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<document id="C46EBBEBE5ECF849CFC15B1C59088112" ID-DOI="10.11646/phytotaxa.629.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="10276047" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701852444489" checkinUser="plazi" docAuthor="Basanta, Diana Wrigley De, Mier, Carlos De &amp; Lado, Carlos" docDate="2023" docId="03B7950C9B0AFFC3FF16028FFCE1F823" docLanguage="en" docName="phytotaxa.629.2.1.pdf" docOrigin="Phytotaxa 629 (2)" docSource="https://phytotaxa.mapress.com/pt/article/download/phytotaxa.629.2.1/51333" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.11:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="11" docTitle="Licea nannengae Pando &amp; Lado, Mycotaxon" docType="treatment" docVersion="2" lastPageNumber="114" masterDocId="FF8EED749B18FFD0FF9E0454FFCCFFA7" masterDocTitle="A taxonomic revision of the species of Licea subg. Licea (Myxomycetes)" masterLastPageNumber="128" masterPageNumber="95" pageNumber="113" updateTime="1701852992442" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1D397D228E8AC907846C5FAF5F6F82DC">A taxonomic revision of the species of Licea subg. Licea (Myxomycetes)</mods:title>
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<mods:namePart id="FB84B6A10D87F46BE4B22F7E8423D60A">Basanta, Diana Wrigley De</mods:namePart>
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<mods:affiliation id="981C1851A6EC36451C2288AA0C8F1EB3">Real Jardín Botánico, CSIC, Plaza de Murillo 2, 28014 Madrid (Spain). &amp; dwbasanta @ gmail. com, https: // orcid. org / 0000 - 0002 - 7700 - 8399</mods:affiliation>
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<mods:namePart id="BDCC62063E9D895D602AE67C10580ED8">Mier, Carlos De</mods:namePart>
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<mods:affiliation id="B58E5D4F802417C9759655782FC437D7">Real Jardín Botánico, CSIC, Plaza de Murillo 2, 28014 Madrid (Spain). &amp; cmier @ rjb. csic. es, https: // orcid. org / 0009 - 0008 - 1431 - 4410</mods:affiliation>
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<mods:namePart id="FF3A7A217BC730301E368AB108F1E1CA">Lado, Carlos</mods:namePart>
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<mods:affiliation id="6855B155E1B40022016A50C366340663">Real Jardín Botánico, CSIC, Plaza de Murillo 2, 28014 Madrid (Spain). &amp; lado @ rjb. csic. es, https: // orcid. org / 0000 - 0002 - 6135 - 2873</mods:affiliation>
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<paragraph id="8BA1241A9B0AFFC2FF16028FFBC1F952" blockId="18.[136,1037,1755,1782]" box="[136,1037,1755,1782]" pageId="18" pageNumber="113">
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<emphasis id="B96AF8089B0AFFC2FF16028FFF67F952" bold="true" box="[136,171,1755,1781]" pageId="18" pageNumber="113">17.</emphasis>
<taxonomicName id="4C1E5F999B0AFFC2FF2C028FFD7EF951" authority="Pando &amp; Lado, Mycotaxon" authorityName="Pando &amp; Lado, Mycotaxon" box="[178,690,1755,1782]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="18" pageNumber="113" phylum="Mycetozoa" rank="species" species="nannengae">
<treatmentCitation id="0ABF020B9B0AFFC2FF2C028FFEB6F951" author="Pando &amp; Lado" box="[178,378,1755,1782]" issue="2" journal="Mycotaxon" page="299" pageId="18" pageNumber="113" score="9.729594737742866" volume="31" year="1988">
<emphasis id="B96AF8089B0AFFC2FF2C028FFEB6F951" bold="true" box="[178,378,1755,1782]" italics="true" pageId="18" pageNumber="113">Licea nannengae</emphasis>
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Pando &amp; Lado, Mycotaxon
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31(2): 299 (1988) 
<figureCitation id="1325389F9B0AFFC2FC19028FFBC1F952" box="[903,1037,1755,1781]" captionStart="FIGURE 14" captionStartId="20.[136,229,1504,1526]" captionTargetBox="[151,1436,192,1477]" captionTargetId="figure-22@20.[151,1436,192,1477]" captionTargetPageId="20" captionText="FIGURE 14. Licea nannengae. A–I (246 Pando, MA-Fungi 16056, Holotype). A. Herbarium box with holotype indicated with arrow. B–C. Sessile sporocarps. D–E. Peridial platelets by TL. F. spore by TL. G. Dehisced sporocarp by SEM. H. Detail of inner peridium surface at platelet edges by SEM. I. Spore by SEM. Scale bars: B–C = 0.1 mm, D–F, H = 10 µm, G = 50 µm, I = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276077" httpUri="https://zenodo.org/record/10276077/files/figure.png" pageId="18" pageNumber="113">Figs. 14A–I</figureCitation>
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<paragraph id="8BA1241A9B0AFFC2FF160349FCDFF7A8" blockId="18.[136,1452,1820,2063]" pageId="18" pageNumber="113">
Sporophores sporocarpic, scattered to gregarious, sessile. Sporocarps subglosose, 
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diam., ridged, dark brown to yellow ochraceous. Peridium double, outer layer gelatinous, covered with refuse material, inner translucent and iridescent very thin, pale olive, smooth; dehiscence along ridges into platelets and lobes leaving a shiny base. Spores free, dark brown in mass, olivaceous brown by TL (
<figureCitation id="1325389F9B0AFFC2FCB303DDFC5DF804" box="[813,913,1929,1955]" captionStart="FIGURE 14" captionStartId="20.[136,229,1504,1526]" captionTargetBox="[151,1436,192,1477]" captionTargetId="figure-22@20.[151,1436,192,1477]" captionTargetPageId="20" captionText="FIGURE 14. Licea nannengae. A–I (246 Pando, MA-Fungi 16056, Holotype). A. Herbarium box with holotype indicated with arrow. B–C. Sessile sporocarps. D–E. Peridial platelets by TL. F. spore by TL. G. Dehisced sporocarp by SEM. H. Detail of inner peridium surface at platelet edges by SEM. I. Spore by SEM. Scale bars: B–C = 0.1 mm, D–F, H = 10 µm, G = 50 µm, I = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276077" httpUri="https://zenodo.org/record/10276077/files/figure.png" pageId="18" pageNumber="113">Fig. 14F</figureCitation>
), with a pale area, wall thick with thinner area, globose, 9.5–13.5 µm diam., smooth. By SEM the inner peridium surface is densely warted, with some warts joined in a loose reticulum, right up to the platelet margins (
<figureCitation id="1325389F9B0AFFC2FD700385FC4AF84C" box="[750,902,2001,2027]" captionStart="FIGURE 14" captionStartId="20.[136,229,1504,1526]" captionTargetBox="[151,1436,192,1477]" captionTargetId="figure-22@20.[151,1436,192,1477]" captionTargetPageId="20" captionText="FIGURE 14. Licea nannengae. A–I (246 Pando, MA-Fungi 16056, Holotype). A. Herbarium box with holotype indicated with arrow. B–C. Sessile sporocarps. D–E. Peridial platelets by TL. F. spore by TL. G. Dehisced sporocarp by SEM. H. Detail of inner peridium surface at platelet edges by SEM. I. Spore by SEM. Scale bars: B–C = 0.1 mm, D–F, H = 10 µm, G = 50 µm, I = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276077" httpUri="https://zenodo.org/record/10276077/files/figure.png" pageId="18" pageNumber="113">Figs. 14G–H</figureCitation>
), with no prominent outgrowths; the epispore is smooth with a thinner area appearing wrinkled (
<figureCitation id="1325389F9B0AFFC2FD3C03A1FCC8F7A8" box="[674,772,2037,2063]" captionStart="FIGURE 14" captionStartId="20.[136,229,1504,1526]" captionTargetBox="[151,1436,192,1477]" captionTargetId="figure-22@20.[151,1436,192,1477]" captionTargetPageId="20" captionText="FIGURE 14. Licea nannengae. A–I (246 Pando, MA-Fungi 16056, Holotype). A. Herbarium box with holotype indicated with arrow. B–C. Sessile sporocarps. D–E. Peridial platelets by TL. F. spore by TL. G. Dehisced sporocarp by SEM. H. Detail of inner peridium surface at platelet edges by SEM. I. Spore by SEM. Scale bars: B–C = 0.1 mm, D–F, H = 10 µm, G = 50 µm, I = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276077" httpUri="https://zenodo.org/record/10276077/files/figure.png" pageId="18" pageNumber="113">Fig. 14 I</figureCitation>
).
</paragraph>
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<emphasis id="B96AF8089B0BFFC3FF16002DFEC7FB29" bold="true" box="[136,267,1145,1167]" pageId="19" pageNumber="114">FIGURE 13.</emphasis>
<emphasis id="B96AF8089B0BFFC3FE8F002EFE10FB29" box="[273,476,1145,1167]" italics="true" pageId="19" pageNumber="114">
<taxonomicName id="4C1E5F999B0BFFC3FE8F002EFE14FB29" box="[273,472,1145,1167]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="morchelloides">Licea morchelloides</taxonomicName>
.
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A–D (NENB 13887, BR5020071552630, Isotype). A. Slide. B–D. platelets and spores by TL. Scale bars: B–D = 10 µm.
</paragraph>
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<paragraph id="8BA1241A9B0BFFC3FF2300B5FC02FA87" blockId="19.[136,1452,1249,1924]" pageId="19" pageNumber="114">
<emphasis id="B96AF8089B0BFFC3FF2300B5FE58FB5C" box="[189,404,1249,1275]" italics="true" pageId="19" pageNumber="114">Material examined</emphasis>
: 
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. 
<collectingCountry id="F309648A9B0BFFC3FDB400B6FDB0FB5B" box="[554,636,1250,1276]" name="Spain" pageId="19" pageNumber="114">SPAIN</collectingCountry>
. Soria, Cubillos, 30TWM0421, 
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, corteza de 
<taxonomicName id="4C1E5F999B0BFFC3FB4E00B6FA60FB5C" box="[1232,1452,1249,1275]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="19" pageNumber="114" phylum="Tracheophyta" rank="species" species="thurifera">
<emphasis id="B96AF8089B0BFFC3FB4E00B6FA60FB5C" box="[1232,1452,1249,1275]" italics="true" pageId="19" pageNumber="114">Juniperus thurifera</emphasis>
</taxonomicName>
cultivado en cámara húmeda, 
<date id="FFA002DA9B0BFFC3FE490151FD91FAB8" box="[471,605,1285,1311]" pageId="19" pageNumber="114" value="1986-03-26">26-III-1986</date>
, 246 Pando (MA-Fungi 16056)!.
</paragraph>
<paragraph id="8BA1241A9B0BFFC3FF23017DFE33FAE3" blockId="19.[136,1452,1249,1924]" box="[189,511,1321,1348]" pageId="19" pageNumber="114">
<emphasis id="B96AF8089B0BFFC3FF23017DFED0FAE4" box="[189,284,1321,1347]" italics="true" pageId="19" pageNumber="114">Habitat:</emphasis>
bark of living trees.
</paragraph>
<paragraph id="8BA1241A9B0BFFC3FF230119FF33FA2C" blockId="19.[136,1452,1249,1924]" pageId="19" pageNumber="114">
<emphasis id="B96AF8089B0BFFC3FF230119FE83FAC0" box="[189,335,1357,1383]" italics="true" pageId="19" pageNumber="114">Distribution:</emphasis>
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(?), 
<collectingCountry id="F309648A9B0BFFC3FE740119FD94FAC0" box="[490,600,1357,1383]" name="Lithuania" pageId="19" pageNumber="114">Lithuania</collectingCountry>
, 
<collectingCountry id="F309648A9B0BFFC3FDFB0119FC8DFAC0" box="[613,833,1357,1383]" name="Russia" pageId="19" pageNumber="114">Russian Federation</collectingCountry>
, 
<collectingCountry id="F309648A9B0BFFC3FCD30119FC53FAC0" box="[845,927,1357,1383]" name="Ireland" pageId="19" pageNumber="114">Ireland</collectingCountry>
, 
<collectingCountry id="F309648A9B0BFFC3FC32011AFBD9FACF" box="[940,1045,1358,1384]" name="Germany" pageId="19" pageNumber="114">Germany</collectingCountry>
, 
<collectingCountry id="F309648A9B0BFFC3FBBC0119FBA9FAC0" box="[1058,1125,1357,1383]" name="Spain" pageId="19" pageNumber="114">Spain</collectingCountry>
, 
<collectingCountry id="F309648A9B0BFFC3FBEC0119FB34FAC0" box="[1138,1272,1357,1383]" name="Kazakhstan" pageId="19" pageNumber="114">Kazakhstan</collectingCountry>
, 
<collectingCountry id="F309648A9B0BFFC3FA9A0119FA90FAC0" box="[1284,1372,1357,1383]" name="Mexico" pageId="19" pageNumber="114">Mexico</collectingCountry>
, 
<collectingCountry id="F309648A9B0BFFC3FAF70119FA6BFAC0" box="[1385,1447,1357,1383]" name="Cuba" pageId="19" pageNumber="114">Cuba</collectingCountry>
, 
<collectingCountry id="F309648A9B0BFFC3FF160125FF37FA2C" box="[136,251,1393,1419]" name="Argentina" pageId="19" pageNumber="114">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8BA1241A9B0BFFC3FF2301C2FB98FA73" blockId="19.[136,1452,1249,1924]" pageId="19" pageNumber="114">
<emphasis id="B96AF8089B0BFFC3FF2301C2FECCFA08" box="[189,256,1430,1455]" italics="true" pageId="19" pageNumber="114">Icon.:</emphasis>
Pando &amp; Lado (1988: 300, 
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), 
<bibRefCitation id="EF8F59EB9B0BFFC3FD1101C1FC78FA17" author="Lado, C. &amp; Pando, F." box="[655,948,1429,1456]" pageId="19" pageNumber="114" pagination="1 - 323" refId="ref20332" refString="Lado, C. &amp; Pando, F. (1997) Myxomycetes, I. Ceratiomyxales, Echinosteliales, Licelaes, Trichiales. Flora Mycologica Iberica 2: 1 - 323." type="journal article" year="1997">Lado &amp; Pando (1997: 122</bibRefCitation>
, Figs. 32a–b), 
<bibRefCitation id="EF8F59EB9B0BFFC3FBC701C1FF6AFA73" author="Johannesen, E. W. &amp; Vetlesen, P." pageId="19" pageNumber="114" pagination="1 - 138" refId="ref19773" refString="Johannesen, E. W. &amp; Vetlesen, P. (2020) New and rare myxomycetes (Mycetozoa, Myxogastria) in Norway, including a complete checklist of Norwegian myxomycetes. Agarica 40: 1 - 138." type="journal article" year="2020">Johannesen &amp; Vetlesen (2020: 77</bibRefCitation>
, Figs. 39A–C as 
<taxonomicName id="4C1E5F999B0BFFC3FEF901EEFD86FA74" box="[359,586,1465,1491]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" isUncertain="true" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="nannengae">
<emphasis id="B96AF8089B0BFFC3FEF901EEFE69FA74" box="[359,421,1466,1491]" italics="true" pageId="19" pageNumber="114">Licea</emphasis>
cf. 
<emphasis id="B96AF8089B0BFFC3FE5101EEFD86FA74" box="[463,586,1466,1491]" italics="true" pageId="19" pageNumber="114">nannengae</emphasis>
</taxonomicName>
), 
<bibRefCitation id="EF8F59EB9B0BFFC3FDFC01EDFC25FA73" author="Nannenga-Bremekamp, N. E." box="[610,1001,1465,1492]" pageId="19" pageNumber="114" refId="ref21685" refString="Nannenga-Bremekamp, N. E. (2022) Descriptive, illustrated keys to the world's Myxomycetes. A posthumous publication with foreword, annotations, updates and editing by C. Lado. Editorial CSIC. Madrid, Espana." type="book" year="2022">Nannenga-Bremekamp (2022: 533</bibRefCitation>
, 
<figureCitation id="1325389F9B0BFFC3FC6A01EDFB8BFA73" box="[1012,1095,1465,1492]" captionStart="FIGURE 17" captionStartId="24.[136,229,1392,1414]" captionTargetBox="[151,1436,187,1364]" captionTargetId="figure-22@24.[151,1436,187,1364]" captionTargetPageId="24" captionText="FIGURE 17. Licea pygmaea.A–L (Myxo 055, LAU, Lectotype).A. Herbarium box. B–E. Sessile sporocarps. F–G. Peridial platelets and spore by TL. H. Whole open sporocarp by SEM. I–J. Peridial platelet margins by SEM. K–L. Spores by SEM. Scale bars: B = 0.1 mm, C–E, H = 50 µm, F–G = 10 µm, I–L = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276083" httpUri="https://zenodo.org/record/10276083/files/figure.png" pageId="19" pageNumber="114">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8BA1241A9B0BFFC3FF23018AFCE1F823" blockId="19.[136,1452,1249,1924]" pageId="19" pageNumber="114">
Notes. This species was described as having a smooth inner surface of the inner peridium by TL. However, by SEM, which was not available at the time of the description, the inner peridium looks densely warted (
<figureCitation id="1325389F9B0BFFC3FAAA0255FA53F9BB" box="[1332,1439,1537,1564]" captionStart="FIGURE 14" captionStartId="20.[136,229,1504,1526]" captionTargetBox="[151,1436,192,1477]" captionTargetId="figure-22@20.[151,1436,192,1477]" captionTargetPageId="20" captionText="FIGURE 14. Licea nannengae. A–I (246 Pando, MA-Fungi 16056, Holotype). A. Herbarium box with holotype indicated with arrow. B–C. Sessile sporocarps. D–E. Peridial platelets by TL. F. spore by TL. G. Dehisced sporocarp by SEM. H. Detail of inner peridium surface at platelet edges by SEM. I. Spore by SEM. Scale bars: B–C = 0.1 mm, D–F, H = 10 µm, G = 50 µm, I = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276077" httpUri="https://zenodo.org/record/10276077/files/figure.png" pageId="19" pageNumber="114">Fig. 14H</figureCitation>
). In the collection of the 
<typeStatus id="54A59AB89B0BFFC3FE0D0271FE34F998" box="[403,504,1573,1599]" pageId="19" pageNumber="114" type="holotype">holotype</typeStatus>
, that is marked with arrows (14A), we have found sporocarps of a different species mixed in with 
<taxonomicName id="4C1E5F999B0BFFC3FEB4021EFDBCF9C4" authority="Pando &amp; Lado" authorityName="Pando &amp; Lado" box="[298,624,1609,1636]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="nannengae">
<emphasis id="B96AF8089B0BFFC3FEB4021EFE0FF9C4" box="[298,451,1610,1635]" italics="true" pageId="19" pageNumber="114">L. nannengae</emphasis>
Pando &amp; Lado
</taxonomicName>
, with an almost smooth peridium and spiny spores. This may have lead to misidentification, such as reports from Colorado (
<bibRefCitation id="EF8F59EB9B0BFFC3FD2B0239FC77F92F" author="Novozhilov, Y. K. &amp; Mitchell, D. W. &amp; Schnittler, M." box="[693,955,1645,1672]" pageId="19" pageNumber="114" pagination="243 - 258" refId="ref21890" refString="Novozhilov, Y. K., Mitchell, D. W. &amp; Schnittler, M. (2003) Myxomycete biodiversity of the Colorado Plateau. Mycological Progress 2 (4): 243 - 258. https: // doi. org / 10.1007 / s 11557 - 006 - 0062 - 3" type="journal article" year="2003">
Novozhilov 
<emphasis id="B96AF8089B0BFFC3FCDC023AFCB7F920" box="[834,891,1645,1671]" italics="true" pageId="19" pageNumber="114">et al.</emphasis>
2003
</bibRefCitation>
) and 
<collectingCountry id="F309648A9B0BFFC3FC670239FB4CF920" box="[1017,1152,1645,1671]" name="Madagascar" pageId="19" pageNumber="114">Madagascar</collectingCountry>
(Wrigley de Basanta 
<emphasis id="B96AF8089B0BFFC3FAED023AFA60F920" box="[1395,1452,1645,1671]" italics="true" pageId="19" pageNumber="114">et al.</emphasis>
2013) with a SEM image showing ornamented spores, and could affect distribution data. 
<taxonomicName id="4C1E5F999B0BFFC3FBEA02C6FAF8F90C" box="[1140,1332,1682,1707]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="nannengae">
<emphasis id="B96AF8089B0BFFC3FBEA02C6FAF8F90C" box="[1140,1332,1682,1707]" italics="true" pageId="19" pageNumber="114">Licea nannengae</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C1E5F999B0BFFC3FF1602E2FE4CF968" authority="Nann." authorityName="Nann." box="[136,384,1717,1743]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="belmontiana">
<emphasis id="B96AF8089B0BFFC3FF1602E2FEF8F968" box="[136,308,1717,1743]" italics="true" pageId="19" pageNumber="114">L. belmontiana</emphasis>
Nann.
</taxonomicName>
-Bremek., however that species has darker brown sporocarps, not yellowish brown (compare 
<figureCitation id="1325389F9B0BFFC3FF16028DFF12F953" box="[136,222,1753,1780]" captionStart="FIGURE 4" captionStartId="7.[136,229,1428,1450]" captionTargetBox="[151,1436,187,1401]" captionTargetId="figure-19@7.[151,1436,187,1401]" captionTargetPageId="7" captionText="FIGURE 4. Licea belmontiana. A–J (NENB-5879, BR-MYC 5020057898868, Holotype). A. Herbarium box. B–C. Sessile sporocarps. D. Peridial platelets by TL. E. Spores by TL. F. Dehisced sporocarp by SEM. G–H. Peridial platelets by SEM showing margins without outgrowths (arrows). I–J. Warted spores by SEM. Scale bars: B–C = 0.1 mm, D–G = 10 µm, H–J = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276055" httpUri="https://zenodo.org/record/10276055/files/figure.png" pageId="19" pageNumber="114">Fig. 4C</figureCitation>
with 
<figureCitation id="1325389F9B0BFFC3FE81028DFE4EF953" box="[287,386,1753,1780]" captionStart="FIGURE 14" captionStartId="20.[136,229,1504,1526]" captionTargetBox="[151,1436,192,1477]" captionTargetId="figure-22@20.[151,1436,192,1477]" captionTargetPageId="20" captionText="FIGURE 14. Licea nannengae. A–I (246 Pando, MA-Fungi 16056, Holotype). A. Herbarium box with holotype indicated with arrow. B–C. Sessile sporocarps. D–E. Peridial platelets by TL. F. spore by TL. G. Dehisced sporocarp by SEM. H. Detail of inner peridium surface at platelet edges by SEM. I. Spore by SEM. Scale bars: B–C = 0.1 mm, D–F, H = 10 µm, G = 50 µm, I = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.10276077" httpUri="https://zenodo.org/record/10276077/files/figure.png" pageId="19" pageNumber="114">Fig. 14B</figureCitation>
taken with the same optical instruments). The spores of 
<taxonomicName id="4C1E5F999B0BFFC3FB9B028EFB7CF954" box="[1029,1200,1753,1779]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="belmontiana">
<emphasis id="B96AF8089B0BFFC3FB9B028EFB7CF954" box="[1029,1200,1753,1779]" italics="true" pageId="19" pageNumber="114">L. belmontiana</emphasis>
</taxonomicName>
are lighter brown and not olivaceous and the inner peridial markings, as seen by SEM, are different in each species. In addition, the spores of 
<taxonomicName id="4C1E5F999B0BFFC3FF160376FEFEF89C" box="[136,306,1825,1851]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="belmontiana">
<emphasis id="B96AF8089B0BFFC3FF160376FEFEF89C" box="[136,306,1825,1851]" italics="true" pageId="19" pageNumber="114">L. belmontiana</emphasis>
</taxonomicName>
are ornamented, as seen by SEM, whereas those of 
<taxonomicName id="4C1E5F999B0BFFC3FCE40376FBDFF89C" box="[890,1043,1826,1851]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="nannengae">
<emphasis id="B96AF8089B0BFFC3FCE40376FBDFF89C" box="[890,1043,1826,1851]" italics="true" pageId="19" pageNumber="114">L. nannengae</emphasis>
</taxonomicName>
are entirely smooth. 
<taxonomicName id="4C1E5F999B0BFFC3FA9E0376FE73F8F8" authority="H. W. Keller &amp; T. E. Brooks" authorityName="H. W. Keller &amp; T. E. Brooks" authorityYear="1977" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="denudescens">
<emphasis id="B96AF8089B0BFFC3FA9E0376FA60F89C" box="[1280,1452,1825,1851]" italics="true" pageId="19" pageNumber="114">L. denudescens</emphasis>
H.W. Keller &amp; T.E. Brooks
</taxonomicName>
differs from 
<taxonomicName id="4C1E5F999B0BFFC3FDC90312FD3DF8F8" box="[599,753,1862,1887]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Licea" kingdom="Protozoa" order="Dictyosteliales" pageId="19" pageNumber="114" phylum="Mycetozoa" rank="species" species="nannengae">
<emphasis id="B96AF8089B0BFFC3FDC90312FD3DF8F8" box="[599,753,1862,1887]" italics="true" pageId="19" pageNumber="114">L. nannengae</emphasis>
</taxonomicName>
by darker, larger sporocarps (
<quantity id="4CE689FF9B0BFFC3FBD90312FB1DF8C7" box="[1095,1233,1862,1888]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="19" pageNumber="114" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 mm</quantity>
diam. vs. 
<quantity id="4CE689FF9B0BFFC3FAD70311FF78F823" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="19" pageNumber="114" unit="mm" value="0.125" valueMax="0.2" valueMin="0.05">0.05–0.2 mm</quantity>
diam.), sporocarps lacking ridges, and by darker spores.
</paragraph>
</subSubSection>
</treatment>
</document>