
(2011: 70), Ronikier 
<emphasis id="B96AF8089B0FFFC7FABA01C0FA90FA0A" box="[1316,1372,1427,1453]" italics="true" pageId="23" pageNumber="118">et al.</emphasis>
(2017: 253, 
<figureCitation id="1325389F9B0FFFC7FF5F01E2FE85FA76" box="[193,329,1462,1489]" captionStart="FIGURE 4" captionStartId="7.[136,229,1428,1450]" captionTargetBox="[151,1436,187,1401]" captionTargetId="figure-19@7.[151,1436,187,1401]" captionTargetPageId="7" captionText="FIGURE 4. Licea belmontiana. A–J (NENB-5879, BR-MYC 5020057898868, Holotype). A. Herbarium box. B–C. Sessile sporocarps. D. Peridial platelets by TL. E. Spores by TL. F. Dehisced sporocarp by SEM. G–H. Peridial platelets by SEM showing margins without outgrowths (arrows). I–J. Warted spores by SEM. Scale bars: B–C = 0.1 mm, D–G = 10 µm, H–J = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276055" httpUri="https://zenodo.org/record/10276055/files/figure.png" pageId="23" pageNumber="118">Figs. 4G–K</figureCitation>
), López-Villalba &amp; Moreno (2020: 149, 
<figureCitation id="1325389F9B0FFFC7FC8601E2FC5EFA77" box="[792,914,1462,1488]" captionStart="FIGURE 3" captionStartId="5.[136,229,1807,1829]" captionTargetBox="[151,1436,566,1780]" captionTargetId="figure-149@5.[151,1436,566,1780]" captionTargetPageId="5" captionText="FIGURE 3. Licea aurea. A–G. (dwb 3815, MA-Fungi 91750, Holotype), H–J. (dwb 3533). A. Herbarium box. B–D. Sporocarps. E. Whole sporocarp and spores by TL. F. Fragment of the inner surface of the peridium and spores. G. Spores by TL with thick walls and thinner germination area. H. Whole open sporocarp showing spores by SEM. I. Group of spores by SEM. J. Spore by SEM. Scale bars: B–D = 0.1 mm, E–G, I = 10 µm, H = 50 µm, J = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276053" httpUri="https://zenodo.org/record/10276053/files/figure.png" pageId="23" pageNumber="118">Figs. 3a–d</figureCitation>
), 
<bibRefCitation id="EF8F59EB9B0FFFC7FC3901E2FAE9FA76" author="Johannesen, E. W. &amp; Vetlesen, P." box="[935,1317,1462,1489]" pageId="23" pageNumber="118" pagination="1 - 138" refId="ref19773" refString="Johannesen, E. W. &amp; Vetlesen, P. (2020) New and rare myxomycetes (Mycetozoa, Myxogastria) in Norway, including a complete checklist of Norwegian myxomycetes. Agarica 40: 1 - 138." type="journal article" year="2020">Johannesen &amp; Vetlesen (2020: 79</bibRefCitation>
, Figs 40A), 
<bibRefCitation id="EF8F59EB9B0FFFC7FF16018EFE62FA53" author="Bortnikov, F. M. &amp; Gmoshinskiy, V. I. &amp; Novozhilov, Y. K." box="[136,430,1498,1524]" pageId="23" pageNumber="118" pagination="21 - 48" refId="ref19320" refString="Bortnikov, F. M., Gmoshinskiy, V. I. &amp; Novozhilov, Y. K. (2022) Species of Licea Schrad. (Myxomycetes) in Kedrovaya Pad State Nature Biosphere Reserve (Far East, Ruassia), including two new species. Phytotaxa 541: 21 - 48. https: // doi. org / 10.11646 / phytotaxa. 541.1.3" type="journal article" year="2022">
Bortnikov 
<emphasis id="B96AF8089B0FFFC7FE9F018EFEF5FA53" box="[257,313,1498,1524]" italics="true" pageId="23" pageNumber="118">et al.</emphasis>
(2022: 39
</bibRefCitation>
, 
<figureCitation id="1325389F9B0FFFC7FE24018EFD8CFA53" box="[442,576,1497,1524]" captionStart="FIGURE 15" captionStartId="21.[136,229,1406,1428]" captionTargetBox="[151,1436,530,1379]" captionTargetId="figure-149@21.[151,1436,530,1379]" captionTargetPageId="21" captionText="FIGURE 15. Licea nigromarginata.A–D (NENB 13886, BR5020071551626, Isotype).A.Isotype slide.B–D. Spores and peridial platelets by TL. Scale bars: B–C = 50 µm, D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.10276079" httpUri="https://zenodo.org/record/10276079/files/figure.png" pageId="23" pageNumber="118">Figs. 15a–k</figureCitation>
).
</paragraph>
<paragraph id="8BA1241A9B0FFFC7FF2301A9FBE8F96F" blockId="23.[136,1452,864,1736]" pageId="23" pageNumber="118">
Notes. This species was published by 
<bibRefCitation id="EF8F59EB9B0FFFC7FDED01A9FCD2F9B0" author="Meylan, C." box="[627,798,1533,1559]" pageId="23" pageNumber="118" pagination="81 - 90" refId="ref21369" refString="Meylan, C. (1933) Recherches sur les Myxomycetes du Jura 1930 - 31 - 32. Bulletin Societe Vaudoise Sciences Naturelles 58: 81 - 90. https: // doi. org / 10.5169 / seals- 271283" type="journal article" year="1933">Meylan (1933)</bibRefCitation>
, as a variety of 
<taxonomicName id="4C1E5F999B0FFFC7FC4701AAFADFF9B0" authority="Schrad. However" authorityName="Schrad. However" box="[985,1299,1533,1559]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="species" species="pusilla">
<emphasis id="B96AF8089B0FFFC7FC4701AAFB89F9B0" box="[985,1093,1533,1559]" italics="true" pageId="23" pageNumber="118">L. pusilla</emphasis>
Schrad. However
</taxonomicName>
based on the number of differences between these taxa, such as spore size, spore ornamentation and the peridium, 
<bibRefCitation id="EF8F59EB9B0FFFC7FB720275FAAEF99D" author="Ing, B." box="[1260,1378,1568,1595]" pageId="23" pageNumber="118" pagination="439 - 446" refId="ref19729" refString="Ing, B. (1982) Notes on Myxomycetes III. Transactions British Mycological Society 78 (3): 439 - 446. https: // doi. org / 10.1016 / S 0007 - 1536 (82) 80150 - 2" type="journal article" year="1982">Ing (1982)</bibRefCitation>
raised the variety to specific rank (see above). The minute size of sporocarps and the fact that it has been often included as a variety in the species 
<taxonomicName id="4C1E5F999B0FFFC7FE080233FDCDF927" box="[406,513,1638,1664]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="species" species="pusilla">
<emphasis id="B96AF8089B0FFFC7FE080233FDCDF927" box="[406,513,1638,1664]" italics="true" pageId="23" pageNumber="118">L. pusilla</emphasis>
</taxonomicName>
makes the distribution of 
<taxonomicName id="4C1E5F999B0FFFC7FCB00233FC7EF927" box="[814,946,1639,1664]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="species" species="pygmaea">
<emphasis id="B96AF8089B0FFFC7FCB00233FC7EF927" box="[814,946,1639,1664]" italics="true" pageId="23" pageNumber="118">L. pygmaea</emphasis>
</taxonomicName>
unclear. As stated above, we agree with 
<bibRefCitation id="EF8F59EB9B0FFFC7FA180233FF1FF903" author="Ing, B." pageId="23" pageNumber="118" pagination="439 - 446" refId="ref19729" refString="Ing, B. (1982) Notes on Myxomycetes III. Transactions British Mycological Society 78 (3): 439 - 446. https: // doi. org / 10.1016 / S 0007 - 1536 (82) 80150 - 2" type="journal article" year="1982">Ing (1982)</bibRefCitation>
and consider 
<taxonomicName id="4C1E5F999B0FFFC7FEED02DEFE12F903" box="[371,478,1674,1700]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="species" species="pusilla">
<emphasis id="B96AF8089B0FFFC7FEED02DEFE12F903" box="[371,478,1674,1700]" italics="true" pageId="23" pageNumber="118">L. pusilla</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1E5F999B0FFFC7FD8902DEFD57F904" box="[535,667,1674,1699]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="23" pageNumber="118" phylum="Mycetozoa" rank="species" species="pygmaea">
<emphasis id="B96AF8089B0FFFC7FD8902DEFD57F904" box="[535,667,1674,1699]" italics="true" pageId="23" pageNumber="118">L. pygmaea</emphasis>
</taxonomicName>
to be separate species based on the differences found by SEM in the ornamentation of the peridium and platelet edges, as well as spore colour and size.
</paragraph>
</subSubSection>
</treatment>
</document>