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<document id="C8B2C4B68008340C0880D625AAFBDF4B" ID-DOI="10.11646/zootaxa.3734.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5275595" ID-ZooBank="36ED88E6-2CEB-4071-8429-A39901B8B9BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629926016628" checkinUser="felipe" docAuthor="Simaiakis, Stylianos Michail &amp; Edgecombe, Gregory D." docDate="2013" docId="03B68458FFBEFFF9FF56A9DBFB6AF8CF" docLanguage="en" docName="zt03734p198.pdf" docOrigin="Zootaxa 3734 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhysida stuhlmanni subsp. stuhlmanni stuhlmanni Kraepelin 1903" docType="treatment" docVersion="4" lastPageNumber="183" masterDocId="FF8FFC20FFB0FFF7FFC1AA2DFF95FFA1" masterDocTitle="Scolopendromorph centipedes (Chilopoda: Scolopendromorpha) in the Natural History Museum (London): A review of the hitherto unidentified species collected in Africa, with remarks on taxonomy and distribution, and a new species of Otostigmus (Parotostigmus)" masterLastPageNumber="198" masterPageNumber="169" pageNumber="183" updateTime="1699128448735" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7E16F3384E8ED174BDA9DCC775F44B70">Scolopendromorph centipedes (Chilopoda: Scolopendromorpha) in the Natural History Museum (London): A review of the hitherto unidentified species collected in Africa, with remarks on taxonomy and distribution, and a new species of Otostigmus (Parotostigmus)</mods:title>
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<mods:namePart id="2C5D0E4247DD9BAEBB64E6BC401B9D2F">Simaiakis, Stylianos Michail</mods:namePart>
<mods:affiliation id="A9EB20CC4382E5D13C96202CC4F7D79F">Natural History Museum of Crete, University of Crete, Knossos Av., Herakleion 71409, Crete, Greece.</mods:affiliation>
<mods:nameIdentifier id="83DEA27E9A3516ACA48BE3346180DFED" type="email">ssimaiakis@yahoo.com</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="2DC05503444081376D54CDF542161551">Edgecombe, Gregory D.</mods:namePart>
<mods:affiliation id="5202A79AFE6D19CC4525BCBC2A096B5F">Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom.</mods:affiliation>
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<mods:title id="A281BBB593CDF3592B734D989FAA10E0">Zootaxa</mods:title>
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<mods:date id="A818377688326E70AE204837CA9D232C">2013</mods:date>
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<mods:number id="D40033E1F69973987BAEE0809F6F80C7">2013-11-05</mods:number>
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<mods:number id="AF0013CD1AB70A9744E25450D680E01B">3734</mods:number>
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<treatment id="03B68458FFBEFFF9FF56A9DBFB6AF8CF" ID-DOI="http://doi.org/10.5281/zenodo.5271303" ID-Zenodo-Dep="5271303" LSID="urn:lsid:plazi:treatment:03B68458FFBEFFF9FF56A9DBFB6AF8CF" httpUri="http://treatment.plazi.org/id/03B68458FFBEFFF9FF56A9DBFB6AF8CF" lastPageNumber="183" pageId="14" pageNumber="183">
<subSubSection id="C30566C5FFBEFFF9FF56A9DBFC80FBB1" box="[151,789,1014,1040]" pageId="14" pageNumber="183" type="nomenclature">
<paragraph id="8BA0354EFFBEFFF9FF56A9DBFC80FBB1" blockId="14.[151,789,1014,1074]" box="[151,789,1014,1040]" pageId="14" pageNumber="183">
<heading id="D0E88222FFBEFFF9FF56A9DBFC80FBB1" bold="true" box="[151,789,1014,1040]" fontSize="11" level="1" pageId="14" pageNumber="183" reason="1">
<emphasis id="B96BE95CFFBEFFF9FF56A9DBFC80FBB1" bold="true" box="[151,789,1014,1040]" pageId="14" pageNumber="183">
15.
<taxonomicName id="4C1F4ECDFFBEFFF9FF02A9DBFC80FBB1" ID-CoL="7J363" authority="stuhlmanni Kraepelin, 1903" authorityName="stuhlmanni Kraepelin" authorityYear="1903" box="[195,789,1014,1040]" class="Chilopoda" family="Scolopendridae" genus="Rhysida" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="183" phylum="Arthropoda" rank="subSpecies" species="stuhlmanni" subSpecies="stuhlmanni">
<emphasis id="B96BE95CFFBEFFF9FF02A9DBFDDDFBB1" bold="true" box="[195,584,1014,1040]" italics="true" pageId="14" pageNumber="183">Rhysida stuhlmanni stuhlmanni</emphasis>
<bibRefCitation id="EF8E48BFFFBEFFF9FD8EA9DBFC80FBB1" author="Kraepelin, K." box="[591,789,1014,1040]" pageId="14" pageNumber="183" pagination="1 - 276" refId="ref17381" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 20 (2), 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30566C5FFBEFFF9FF56AE34FEB2FB93" box="[151,295,1049,1074]" pageId="14" pageNumber="183" type="description">
<paragraph id="8BA0354EFFBEFFF9FF56AE34FEB2FB93" blockId="14.[151,789,1014,1074]" box="[151,295,1049,1074]" pageId="14" pageNumber="183">
(
<figureCitation id="132429CBFFBEFFF9FF5EAE34FE8AFB93" box="[159,287,1049,1074]" captionStart="FIGURES 2329" captionStartId="15.[151,264,1860,1883]" captionTargetBox="[151,1436,195,1839]" captionTargetId="figure-122@15.[151,1436,195,1839]" captionTargetPageId="15" captionText="FIGURES 2329. Figs 2326. Rhysida paucidens Pocock, 1897. BMNH 1953.2.14.515 (Ethiopia): Figure 23, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 24, forcipular segment, ventral view, scale bar 0.5 mm; Figure 25, ultimate leg prefemur, scale bar 0.5 mm; Figure 26, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm. Figs 2729. Rhysida stuhlmanni stuhlmanni Kraepelin, 1903. BMNH 94.1.15.39 (Malawi): Figure 27, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 28, forcipular segment, ventral view, scale bar 0.5 mm; Figure 29, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5275615" httpUri="https://zenodo.org/record/5275615/files/figure.png" pageId="14" pageNumber="183">Figs 2729</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30566C5FFBEFFF9FF56AE4CFAF9FA8F" pageId="14" pageNumber="183" type="materials_examined">
<paragraph id="8BA0354EFFBEFFF9FF56AE4CFD46FB63" blockId="14.[151,1437,1121,1902]" pageId="14" pageNumber="183">
<emphasis id="B96BE95CFFBEFFF9FF56AE4CFE15FBDB" bold="true" box="[151,384,1121,1146]" inLineHeading="true" pageId="14" pageNumber="183" reason="1">Material examined.</emphasis>
<materialsCitation id="3B773F13FFBEFFF9FE47AE4CFA0DFBDB" box="[390,1432,1121,1146]" collectingDate="1892-08" collectingDateMax="1892-11-31" collectingDateMin="1892-08" collectionCode="BMNH" collectorName="H. H. Johnson" country="Malawi" location="Zomba" pageId="14" pageNumber="183" specimenCount="1" stateProvince="Zomba">
<collectingCountry id="F30875DEFFBEFFF9FE47AE4CFE75FBDB" box="[390,480,1121,1146]" name="Malawi" pageId="14" pageNumber="183">Malawi</collectingCountry>
:
<collectingRegion id="49DBFBACFFBEFFF9FE2AAE4CFDA8FBDB" box="[491,573,1121,1146]" country="Malawi" name="Zomba" pageId="14" pageNumber="183">Zomba</collectingRegion>
, Nyasaland,
<date id="FFA1138EFFBEFFF9FD0DAE4CFC08FBDB" box="[716,925,1121,1146]" pageId="14" pageNumber="183" value="1892-08" valueMax="1892-11-31" valueMin="1892-08-00">
<collectingDate id="EFE5EA66FFBEFFF9FD0DAE4CFC08FBDB" box="[716,925,1121,1146]" pageId="14" pageNumber="183" value="1892-08" valueMax="1892-11-31" valueMin="1892-08-00">Aug. to Nov. 1892</collectingDate>
</date>
, leg.
<collectorName id="26EA5098FFBEFFF9FC19AE4FFBE6FBDB" box="[984,1139,1121,1146]" pageId="14" pageNumber="183">H.H. Johnson</collectorName>
,
<specimenCount id="9D19FEC7FFBEFFF9FBBEAE4CFB22FBD8" box="[1151,1207,1121,1146]" pageId="14" pageNumber="183" type="generic">1 ex.</specimenCount>
,
<collectionCode id="ED0EAD8BFFBEFFF9FB03AE4FFA83FBD8" box="[1218,1302,1122,1145]" pageId="14" pageNumber="183">BMNH</collectionCode>
94.1.15.39
</materialsCitation>
;
<materialsCitation id="3B773F13FFBEFFF9FF56AEA8FD5AFB63" collectionCode="R, BMNH" collectorName="Prof. P. A. Buxter" country="Malawi" location="London School of Tropical Medicine" municipality="Lake Nyassa" pageId="14" pageNumber="183" specimenCount="1" stateProvince="Likoma">
<collectingCountry id="F30875DEFFBEFFF9FF56AEA8FF64FB3F" box="[151,241,1157,1182]" name="Malawi" pageId="14" pageNumber="183">Malawi</collectingCountry>
:
<collectingMunicipality id="6BC4AF34FFBEFFF9FF3EAEA8FE00FB3F" box="[255,405,1157,1182]" pageId="14" pageNumber="183">Lake Nyassa</collectingMunicipality>
,
<collectingRegion id="49DBFBACFFBEFFF9FE65AEA8FE6EFB3F" box="[420,507,1157,1182]" country="Malawi" name="Likoma" pageId="14" pageNumber="183">Likoma</collectingRegion>
Isl.,
<location id="8EC06395FFBEFFF9FDFAAEA8FC7BFB3F" LSID="urn:lsid:plazi:treatment:03B68458FFBEFFF9FF56A9DBFB6AF8CF:8EC06395FFBEFFF9FDFAAEA8FC7BFB3F" box="[571,1006,1157,1182]" country="Malawi" municipality="Lake Nyassa" name="London School of Tropical Medicine" pageId="14" pageNumber="183" stateProvince="Likoma">London School of Tropical Medicine</location>
, Presd by
<collectorName id="26EA5098FFBEFFF9FBB7AEA8FAAAFB3C" box="[1142,1343,1157,1182]" pageId="14" pageNumber="183">Prof. P.A. Buxter</collectorName>
, leg.
<collectionCode id="ED0EAD8BFFBEFFF9FA43AEABFA0DFB3C" box="[1410,1432,1158,1181]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="183">R</collectionCode>
. Howard,
<specimenCount id="9D19FEC7FFBEFFF9FF3EAE84FEADFB60" box="[255,312,1193,1218]" pageId="14" pageNumber="183" type="generic">1 ex.</specimenCount>
,
<collectionCode id="ED0EAD8BFFBEFFF9FE82AE87FE02FB60" box="[323,407,1194,1217]" pageId="14" pageNumber="183">BMNH</collectionCode>
1950.4.19.2829. (
<figureCitation id="132429CBFFBEFFF9FDB3AE84FD53FB63" box="[626,710,1193,1218]" captionStart="FIGURE 22" captionStartId="13.[151,249,1925,1947]" captionTargetBox="[151,1436,844,1904]" captionTargetId="figure-344@13.[150,1436,844,1904]" captionTargetPageId="13" captionText="FIGURE 22. Rhysida afra, R. longicornis, R. longipes longipes, R. paucidens, R. stuhlmanni stuhlmanni and R. togoensis, distribution records of specimens from Africa housed in the NHM collection." figureDoi="http://doi.org/10.5281/zenodo.5275613" httpUri="https://zenodo.org/record/5275613/files/figure.png" pageId="14" pageNumber="183">Fig. 22</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA0354EFFBEFFF9FF06AEE0FBFEFB47" blockId="14.[151,1437,1121,1902]" box="[199,1131,1229,1254]" pageId="14" pageNumber="183">
<materialsCitation id="3B773F13FFBEFFF9FF06AEE0FBF2FB47" box="[199,1127,1229,1254]" country="Tanzania" county="Zambesi'" location="Bagamoyo" municipality="Rio Quaqua" pageId="14" pageNumber="183" specimenCount="1" typeStatus="holotype">
<emphasis id="B96BE95CFFBEFFF9FF06AEE0FEF0FB47" bold="true" box="[199,357,1229,1254]" inLineHeading="true" pageId="14" pageNumber="183" reason="1">
<typeStatus id="54A48BECFFBEFFF9FF06AEE0FE97FB47" box="[199,258,1229,1254]" pageId="14" pageNumber="183">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F30875DEFFBEFFF9FEADAEE0FE44FB47" box="[364,465,1229,1254]" name="Tanzania" pageId="14" pageNumber="183">Tanzania</collectingCountry>
(
<location id="8EC06395FFBEFFF9FE1EAEE0FDC8FB47" LSID="urn:lsid:plazi:treatment:03B68458FFBEFFF9FF56A9DBFB6AF8CF:8EC06395FFBEFFF9FE1EAEE0FDC8FB47" box="[479,605,1229,1254]" country="Tanzania" county="Zambesi'" municipality="Rio Quaqua" name="Bagamoyo" pageId="14" pageNumber="183">Bagamoyo</location>
,
<collectingMunicipality id="6BC4AF34FFBEFFF9FDA9AEE0FD67FB47" box="[616,754,1229,1254]" pageId="14" pageNumber="183">Rio Quaqua</collectingMunicipality>
,
<collectingCounty id="62C14DC2FFBEFFF9FCC0AEE0FCE4FB47" box="[769,881,1229,1254]" pageId="14" pageNumber="183">Zambesi</collectingCounty>
) (
<bibRefCitation id="EF8E48BFFFBEFFF9FC49AEE0FBCAFB47" author="Minelli, A. &amp; Bonato, L. &amp; Dioguardi, R. &amp; Chagas, J. A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Uliana, M. &amp; Zapparoli, M." box="[904,1119,1229,1254]" pageId="14" pageNumber="183" refId="ref18465" refString="Minelli, A., Bonato, L., Dioguardi, R., Chagas, J. A., Edgecombe, G. D., Lewis, J. G. E., Pereira, L. A., Shelley, R. M., Stoev, P., Uliana, M. &amp; Zapparoli, M. (2006 and onwards). CHILOBASE. A web resource for Chilopoda taxonomy." type="book" year="2006">
Minelli
<emphasis id="B96BE95CFFBEFFF9FC22AEE3FB81FB47" box="[995,1044,1229,1254]" italics="true" pageId="14" pageNumber="183">et al</emphasis>
. 2006
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA0354EFFBEFFF9FF06AEDCFAF9FA8F" blockId="14.[151,1437,1121,1902]" pageId="14" pageNumber="183">
<emphasis id="B96BE95CFFBEFFF9FF06AEDCFE5DFAAB" bold="true" box="[199,456,1265,1290]" inLineHeading="true" pageId="14" pageNumber="183" reason="1">General distribution.</emphasis>
East Tropical Africa:
<collectingCountry id="F30875DEFFBEFFF9FD14AEDCFCACFAAB" box="[725,825,1265,1290]" name="Tanzania" pageId="14" pageNumber="183">Tanzania</collectingCountry>
(
<collectionCode id="ED0EAD8BFFBEFFF9FC8FAEDCFCE0FAAB" box="[846,885,1265,1290]" pageId="14" pageNumber="183">TZ</collectionCode>
); South Tropical Africa:
<collectingCountry id="F30875DEFFBEFFF9FB61AEDCFB60FAAB" box="[1184,1269,1265,1290]" name="Malawi" pageId="14" pageNumber="183">Malawi</collectingCountry>
(
<collectionCode id="ED0EAD8BFFBEFFF9FAC8AEDCFAD7FAAB" box="[1289,1346,1265,1290]" country="Netherlands" name="Museum Wasmann" pageId="14" pageNumber="183">MW</collectionCode>
),
<collectingCountry id="F30875DEFFBEFFF9FA9BAEDCFF4AFA8F" name="South Africa" pageId="14" pageNumber="183">South Africa</collectingCountry>
(ZA),
<collectingCountry id="F30875DEFFBEFFF9FEEAAF38FE36FA8F" box="[299,419,1301,1326]" name="Zimbabwe" pageId="14" pageNumber="183">Zimbabwe</collectingCountry>
(
<collectionCode id="ED0EAD8BFFBEFFF9FE73AF38FE74FA8F" box="[434,481,1301,1326]" pageId="14" pageNumber="183">ZW</collectionCode>
) (
<bibRefCitation id="EF8E48BFFFBEFFF9FE37AF38FD3CFA8F" author="Lawrence, R. F." box="[502,681,1301,1326]" pageId="14" pageNumber="183" pagination="4 - 56" refId="ref17522" refString="Lawrence, R. F. (1955) Chilopoda. In: Hanstrom, B., Brink, P. &amp; Rudebeck, G. (Eds.), South African Animal Life. Results of the Lund University Expedition in 1950 - 1951. Vol. II. Almqvist &amp; Wiksell, Stockholm, 4 - 56." type="book chapter" year="1955">Lawrence 1955</bibRefCitation>
;
<bibRefCitation id="EF8E48BFFFBEFFF9FD75AF38FC1FFA8F" author="Minelli, A. &amp; Bonato, L. &amp; Dioguardi, R. &amp; Chagas, J. A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Uliana, M. &amp; Zapparoli, M." box="[692,906,1301,1326]" pageId="14" pageNumber="183" refId="ref18465" refString="Minelli, A., Bonato, L., Dioguardi, R., Chagas, J. A., Edgecombe, G. D., Lewis, J. G. E., Pereira, L. A., Shelley, R. M., Stoev, P., Uliana, M. &amp; Zapparoli, M. (2006 and onwards). CHILOBASE. A web resource for Chilopoda taxonomy." type="book" year="2006">
Minelli
<emphasis id="B96BE95CFFBEFFF9FCCFAF3BFCAAFA8F" box="[782,831,1301,1326]" italics="true" pageId="14" pageNumber="183">et al</emphasis>
. 2006
</bibRefCitation>
); Asia Tropical:
<collectingCountry id="F30875DEFFBEFFF9FB85AF38FBEBFA8F" box="[1092,1150,1301,1326]" name="India" pageId="14" pageNumber="183">India</collectingCountry>
(
<collectionCode id="ED0EAD8BFFBEFFF9FB4CAF38FB38FA8F" box="[1165,1197,1301,1326]" pageId="14" pageNumber="183">IN</collectionCode>
) (
<bibRefCitation id="EF8E48BFFFBEFFF9FB02AF38FACAFA8F" author="Khanna, V." box="[1219,1375,1301,1326]" pageId="14" pageNumber="183" pagination="459 - 472" refId="ref17286" refString="Khanna, V. (1994) Centipede fauna of NE States, India (Chilopoda: Scolopendromorpha). Records of the Zoological Survey of India, 94, 459 - 472." type="journal article" year="1994">Khanna 1994</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30566C5FFBEFFF9FF06AF14FB6AF8CF" pageId="14" pageNumber="183" type="discussion">
<paragraph id="8BA0354EFFBEFFF9FF06AF14FB6AF8CF" blockId="14.[151,1437,1121,1902]" pageId="14" pageNumber="183">
<emphasis id="B96BE95CFFBEFFF9FF06AF14FEAEFAF3" bold="true" box="[199,315,1337,1362]" inLineHeading="true" pageId="14" pageNumber="183" reason="1">Remarks.</emphasis>
The smaller specimen from
<collectingRegion id="49DBFBACFFBEFFF9FD45AF14FD46FAF3" box="[644,723,1337,1362]" country="Malawi" name="Zomba" pageId="14" pageNumber="183">Zomba</collectingRegion>
(
<quantity id="4CE798ABFFBEFFF9FD25AF14FCADFAF3" box="[740,824,1337,1362]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="14" pageNumber="183" unit="mm" value="22.0">22 mm</quantity>
) has 17 antennal articles, of which three are glabrous dorsally (
<figureCitation id="132429CBFFBEFFF9FEC5AF70FECDFAD7" box="[260,344,1373,1398]" captionStart="FIGURES 2329" captionStartId="15.[151,264,1860,1883]" captionTargetBox="[151,1436,195,1839]" captionTargetId="figure-122@15.[151,1436,195,1839]" captionTargetPageId="15" captionText="FIGURES 2329. Figs 2326. Rhysida paucidens Pocock, 1897. BMNH 1953.2.14.515 (Ethiopia): Figure 23, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 24, forcipular segment, ventral view, scale bar 0.5 mm; Figure 25, ultimate leg prefemur, scale bar 0.5 mm; Figure 26, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm. Figs 2729. Rhysida stuhlmanni stuhlmanni Kraepelin, 1903. BMNH 94.1.15.39 (Malawi): Figure 27, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 28, forcipular segment, ventral view, scale bar 0.5 mm; Figure 29, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5275615" httpUri="https://zenodo.org/record/5275615/files/figure.png" pageId="14" pageNumber="183">Fig. 27</figureCitation>
) and 5+5 forcipular teeth (
<figureCitation id="132429CBFFBEFFF9FD57AF70FD79FAD7" box="[662,748,1373,1398]" captionStart="FIGURES 2329" captionStartId="15.[151,264,1860,1883]" captionTargetBox="[151,1436,195,1839]" captionTargetId="figure-122@15.[151,1436,195,1839]" captionTargetPageId="15" captionText="FIGURES 2329. Figs 2326. Rhysida paucidens Pocock, 1897. BMNH 1953.2.14.515 (Ethiopia): Figure 23, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 24, forcipular segment, ventral view, scale bar 0.5 mm; Figure 25, ultimate leg prefemur, scale bar 0.5 mm; Figure 26, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm. Figs 2729. Rhysida stuhlmanni stuhlmanni Kraepelin, 1903. BMNH 94.1.15.39 (Malawi): Figure 27, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 28, forcipular segment, ventral view, scale bar 0.5 mm; Figure 29, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5275615" httpUri="https://zenodo.org/record/5275615/files/figure.png" pageId="14" pageNumber="183">Fig. 28</figureCitation>
); the larger specimen from
<collectingRegion id="49DBFBACFFBEFFF9FBF0AF70FB1DFAD7" box="[1073,1160,1373,1398]" country="Malawi" name="Likoma" pageId="14" pageNumber="183">Likoma</collectingRegion>
Island (
<quantity id="4CE798ABFFBEFFF9FB28AF70FAABFAD7" box="[1257,1342,1373,1398]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="14" pageNumber="183" unit="mm" value="58.0">58 mm</quantity>
) has 18 antennal articles, 2.66 of which are glabrous dorsally. Both specimens from
<collectingCountry id="F30875DEFFBEFFF9FBD9AFACFBFBFA3B" box="[1048,1134,1409,1434]" name="Malawi" pageId="14" pageNumber="183">Malawi</collectingCountry>
have the diagnostic short paramedian sutures on the tergites (confined to the posterior part of TT420), only the tergite of the ultimate legbearing segment being marginate, the strongly embayed posterior margin and approximately quadratic shape of the sternite of the ultimate leg-bearing segment (
<figureCitation id="132429CBFFBEFFF9FD4FAFC0FD75F9A7" box="[654,736,1517,1542]" captionStart="FIGURES 2329" captionStartId="15.[151,264,1860,1883]" captionTargetBox="[151,1436,195,1839]" captionTargetId="figure-122@15.[151,1436,195,1839]" captionTargetPageId="15" captionText="FIGURES 2329. Figs 2326. Rhysida paucidens Pocock, 1897. BMNH 1953.2.14.515 (Ethiopia): Figure 23, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 24, forcipular segment, ventral view, scale bar 0.5 mm; Figure 25, ultimate leg prefemur, scale bar 0.5 mm; Figure 26, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm. Figs 2729. Rhysida stuhlmanni stuhlmanni Kraepelin, 1903. BMNH 94.1.15.39 (Malawi): Figure 27, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 28, forcipular segment, ventral view, scale bar 0.5 mm; Figure 29, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5275615" httpUri="https://zenodo.org/record/5275615/files/figure.png" pageId="14" pageNumber="183">Fig. 29</figureCitation>
, cf.
<bibRefCitation id="EF8E48BFFFBEFFF9FCD1AFC0FC54F9A7" author="Kraepelin, K." box="[784,961,1517,1542]" pageId="14" pageNumber="183" pagination="1 - 276" refId="ref17381" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 20 (2), 1 - 276." type="journal article" year="1903">Kraepelin 1903</bibRefCitation>
: fig. 97), and three spines (two apical, one subapical, no lateral or spine spine) on the coxopleural process (
<figureCitation id="132429CBFFBEFFF9FCA0AC3CFC21F98B" box="[865,948,1553,1578]" captionStart="FIGURES 2329" captionStartId="15.[151,264,1860,1883]" captionTargetBox="[151,1436,195,1839]" captionTargetId="figure-122@15.[151,1436,195,1839]" captionTargetPageId="15" captionText="FIGURES 2329. Figs 2326. Rhysida paucidens Pocock, 1897. BMNH 1953.2.14.515 (Ethiopia): Figure 23, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 24, forcipular segment, ventral view, scale bar 0.5 mm; Figure 25, ultimate leg prefemur, scale bar 0.5 mm; Figure 26, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm. Figs 2729. Rhysida stuhlmanni stuhlmanni Kraepelin, 1903. BMNH 94.1.15.39 (Malawi): Figure 27, cephalic plate and T1, dorsal view, scale bar 1 mm; Figure 28, forcipular segment, ventral view, scale bar 0.5 mm; Figure 29, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5275615" httpUri="https://zenodo.org/record/5275615/files/figure.png" pageId="14" pageNumber="183">Fig. 29</figureCitation>
), though these characters are also shared by
<taxonomicName id="4C1F4ECDFFBEFFF9FF56AC1BFE74F9EF" authority="Attems, 1910" authorityName="Attems" authorityYear="1910" box="[151,481,1589,1614]" class="Chilopoda" family="Scolopendridae" genus="Rhysida" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="183" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B96BE95CFFBEFFF9FF56AC1BFEA0F9EF" box="[151,309,1589,1614]" italics="true" pageId="14" pageNumber="183">R. intermedia</emphasis>
<bibRefCitation id="EF8E48BFFFBEFFF9FE83AC18FE74F9EF" author="Attems, C." box="[322,481,1589,1614]" pageId="14" pageNumber="183" pagination="73 - 115" refId="ref16598" refString="Attems, C. (1910) Myriopoden von Madagaskar den Comoren und den Inseln Ostafrikas. In: Voeltzkow A. (ed.) Reise in Ostafrika. Schweizerbart'sche E. Nagele &amp; Dr. Sproessner, Stuttgart, 3, pp. 73 - 115." type="journal article" year="1910">Attems, 1910</bibRefCitation>
</taxonomicName>
, from
<collectingRegion id="49DBFBACFFBEFFF9FDF4AC18FD40F9EF" box="[565,725,1589,1614]" country="Tanzania" name="Kusini Pemba" pageId="14" pageNumber="183">Pemba Island</collectingRegion>
,
<collectingCountry id="F30875DEFFBEFFF9FD27AC18FCD9F9EF" box="[742,844,1589,1614]" name="Tanzania" pageId="14" pageNumber="183">Zanzibar</collectingCountry>
. 17 antennal articles in
<taxonomicName id="4C1F4ECDFFBEFFF9FBB0AC1BFA09F9EF" authority="stuhlmanni" authorityName="stuhlmanni Kraepelin" authorityYear="1903" box="[1137,1436,1589,1614]" class="Chilopoda" family="Scolopendridae" genus="Rhysida" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="183" phylum="Arthropoda" rank="subSpecies" species="stuhlmanni" subSpecies="stuhlmanni">
<emphasis id="B96BE95CFFBEFFF9FBB0AC1BFA09F9EF" box="[1137,1436,1589,1614]" italics="true" pageId="14" pageNumber="183">R. stuhlmanni stuhlmanni</emphasis>
</taxonomicName>
corresponds to the description of
<taxonomicName id="4C1F4ECDFFBEFFF9FDCBAC77FD34F9D3" authorityName="Attems" authorityYear="1910" box="[522,673,1625,1650]" class="Chilopoda" family="Scolopendridae" genus="Rhysida" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="183" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B96BE95CFFBEFFF9FDCBAC77FD34F9D3" box="[522,673,1625,1650]" italics="true" pageId="14" pageNumber="183">R. intermedia</emphasis>
</taxonomicName>
but previous descriptions have not indicated the number of glabrous antennal articles in
<emphasis id="B96BE95CFFBEFFF9FEB2AC53FDBDF937" box="[371,552,1661,1686]" italics="true" pageId="14" pageNumber="183">
R. s.
<taxonomicName id="4C1F4ECDFFBEFFF9FE6DAC50FDBDF937" authorityName="Kraepelin" authorityYear="1903" box="[428,552,1661,1686]" class="Chilopoda" family="Scolopendridae" genus="Rhysida" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="183" phylum="Arthropoda" rank="species" species="stuhlmanni">stuhlmanni</taxonomicName>
</emphasis>
, so it is not clear whether this might serve as a basis to recognize two species. In the larger specimen, legs 119 bear two tarsal spurs (118 fide
<bibRefCitation id="EF8E48BFFFBEFFF9FCBBAC8CFBBFF91B" author="Kraepelin, K." box="[890,1066,1697,1722]" pageId="14" pageNumber="183" pagination="1 - 276" refId="ref17381" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 20 (2), 1 - 276." type="journal article" year="1903">Kraepelin 1903</bibRefCitation>
), and legs 117 bear a tibial spur (leg 18 is missing, and leg 19 lacks a tibial spur). The smaller specimen has two tarsal spurs to at least leg 17 (the following legs are missing) and a tibial spur to at least leg 13 but lacking on leg 17. The main differences between the
<collectingCountry id="F30875DEFFBEFFF9FF01AD20FE80F887" box="[192,277,1805,1830]" name="Malawi" pageId="14" pageNumber="183">Malawi</collectingCountry>
specimens and
<taxonomicName id="4C1F4ECDFFBEFFF9FE07AD23FDCBF887" authorityName="Attems" authorityYear="1910" box="[454,606,1805,1830]" class="Chilopoda" family="Scolopendridae" genus="Rhysida" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="183" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B96BE95CFFBEFFF9FE07AD23FDCBF887" box="[454,606,1805,1830]" italics="true" pageId="14" pageNumber="183">R. intermedia</emphasis>
</taxonomicName>
are the presence of 1718 antennal articles (versus 19), two tarsal spurs to legs 17+/19 (versus 15) and a gentle curve to the outer margin of the coxopleural pore field, corresponding to “ziemlich geradlinig” (
<taxonomicName id="4C1F4ECDFFBEFFF9FE58AD7BFDA0F8CF" authorityName="Kraepelin" authorityYear="1903" box="[409,565,1877,1902]" class="Chilopoda" family="Scolopendridae" genus="Rhysida" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="183" phylum="Arthropoda" rank="species" species="stuhlmanni">
<emphasis id="B96BE95CFFBEFFF9FE58AD7BFDA0F8CF" box="[409,565,1877,1902]" italics="true" pageId="14" pageNumber="183">R. stuhlmanni</emphasis>
</taxonomicName>
in Attems key) rather than “tief eingebuchtet” (
<taxonomicName id="4C1F4ECDFFBEFFF9FB97AD7BFB7BF8CF" authorityName="Attems" authorityYear="1910" box="[1110,1262,1877,1902]" class="Chilopoda" family="Scolopendridae" genus="Rhysida" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="183" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B96BE95CFFBEFFF9FB97AD7BFB7BF8CF" box="[1110,1262,1877,1902]" italics="true" pageId="14" pageNumber="183">R. intermedia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>