treatments-xml/data/03/A7/87/03A787BAFFC2E53A50988027669FFC28.xml

<document id="742A785FC0E31F0AEDE4B6A0A2788E04" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFC2E53A50988027669FFC28" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras STEPHENS 1835" docType="key" docVersion="10" lastPageNumber="125" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="120" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="40A8E0178BCA87A92A82153C4CF0EEEA">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<treatment id="03A787BAFFC2E53A50988027669FFC28" ID-DOI="http://doi.org/10.5281/zenodo.5885203" ID-GBIF-Taxon="192498147" ID-Zenodo-Dep="5885203" LSID="urn:lsid:plazi:treatment:03A787BAFFC2E53A50988027669FFC28" httpUri="http://treatment.plazi.org/id/03A787BAFFC2E53A50988027669FFC28" lastPageId="8" lastPageNumber="125" pageId="3" pageNumber="120">
<subSubSection id="C3146527FFC2E5315098802767BFF8FB" box="[189,1204,1839,1871]" pageId="3" pageNumber="120" type="nomenclature">
<paragraph id="8BB136ACFFC2E5315098802767BFF8FB" blockId="3.[189,1204,1839,1871]" box="[189,1204,1839,1871]" pageId="3" pageNumber="120">
<heading id="D0F981C0FFC2E5315098802767BFF8FB" box="[189,1204,1839,1871]" centered="true" fontSize="12" level="2" pageId="3" pageNumber="120" reason="2">
3.2 Key to the species of the subgenus 
<taxonomicName id="4C0E4D2FFFC2E5315286802761E8F8FB" ID-CoL="8VS6V" authority="STEPHENS, 1835" authorityName="STEPHENS" authorityYear="1835" box="[675,739,1839,1871]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEABEFFC2E5315286802761E8F8FB" box="[675,739,1839,1871]" italics="true" pageId="3" pageNumber="120">Zyras</emphasis>
</taxonomicName>
of the Himalaya, 
<collectingCountry id="F319763CFFC2E5315398802760F4F8FB" box="[957,1023,1839,1871]" name="India" pageId="3" pageNumber="120">India</collectingCountry>
, and 
<collectingCountry id="F319763CFFC2E5315464802767BFF8FB" box="[1089,1204,1839,1871]" name="Sri Lanka" pageId="3" pageNumber="120">Sri Lanka</collectingCountry>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFC2E53150B2806263F2F875" pageId="3" pageNumber="120" type="distribution">
<paragraph id="8BB136ACFFC2E53150B2806263F2F875" blockId="3.[151,1398,1897,1985]" pageId="3" pageNumber="120">
In total, 25 named and two undescribed species are currently known from the region including the Himalaya (exclusive of 
<collectingCountry id="F319763CFFC2E531510380816266F815" box="[294,365,1929,1953]" name="China" pageId="3" pageNumber="120">China</collectingCountry>
), 
<collectingCountry id="F319763CFFC2E53151A5808162B7F815" box="[384,444,1929,1953]" name="India" pageId="3" pageNumber="120">India</collectingCountry>
, and 
<collectingCountry id="F319763CFFC2E53151D38081616AF815" box="[502,609,1929,1953]" name="Sri Lanka" pageId="3" pageNumber="120">Sri Lanka</collectingCountry>
. 
<taxonomicName id="4C0E4D2FFFC2E531524980826006F815" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[620,781,1929,1953]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFC2E531524980826006F815" box="[620,781,1929,1953]" italics="true" pageId="3" pageNumber="120">Zyras pindarae</emphasis>
</taxonomicName>
, a species of highly variable coloration, keys out at three couplets.
</paragraph>
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<key id="FF9F619DFFC5E53450E987F06686F8C6" lastPageId="6" lastPageNumber="123" pageId="4" pageNumber="121">
<key id="FF9F619DFFC5E53650E987F06686F845" pageId="4" pageNumber="121">
<keyStep id="30FA2E09FFC5E53650E987F06686FE4B" pageId="4" pageNumber="121">
<paragraph id="8BB136ACFFC5E53650E987F06686FE24" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E987F06686FE24" pageId="4" pageNumber="121">
1. Whole body with dense and long and sub-erect to erect pubescence (in dorsal view best visible on the posterolateral contours of the head, on the lateral margins of the elytra, and on the posterior margins of the sternites) (e.g., 
<figureCitation id="13352A29FFC5E53650D786306286FEE4" box="[242,397,312,336]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="4" pageNumber="121">Figs 59, 62–64</figureCitation>
). Body broad and with rather large abdomen (e.g., 
<figureCitation id="13352A29FFC5E53653B986306718FEE4" box="[924,1043,312,336]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="4" pageNumber="121">Figs 95–97</figureCitation>
). Antennomere XI very short, much shorter than the combined length of antennomeres IX and X, and of broadly conical shape (e.g., 
<figureCitation id="13352A29FFC5E53654C08650668AFEC4" box="[1253,1409,344,368]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="4" pageNumber="121">Figs 12–13, 21</figureCitation>
). Apical lobe of paramere very short (e.g., 
<figureCitation id="13352A29FFC5E5365284867061F2FE24" box="[673,761,376,400]" captionStart="Figs 174–190" captionStartId="58.[189,229,2020,2042]" captionTargetBox="[190,1430,246,1968]" captionTargetId="figure-107@58.[189,1436,237,1976]" captionText="Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888372" httpUri="https://zenodo.org/record/5888372/files/figure.png" pageId="4" pageNumber="121">Fig. 184</figureCitation>
). 
<taxonomicName id="4C0E4D2FFFC5E536532F86716083FE24" ID-CoL="5DDCZ" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[778,904,376,400]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B97AEABEFFC5E536532F86716083FE24" box="[778,904,376,400]" italics="true" pageId="4" pageNumber="121">Zyras hirtus</emphasis>
</taxonomicName>
group. .................................................................... 2
</keyLead>
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<paragraph id="8BB136ACFFC5E53650E986A06686FE4B" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E986A06686FE4B" pageId="4" pageNumber="121">– At least the forebody with distinctly less dense and less conspicuous pubescence. Habitus more slender. Antennomere XI mostly more elongate and not of distinctly conical shape. Apical lobe of paramere usually longer and more slender. ...................................................................................................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC5E53650E9851E6686FDEA" pageId="4" pageNumber="121">
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<keyLead id="30FF9599FFC5E53650E9851E6686FD9A" box="[204,1421,534,558]" pageId="4" pageNumber="121">2. Forebody black; abdomen blackish-brown to black. .................................................................................................... 3</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC5E53650E9854E6686FDEA" blockId="4.[204,1422,248,2033]" box="[204,1421,581,606]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E9854E6686FDEA" box="[204,1421,581,606]" pageId="4" pageNumber="121">– Pale-coloured or bicoloured species. .............................................................................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC5E53650E9857D6686FCCF" pageId="4" pageNumber="121">
<paragraph id="8BB136ACFFC5E53650E9857D6686FD58" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E9857D6686FD58" pageId="4" pageNumber="121">
3. Meso- and metafemora distinctly bicoloured, basally yellow and apically infuscate; antennomere XI blackish, not distinctly paler than antennomeres IV–X (
<figureCitation id="13352A29FFC5E536528F859C61FEFD19" box="[682,757,660,685]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="4" pageNumber="121">Fig. 13</figureCitation>
). Tergites III–V without distinct micropunctation. Median lobe of aedeagus subapically more strongly angled in lateral view and apically less acute (
<figureCitation id="13352A29FFC5E536544585BC67F9FD79" box="[1120,1266,692,717]" captionStart="Figs 174–190" captionStartId="58.[189,229,2020,2042]" captionTargetBox="[190,1430,246,1968]" captionTargetId="figure-107@58.[189,1436,237,1976]" captionText="Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888372" httpUri="https://zenodo.org/record/5888372/files/figure.png" pageId="4" pageNumber="121">Figs 178–179</figureCitation>
). Distribution: North 
<collectingCountry id="F319763CFFC5E536511C85DC6278FD58" box="[313,371,724,748]" name="India" pageId="4" pageNumber="121">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC5E536515885DC62B7FD58" box="[381,444,724,748]" name="Nepal" pageId="4" pageNumber="121">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC5E53651EF85DD611AFD59" box="[458,529,725,749]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="4" pageNumber="121">Map 7</figureCitation>
). ...................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC5E53654DB85DC6686FD58" box="[1278,1421,724,748]" class="Insecta" family="Staphylinidae" genus="Myrmedonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B97AEABEFFC5E53654DB85DC6686FD58" bold="true" box="[1278,1421,724,748]" italics="true" pageId="4" pageNumber="121">hirsutiventris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC5E53650E9840C6686FCCF" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E9840C6686FCCF" pageId="4" pageNumber="121">
– Meso- and metafemora uniformly dark-yellowish; apices very indistinctly darker at most; antennomere XI dark-reddish, somewhat contrasting with the blackish antennomeres IV–X (
<figureCitation id="13352A29FFC5E5365434842C6756FC88" box="[1041,1117,804,828]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="4" pageNumber="121">Fig. 23</figureCitation>
). Tergites III–V with distinct micropunctation. Median lobe of aedeagus subapically weakly angled and apically very acute both in lateral and in ventral view (
<figureCitation id="13352A29FFC5E53651B8846C6124FCC8" box="[413,559,868,892]" captionStart="Figs 174–190" captionStartId="58.[189,229,2020,2042]" captionTargetBox="[190,1430,246,1968]" captionTargetId="figure-107@58.[189,1436,237,1976]" captionText="Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888372" httpUri="https://zenodo.org/record/5888372/files/figure.png" pageId="4" pageNumber="121">Figs 182–183</figureCitation>
). Distribution: Northeast 
<collectingCountry id="F319763CFFC5E5365365846C6073FCC8" box="[832,888,868,892]" name="India" pageId="4" pageNumber="121">India</collectingCountry>
(
<figureCitation id="13352A29FFC5E53653A3846C60C7FCC8" box="[902,972,868,892]" captionStart="Map 8" captionStartId="68.[189,234,1235,1256]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-71@68.[189,1436,232,1226]" captionTargetPageId="68" captionText="Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white" figureDoi="http://doi.org/10.5281/zenodo.5742417" httpUri="https://zenodo.org/record/5742417/files/figure.png" pageId="4" pageNumber="121">Map 8</figureCitation>
). ....................................................... 
<taxonomicName id="4C0E4D2FFFC5E5365519846B6686FCCF" ID-CoL="5DDDL" authorityName="Assing" authorityYear="2017" box="[1340,1421,867,891]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="B97AEABEFFC5E5365519846B6686FCCF" bold="true" box="[1340,1421,867,891]" italics="true" pageId="4" pageNumber="121">luteipes</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC5E53650E9849B6686FB8E" pageId="4" pageNumber="121">
<paragraph id="8BB136ACFFC5E53650E9849B6686FC5E" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E9849B6686FC5E" pageId="4" pageNumber="121">
4. Meso- and metafemora distinctly bicoloured, basally yellowish and apically infuscate. Elytra bicoloured, darkyellowish with the postero-lateral portion blackish (
<figureCitation id="13352A29FFC5E536533284BB606FFC7F" box="[791,868,947,971]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="4" pageNumber="121">Fig. 60</figureCitation>
). Abdomen bicoloured, with segments II–V reddish and VI–X blackish (
<figureCitation id="13352A29FFC5E53651E384DB612BFC5F" box="[454,544,979,1003]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="4" pageNumber="121">Fig. 101</figureCitation>
). Distribution: South 
<collectingCountry id="F319763CFFC5E536532384DB6035FC5F" box="[774,830,979,1003]" name="India" pageId="4" pageNumber="121">India</collectingCountry>
(Map 79). ........................................................ 
<taxonomicName id="4C0E4D2FFFC5E536553284DA6686FC5E" ID-CoL="7GHYS" authorityName="CAMERON" authorityYear="1939" box="[1303,1421,978,1002]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="nilgiriensis">
<emphasis id="B97AEABEFFC5E536553284DA6686FC5E" bold="true" box="[1303,1421,978,1002]" italics="true" pageId="4" pageNumber="121">nilgiriensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC5E53650E9830A6686FB8E" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E9830A6686FB8E" pageId="4" pageNumber="121">– Meso- and metafemora uniformly dark-yellowish to reddish; apices very indistinctly darker at most. Coloration of body different. ............................................................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC5E53650E983596686FACC" pageId="4" pageNumber="121">
<paragraph id="8BB136ACFFC5E53650E983596686FB5D" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E983596686FB5D" pageId="4" pageNumber="121">
5. Antennae distinctly bicoloured, blackish with the apical three antennomeres pale yellowish (
<figureCitation id="13352A29FFC5E53654E28359661FFBDD" box="[1223,1300,1105,1129]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="4" pageNumber="121">Fig. 40</figureCitation>
). Head and pronotum dark-brown to blackish; elytra reddish with the postero-lateral portions infuscate (
<figureCitation id="13352A29FFC5E53654E08379661AFB3D" box="[1221,1297,1137,1161]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="4" pageNumber="121">Fig. 62</figureCitation>
). Abdomen bicoloured, reddish with segments V–VIII blackish (
<figureCitation id="13352A29FFC5E536530D8399607FFB1D" box="[808,884,1169,1193]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="4" pageNumber="121">Fig. 60</figureCitation>
). Antenna very long (approximately 3.0 mm) and slender (
<figureCitation id="13352A29FFC5E536516883B96290FB7D" box="[333,411,1201,1225]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="4" pageNumber="121">Fig. 40</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC5E53652D583B96074FB7D" box="[752,895,1201,1225]" captionStart="Figs 174–190" captionStartId="58.[189,229,2020,2042]" captionTargetBox="[190,1430,246,1968]" captionTargetId="figure-107@58.[189,1436,237,1976]" captionText="Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888372" httpUri="https://zenodo.org/record/5888372/files/figure.png" pageId="4" pageNumber="121">Figs 174–175</figureCitation>
. Distribution: North 
<collectingCountry id="F319763CFFC5E536544583B96791FB7D" box="[1120,1178,1201,1225]" name="India" pageId="4" pageNumber="121">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC5E536548183B967E8FB7D" box="[1188,1251,1201,1225]" name="Nepal" pageId="4" pageNumber="121">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC5E53654D483B96633FB7D" box="[1265,1336,1201,1225]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="4" pageNumber="121">Map 7</figureCitation>
). .......... ................................................................................................................................................................................. 
<taxonomicName id="4C0E4D2FFFC5E536551483D96686FB5D" ID-CoL="5DDCJ" authorityName="Cameron" authorityYear="1939" box="[1329,1421,1233,1257]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="gardneri">
<emphasis id="B97AEABEFFC5E536551483D96686FB5D" bold="true" box="[1329,1421,1233,1257]" italics="true" pageId="4" pageNumber="121">gardneri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC5E53650E982096686FACC" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E982096686FACC" pageId="4" pageNumber="121">
– Antennae blackish-brown to blackish, with usually only antennomere XI, rarely X–XI or IX–XI, slightly paler (
<figureCitation id="13352A29FFC5E53650DC8228624FFA8C" box="[249,324,1312,1336]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="4" pageNumber="121">Fig. 21</figureCitation>
). Body more or less uniformly reddish to reddish-brown, with abdominal segments VI–VII slightly darker at most (
<figureCitation id="13352A29FFC5E5365177824862A9FAEC" box="[338,418,1344,1368]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="4" pageNumber="121">Figs 59</figureCitation>
, 
<figureCitation id="13352A29FFC5E536518B824862C0FAEC" box="[430,459,1344,1368]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="4" pageNumber="121">95</figureCitation>
). Antenna much shorter (approximately 2.0 mm) and more strongly incrassate (
<figureCitation id="13352A29FFC5E53655118248668AFAEC" box="[1332,1409,1344,1368]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="4" pageNumber="121">Fig. 21</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC5E5365213826861CEFACC" box="[566,709,1376,1400]" captionStart="Figs 156–173" captionStartId="57.[151,191,2014,2036]" captionTargetBox="[154,1376,242,1969]" captionTargetId="figure-115@57.[151,1398,233,1972]" captionText="Figs 156–173: Zyras longilobatus (156–160), Z. latilobatus (161–165), Z. alternans (166–169), and Z. hirtus (170–173; 170–171: Sri Lanka; 172–173: South India): median lobe of aedeagus in lateral and in ventral view (156–157, 161–162, 166–167, 170–173); paramere (158, 163); male tergite VIII (159, 164, 168); male sternite VIII (160, 165, 169). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888370" httpUri="https://zenodo.org/record/5888370/files/figure.png" pageId="4" pageNumber="121">Figs 170–171</figureCitation>
. Distribution: South 
<collectingCountry id="F319763CFFC5E5365381826860D5FACC" box="[932,990,1376,1400]" name="India" pageId="4" pageNumber="121">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC5E53653CD82686745FACC" box="[1000,1102,1376,1400]" name="Sri Lanka" pageId="4" pageNumber="121">Sri Lanka</collectingCountry>
(
<figureCitation id="13352A29FFC5E536547E826867A9FACC" box="[1115,1186,1376,1400]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="4" pageNumber="121">Map 7</figureCitation>
). ....................... 
<taxonomicName id="4C0E4D2FFFC5E536556882686686FACC" ID-CoL="5DDCZ" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[1357,1421,1376,1400]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B97AEABEFFC5E536556882686686FACC" bold="true" box="[1357,1421,1376,1400]" italics="true" pageId="4" pageNumber="121">hirtus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC5E53650E982876686FA42" pageId="4" pageNumber="121">
<paragraph id="8BB136ACFFC5E53650E982876686FA73" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E982876686FA73" pageId="4" pageNumber="121">6. Forebody and abdomen completely blackish. Legs bicoloured with the femora blackish-brown to black and the tibiae yellowish. ................................................................................................................................................................. 7</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC5E53650E982D76686FA42" blockId="4.[204,1422,248,2033]" box="[204,1421,1502,1526]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E982D76686FA42" box="[204,1421,1502,1526]" pageId="4" pageNumber="121">– Coloration different, at least the abdomen not completely blackish. ......................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC5E53650E981066686F921" pageId="4" pageNumber="121">
<paragraph id="8BB136ACFFC5E53650E981066686F9F1" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E981066686F9F1" pageId="4" pageNumber="121">
7. Forebody with blueish hue (
<figureCitation id="13352A29FFC5E536520781056165F991" box="[546,622,1549,1573]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="4" pageNumber="121">Fig. 50</figureCitation>
). Antennae blackish with antennomeres I–II and the base of III yellowish (
<figureCitation id="13352A29FFC5E53650DC8125624DF9F2" box="[249,326,1581,1606]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="4" pageNumber="121">Fig. 22</figureCitation>
). Distribution: North 
<collectingCountry id="F319763CFFC5E536520A8125616CF9F1" box="[559,615,1581,1605]" name="India" pageId="4" pageNumber="121">India</collectingCountry>
(
<figureCitation id="13352A29FFC5E5365250812661B7F9F2" box="[629,700,1582,1606]" captionStart="Map 2" captionStartId="64.[189,234,1456,1477]" captionTargetBox="[189,1436,891,1446]" captionTargetId="figure-96@64.[189,1436,891,1446]" captionTargetPageId="64" captionText="Map 2: Revised distributions of Zyras pindarae (black circles) and Z. nigroaeneus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742395" httpUri="https://zenodo.org/record/5742395/files/figure.png" pageId="4" pageNumber="121">Map 2</figureCitation>
). ............................................................................................ 
<taxonomicName id="4C0E4D2FFFC5E536552E81256686F9F1" ID-CoL="7GHYR" authorityName="Cameron" authorityYear="1939" box="[1291,1421,1581,1605]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="nigroaeneus">
<emphasis id="B97AEABEFFC5E536552E81256686F9F1" bold="true" box="[1291,1421,1581,1605]" italics="true" pageId="4" pageNumber="121">nigroaeneus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC5E53650E981556686F921" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E981556686F921" pageId="4" pageNumber="121">– Forebody without blueish hue. Antennae, including the basal antennomeres, uniformly blackish, or nearly so. ...................................................................................................................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC5E53650E981A46686F8C6" pageId="4" pageNumber="121">
<paragraph id="8BB136ACFFC5E53650E981A46687F8B7" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E981A46687F8B7" pageId="4" pageNumber="121">
8. Tibiae blackish, as dark as femora. Ventral process of aedeagus basally broader in ventral view (
<figureCitation id="13352A29FFC5E53654CA81A46689F970" box="[1263,1410,1708,1732]" captionStart="Figs 260–277" captionStartId="63.[151,191,1987,2009]" captionTargetBox="[157,1395,232,1954]" captionTargetId="figure-126@63.[151,1398,232,1971]" captionText="Figs 260–277: Zyras morulus (260–265), Z. variolatus (266–270), Z. montanus (271–275), and Z. preangeranus from Java (276–277): forebody (260); antenna (261); abdomen (262); median lobe of aedeagus in lateral and in ventral view (263–264, 266–267, 271–272, 276–277); paramere (265, 268, 273); male tergite VIII (269, 274); male sternite VIII (270, 275). Scale bars: 260–262: 1.0 mm; 263–277: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888380" httpUri="https://zenodo.org/record/5888380/files/figure.png" pageId="4" pageNumber="121">Figs 263–264</figureCitation>
). Paramere much longer than median lobe and with rather long and slender apical lobe (
<figureCitation id="13352A29FFC5E53654B281C467FAF950" box="[1175,1265,1740,1764]" captionStart="Figs 260–277" captionStartId="63.[151,191,1987,2009]" captionTargetBox="[157,1395,232,1954]" captionTargetId="figure-126@63.[151,1398,232,1971]" captionText="Figs 260–277: Zyras morulus (260–265), Z. variolatus (266–270), Z. montanus (271–275), and Z. preangeranus from Java (276–277): forebody (260); antenna (261); abdomen (262); median lobe of aedeagus in lateral and in ventral view (263–264, 266–267, 271–272, 276–277); paramere (265, 268, 273); male tergite VIII (269, 274); male sternite VIII (270, 275). Scale bars: 260–262: 1.0 mm; 263–277: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888380" httpUri="https://zenodo.org/record/5888380/files/figure.png" pageId="4" pageNumber="121">Fig. 265</figureCitation>
). Distribution: Central 
<collectingCountry id="F319763CFFC5E536516D81E4628CF8B0" box="[328,391,1772,1796]" name="Nepal" pageId="4" pageNumber="121">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC5E53651B081E462D6F8B0" box="[405,477,1772,1796]" captionStart="Map 4" captionStartId="65.[154,199,794,815]" captionTargetBox="[151,1398,230,785]" captionTargetId="figure-63@65.[151,1398,230,785]" captionTargetPageId="65" captionText="Map 4: Revised distributions of Zyras condignus (black circles) and Z. morulus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742401" httpUri="https://zenodo.org/record/5742401/files/figure.png" pageId="4" pageNumber="121">Map 4</figureCitation>
). ........................................................................................................................................ 
<taxonomicName id="4C0E4D2FFFC5E536551681E36687F8B7" ID-CoL="7GHYG" authorityName="Assing" authorityYear="2017" box="[1331,1420,1771,1795]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="morulus">
<emphasis id="B97AEABEFFC5E536551681E36687F8B7" bold="true" box="[1331,1420,1771,1795]" italics="true" pageId="4" pageNumber="121">morulus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC5E53650E980136686F8C6" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E980136686F8C6" pageId="4" pageNumber="121">
– Tibiae distinctly paler than femora. Ventral process of aedeagus basally more slender in ventral view (
<figureCitation id="13352A29FFC5E53654D680136689F887" box="[1267,1410,1819,1843]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="4" pageNumber="121">Figs 125–128</figureCitation>
). Paramere approximately as long as median lobe and with short apical lobe (
<figureCitation id="13352A29FFC5E53654288033676CF8E7" box="[1037,1127,1851,1875]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="4" pageNumber="121">Fig. 129</figureCitation>
). Distribution: North 
<collectingCountry id="F319763CFFC5E536556A80336682F8E7" box="[1359,1417,1851,1875]" name="India" pageId="4" pageNumber="121">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC5E53650D78053623AF8C7" box="[242,305,1883,1907]" name="Nepal" pageId="4" pageNumber="121">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC5E536511B8053628DF8C7" box="[318,390,1883,1907]" captionStart="Map 2" captionStartId="64.[189,234,1456,1477]" captionTargetBox="[189,1436,891,1446]" captionTargetId="figure-96@64.[189,1436,891,1446]" captionTargetPageId="64" captionText="Map 2: Revised distributions of Zyras pindarae (black circles) and Z. nigroaeneus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742395" httpUri="https://zenodo.org/record/5742395/files/figure.png" pageId="4" pageNumber="121">Map 2</figureCitation>
). ..................................................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC5E536550980526686F8C6" ID-CoL="7GHXX" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[1324,1421,1882,1906]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFC5E536550980526686F8C6" bold="true" box="[1324,1421,1882,1906]" italics="true" pageId="4" pageNumber="121">pindarae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC5E53650E980826686F845" pageId="4" pageNumber="121">
<paragraph id="8BB136ACFFC5E53650E980826686F876" blockId="4.[204,1422,248,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E980826686F876" pageId="4" pageNumber="121">9. Abdomen black with the apex (segments VII–X or VIII–X) reddish, strongly contrasting with the anterior segments; forebody black. .............................................................................................................................................. 10</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC5E53650E980D26686F845" blockId="4.[204,1422,248,2033]" box="[204,1421,2009,2033]" pageId="4" pageNumber="121">
<keyLead id="30FF9599FFC5E53650E980D26686F845" box="[204,1421,2009,2033]" pageId="4" pageNumber="121">– Abdomen of different coloration; forebody usually not uniformly black. .............................................................. 12</keyLead>
</paragraph>
</keyStep>
</key>
<key id="FF9F619DFFC4E537508387F1666CF786" pageId="5" pageNumber="122">
<keyStep id="30FA2E09FFC4E537508387F16663FE14" pageId="5" pageNumber="122">
<paragraph id="8BB136ACFFC4E537508387F16663FEE5" blockId="5.[166,1384,249,417]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E537508387F16663FEE5" pageId="5" pageNumber="122">
10. Segment VII black or reddish (
<figureCitation id="13352A29FFC4E537523787F16180FEA5" box="[530,651,249,273]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="5" pageNumber="122">Figs 82–85</figureCitation>
). Pronotum with slightly irregular and denser punctation; elytra with dense and nearly regularly distributed punctation (
<figureCitation id="13352A29FFC4E53752D486116034FE85" box="[753,831,281,305]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="5" pageNumber="122">Fig. 44</figureCitation>
). Aedeagus as in 
<figureCitation id="13352A29FFC4E53753DF86116781FE85" box="[1018,1162,281,305]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="5" pageNumber="122">Figs 125–129</figureCitation>
. Distribution: North 
<collectingCountry id="F319763CFFC4E53750E98631620DFEE5" box="[204,262,313,337]" name="India" pageId="5" pageNumber="122">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC4E537513586316244FEE5" box="[272,335,313,337]" name="Nepal" pageId="5" pageNumber="122">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC4E5375178863162AFFEE5" box="[349,420,313,337]" captionStart="Map 2" captionStartId="64.[189,234,1456,1477]" captionTargetBox="[189,1436,891,1446]" captionTargetId="figure-96@64.[189,1436,891,1446]" captionTargetPageId="64" captionText="Map 2: Revised distributions of Zyras pindarae (black circles) and Z. nigroaeneus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742395" httpUri="https://zenodo.org/record/5742395/files/figure.png" pageId="5" pageNumber="122">Map 2</figureCitation>
). .......................................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC4E537552286316663FEE5" ID-CoL="7GHXX" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[1287,1384,313,337]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFC4E537552286316663FEE5" bold="true" box="[1287,1384,313,337]" italics="true" pageId="5" pageNumber="122">pindarae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC4E537508386616663FE14" blockId="5.[166,1384,249,417]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E537508386616663FE14" pageId="5" pageNumber="122">
– Segment VII reddish (
<figureCitation id="13352A29FFC4E537519A86606100FE34" box="[447,523,360,384]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="5" pageNumber="122">Fig. 79</figureCitation>
). Pronotum with sparse and distinctly irregular punctation; elytra with sparser (posteriorly very sparse) and more distinctly irregular punctation (
<figureCitation id="13352A29FFC4E53753B5868060D6FE14" box="[912,989,392,416]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="5" pageNumber="122">Fig. 42</figureCitation>
). ........................................................ 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC4E537508386B0666CFDEB" pageId="5" pageNumber="122">
<paragraph id="8BB136ACFFC4E537508386B0666CFE5B" blockId="5.[166,1384,439,607]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E537508386B0666CFE5B" pageId="5" pageNumber="122">
11. Legs uniformly pale-yellowish. Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC4E537534886BF60F7FE64" box="[877,1020,439,464]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="5" pageNumber="122">Figs 117–118</figureCitation>
. Distribution: North 
<collectingCountry id="F319763CFFC4E53754C786BF6617FE7B" box="[1250,1308,439,463]" name="India" pageId="5" pageNumber="122">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC4E537550D86BF666CFE7B" box="[1320,1383,439,463]" name="Nepal" pageId="5" pageNumber="122">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC4E53750F686D06213FE44" box="[211,280,472,496]" captionStart="Map 1" captionStartId="64.[189,234,822,843]" captionTargetBox="[189,1436,232,813]" captionTargetId="figure-76@64.[189,1436,232,813]" captionTargetPageId="64" captionText="Map 1: Revised distributions of Zyras kraatzi (black and white circles; white circle: type locality), Z. exasperatus (white circle), Z. iniquus (triangles), Z. pallipes (diamonds), and Z. championi (white stars) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742393" httpUri="https://zenodo.org/record/5742393/files/figure.png" pageId="5" pageNumber="122">Map 1</figureCitation>
). ................................................................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC4E537553E86DF666CFE5B" ID-CoL="5DDDC" authorityName="SCHUBERT" authorityYear="1908" box="[1307,1383,471,495]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="kraatzi">
<emphasis id="B97AEABEFFC4E537553E86DF666CFE5B" bold="true" box="[1307,1383,471,495]" italics="true" pageId="5" pageNumber="122">kraatzi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC4E5375083850F666CFDEB" blockId="5.[166,1384,439,607]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E5375083850F666CFDEB" pageId="5" pageNumber="122">
– Legs bicoloured, with the femora blackish-brown to black and the tibiae and tarsi distinctly paler, yellowish to brown. Median lobe of aedeagus as in 
<bibRefCitation id="EF9F4B5DFFC4E5375250852F6013FD8B" author="ASSING, V." box="[629,792,551,575]" pageId="5" pageNumber="122" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 268–269). Distribution: 
<collectingCountry id="F319763CFFC4E537545E852F67D3FD8B" box="[1147,1240,551,575]" name="Pakistan" pageId="5" pageNumber="122">Pakistan</collectingCountry>
, 
<collectingCountry id="F319763CFFC4E53754C0852F666CFD8B" box="[1253,1383,551,575]" name="Afghanistan" pageId="5" pageNumber="122">Afghanistan</collectingCountry>
(
<figureCitation id="13352A29FFC4E53750F6854F6213FDEB" box="[211,280,583,607]" captionStart="Map 1" captionStartId="64.[189,234,822,843]" captionTargetBox="[189,1436,232,813]" captionTargetId="figure-76@64.[189,1436,232,813]" captionTargetPageId="64" captionText="Map 1: Revised distributions of Zyras kraatzi (black and white circles; white circle: type locality), Z. exasperatus (white circle), Z. iniquus (triangles), Z. pallipes (diamonds), and Z. championi (white stars) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742393" httpUri="https://zenodo.org/record/5742393/files/figure.png" pageId="5" pageNumber="122">Map 1</figureCitation>
). .................................................................................................................................................................. 
<taxonomicName id="4C0E4D2FFFC4E5375532854F666CFDEB" ID-CoL="5DDD7" authorityName="ASSING" authorityYear="2016" box="[1303,1383,583,607]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="iniquus">
<emphasis id="B97AEABEFFC4E5375532854F666CFDEB" bold="true" box="[1303,1383,583,607]" italics="true" pageId="5" pageNumber="122">iniquus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC4E5375083857E6663FD09" pageId="5" pageNumber="122">
<paragraph id="8BB136ACFFC4E5375083857E6663FD3A" blockId="5.[166,1384,630,701]" box="[166,1384,630,654]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E5375083857E6663FD3A" box="[166,1384,630,654]" pageId="5" pageNumber="122">12. Pronotum bright-reddish, strongly contrasting with the blackish head. ................................................................. 13</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC4E537508385AD6663FD09" blockId="5.[166,1384,630,701]" box="[166,1384,677,701]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E537508385AD6663FD09" box="[166,1384,677,701]" pageId="5" pageNumber="122">– Pronotum brown to black. ............................................................................................................................................. 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC4E537508385DC6663FCAF" pageId="5" pageNumber="122">
<paragraph id="8BB136ACFFC4E537508385DC6663FD58" blockId="5.[166,1384,724,795]" box="[166,1384,724,748]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E537508385DC6663FD58" box="[166,1384,724,748]" pageId="5" pageNumber="122">13. Elytra bicoloured. ............................................................................................................................................................ 14</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC4E5375083840C6663FCAF" blockId="5.[166,1384,724,795]" box="[166,1384,771,795]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E5375083840C6663FCAF" box="[166,1384,771,795]" pageId="5" pageNumber="122">– Elytra uniformly blackish-brown to black, sometimes with blueish hue. ............................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC4E5375083843A6663FC4D" pageId="5" pageNumber="122">
<paragraph id="8BB136ACFFC4E5375083843A666CFC3E" blockId="5.[166,1384,818,1017]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E5375083843A666CFC3E" pageId="5" pageNumber="122">
14. Elytra blackish with the suture and the area near the scutellum extensively yellowish-red (
<figureCitation id="13352A29FFC4E5375448843A67B2FCFE" box="[1133,1209,818,842]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="5" pageNumber="122">Fig. 54</figureCitation>
). Antennomeres VIII–X distinctly transverse (
<figureCitation id="13352A29FFC4E5375226845A6144FCDE" box="[515,591,850,874]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="5" pageNumber="122">Fig. 16</figureCitation>
). Male sexual characters unknown. Distribution: South 
<collectingCountry id="F319763CFFC4E5375485845A67D3FCDE" box="[1184,1240,850,874]" name="India" pageId="5" pageNumber="122">India</collectingCountry>
(
<figureCitation id="13352A29FFC4E53754C0845A6626FCDE" box="[1253,1325,850,874]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="5" pageNumber="122">Map 7</figureCitation>
). ...... ................................................................................................................................................................................. 
<taxonomicName id="4C0E4D2FFFC4E5375528847A666CFC3E" ID-CoL="5DDCS" authorityName="Fauvel" authorityYear="1904" box="[1293,1383,882,906]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B97AEABEFFC4E5375528847A666CFC3E" bold="true" box="[1293,1383,882,906]" italics="true" pageId="5" pageNumber="122">hastatus</emphasis>
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</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC4E537508384AA6663FC4D" blockId="5.[166,1384,818,1017]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E537508384AA6663FC4D" pageId="5" pageNumber="122">
– Elytra blackish with the anterior portion more or less extensively yellowish-red; posterior portion of suture blackish (
<figureCitation id="13352A29FFC4E537511184C9628BFC6D" box="[308,384,961,985]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="5" pageNumber="122">Fig. 57</figureCitation>
). Antennomeres VIII–X oblong (
<figureCitation id="13352A29FFC4E53752C984C96032FC6D" box="[748,825,961,985]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="5" pageNumber="122">Fig. 19</figureCitation>
). Aedeagus as in 
<figureCitation id="13352A29FFC4E53753DF84C96781FC6D" box="[1018,1162,961,985]" captionStart="Figs 156–173" captionStartId="57.[151,191,2014,2036]" captionTargetBox="[154,1376,242,1969]" captionTargetId="figure-115@57.[151,1398,233,1972]" captionText="Figs 156–173: Zyras longilobatus (156–160), Z. latilobatus (161–165), Z. alternans (166–169), and Z. hirtus (170–173; 170–171: Sri Lanka; 172–173: South India): median lobe of aedeagus in lateral and in ventral view (156–157, 161–162, 166–167, 170–173); paramere (158, 163); male tergite VIII (159, 164, 168); male sternite VIII (160, 165, 169). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888370" httpUri="https://zenodo.org/record/5888370/files/figure.png" pageId="5" pageNumber="122">Figs 161–163</figureCitation>
. Distribution: South 
<collectingCountry id="F319763CFFC4E53750E984E9620FFC4D" box="[204,260,993,1017]" name="India" pageId="5" pageNumber="122">India</collectingCountry>
(
<figureCitation id="13352A29FFC4E537513484E96252FC4D" box="[273,345,993,1017]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="5" pageNumber="122">Map 7</figureCitation>
). ................................................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC4E53754D084E96663FC4D" ID-CoL="7GHYF" authorityName="Assing" authorityYear="2017" box="[1269,1384,993,1017]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="latilobatus">
<emphasis id="B97AEABEFFC4E53754D084E96663FC4D" bold="true" box="[1269,1384,993,1017]" italics="true" pageId="5" pageNumber="122">latilobatus</emphasis>
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</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC4E537508383186663FB43" pageId="5" pageNumber="122">
<paragraph id="8BB136ACFFC4E53750838318666CFB3C" blockId="5.[166,1384,1040,1272]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E53750838318666CFB3C" pageId="5" pageNumber="122">
15. Antennae short, 
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long, and distinctly incrassate apically; antennomere X at least twice as broad as long (
<figureCitation id="13352A29FFC4E537512283386274FBFD" box="[263,383,1072,1097]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="5" pageNumber="122">Figs 17–18</figureCitation>
). Pronotum approximately 1.15 times as broad as long (
<figureCitation id="13352A29FFC4E53753E483386730FBFC" box="[961,1083,1072,1096]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="5" pageNumber="122">Figs 55–56</figureCitation>
). Aedeagus as in 
<bibRefCitation id="EF9F4B5DFFC4E53754CF8339666FFBFD" author="PACE, R." box="[1258,1380,1073,1097]" pageId="5" pageNumber="122" pagination="23 - 40" refId="ref58918" refString="PACE, R. 2011: Aleocharinae from India gathered by Guillaume de Rougemont (Coleoptera, Staphylinidae). - Lavori della Societa Veneziana di Scienze Naturali 36: 23 - 40." type="journal article" year="2011">PACE (2011</bibRefCitation>
: figures 39–40). Small species; length of forebody 2.0– 
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. Distribution: 
<collectingCountry id="F319763CFFC4E53753CA83586722FBDC" box="[1007,1065,1104,1128]" name="India" pageId="5" pageNumber="122">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC4E537541183586792FBDC" box="[1076,1177,1104,1128]" name="Sri Lanka" pageId="5" pageNumber="122">Sri Lanka</collectingCountry>
(
<figureCitation id="13352A29FFC4E5375482835967E5FBDD" box="[1191,1262,1105,1129]" captionStart="Map 7" captionStartId="67.[151,196,1246,1267]" captionTargetBox="[351,1198,232,1238]" captionTargetId="figure-79@67.[351,1198,232,1238]" captionTargetPageId="67" captionText="Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions." figureDoi="http://doi.org/10.5281/zenodo.5742413" httpUri="https://zenodo.org/record/5742413/files/figure.png" pageId="5" pageNumber="122">Map 7</figureCitation>
). ................ .......................................................................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC4E53754F98378666CFB3C" ID-CoL="7GHXL" authorityName="PACE" authorityYear="1998" box="[1244,1383,1136,1160]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="parageminus">
<emphasis id="B97AEABEFFC4E53754F98378666CFB3C" bold="true" box="[1244,1383,1136,1160]" italics="true" pageId="5" pageNumber="122">parageminus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC4E537508383A86663FB43" blockId="5.[166,1384,1040,1272]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E537508383A86663FB43" pageId="5" pageNumber="122">
– Antennae longer and more slender,&gt; 
<quantity id="4CF69B49FFC4E537524583A861B9FB03" box="[608,690,1184,1208]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="122" unit="mm" value="1.7">1.7 mm</quantity>
long; antennomere X less than twice as broad as long. Pronotum more slender, or if similarly transverse, pronotum, elytra, tergite II, and anterior impressions of tergites III–V coarsely punctate. Larger species, length of forebody&gt; 
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. ............................................................................. 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC4E53750838207666CF9A2" pageId="5" pageNumber="122">
<paragraph id="8BB136ACFFC4E53750838207666CFA32" blockId="5.[166,1384,1295,1558]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E53750838207666CFA32" pageId="5" pageNumber="122">
16. Antennae blackish (
<figureCitation id="13352A29FFC4E5375183820762FFFA93" box="[422,500,1295,1319]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="5" pageNumber="122">Fig. 20</figureCitation>
). Pronotum, elytra, abdominal tergite II, and anterior impressions of abdominal tergites III–V with coarse punctation (
<figureCitation id="13352A29FFC4E5375244822761A4FAF3" box="[609,687,1327,1351]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="5" pageNumber="122">Figs 58</figureCitation>
, 
<figureCitation id="13352A29FFC4E537529F822761DCFAF3" box="[698,727,1327,1351]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="5" pageNumber="122">99</figureCitation>
). Pronotum more transverse, 1.15–1.18 times as broad as long (
<figureCitation id="13352A29FFC4E53750F18247622BFAD3" box="[212,288,1359,1383]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="5" pageNumber="122">Fig. 58</figureCitation>
). Rather large and robust species; length of forebody 3.0– 
<quantity id="4CF69B49FFC4E5375356824760CDFAD2" box="[883,966,1359,1383]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="122" unit="mm" value="3.3">3.3 mm</quantity>
. Aedeagus as in 
<figureCitation id="13352A29FFC4E537544B824767F6FAD3" box="[1134,1277,1359,1383]" captionStart="Figs 156–173" captionStartId="57.[151,191,2014,2036]" captionTargetBox="[154,1376,242,1969]" captionTargetId="figure-115@57.[151,1398,233,1972]" captionText="Figs 156–173: Zyras longilobatus (156–160), Z. latilobatus (161–165), Z. alternans (166–169), and Z. hirtus (170–173; 170–171: Sri Lanka; 172–173: South India): median lobe of aedeagus in lateral and in ventral view (156–157, 161–162, 166–167, 170–173); paramere (158, 163); male tergite VIII (159, 164, 168); male sternite VIII (160, 165, 169). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888370" httpUri="https://zenodo.org/record/5888370/files/figure.png" pageId="5" pageNumber="122">Figs 166–167</figureCitation>
. Distribution: Oriental and southern East Palaearctic regions (
<figureCitation id="13352A29FFC4E53752DB8267604EFA33" box="[766,837,1391,1415]" captionStart="Map 8" captionStartId="68.[189,234,1235,1256]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-71@68.[189,1436,232,1226]" captionTargetPageId="68" captionText="Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white" figureDoi="http://doi.org/10.5281/zenodo.5742417" httpUri="https://zenodo.org/record/5742417/files/figure.png" pageId="5" pageNumber="122">Map 8</figureCitation>
). .................................................................... 
<taxonomicName id="4C0E4D2FFFC4E53755278266666CFA32" ID-CoL="7GHVZ" baseAuthorityName="Cameron" baseAuthorityYear="1925" box="[1282,1383,1390,1414]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B97AEABEFFC4E53755278266666CFA32" bold="true" box="[1282,1383,1390,1414]" italics="true" pageId="5" pageNumber="122">alternans</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC4E53750838296666CF9A2" blockId="5.[166,1384,1295,1558]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E53750838296666CF9A2" pageId="5" pageNumber="122">
– Antennae brown to blackish-brown with the apical 1–4 antennomeres yellowish (
<bibRefCitation id="EF9F4B5DFFC4E537540C829667B4FA02" author="ASSING, V." box="[1065,1215,1438,1462]" pageId="5" pageNumber="122" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figures 33–35). Pronotum, elytra, and anterior abdominal tergites with fine shallow punctation (
<bibRefCitation id="EF9F4B5DFFC4E537540082B667B0FA62" author="ASSING, V." box="[1061,1211,1470,1494]" pageId="5" pageNumber="122" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figures 99–100, 109, 157–158, 165). Pronotum more slender, 1.03–1.10 times as broad as long. Aedeagus: 
<bibRefCitation id="EF9F4B5DFFC4E537549882D6666FFA42" author="ASSING, V." box="[1213,1380,1502,1526]" pageId="5" pageNumber="122" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 274–281). Distribution: Oriental and southern East Palaearctic regions (
<figureCitation id="13352A29FFC4E537543382F66757F9A2" box="[1046,1116,1534,1558]" captionStart="Map 8" captionStartId="68.[189,234,1235,1256]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-71@68.[189,1436,232,1226]" captionTargetPageId="68" captionText="Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white" figureDoi="http://doi.org/10.5281/zenodo.5742417" httpUri="https://zenodo.org/record/5742417/files/figure.png" pageId="5" pageNumber="122">Map 8</figureCitation>
). ........................ 
<taxonomicName id="4C0E4D2FFFC4E537552B82F6666CF9A2" ID-CoL="5DDCK" baseAuthorityName="Kraatz" baseAuthorityYear="1859" box="[1294,1383,1534,1558]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="geminus">
<emphasis id="B97AEABEFFC4E537552B82F6666CF9A2" bold="true" box="[1294,1383,1534,1558]" italics="true" pageId="5" pageNumber="122">geminus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC4E537508381256663F9C0" pageId="5" pageNumber="122">
<paragraph id="8BB136ACFFC4E537508381256663F9F1" blockId="5.[166,1384,1581,1652]" box="[166,1384,1581,1605]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E537508381256663F9F1" box="[166,1384,1581,1605]" pageId="5" pageNumber="122">17. Metafemora bicoloured, yellowish with distinctly infuscate apices. ........................................................................ 18</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC4E537508381556663F9C0" blockId="5.[166,1384,1581,1652]" box="[166,1384,1628,1652]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E537508381556663F9C0" box="[166,1384,1628,1652]" pageId="5" pageNumber="122">– Metafemora of uniformly pale or dark coloration. ..................................................................................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC4E53750838183666CF786" pageId="5" pageNumber="122">
<paragraph id="8BB136ACFFC4E53750838183666CF8D7" blockId="5.[166,1384,1675,2098]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E53750838183666CF8D7" pageId="5" pageNumber="122">
18. At least profemora, often also mesofemora of uniformly yellowish coloration. Antennomeres IV–VII distinctly oblong (
<figureCitation id="13352A29FFC4E537510081A36268F977" box="[293,355,1707,1731]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="5" pageNumber="122">Fig. 9</figureCitation>
). Eyes approximately as long as distance from posterior margin of eye to posterior constriction of head (
<figureCitation id="13352A29FFC4E537510C81C362A8F957" box="[297,419,1739,1763]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="5" pageNumber="122">Figs 48–49</figureCitation>
). Pronotum&gt; 1.1 times as broad as long and with straight lateral margins in posterior half (
<figureCitation id="13352A29FFC4E53750F681E36245F8B7" box="[211,334,1771,1795]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="5" pageNumber="122">Figs 48–49</figureCitation>
). Punctation of pronotum and elytra irregular and moderately coarse (
<figureCitation id="13352A29FFC4E537547A81E367D2F8B7" box="[1119,1241,1771,1795]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="5" pageNumber="122">Figs 48–49</figureCitation>
). Abdominal tergites III–VI with few setiferous punctures at posterior margins (
<figureCitation id="13352A29FFC4E537535F800360CDF897" box="[890,966,1803,1827]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="5" pageNumber="122">Fig. 94</figureCitation>
). Male sternite VIII conspicuously acute posteriorly (
<figureCitation id="13352A29FFC4E5375168802362ACF8F7" box="[333,423,1835,1859]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="5" pageNumber="122">Fig. 147</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC4E53752D880236086F8F7" box="[765,909,1835,1859]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="5" pageNumber="122">Figs 142–143</figureCitation>
. Distribution: North 
<collectingCountry id="F319763CFFC4E5375448802367ACF8F7" box="[1133,1191,1835,1859]" name="India" pageId="5" pageNumber="122">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC4E5375494802367FBF8F7" box="[1201,1264,1835,1859]" name="Nepal" pageId="5" pageNumber="122">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC4E53754DB8023664DF8F7" box="[1278,1350,1835,1859]" captionStart="Map 4" captionStartId="65.[154,199,794,815]" captionTargetBox="[151,1398,230,785]" captionTargetId="figure-63@65.[151,1398,230,785]" captionTargetPageId="65" captionText="Map 4: Revised distributions of Zyras condignus (black circles) and Z. morulus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742401" httpUri="https://zenodo.org/record/5742401/files/figure.png" pageId="5" pageNumber="122">Map 4</figureCitation>
). .. .............................................................................................................................................................................. 
<taxonomicName id="4C0E4D2FFFC4E53754DF8043666CF8D7" ID-CoL="5DDBJ" authorityName="Last" authorityYear="1969" box="[1274,1383,1867,1891]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="condignus">
<emphasis id="B97AEABEFFC4E53754DF8043666CF8D7" bold="true" box="[1274,1383,1867,1891]" italics="true" pageId="5" pageNumber="122">condignus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC4E53750838073666CF786" blockId="5.[166,1384,1675,2098]" pageId="5" pageNumber="122">
<keyLead id="30FF9599FFC4E53750838073666CF786" pageId="5" pageNumber="122">
– Apices of all femora blackish. Antennomeres VI–VII transverse (
<figureCitation id="13352A29FFC4E537534E807260B3F826" box="[875,952,1914,1938]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="5" pageNumber="122">Fig. 25</figureCitation>
). Eyes more than twice as long as distance from posterior margin of eye to posterior constriction of head (
<figureCitation id="13352A29FFC4E537534B809260B0F806" box="[878,955,1946,1970]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="5" pageNumber="122">Fig. 51</figureCitation>
). Pronotum &lt;1.1 times as broad as long and with sinuate lateral margins in posterior half (
<figureCitation id="13352A29FFC4E53752C680B26024F866" box="[739,815,1978,2002]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="5" pageNumber="122">Fig. 51</figureCitation>
). Punctation of pronotum and elytra regular and fine (
<figureCitation id="13352A29FFC4E53750F180D2622BF846" box="[212,288,2010,2034]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="5" pageNumber="122">Fig. 51</figureCitation>
). Abdominal tergites III–VI with numerous setiferous punctures at posterior margins. Male sternite VIII convex posteriorly (
<figureCitation id="13352A29FFC4E537518580F262F1F7A6" box="[416,506,2042,2066]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="5" pageNumber="122">Fig. 149</figureCitation>
). Median lobe of aedeagus as in PACE (1986: figures 54–55). Distribution: Central 
<collectingCountry id="F319763CFFC4E53750E98F126200F786" box="[204,267,2074,2098]" name="Nepal" pageId="5" pageNumber="122">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC4E537513D8F126254F786" box="[280,351,2074,2098]" captionStart="Map 3" captionStartId="64.[189,233,2058,2079]" captionTargetBox="[189,1436,1492,2047]" captionTargetId="figure-120@64.[189,1436,1492,2047]" captionTargetPageId="64" captionText="Map 3: Revised distributionsof Zyras perforatus (blackcircles),Z. morvani(white circles),andZ. truncatus(white star)in theHimalaya." figureDoi="http://doi.org/10.5281/zenodo.5742399" httpUri="https://zenodo.org/record/5742399/files/figure.png" pageId="5" pageNumber="122">Map 3</figureCitation>
). ...................................................................................................................................................... morvani
</keyLead>
</paragraph>
</keyStep>
</key>
<key id="FF9F619DFFC7E53450E987F06686F8C6" pageId="6" pageNumber="123">
<keyStep id="30FA2E09FFC7E53450E987F06686FE6B" pageId="6" pageNumber="123">
<paragraph id="8BB136ACFFC7E53450E987F06686FEC4" blockId="6.[204,1421,248,479]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E987F06686FEC4" pageId="6" pageNumber="123">
19. Pronotum with uneven surface and conspicuously irregular punctation more or less grouped in clusters, these clusters at least partly situated in more or less distinct impressions (
<figureCitation id="13352A29FFC7E53453E986106713FE84" box="[972,1048,280,304]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="6" pageNumber="123">Fig. 65</figureCitation>
). Male tergite VIII with four blunt projections posteriorly (
<figureCitation id="13352A29FFC7E53451CC86306149FEE4" box="[489,578,312,336]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="6" pageNumber="123">Fig. 196</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC7E53453A286306718FEE4" box="[903,1043,312,336]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="6" pageNumber="123">Figs 191–194</figureCitation>
. Distribution: Oriental and southern East Palaearctic regions (
<figureCitation id="13352A29FFC7E53451DF8650614AFEC4" box="[506,577,344,368]" captionStart="Map 6" captionStartId="66.[189,234,1236,1257]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-64@66.[189,1436,232,1226]" captionTargetPageId="66" captionText="Map 6: Revised distributions of Zyras geminus (black circles) and Z. proximus (white circles) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742411" httpUri="https://zenodo.org/record/5742411/files/figure.png" pageId="6" pageNumber="123">Map 6</figureCitation>
). ..................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC7E534550D86506686FEC4" ID-CoL="5DDF4" authorityName="CAMERON" authorityYear="1939" box="[1320,1421,344,368]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="B97AEABEFFC7E534550D86506686FEC4" bold="true" box="[1320,1421,344,368]" italics="true" pageId="6" pageNumber="123">proximus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC7E53450E9868F6686FE6B" blockId="6.[204,1421,248,479]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E9868F6686FE6B" pageId="6" pageNumber="123">– Pronotum with smooth surface without impressions (except the usual postero-median impression). Punctation of pronotum regular or less conspicuously irregular. Male tergite VIII often with a median concavity, but without four blunt projections. .................................................................................................................................................... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC7E53450E986FE6686FDC9" pageId="6" pageNumber="123">
<paragraph id="8BB136ACFFC7E53450E986FE6686FD9A" blockId="6.[204,1421,502,637]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E986FE6686FD9A" pageId="6" pageNumber="123">20. Male sternite VIII distinctly truncate posteriorly. Ventral process of aedeagus with long, slender, and acute apex in lateral view. Apical lobe of aedeagus long and slender. ......................................................................................... 21</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC7E53450E9854E6686FDC9" blockId="6.[204,1421,502,637]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E9854E6686FDC9" pageId="6" pageNumber="123">– Male sternite VIII convex posteriorly. Ventral process of aedeagus with shorter, less slender, and less acute apex in lateral view. Apical lobe of aedeagus distinctly shorter. ......................................................................................... 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC7E53450E9859C6686FC6F" pageId="6" pageNumber="123">
<paragraph id="8BB136ACFFC7E53450E9859C6686FC98" blockId="6.[204,1422,660,987]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E9859C6686FC98" pageId="6" pageNumber="123">
21. Antennae approximately 2.0 mm long and not particularly slender, antennomeres VI–IX transverse (
<figureCitation id="13352A29FFC7E5345511859C668AFD18" box="[1332,1409,660,684]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="6" pageNumber="123">Fig. 24</figureCitation>
). Pronotum broader, approximately 1.15 times as broad as long, lateral margins straight or weakly convex in posterior half in dorsal view; pronotal punctation moderately irregularly distributed (
<figureCitation id="13352A29FFC7E53454AB85DC67D1FD58" box="[1166,1242,724,748]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="6" pageNumber="123">Fig. 53</figureCitation>
). Non-setiferous punctation of the anterior impressions of tergites III–V fine (
<figureCitation id="13352A29FFC7E534535085FC60C9FCB8" box="[885,962,756,780]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="6" pageNumber="123">Fig. 89</figureCitation>
). Median lobe of aedeagus with moderately slender and acute apex in lateral view (
<figureCitation id="13352A29FFC7E53452A9841C6015FC98" box="[652,798,788,812]" captionStart="Figs 156–173" captionStartId="57.[151,191,2014,2036]" captionTargetBox="[154,1376,242,1969]" captionTargetId="figure-115@57.[151,1398,233,1972]" captionText="Figs 156–173: Zyras longilobatus (156–160), Z. latilobatus (161–165), Z. alternans (166–169), and Z. hirtus (170–173; 170–171: Sri Lanka; 172–173: South India): median lobe of aedeagus in lateral and in ventral view (156–157, 161–162, 166–167, 170–173); paramere (158, 163); male tergite VIII (159, 164, 168); male sternite VIII (160, 165, 169). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888370" httpUri="https://zenodo.org/record/5888370/files/figure.png" pageId="6" pageNumber="123">Figs 156–158</figureCitation>
). Distribution: Northeast 
<collectingCountry id="F319763CFFC7E534540A841C676CFC98" box="[1071,1127,788,812]" name="India" pageId="6" pageNumber="123">India</collectingCountry>
(
<figureCitation id="13352A29FFC7E5345450841C67B7FC98" box="[1141,1212,788,812]" captionStart="Map 9" captionStartId="69.[151,196,1235,1256]" captionTargetBox="[151,1398,232,1226]" captionTargetId="figure-70@69.[151,1398,232,1226]" captionTargetPageId="69" captionText="Map 9: Revised distributions of Zyras bettotanus (black circles), Z. nilgiriensis (star), and Z. longilobatus (white circles) in the" figureDoi="http://doi.org/10.5281/zenodo.5742419" httpUri="https://zenodo.org/record/5742419/files/figure.png" pageId="6" pageNumber="123">Map 9</figureCitation>
). ........ 
<taxonomicName id="4C0E4D2FFFC7E534552C841C6686FC98" ID-CoL="5DDDH" authorityName="Assing" authorityYear="2017" box="[1289,1421,788,812]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="longilobatus">
<emphasis id="B97AEABEFFC7E534552C841C6686FC98" bold="true" box="[1289,1421,788,812]" italics="true" pageId="6" pageNumber="123">longilobatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC7E53450E9844C6686FC6F" blockId="6.[204,1422,660,987]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E9844C6686FC6F" pageId="6" pageNumber="123">
– Antennae very long and slender, 
<quantity id="4CF69B49FFC7E534527E844B61D1FCEF" box="[603,730,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="6" pageNumber="123" unit="mm" value="2.5" valueMax="2.6" valueMin="2.4">2.4–2.6 mm</quantity>
long; antennomeres VI–XI oblong (
<figureCitation id="13352A29FFC7E5345440844B67BBFCEF" box="[1125,1200,835,859]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="6" pageNumber="123">Fig. 11</figureCitation>
). Pronotum slender, approximately 1.1 times as broad as long, lateral margins sinuate in posterior half in dorsal view; lateral portions of disc with a cluster of punctures slightly behind middle, otherwise nearly impunctate (
<figureCitation id="13352A29FFC7E53454B5848B67D6FC2F" box="[1168,1245,899,923]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="6" pageNumber="123">Fig. 52</figureCitation>
). Non-setiferous punctation of anterior impressions of tergites III–V coarse (
<figureCitation id="13352A29FFC7E534537B84AB60A0FC0F" box="[862,939,931,955]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="6" pageNumber="123">Fig. 91</figureCitation>
). Median lobe of aedeagus with conspicuously slender and acute apex in lateral view (
<figureCitation id="13352A29FFC7E53452A984CB6015FC6F" box="[652,798,963,987]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="6" pageNumber="123">Figs 150–151</figureCitation>
). Distribution: Central 
<collectingCountry id="F319763CFFC7E534543384CB675EFC6F" box="[1046,1109,963,987]" name="Nepal" pageId="6" pageNumber="123">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC7E534544784CB67A2FC6F" box="[1122,1193,963,987]" captionStart="Map 3" captionStartId="64.[189,233,2058,2079]" captionTargetBox="[189,1436,1492,2047]" captionTargetId="figure-120@64.[189,1436,1492,2047]" captionTargetPageId="64" captionText="Map 3: Revised distributionsof Zyras perforatus (blackcircles),Z. morvani(white circles),andZ. truncatus(white star)in theHimalaya." figureDoi="http://doi.org/10.5281/zenodo.5742399" httpUri="https://zenodo.org/record/5742399/files/figure.png" pageId="6" pageNumber="123">Map 3</figureCitation>
). ................ 
<taxonomicName id="4C0E4D2FFFC7E534550384CC6686FC6F" ID-CoL="5DDGC" authorityName="Assing" authorityYear="2017" box="[1318,1421,964,987]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B97AEABEFFC7E534550384CC6686FC6F" bold="true" box="[1318,1421,964,987]" italics="true" pageId="6" pageNumber="123">truncatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC7E53450E984FB6686FBCE" pageId="6" pageNumber="123">
<paragraph id="8BB136ACFFC7E53450E984FB6686FBFF" blockId="6.[204,1422,1010,1146]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E984FB6686FBFF" pageId="6" pageNumber="123">22. Anterior impressions of tergites III–V with few fine non-setiferous punctures confined to the middle. Species of rather large size with a rather large, but weakly transverse pronotum (approximately 1.1 times as broad as long). ............................................................................................................................................................................................ 23</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC7E53450E9836A6686FBCE" blockId="6.[204,1422,1010,1146]" box="[204,1421,1122,1146]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E9836A6686FBCE" box="[204,1421,1122,1146]" pageId="6" pageNumber="123">– Anterior impressions of tergite III–V with more numerous non-setiferous punctures everywhere. .................. 24</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC7E53450E983996686FA23" pageId="6" pageNumber="123">
<paragraph id="8BB136ACFFC7E53450E983996686FABC" blockId="6.[204,1421,1169,1432]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E983996686FABC" pageId="6" pageNumber="123">
23. Legs of uniformly pale-brown coloration. Pronotum sparsely and distinctly irregularly punctate; lateral margins not sinuate near posterior angles (
<figureCitation id="13352A29FFC7E534527583B96197FB7D" box="[592,668,1201,1225]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="6" pageNumber="123">Fig. 43</figureCitation>
). Punctation of elytra coarse and irregularly distributed, anteriorly dense and somewhat asperate, posteriorly very sparse (
<figureCitation id="13352A29FFC7E53452D783D96035FB5D" box="[754,830,1233,1257]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="6" pageNumber="123">Fig. 43</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC7E53454B783D96629FB5D" box="[1170,1314,1233,1257]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="6" pageNumber="123">Figs 121–122</figureCitation>
. Distribution: Northwest 
<collectingCountry id="F319763CFFC7E53451BA83F962DCFABD" box="[415,471,1265,1289]" name="India" pageId="6" pageNumber="123">India</collectingCountry>
(
<figureCitation id="13352A29FFC7E53451C183F96122FABD" box="[484,553,1265,1289]" captionStart="Map 1" captionStartId="64.[189,234,822,843]" captionTargetBox="[189,1436,232,813]" captionTargetId="figure-76@64.[189,1436,232,813]" captionTargetPageId="64" captionText="Map 1: Revised distributions of Zyras kraatzi (black and white circles; white circle: type locality), Z. exasperatus (white circle), Z. iniquus (triangles), Z. pallipes (diamonds), and Z. championi (white stars) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742393" httpUri="https://zenodo.org/record/5742393/files/figure.png" pageId="6" pageNumber="123">Map 1</figureCitation>
). ..................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC7E534552B83F96686FABC" ID-CoL="5DDBZ" authorityName="SCHUBERT" authorityYear="1908" box="[1294,1421,1265,1288]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFC7E534552B83F96686FABC" bold="true" box="[1294,1421,1265,1288]" italics="true" pageId="6" pageNumber="123">exasperatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC7E53450E982286686FA23" blockId="6.[204,1421,1169,1432]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E982286686FA23" pageId="6" pageNumber="123">
– Legs bicoloured, with dark-brown to blackish-brown femora and with paler tibiae and tarsi. Pronotum with nearly regularly distributed punctation (
<figureCitation id="13352A29FFC7E534527C824861ACFAEC" box="[601,679,1344,1368]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="6" pageNumber="123">Fig. 44</figureCitation>
). Elytra with much finer, much denser, and not distinctly irregularly distributed punctation, only slightly sparser posteriorly than anteriorly (
<figureCitation id="13352A29FFC7E53453CA82686737FACC" box="[1007,1084,1376,1400]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="6" pageNumber="123">Fig. 44</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC7E53450D78288628AFA2C" box="[242,385,1408,1432]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="6" pageNumber="123">Figs 125–129</figureCitation>
. Distribution: North 
<collectingCountry id="F319763CFFC7E534524482886190FA2C" box="[609,667,1408,1432]" name="India" pageId="6" pageNumber="123">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC7E5345283828861EEFA2C" box="[678,741,1408,1432]" name="Nepal" pageId="6" pageNumber="123">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC7E53452D782886031FA2C" box="[754,826,1408,1432]" captionStart="Map 2" captionStartId="64.[189,234,1456,1477]" captionTargetBox="[189,1436,891,1446]" captionTargetId="figure-96@64.[189,1436,891,1446]" captionTargetPageId="64" captionText="Map 2: Revised distributions of Zyras pindarae (black circles) and Z. nigroaeneus (white circles) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742395" httpUri="https://zenodo.org/record/5742395/files/figure.png" pageId="6" pageNumber="123">Map 2</figureCitation>
). ............................................................................. 
<taxonomicName id="4C0E4D2FFFC7E534550982776686FA23" ID-CoL="7GHXX" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[1324,1421,1407,1431]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFC7E534550982776686FA23" bold="true" box="[1324,1421,1407,1431]" italics="true" pageId="6" pageNumber="123">pindarae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC7E53450E982A76686F9E2" pageId="6" pageNumber="123">
<paragraph id="8BB136ACFFC7E53450E982A76686F992" blockId="6.[204,1421,1455,1622]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E982A76686F992" pageId="6" pageNumber="123">
24. Small species; body length 4.3–5.0 mm; length of forebody 
<quantity id="4CF69B49FFC7E534537082A760DDFA72" box="[853,982,1455,1479]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="6" pageNumber="123" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
. Antennomere XI conspicuously elongate, approximately as long as the combined length of VIII–X (
<figureCitation id="13352A29FFC7E534537182C760ABFA53" box="[852,928,1487,1511]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="6" pageNumber="123">Fig. 27</figureCitation>
). Head relatively large in relation to pronotum (
<bibRefCitation id="EF9F4B5DFFC7E53450DE82E7629BF9B3" author="ASSING, V." box="[251,400,1519,1543]" pageId="6" pageNumber="123" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 97). Aedeagus: 
<bibRefCitation id="EF9F4B5DFFC7E534525882E76017F9B3" author="ASSING, V." box="[637,796,1519,1543]" pageId="6" pageNumber="123" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 272–273). Distribution: Oriental and southern East Palaearctic regions (
<figureCitation id="13352A29FFC7E53451E281076105F993" box="[455,526,1551,1575]" captionStart="Map 5" captionStartId="65.[151,196,1829,1850]" captionTargetBox="[151,1398,827,1821]" captionTargetId="figure-87@65.[151,1398,827,1821]" captionTargetPageId="65" captionText="Map 5: Revised distributions of Zyras castaneus (black circles) and Z. novinversus (white circles) in the southern East Palaearctic" figureDoi="http://doi.org/10.5281/zenodo.5742405" httpUri="https://zenodo.org/record/5742405/files/figure.png" pageId="6" pageNumber="123">Map 5</figureCitation>
). ............................................................................................................................. 
<taxonomicName id="4C0E4D2FFFC7E534550081076686F992" ID-CoL="5DDBD" baseAuthorityName="Motschulsky" baseAuthorityYear="1861" box="[1317,1421,1551,1574]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="B97AEABEFFC7E534550081076686F992" bold="true" box="[1317,1421,1551,1574]" italics="true" pageId="6" pageNumber="123">castaneus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC7E53450E981376686F9E2" blockId="6.[204,1421,1455,1622]" box="[204,1421,1598,1622]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E981376686F9E2" box="[204,1421,1598,1622]" pageId="6" pageNumber="123">– Distinctly larger species. Antennomere XI less elongate, shorter than the combined length of VIII–X. ........... 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC7E53450E981656686F940" pageId="6" pageNumber="123">
<paragraph id="8BB136ACFFC7E53450E981656686F911" blockId="6.[204,1422,1645,1780]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E981656686F911" pageId="6" pageNumber="123">
25. Anterior third to anterior half of elytra or only humeral portions yellowish, posterior half (including suture) blackish (
<figureCitation id="13352A29FFC7E5345172818562A8F911" box="[343,419,1677,1701]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="6" pageNumber="123">Fig. 45</figureCitation>
). Aedeagus as in 
<figureCitation id="13352A29FFC7E534527C818561E2F911" box="[601,745,1677,1701]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="6" pageNumber="123">Figs 132–133</figureCitation>
. Distribution: North 
<collectingCountry id="F319763CFFC7E53453EC81856708F911" box="[969,1027,1677,1701]" name="India" pageId="6" pageNumber="123">India</collectingCountry>
, 
<collectingCountry id="F319763CFFC7E534542881856747F911" box="[1037,1100,1677,1701]" name="Nepal" pageId="6" pageNumber="123">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC7E534547F818567ABF911" box="[1114,1184,1677,1701]" captionStart="Map 3" captionStartId="64.[189,233,2058,2079]" captionTargetBox="[189,1436,1492,2047]" captionTargetId="figure-120@64.[189,1436,1492,2047]" captionTargetPageId="64" captionText="Map 3: Revised distributionsof Zyras perforatus (blackcircles),Z. morvani(white circles),andZ. truncatus(white star)in theHimalaya." figureDoi="http://doi.org/10.5281/zenodo.5742399" httpUri="https://zenodo.org/record/5742399/files/figure.png" pageId="6" pageNumber="123">Map 3</figureCitation>
). ............... 
<taxonomicName id="4C0E4D2FFFC7E534553B81856686F911" ID-CoL="7GHXD" baseAuthorityName="Champion" baseAuthorityYear="1921" box="[1310,1421,1677,1701]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="perforatus">
<emphasis id="B97AEABEFFC7E534553B81856686F911" bold="true" box="[1310,1421,1677,1701]" italics="true" pageId="6" pageNumber="123">perforatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC7E53450E981B56686F940" blockId="6.[204,1422,1645,1780]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E981B56686F940" pageId="6" pageNumber="123">
– Elytra yellowish with the postero-lateral portions more or less triangularly blackish; yellowish coloration extending posteriad along suture to posterior margin, or nearly so (
<figureCitation id="13352A29FFC7E534536E81D460CFF940" box="[843,964,1756,1780]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="6" pageNumber="123">Figs 46–47</figureCitation>
). ................................................................... 26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC7E53450E980036686F8C6" pageId="6" pageNumber="123">
<paragraph id="8BB136ACFFC7E53450E980036686F897" blockId="6.[204,1422,1803,1907]" box="[204,1421,1803,1828]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E980036686F897" box="[204,1421,1803,1828]" pageId="6" pageNumber="123">
26. Tergites VII–X reddish (
<figureCitation id="13352A29FFC7E53451D68003614BF890" box="[499,576,1803,1828]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="6" pageNumber="123">Fig. 87</figureCitation>
). Distribution: North 
<collectingCountry id="F319763CFFC7E534530C8003606AF897" box="[809,865,1803,1827]" name="India" pageId="6" pageNumber="123">India</collectingCountry>
(
<figureCitation id="13352A29FFC7E534534A800460B8F890" box="[879,947,1804,1828]" captionStart="Map 1" captionStartId="64.[189,234,822,843]" captionTargetBox="[189,1436,232,813]" captionTargetId="figure-76@64.[189,1436,232,813]" captionTargetPageId="64" captionText="Map 1: Revised distributions of Zyras kraatzi (black and white circles; white circle: type locality), Z. exasperatus (white circle), Z. iniquus (triangles), Z. pallipes (diamonds), and Z. championi (white stars) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742393" httpUri="https://zenodo.org/record/5742393/files/figure.png" pageId="6" pageNumber="123">Map 1</figureCitation>
). ...................................................... 
<taxonomicName id="4C0E4D2FFFC7E534553E80036686F897" ID-CoL="7GHYM" authorityName="CAMERON" authorityYear="1939" box="[1307,1421,1803,1827]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B97AEABEFFC7E534553E80036686F897" bold="true" box="[1307,1421,1803,1827]" italics="true" pageId="6" pageNumber="123">championi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC7E53450E980336686F8C6" blockId="6.[204,1422,1803,1907]" pageId="6" pageNumber="123">
<keyLead id="30FF9599FFC7E53450E980336686F8C6" pageId="6" pageNumber="123">
– Tergite VII blackish; posterior portion of tergite VIII somewhat infuscate (
<figureCitation id="13352A29FFC7E53453D980336742F8E7" box="[1020,1097,1851,1875]" captionStart="Figs 82–99" captionStartId="53.[151,191,2018,2040]" captionTargetBox="[151,1398,232,1962]" captionTargetId="figure-116@53.[151,1398,232,1971]" captionTargetPageId="53" captionText="Figs 82–99: Abdomen of Zyras spp.: pindarae (82–85), perforatus (86), championi (87), pallipes (88), longilobatus (89), hastatus (90), truncatus (91), latilobatus (92), morvani (93), condignus (94), hirtus (95), gardneri (96), hirsutiventris (97), parageminus (98), alternans (99). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742383" httpUri="https://zenodo.org/record/5742383/files/figure.png" pageId="6" pageNumber="123">Fig. 88</figureCitation>
). Aedeagus as in 
<figureCitation id="13352A29FFC7E53454DE80336681F8E6" box="[1275,1418,1850,1875]" captionStart="Figs 134–155" captionStartId="56.[189,229,1989,2011]" captionTargetBox="[193,1432,245,1957]" captionTargetId="figure-136@56.[189,1436,232,1971]" captionText="Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888366" httpUri="https://zenodo.org/record/5888366/files/figure.png" pageId="6" pageNumber="123">Figs 136–137</figureCitation>
. Distribution: 
<collectingCountry id="F319763CFFC7E53451A7805362CAF8C7" box="[386,449,1883,1907]" name="Nepal" pageId="6" pageNumber="123">Nepal</collectingCountry>
(
<figureCitation id="13352A29FFC7E53451EB80536118F8C7" box="[462,531,1883,1907]" captionStart="Map 1" captionStartId="64.[189,234,822,843]" captionTargetBox="[189,1436,232,813]" captionTargetId="figure-76@64.[189,1436,232,813]" captionTargetPageId="64" captionText="Map 1: Revised distributions of Zyras kraatzi (black and white circles; white circle: type locality), Z. exasperatus (white circle), Z. iniquus (triangles), Z. pallipes (diamonds), and Z. championi (white stars) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742393" httpUri="https://zenodo.org/record/5742393/files/figure.png" pageId="6" pageNumber="123">Map 1</figureCitation>
). ............................................................................................................................... pallipes
</keyLead>
</paragraph>
</keyStep>
</key>
</key>
</subSubSection>
<subSubSection id="C3146527FFC6E535509887E761ADFE35" pageId="7" pageNumber="124" type="discussion">
<paragraph id="8BB136ACFFC6E535509887E76700FEBB" blockId="7.[189,1035,239,271]" box="[189,1035,239,271]" pageId="7" pageNumber="124">
<heading id="D0F981C0FFC6E535509887E76700FEBB" box="[189,1035,239,271]" centered="true" fontSize="12" level="2" pageId="7" pageNumber="124" reason="2">
3.3 Key to the species of the subgenus 
<taxonomicName id="4C0E4D2FFFC6E535528687E761E8FEBB" box="[675,739,239,271]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEABEFFC6E535528687E761E8FEBB" box="[675,739,239,271]" italics="true" pageId="7" pageNumber="124">Zyras</emphasis>
</taxonomicName>
recorded from 
<collectingCountry id="F319763CFFC6E535538487E76700FEBB" box="[929,1035,239,271]" name="Thailand" pageId="7" pageNumber="124">Thailand</collectingCountry>
</heading>
</paragraph>
<paragraph id="8BB136ACFFC6E53550B2862261ADFE35" blockId="7.[151,1399,297,385]" pageId="7" pageNumber="124">
Up to today, 13 named species of 
<taxonomicName id="4C0E4D2FFFC6E53552318622619FFEF5" authority="sensu" authorityName="SenSU" box="[532,660,298,321]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEABEFFC6E535523186226145FEF5" box="[532,590,298,321]" italics="true" pageId="7" pageNumber="124">Zyras</emphasis>
sensu
</taxonomicName>
strictu have been recorded from 
<collectingCountry id="F319763CFFC6E535542F86216767FEF5" box="[1034,1132,297,321]" name="Thailand" pageId="7" pageNumber="124">Thailand</collectingCountry>
, four of them exclusive. In addition, several unnamed species have been examined, one of them represented by a male without antennae and the remainder represented exclusively by females.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFC6E53A50BA86A4669FFC28" lastPageId="8" lastPageNumber="125" pageId="7" pageNumber="124" type="key">
<key id="FF9F619DFFC6E53550BA86A46664F870" pageId="7" pageNumber="124">
<keyStep id="30FA2E09FFC6E53550BA86A46664FE47" pageId="7" pageNumber="124">
<paragraph id="8BB136ACFFC6E53550BA86A46664FE70" blockId="7.[159,1391,428,499]" box="[159,1391,428,452]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA86A46664FE70" box="[159,1391,428,452]" pageId="7" pageNumber="124">1. Pronotum red. ...................................................................................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC6E53550BA86D36664FE47" blockId="7.[159,1391,428,499]" box="[159,1391,475,499]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA86D36664FE47" box="[159,1391,475,499]" pageId="7" pageNumber="124">– Pronotum dark-brown to black. ........................................................................................................................................ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC6E53550BA85026664FDC5" pageId="7" pageNumber="124">
<paragraph id="8BB136ACFFC6E53550BA85026664FDF6" blockId="7.[159,1391,522,625]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA85026664FDF6" pageId="7" pageNumber="124">
2. Head red, of similar coloration as pronotum (
<figureCitation id="13352A29FFC6E53552928502600FFD96" box="[695,772,522,546]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="7" pageNumber="124">Fig. 61</figureCitation>
). Elytra with very coarse and dense punctation (
<figureCitation id="13352A29FFC6E535553385026668FD96" box="[1302,1379,522,546]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="7" pageNumber="124">Fig. 61</figureCitation>
). Aedeagus as in 
<figureCitation id="13352A29FFC6E535514D852262FCFDF6" box="[360,503,554,578]" captionStart="Figs 174–190" captionStartId="58.[189,229,2020,2042]" captionTargetBox="[190,1430,246,1968]" captionTargetId="figure-107@58.[189,1436,237,1976]" captionText="Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888372" httpUri="https://zenodo.org/record/5888372/files/figure.png" pageId="7" pageNumber="124">Figs 187–188</figureCitation>
. Distribution: 
<collectingCountry id="F319763CFFC6E53552B4852261F8FDF6" box="[657,755,554,578]" name="Thailand" pageId="7" pageNumber="124">Thailand</collectingCountry>
, 
<collectingCountry id="F319763CFFC6E53552D885226055FDF6" box="[765,862,554,578]" name="Malaysia" pageId="7" pageNumber="124">Malaysia</collectingCountry>
. .................................................................... 
<taxonomicName id="4C0E4D2FFFC6E535553585226664FDF6" ID-CoL="5DDFK" authorityName="Assing" authorityYear="2017" box="[1296,1391,554,578]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="russiceps">
<emphasis id="B97AEABEFFC6E535553585226664FDF6" bold="true" box="[1296,1391,554,578]" italics="true" pageId="7" pageNumber="124">russiceps</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC6E53550BA85516664FDC5" blockId="7.[159,1391,522,625]" box="[159,1391,601,625]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA85516664FDC5" box="[159,1391,601,625]" pageId="7" pageNumber="124">– Head blackish. ...................................................................................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC6E53550BA85806664FCFB" pageId="7" pageNumber="124">
<paragraph id="8BB136ACFFC6E53550BA85806665FD4B" blockId="7.[159,1391,648,847]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA85806665FD4B" pageId="7" pageNumber="124">
3. Very large and robust species; length of forebody&gt; 
<quantity id="4CF69B49FFC6E53552DD85806047FD14" box="[760,844,648,672]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="124" unit="mm" value="3.5">3.5 mm</quantity>
; width of pronotum approximately 
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. Elytra reddish with the postero-lateral portion more or less distinctly and more or less extensively infuscate. Legs usually uniformly yellowish. Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC6E53552C185C06078FD54" box="[740,883,712,736]" captionStart="Figs 260–277" captionStartId="63.[151,191,1987,2009]" captionTargetBox="[157,1395,232,1954]" captionTargetId="figure-126@63.[151,1398,232,1971]" captionText="Figs 260–277: Zyras morulus (260–265), Z. variolatus (266–270), Z. montanus (271–275), and Z. preangeranus from Java (276–277): forebody (260); antenna (261); abdomen (262); median lobe of aedeagus in lateral and in ventral view (263–264, 266–267, 271–272, 276–277); paramere (265, 268, 273); male tergite VIII (269, 274); male sternite VIII (270, 275). Scale bars: 260–262: 1.0 mm; 263–277: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888380" httpUri="https://zenodo.org/record/5888380/files/figure.png" pageId="7" pageNumber="124">Figs 276–277</figureCitation>
. Distribution: Oriental and southern East Palaearctic regions (
<figureCitation id="13352A29FFC6E535514485E062BCFCB4" box="[353,439,744,768]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="7" pageNumber="124">Map 10</figureCitation>
). ................................................................................................................................ 
<taxonomicName id="4C0E4D2FFFC6E53554F885E06665FD4B" ID-CoL="5DDF3" authorityName="Cameron" authorityYear="1939" box="[1245,1390,744,767]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="preangeranus">
<emphasis id="B97AEABEFFC6E53554F885E06665FD4B" bold="true" box="[1245,1390,744,767]" italics="true" pageId="7" pageNumber="124">preangeranus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC6E53550BA84106664FCFB" blockId="7.[159,1391,648,847]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA84106664FCFB" pageId="7" pageNumber="124">
– Smaller and more slender species; length of forebody &lt;
<quantity id="4CF69B49FFC6E535533A841F6078FC9B" box="[799,883,791,815]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="124" unit="mm" value="3.5">3.5 mm</quantity>
; width of pronotum &lt;
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. Elytra blackish, often with slight bluish hue. At least the apices of the femora usually infuscate. ....................................................... 4
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</paragraph>
</keyStep>
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<paragraph id="8BB136ACFFC6E53550BA846E6664FC0A" blockId="7.[159,1392,870,1070]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA846E6664FC0A" pageId="7" pageNumber="124">
4. Profemora and the apical halves of the meso- and metafemora blackish-brown to black. Antennae very slender, approximately 
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long; antennomeres IV–VII distinctly oblong and X very weakly transverse. Elytra with rather coarse punctation. Distribution: 
<collectingCountry id="F319763CFFC6E535524384AE61CDFC0A" box="[614,710,934,958]" name="Thailand" pageId="7" pageNumber="124">Thailand</collectingCountry>
(
<figureCitation id="13352A29FFC6E53552F684AE6011FC0A" box="[723,794,934,958]" captionStart="Map 8" captionStartId="68.[189,234,1235,1256]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-71@68.[189,1436,232,1226]" captionTargetPageId="68" captionText="Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white" figureDoi="http://doi.org/10.5281/zenodo.5742417" httpUri="https://zenodo.org/record/5742417/files/figure.png" pageId="7" pageNumber="124">Map 8</figureCitation>
). ............................................................................ 
<taxonomicName id="4C0E4D2FFFC6E535552C84AE6664FC0A" ID-CoL="5DDG9" authorityName="PACE" authorityYear="1986" box="[1289,1391,934,958]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="thaiorum">
<emphasis id="B97AEABEFFC6E535552C84AE6664FC0A" bold="true" box="[1289,1391,934,958]" italics="true" pageId="7" pageNumber="124">thaiorum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC6E53550BA84DE6664FB99" blockId="7.[159,1392,870,1070]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA84DE6664FB99" pageId="7" pageNumber="124">
– Femora yellowish with the apices usually narrowly infuscate. Antennae much shorter, &lt;
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long; antennomeres V–VII not oblong and X distinctly transverse. Elytra with very fine punctation. Aedeagus: 
<bibRefCitation id="EF9F4B5DFFC6E53554E984FE6667FBB9" author="ASSING, V." box="[1228,1388,1013,1038]" pageId="7" pageNumber="124" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 274–281). Distribution: Oriental and southern East Palaearctic regions (
<figureCitation id="13352A29FFC6E535542B831E675EFB9A" box="[1038,1109,1046,1070]" captionStart="Map 8" captionStartId="68.[189,234,1235,1256]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-71@68.[189,1436,232,1226]" captionTargetPageId="68" captionText="Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white" figureDoi="http://doi.org/10.5281/zenodo.5742417" httpUri="https://zenodo.org/record/5742417/files/figure.png" pageId="7" pageNumber="124">Map 8</figureCitation>
). ........................... 
<taxonomicName id="4C0E4D2FFFC6E5355533831D6664FB99" ID-CoL="5DDCK" baseAuthorityName="Kraatz" baseAuthorityYear="1859" box="[1302,1391,1045,1069]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="geminus">
<emphasis id="B97AEABEFFC6E5355533831D6664FB99" bold="true" box="[1302,1391,1045,1069]" italics="true" pageId="7" pageNumber="124">geminus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC6E53550BA834C6664FB3F" pageId="7" pageNumber="124">
<paragraph id="8BB136ACFFC6E53550BA834C6664FBE8" blockId="7.[159,1391,1092,1164]" box="[159,1391,1092,1117]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA834C6664FBE8" box="[159,1391,1092,1117]" pageId="7" pageNumber="124">5. Legs bicoloured, at least the apices of the meso- and metafemora more or less distinctly infuscate. ...................... 6</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC6E53550BA837C6664FB3F" blockId="7.[159,1391,1092,1164]" box="[159,1391,1140,1164]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA837C6664FB3F" box="[159,1391,1140,1164]" pageId="7" pageNumber="124">– Legs uniformly pale-yellowish. .......................................................................................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC6E53550BA83AB6664FADD" pageId="7" pageNumber="124">
<paragraph id="8BB136ACFFC6E53550BA83AB6664FB4E" blockId="7.[159,1391,1187,1386]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA83AB6664FB4E" pageId="7" pageNumber="124">
6. Apices of meso- and metafemora narrowly and weakly infuscate. Elytra short and with fine punctation (
<figureCitation id="13352A29FFC6E535553383AB6668FB0F" box="[1302,1379,1187,1211]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="7" pageNumber="124">Fig. 70</figureCitation>
); hind wings reduced. Segments II–V of abdomen reddish (
<figureCitation id="13352A29FFC6E535530F83CB6088FB6F" box="[810,899,1219,1243]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="7" pageNumber="124">Fig. 107</figureCitation>
). Aedeagus: 
<figureCitation id="13352A29FFC6E535542D83CB679CFB6F" box="[1032,1175,1219,1243]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="7" pageNumber="124">Figs 212–215</figureCitation>
. Distribution: South 
<collectingCountry id="F319763CFFC6E53550E183EB622FFB4F" box="[196,292,1251,1275]" name="Thailand" pageId="7" pageNumber="124">Thailand</collectingCountry>
(
<figureCitation id="13352A29FFC6E535511783EB628CFB4F" box="[306,391,1251,1275]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="7" pageNumber="124">Map 10</figureCitation>
). .............................................................................................................................................. 
<taxonomicName id="4C0E4D2FFFC6E535552083EA6664FB4E" ID-CoL="7GHVN" authorityName="Assing" authorityYear="2017" box="[1285,1391,1250,1274]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="ambulans">
<emphasis id="B97AEABEFFC6E535552083EA6664FB4E" bold="true" box="[1285,1391,1250,1274]" italics="true" pageId="7" pageNumber="124">ambulans</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC6E53550BA821A6664FADD" blockId="7.[159,1391,1187,1386]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BA821A6664FADD" pageId="7" pageNumber="124">
– Profemora dark-brown; apical halves of meso- and metafemora blackish. Elytra longer and with coarser punctation (
<figureCitation id="13352A29FFC6E53550D9823A6243FAFE" box="[252,328,1330,1354]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="7" pageNumber="124">Fig. 66</figureCitation>
). Most of segments II–V blackish (
<figureCitation id="13352A29FFC6E5355295823A6002FAFE" box="[688,777,1330,1354]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="7" pageNumber="124">Fig. 103</figureCitation>
). Male unknown. Distribution: 
<collectingCountry id="F319763CFFC6E5355477823A67BFFAFE" box="[1106,1204,1330,1354]" name="Thailand" pageId="7" pageNumber="124">Thailand</collectingCountry>
, Southwest 
<collectingCountry id="F319763CFFC6E5355508823A6665FAFE" box="[1325,1390,1330,1354]" name="China" pageId="7" pageNumber="124">China</collectingCountry>
(
<figureCitation id="13352A29FFC6E53550E9825A6219FADE" box="[204,274,1362,1386]" captionStart="Map 8" captionStartId="68.[189,234,1235,1256]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-71@68.[189,1436,232,1226]" captionTargetPageId="68" captionText="Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white" figureDoi="http://doi.org/10.5281/zenodo.5742417" httpUri="https://zenodo.org/record/5742417/files/figure.png" pageId="7" pageNumber="124">Map 8</figureCitation>
). ................................................................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC6E535553682596664FADD" ID-CoL="5DDB9" baseAuthorityName="PACE" baseAuthorityYear="1986" box="[1299,1391,1361,1385]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="B97AEABEFFC6E535553682596664FADD" bold="true" box="[1299,1391,1361,1385]" italics="true" pageId="7" pageNumber="124">brignolii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC6E53550BB82896664FA7C" pageId="7" pageNumber="124">
<paragraph id="8BB136ACFFC6E53550BB82896664FA2D" blockId="7.[158,1391,1409,1480]" box="[158,1391,1409,1433]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BB82896664FA2D" box="[158,1391,1409,1433]" pageId="7" pageNumber="124">7. Anterior abdominal tergites of dark coloration (posterior margins of segments may be paler). ............................. 8</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC6E53550BB82B96664FA7C" blockId="7.[158,1391,1409,1480]" box="[158,1391,1456,1480]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BB82B96664FA7C" box="[158,1391,1456,1480]" pageId="7" pageNumber="124">– At least tergites II–V reddish. .......................................................................................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC6E53550BB82D76664F972" pageId="7" pageNumber="124">
<paragraph id="8BB136ACFFC6E53550BB82D76665F9E3" blockId="7.[158,1391,1503,1734]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BB82D76665F9E3" pageId="7" pageNumber="124">
8. Antennae massive; antennomere XI barely as long as the combined length of IX and X (
<bibRefCitation id="EF9F4B5DFFC6E535544482D767FCFA43" author="ASSING, V." box="[1121,1271,1503,1527]" pageId="7" pageNumber="124" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 41). Pronotum distinctly transverse, at least 1.2 times as broad as long and approximately 1.3 times as broad as head, rather finely and regularly punctate. Median lobe of aedeagus: 
<bibRefCitation id="EF9F4B5DFFC6E53553518117671CF983" author="ASSING, V." box="[884,1047,1567,1591]" pageId="7" pageNumber="124" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 215–216). Distribution: 
<collectingCountry id="F319763CFFC6E53550E18137622DF9E3" box="[196,294,1599,1623]" name="Thailand" pageId="7" pageNumber="124">Thailand</collectingCountry>
, 
<collectingCountry id="F319763CFFC6E53551158137627FF9E3" box="[304,372,1599,1623]" name="China" pageId="7" pageNumber="124">China</collectingCountry>
: 
<collectingRegion id="49CAF84EFFC6E535515B814862D9F9E3" box="[382,466,1600,1623]" country="China" name="Yunnan" pageId="7" pageNumber="124">Yunnan</collectingRegion>
(
<figureCitation id="13352A29FFC6E53551FA8137612EF9E3" box="[479,549,1599,1623]" captionStart="Map 8" captionStartId="68.[189,234,1235,1256]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-71@68.[189,1436,232,1226]" captionTargetPageId="68" captionText="Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white" figureDoi="http://doi.org/10.5281/zenodo.5742417" httpUri="https://zenodo.org/record/5742417/files/figure.png" pageId="7" pageNumber="124">Map 8</figureCitation>
). ............................................................................................................... 
<taxonomicName id="4C0E4D2FFFC6E53554CC81376665F9E3" ID-CoL="5DDBC" authorityName="ASSING" authorityYear="2016" box="[1257,1390,1599,1623]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="caloderoides">
<emphasis id="B97AEABEFFC6E53554CC81376665F9E3" bold="true" box="[1257,1390,1599,1623]" italics="true" pageId="7" pageNumber="124">caloderoides</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC6E53550BB81676664F972" blockId="7.[158,1391,1503,1734]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BB81676664F972" pageId="7" pageNumber="124">
– Antennae slender; antennomere XI conspicuously elongate, approximately as long as the combined length of VIII–X (
<figureCitation id="13352A29FFC6E53550EE81866263F912" box="[203,360,1678,1702]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="7" pageNumber="124">Figs 28–29, 33</figureCitation>
). Pronotum slender, much less than 1.2 times as broad as long and only slightly broader than head (
<figureCitation id="13352A29FFC6E53550EE81A66211F972" box="[203,282,1710,1734]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="7" pageNumber="124">Figs 65</figureCitation>
, 
<figureCitation id="13352A29FFC6E535510081A66249F972" box="[293,322,1710,1734]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="7" pageNumber="124">69</figureCitation>
); punctation coarse and irregularly distributed. ........................................................................................ 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC6E53550BB81D66664F870" pageId="7" pageNumber="124">
<paragraph id="8BB136ACFFC6E53550BB81D66664F8E1" blockId="7.[158,1391,1757,1989]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BB81D66664F8E1" pageId="7" pageNumber="124">
9. Antennae more slender (
<figureCitation id="13352A29FFC6E53551ED81D5614AF942" box="[456,577,1757,1782]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="7" pageNumber="124">Figs 28–29</figureCitation>
). Pronotum with very uneven surface (i.e., with more or less irregular impressions); punctation conspicuously irregularly distributed (
<figureCitation id="13352A29FFC6E535530F81F5607CF8A1" box="[810,887,1789,1813]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="7" pageNumber="124">Fig. 65</figureCitation>
). Male tergite VIII with four blunt projections posteriorly (
<figureCitation id="13352A29FFC6E535516F801562AEF881" box="[330,421,1821,1845]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="7" pageNumber="124">Fig. 196</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC6E5355337801560A8F882" box="[786,931,1821,1846]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="7" pageNumber="124">Figs 191–194</figureCitation>
. Distribution: Oriental and southern East Palaearctic regions (
<figureCitation id="13352A29FFC6E53551BC803662EAF8E1" box="[409,481,1853,1878]" captionStart="Map 6" captionStartId="66.[189,234,1236,1257]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-64@66.[189,1436,232,1226]" captionTargetPageId="66" captionText="Map 6: Revised distributions of Zyras geminus (black circles) and Z. proximus (white circles) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742411" httpUri="https://zenodo.org/record/5742411/files/figure.png" pageId="7" pageNumber="124">Map 6</figureCitation>
). ................................................................................................................................ 
<taxonomicName id="4C0E4D2FFFC6E535552F80356664F8E1" ID-CoL="5DDF4" authorityName="CAMERON" authorityYear="1939" box="[1290,1391,1853,1877]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="B97AEABEFFC6E535552F80356664F8E1" bold="true" box="[1290,1391,1853,1877]" italics="true" pageId="7" pageNumber="124">proximus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC6E53550BB80656664F870" blockId="7.[158,1391,1757,1989]" pageId="7" pageNumber="124">
<keyLead id="30FF9599FFC6E53550BB80656664F870" pageId="7" pageNumber="124">
– Antennae less slender (
<figureCitation id="13352A29FFC6E535519880656101F831" box="[445,522,1901,1925]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="7" pageNumber="124">Fig. 33</figureCitation>
). Pronotum without impressions and with less irregularly distributed punctation (
<figureCitation id="13352A29FFC6E53550E980856213F811" box="[204,280,1933,1957]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="7" pageNumber="124">Fig. 69</figureCitation>
). Male tergite VIII only with a median pair of less distinct projections posteriorly (
<figureCitation id="13352A29FFC6E535545A808567D2F811" box="[1151,1241,1933,1957]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="7" pageNumber="124">Fig. 206</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC6E535515880A56106F871" box="[381,525,1965,1989]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="7" pageNumber="124">Figs 208–211</figureCitation>
. Distribution: 
<collectingCountry id="F319763CFFC6E535528380A5600DF871" box="[678,774,1965,1989]" name="Thailand" pageId="7" pageNumber="124">Thailand</collectingCountry>
(
<figureCitation id="13352A29FFC6E535533680A56062F871" box="[787,873,1965,1989]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="7" pageNumber="124">Map 10</figureCitation>
). .............................................................. parvicollis
</keyLead>
</paragraph>
</keyStep>
</key>
<key id="FF9F619DFFC9E53A50E087F1669FFC28" pageId="8" pageNumber="125">
<keyStep id="30FA2E09FFC9E53A50E087F1669EFED4" pageId="8" pageNumber="125">
<paragraph id="8BB136ACFFC9E53A50E087F1669EFE85" blockId="8.[196,1429,248,924]" pageId="8" pageNumber="125">
<keyLead id="30FF9599FFC9E53A50E087F1669EFE85" pageId="8" pageNumber="125">
10. Antennomere XI conspicuously elongate, approximately as long as the combined length of VIII–X (
<figureCitation id="13352A29FFC9E53A54C887F06681FEA5" box="[1261,1418,248,273]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="8" pageNumber="125">Figs 27, 30–31</figureCitation>
). .............................................................................................................................................................................................. 11
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC9E53A50E08640669EFED4" blockId="8.[196,1429,248,924]" box="[197,1429,327,352]" pageId="8" pageNumber="125">
<keyLead id="30FF9599FFC9E53A50E08640669EFED4" box="[197,1429,327,352]" pageId="8" pageNumber="125">– Antennomere XI less elongate, shorter than the combined length of VIII–X. ......................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FA2E09FFC9E53A50E1867F669EFD29" pageId="8" pageNumber="125">
<paragraph id="8BB136ACFFC9E53A50E1867F669EFE5B" blockId="8.[196,1429,248,924]" pageId="8" pageNumber="125">
<keyLead id="30FF9599FFC9E53A50E1867F669EFE5B" pageId="8" pageNumber="125">
11. Pronotum with uneven surface, more or less irregularly impressed; punctation coarse and irregularly distributed (
<figureCitation id="13352A29FFC9E53A50D7869F6235FE1B" box="[242,318,407,431]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="8" pageNumber="125">Fig. 67</figureCitation>
). Anterior impressions of tergites III–V and anterior portion of tergite VI with coarse and dense nonsetiferous punctation (
<figureCitation id="13352A29FFC9E53A51F686BF6120FE7B" box="[467,555,439,463]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="8" pageNumber="125">Fig. 106</figureCitation>
). Posterior margin of male tergite VIII with four blunt projections (
<figureCitation id="13352A29FFC9E53A54F986BF663FFE7B" box="[1244,1332,439,463]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="8" pageNumber="125">Fig. 201</figureCitation>
). Median lobe of aedeagus as in 
<figureCitation id="13352A29FFC9E53A51F286DF616DFE5B" box="[471,614,471,495]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="8" pageNumber="125">Figs 198–199</figureCitation>
. Distribution: 
<collectingCountry id="F319763CFFC9E53A52DA86DF606AFE5B" box="[767,865,471,495]" name="Thailand" pageId="8" pageNumber="125">Thailand</collectingCountry>
, 
<collectingCountry id="F319763CFFC9E53A534986DF6095FE5A" box="[876,926,471,494]" name="Laos" pageId="8" pageNumber="125">Laos</collectingCountry>
(
<figureCitation id="13352A29FFC9E53A538E86DF60F9FE5A" box="[939,1010,470,495]" captionStart="Map 5" captionStartId="65.[151,196,1829,1850]" captionTargetBox="[151,1398,827,1821]" captionTargetId="figure-87@65.[151,1398,827,1821]" captionTargetPageId="65" captionText="Map 5: Revised distributions of Zyras castaneus (black circles) and Z. novinversus (white circles) in the southern East Palaearctic" figureDoi="http://doi.org/10.5281/zenodo.5742405" httpUri="https://zenodo.org/record/5742405/files/figure.png" pageId="8" pageNumber="125">Map 5</figureCitation>
). ........................................... 
<taxonomicName id="4C0E4D2FFFC9E53A553086DF669EFE5B" ID-CoL="7GHZ5" authorityName="Assing" authorityYear="2017" box="[1301,1429,471,495]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="125" phylum="Arthropoda" rank="species" species="novinversus">
<emphasis id="B97AEABEFFC9E53A553086DF669EFE5B" bold="true" box="[1301,1429,471,495]" italics="true" pageId="8" pageNumber="125">novinversus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB136ACFFC9E53A50E1850E669EFD29" blockId="8.[196,1429,248,924]" pageId="8" pageNumber="125">
<keyLead id="30FF9599FFC9E53A50E1850E669EFD29" pageId="8" pageNumber="125">
– Pronotum with smooth surface, without impressions; punctation fine and more or less regularly distributed (
<bibRefCitation id="EF9F4B5DFFC9E53A5562850E6227FD8A" author="ASSING, V." pageId="8" pageNumber="125" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 97). Anterior impressions of tergites III–V and anterior portion of tergite VI with sparse and very fine non-setiferous punctation (
<bibRefCitation id="EF9F4B5DFFC9E53A522F854E6195FDEA" author="ASSING, V." box="[522,670,582,606]" pageId="8" pageNumber="125" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 154). Posterior margin of male tergite VIII with shallow concavity in the middle. Median lobe of aedeagus: 
<bibRefCitation id="EF9F4B5DFFC9E53A5280856E604CFDCA" author="ASSING, V." box="[677,839,614,638]" pageId="8" pageNumber="125" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 272–273). Distribution: Oriental and southern East Palaearctic regions (
<figureCitation id="13352A29FFC9E53A51D7858E6131FD2A" box="[498,570,646,670]" captionStart="Map 5" captionStartId="65.[151,196,1829,1850]" captionTargetBox="[151,1398,827,1821]" captionTargetId="figure-87@65.[151,1398,827,1821]" captionTargetPageId="65" captionText="Map 5: Revised distributions of Zyras castaneus (black circles) and Z. novinversus (white circles) in the southern East Palaearctic" figureDoi="http://doi.org/10.5281/zenodo.5742405" httpUri="https://zenodo.org/record/5742405/files/figure.png" pageId="8" pageNumber="125">Map 5</figureCitation>
). ....................................................................................................................... 
<taxonomicName id="4C0E4D2FFFC9E53A5508858E669EFD29" ID-CoL="5DDBD" baseAuthorityName="Motschulsky" baseAuthorityYear="1861" box="[1325,1429,646,669]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="125" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="B97AEABEFFC9E53A5508858E669EFD29" bold="true" box="[1325,1429,646,669]" italics="true" pageId="8" pageNumber="125">castaneus</emphasis>
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12. Pronotum strongly transverse, nearly 1.3 times as broad as long and 1.2 times as broad as head (
<figureCitation id="13352A29FFC9E53A54C985BD6633FD79" box="[1260,1336,693,717]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="8" pageNumber="125">Fig. 68</figureCitation>
). Tergite VII nearly completely blackish-brown to black (
<figureCitation id="13352A29FFC9E53A52FB85DD6033FD59" box="[734,824,725,749]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="8" pageNumber="125">Fig. 102</figureCitation>
). Median lobe of aedeagus with very short ventral process (
<figureCitation id="13352A29FFC9E53A50D485FD628FFCB9" box="[241,388,757,781]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="8" pageNumber="125">Figs 203–204</figureCitation>
). Distribution: 
<collectingCountry id="F319763CFFC9E53A520085FD618EFCB9" box="[549,645,757,781]" name="Thailand" pageId="8" pageNumber="125">Thailand</collectingCountry>
(
<figureCitation id="13352A29FFC9E53A52B685FD61E3FCB9" box="[659,744,757,781]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="8" pageNumber="125">Map 10</figureCitation>
). ..................................................................................... 
<taxonomicName id="4C0E4D2FFFC9E53A553585FC669EFCB8" ID-CoL="5DDB8" authorityName="Assing" authorityYear="2017" box="[1296,1429,756,780]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="125" phylum="Arthropoda" rank="species" species="brevilobatus">
<emphasis id="B97AEABEFFC9E53A553585FC669EFCB8" bold="true" box="[1296,1429,756,780]" italics="true" pageId="8" pageNumber="125">brevilobatus</emphasis>
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<paragraph id="8BB136ACFFC9E53A50E1842D669FFC28" blockId="8.[196,1429,248,924]" pageId="8" pageNumber="125">
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– Pronotum weakly transverse, approximately 1.15 times as broad as long (
<bibRefCitation id="EF9F4B5DFFC9E53A53DE842C6799FC88" author="ASSING, V." box="[1019,1170,804,828]" pageId="8" pageNumber="125" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 172). Tergite VII bicoloured with the anterior third reddish and the posterior two-thirds black (
<bibRefCitation id="EF9F4B5DFFC9E53A5401844C67B2FCE8" author="ASSING, V." box="[1060,1209,836,860]" pageId="8" pageNumber="125" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
: figure 174). Median lobe of aedeagus: 
<bibRefCitation id="EF9F4B5DFFC9E53A518D846C6142FCC8" author="ASSING, V." box="[424,585,868,892]" pageId="8" pageNumber="125" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a</bibRefCitation>
: figures 231–232). Distribution: Oriental and southern East Palaearctic regions (
<figureCitation id="13352A29FFC9E53A50D4848C6232FC28" box="[241,313,900,924]" captionStart="Map 9" captionStartId="69.[151,196,1235,1256]" captionTargetBox="[151,1398,232,1226]" captionTargetId="figure-70@69.[151,1398,232,1226]" captionTargetPageId="69" captionText="Map 9: Revised distributions of Zyras bettotanus (black circles), Z. nilgiriensis (star), and Z. longilobatus (white circles) in the" figureDoi="http://doi.org/10.5281/zenodo.5742419" httpUri="https://zenodo.org/record/5742419/files/figure.png" pageId="8" pageNumber="125">Map 9</figureCitation>
). ................................................................................................................................................................ bettotanus
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