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<mods:title id="8A8C9347617E836E0FCB605D12BCC2FB">A Revision Of The Didelphid Marsupial Genus Marmosops, Part 1. Species Of The Subgenus Sciophanes</mods:title>
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<taxonomicName id="4C0F43E79835FFF006AD71E6D02DFE95" ID-CoL="3Y83R" authority="(Thomas, 1896)" baseAuthorityName="Thomas" baseAuthorityYear="1896" box="[210,606,250,274]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769835FFF006AD71E6D3D8FE95" box="[210,427,250,274]" italics="true" pageId="34" pageNumber="35">Marmosops fuscatus</emphasis>
(
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)
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<paragraph id="8BB038649835FFF007497037D389FEC4" blockId="34.[310,506,299,323]" box="[310,506,299,323]" pageId="34" pageNumber="35">
<heading id="D0F88F089835FFF007497037D389FEC4" box="[310,506,299,323]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="35" reason="2">
<figureCitation id="133424E19835FFF007497037D3DFFEC4" box="[310,428,299,323]" captionStart="FIG" captionStartId="8.[144,185,910,931]" captionTargetBox="[144,1248,249,884]" captionTargetId="figure-387@8.[144,1248,248,884]" captionTargetPageId="8" captionText="FIG. 4. Dorsal views of the rostrum of Marmosops invictus (A, USNM 306386) and M. fuscatus (B, FMNH 22174), illustrating taxonomic differences in nasal width. Abbreviations: fro, frontal; max, maxillary; nas, nasal." figureDoi="http://doi.org/10.5281/zenodo.4612626" httpUri="https://zenodo.org/record/4612626/files/figure.png" pageId="34" pageNumber="35">Figures 4B</figureCitation>
, 
<figureCitation id="133424E19835FFF007C77037D3A7FEC4" box="[440,468,299,323]" captionStart="FIG" captionStartId="30.[144,186,1727,1748]" captionTargetBox="[243,1149,248,1701]" captionTargetId="figure-33@30.[243,1149,248,1701]" captionTargetPageId="30" captionText="FIG. 15. Dorsal and ventral views of skulls of Marmosops carri (left, AMNH 259978) and M. fuscatus (right, AMNH 276580). All views about ×2.5." figureDoi="http://doi.org/10.5281/zenodo.4612658" httpUri="https://zenodo.org/record/4612658/files/figure.png" pageId="34" pageNumber="35">15</figureCitation>
, 
<figureCitation id="133424E19835FFF0079F7037D389FEC4" box="[480,506,299,323]" captionStart="FIG" captionStartId="31.[144,186,1347,1368]" captionTargetBox="[282,1110,249,1311]" captionTargetId="figure-159@31.[282,1110,249,1311]" captionTargetPageId="31" captionText="FIG. 16. Lateral views of skulls and mandibles of Marmosops carri (above, AMNH 259978) and M. fuscatus (below, AMNH 276580). All views about ×2.5." figureDoi="http://doi.org/10.5281/zenodo.4612662" httpUri="https://zenodo.org/record/4612662/files/figure.png" pageId="34" pageNumber="35">16</figureCitation>
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<emphasis id="B97BE4769835FFF006EF7043D335FEF1" box="[144,326,350,374]" italics="true" pageId="34" pageNumber="35">Marmosa fuscata</emphasis>
<bibRefCitation id="EF9E45959835FFF007327042D065FEF1" author="Thomas, O." box="[333,534,350,374]" pageId="34" pageNumber="35" pagination="301 - 314" refId="ref39327" refString="Thomas, O. 1896. On new small mammals from the Neotropical region. Annals and Magazine of Natural History (ser. 6) 18: 301 - 314." type="journal article" year="1896">Thomas, 1896: 313</bibRefCitation>
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(original description).
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<treatmentCitation id="0AAE1E759835FFF006EF70BED280FE5B" author="Trouessart, E. - L." page="1240" pageId="34" pageNumber="35" year="1898">
<taxonomicName id="4C0F43E79835FFF006EF70BED280FE5B" authority=": Trouessart, 1898: 1240" authorityName="Trouessart" authorityPageNumber="1240" authorityYear="1898" class="Mammalia" family="Didelphidae" genus="Didelphys" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="fuscata" subGenus="Marmosa">
<emphasis id="B97BE4769835FFF006EF70BED284FE3D" box="[144,247,418,442]" italics="true" pageId="34" pageNumber="35">Didelphys</emphasis>
(
<emphasis id="B97BE4769835FFF0077870BFD31FFE3D" box="[263,364,419,442]" italics="true" pageId="34" pageNumber="35">Marmosa</emphasis>
) 
<emphasis id="B97BE4769835FFF0070370BED3BEFE3D" box="[380,461,418,442]" italics="true" pageId="34" pageNumber="35">fuscata:</emphasis>
<bibRefCitation id="EF9E45959835FFF007AB70BFD280FE5B" author="Trouessart, E. - L." pageId="34" pageNumber="35" refId="ref39499" refString="Trouessart, E. - L. 1898. Catalogus mammalium tam viventium quam fossilium. Fasiculus V. Sirenia, Cetacea, Edentata, Marsupialia, Allotheria, Monotremata. Berolini: R. Friedlander &amp; Sohn." type="book" year="1898">Trouessart, 1898: 1240</bibRefCitation>
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(name combination).
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<treatmentCitation id="0AAE1E759835FFF006EF70FBD0FCFE79" author="Cabrera, A." box="[144,655,486,511]" page="36" pageId="34" pageNumber="35" year="1919">
<taxonomicName id="4C0F43E79835FFF006EF70FBD0FCFE79" authority=": Cabrera, 1919: 36" authorityName="Cabrera" authorityPageNumber="36" authorityYear="1919" box="[144,655,486,511]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="fuscata" subGenus="Marmosa">
<emphasis id="B97BE4769835FFF006EF70FBD286FE79" box="[144,245,487,510]" italics="true" pageId="34" pageNumber="35">Marmosa</emphasis>
(
<emphasis id="B97BE4769835FFF0077A70FBD319FE79" box="[261,362,487,510]" italics="true" pageId="34" pageNumber="35">Marmosa</emphasis>
) 
<emphasis id="B97BE4769835FFF0070570FAD3B9FE79" box="[378,458,486,510]" italics="true" pageId="34" pageNumber="35">fuscata:</emphasis>
<bibRefCitation id="EF9E45959835FFF007AD70FAD0FCFE79" author="Cabrera, A." box="[466,655,486,511]" pageId="34" pageNumber="35" refId="ref35568" refString="Cabrera, A. 1919. Genera mammalium: Monotremata, Marsupialia. Madrid: Museo de Ciencias Naturales." type="book" year="1919">Cabrera, 1919: 36</bibRefCitation>
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(name combination).
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<treatmentCitation id="0AAE1E759835FFF006EF7337D0EFFDC4" author="Handley, C. O., Jr. &amp; L. K Gordon" box="[144,668,555,579]" page="66" pageId="34" pageNumber="35" year="1979">
<taxonomicName id="4C0F43E79835FFF006EF7337D0EFFDC4" authority="Handley and Gordon, 1979: 66" authorityName="Handley and Gordon" authorityPageNumber="66" authorityYear="1979" box="[144,668,555,579]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769835FFF006EF7337D33BFDC5" box="[144,328,555,578]" italics="true" pageId="34" pageNumber="35">Marmosa cracens</emphasis>
<bibRefCitation id="EF9E45959835FFF007307337D0EFFDC4" author="Handley, C. O., Jr. &amp; L. K Gordon" box="[335,668,555,579]" pageId="34" pageNumber="35" pagination="65 - 72" refId="ref36974" refString="Handley, C. O., Jr., and L. K Gordon. 1979. New species of mammals from northern South America: mouse opossums, genus Marmosa Gray. In J. F. Eisenberg (editor), Vertebrate ecology in the northern Neotropics: 65 - 72. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1979">Handley and Gordon, 1979: 66</bibRefCitation>
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(original description). 
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locality “near La Pastora, 
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ENE Mirimire, 
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, 
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.”
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<treatmentCitation id="0AAE1E759835FFF006EF73A8D04EFD4C" author="Reig, O. A. &amp; J. A. W. Kirsch &amp; L. G. Marshall" box="[144,573,691,715]" page="342" pageId="34" pageNumber="35" year="1985">
<taxonomicName id="4C0F43E79835FFF006EF73A8D04EFD4C" authority=": Reig et al., 1985: 342" authorityName="Reig" authorityPageNumber="342" authorityYear="1985" box="[144,573,691,715]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="crascens">
<emphasis id="B97BE4769835FFF006EF73A8D324FD4C" box="[144,343,692,715]" italics="true" pageId="34" pageNumber="35">Marmosa crascens:</emphasis>
<bibRefCitation id="EF9E45959835FFF0072073AFD04EFD4C" author="Reig, O. A. &amp; J. A. W. Kirsch &amp; L. G. Marshall" box="[351,573,691,715]" pageId="34" pageNumber="35" pagination="335 - 343" refId="ref38811" refString="Reig, O. A., J. A. W. Kirsch, and L. G. Marshall. 1985. New conclusions on the relationships of the opossumlike marsupials, with an annotated classification of the Didelphimorphia. Ameghiniana 21: 335 - 343." type="journal article" year="1985">Reig et al., 1985: 342</bibRefCitation>
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(incorrect subsequent spelling of 
<taxonomicName id="4C0F43E79835FFF0079A73CAD39AFC97" authority="Handley and Gordon, 1979" authorityName="Handley and Gordon" authorityYear="1979" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769835FFF0079A73CAD025FD6A" box="[485,598,726,749]" italics="true" pageId="34" pageNumber="35">M. cracens</emphasis>
<bibRefCitation id="EF9E45959835FFF006BF73EBD39AFC97" author="Handley, C. O., Jr. &amp; L. K Gordon" box="[192,489,759,784]" pageId="34" pageNumber="35" pagination="65 - 72" refId="ref36974" refString="Handley, C. O., Jr., and L. K Gordon. 1979. New species of mammals from northern South America: mouse opossums, genus Marmosa Gray. In J. F. Eisenberg (editor), Vertebrate ecology in the northern Neotropics: 65 - 72. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1979">Handley and Gordon, 1979</bibRefCitation>
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).
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<treatmentCitation id="0AAE1E759835FFF006EF7206D37DFCD3" author="Gardner, A. L. &amp; G. K. Creighton" page="4" pageId="34" pageNumber="35" year="1989">
<taxonomicName id="4C0F43E79835FFF006EF7206D37DFCD3" ID-CoL="3Y83P" authority=": Gardner and Creighton, 1989: 4" authorityName="Gardner and Creighton" authorityPageNumber="4" authorityYear="1989" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769835FFF006EF7206D310FCB6" box="[144,355,794,817]" italics="true" pageId="34" pageNumber="35">Marmosops cracens:</emphasis>
<bibRefCitation id="EF9E45959835FFF007147206D37DFCD3" author="Gardner, A. L. &amp; G. K. Creighton" pageId="34" pageNumber="35" pagination="3 - 7" refId="ref36576" refString="Gardner, A. L., and G. K. Creighton. 1989. A new generic name for Tate's (1933) microtarsus group of South American mouse opossums (Marsupialia: Didelphidae). Proceedings of the Biological Society of Washington 102: 3 - 7." type="journal article" year="1989">Gardner and Creighton, 1989: 4</bibRefCitation>
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(name combination).
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<treatmentCitation id="0AAE1E759835FFF006EF7243D37DFC1F" author="Gardner, A. L. &amp; G. K. Creighton" page="4" pageId="34" pageNumber="35" year="1989">
<taxonomicName id="4C0F43E79835FFF006EF7243D37DFC1F" ID-CoL="3Y83R" authority=": Gardner and Creighton, 1989: 4" authorityName=": Gardner and Creighton" authorityPageNumber="4" authorityYear="1989" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769835FFF006EF7243D318FCF1" box="[144,363,862,886]" italics="true" pageId="34" pageNumber="35">Marmosops fuscatus:</emphasis>
<bibRefCitation id="EF9E45959835FFF0070C7242D37DFC1F" author="Gardner, A. L. &amp; G. K. Creighton" pageId="34" pageNumber="35" pagination="3 - 7" refId="ref36576" refString="Gardner, A. L., and G. K. Creighton. 1989. A new generic name for Tate's (1933) microtarsus group of South American mouse opossums (Marsupialia: Didelphidae). Proceedings of the Biological Society of Washington 102: 3 - 7." type="journal article" year="1989">Gardner and Creighton, 1989: 4</bibRefCitation>
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(first use of current binomial).
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</paragraph>
<paragraph id="8BB038649835FFF006EF72BFD002FC5B" blockId="34.[144,673,350,1757]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="AB1F1F4A9835FFF006EF72BFD002FC5B" pageId="34" pageNumber="35">
<treatmentCitation id="0AAE1E759835FFF006EF72BFD3F4FC5B" author="Perez-Hernandez, R. &amp; P. Soriano &amp; D. Lew" page="35" pageId="34" pageNumber="35" year="1994">
<taxonomicName id="4C0F43E79835FFF006EF72BFD3F4FC5B" authority=": Perez-Hernandez et al., 1994: 35" authorityName="Perez-Hernandez" authorityPageNumber="35" authorityYear="1994" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="cracens" subGenus="Marmosops">
<emphasis id="B97BE4769835FFF006EF72BFD286FC3D" box="[144,245,931,954]" italics="true" pageId="34" pageNumber="35">Marmosa</emphasis>
(
<emphasis id="B97BE4769835FFF0077A72BFD30CFC3D" box="[261,383,931,954]" italics="true" pageId="34" pageNumber="35">Marmosops</emphasis>
) 
<emphasis id="B97BE4769835FFF007F072BFD392FC3D" box="[399,481,931,954]" italics="true" pageId="34" pageNumber="35">cracens:</emphasis>
<bibRefCitation id="EF9E45959835FFF0079672BED3F4FC5B" author="Perez-Hernandez, R. &amp; P. Soriano &amp; D. Lew" pageId="34" pageNumber="35" refId="ref38683" refString="Perez-Hernandez, R., P. Soriano, and D. Lew. 1994. Marsupiales de Venezuela. Caracas: Cuadernos Lagoven." type="book" year="1994">Pérez-Hernández et al., 1994: 35</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(name combination).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB038649835FFF006EF72FBD354FBE2" blockId="34.[144,673,350,1757]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="AB1F1F4A9835FFF006EF72FBD354FBE2" pageId="34" pageNumber="35">
<treatmentCitation id="0AAE1E759835FFF006EF72FBD328FBA6" author="Perez-Hernandez, R. &amp; P. Soriano &amp; D. Lew" page="37" pageId="34" pageNumber="35" year="1994">
<taxonomicName id="4C0F43E79835FFF006EF72FBD328FBA6" authority=": Perez-Hernandez et al., 1994: 37" authorityName="Perez-Hernandez" authorityPageNumber="37" authorityYear="1994" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="fuscata" subGenus="Marmosops">
<emphasis id="B97BE4769835FFF006EF72FBD286FC79" box="[144,245,999,1022]" italics="true" pageId="34" pageNumber="35">Marmosa</emphasis>
(
<emphasis id="B97BE4769835FFF0077A72FBD30CFC79" box="[261,383,999,1022]" italics="true" pageId="34" pageNumber="35">Marmosops</emphasis>
) 
<emphasis id="B97BE4769835FFF007F072FAD3ACFC79" box="[399,479,998,1022]" italics="true" pageId="34" pageNumber="35">fuscata:</emphasis>
<bibRefCitation id="EF9E45959835FFF0079972FAD328FBA6" author="Perez-Hernandez, R. &amp; P. Soriano &amp; D. Lew" pageId="34" pageNumber="35" refId="ref38683" refString="Perez-Hernandez, R., P. Soriano, and D. Lew. 1994. Marsupiales de Venezuela. Caracas: Cuadernos Lagoven." type="book" year="1994">Pérez-Hernández et al., 1994: 37</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(name combination); part (not including misidentified specimens of 
<taxonomicName id="4C0F43E79835FFF006BF7552D364FBE2" baseAuthorityName="Allen and Chapman" baseAuthorityYear="1897" box="[192,279,1102,1125]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="carri">
<emphasis id="B97BE4769835FFF006BF7552D364FBE2" box="[192,279,1102,1125]" italics="true" pageId="34" pageNumber="35">M. carri</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB038649835FFF006EF756CD330FB6A" blockId="34.[144,673,350,1757]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="AB1F1F4A9835FFF006EF756CD330FB6A" pageId="34" pageNumber="35">
<treatmentCitation id="0AAE1E759835FFF006EF756CD3E7FB2E" author="Gardner, A. L. &amp; G. K. Creighton" page="66" pageId="34" pageNumber="35" year="2008">
<taxonomicName id="4C0F43E79835FFF006EF756CD3E7FB2E" ID-CoL="5JJQH" authority="fuscatus: Gardner and Creighton, 2008: 66" authorityName="fuscatus: Gardner and Creighton" authorityPageNumber="66" authorityYear="2008" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="subSpecies" species="fuscatus" subSpecies="fuscatus">
<emphasis id="B97BE4769835FFF006EF756CD3B4FB00" box="[144,455,1135,1159]" italics="true" pageId="34" pageNumber="35">Marmosops fuscatus fuscatus:</emphasis>
<bibRefCitation id="EF9E45959835FFF007B07573D3E7FB2E" author="Gardner, A. L. &amp; G. K. Creighton" pageId="34" pageNumber="35" pagination="61 - 74" refId="ref36627" refString="Gardner, A. L., and G. K. Creighton. 2008 (&quot; 2007 &quot;). Genus Marmosops Matschie, 1916. In A. L. Gardner (editor), Mammals of South America, vol. 1: marsupials, xenarthrans, shrews, and bats: 61 - 74. Chicago: University of Chicago Press." type="book chapter" year="2008">Gardner and Creighton, 2008: 66</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(name combination); part (not including misidentified specimens of 
<taxonomicName id="4C0F43E79835FFF006A375CAD340FB6A" baseAuthorityName="Allen and Chapman" baseAuthorityYear="1897" box="[220,307,1238,1261]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="carri">
<emphasis id="B97BE4769835FFF006A375CAD340FB6A" box="[220,307,1238,1261]" italics="true" pageId="34" pageNumber="35">M. carri</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3156BEF9835FFF006CF75EBD330F990" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BB038649835FFF006CF75EBD330F990" blockId="34.[144,673,350,1757]" pageId="34" pageNumber="35">
<collectionCode id="ED1EA0A19835FFF006CF75EBD317FA88" box="[176,356,1271,1295]" pageId="34" pageNumber="35">TYPE MATERIAL</collectionCode>
: The 
<typeStatus id="54B486C69835FFF007D475EAD07FFA89" box="[427,524,1270,1294]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
(by original designation) consists of the skin and skull of an old adult female (
<collectionCode id="ED1EA0A19835FFF0072A7425D3DBFAD6" box="[341,424,1337,1361]" pageId="34" pageNumber="35">BMNH</collectionCode>
1896.11.1.6) collected by “S. Briceno” (= Salomón Briceño-Gabaldón, a commercial collector) at 
<quantity id="4CF795819835FFF007DD7467D380FA15" box="[418,499,1403,1427]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="34" pageNumber="35" unit="m" value="1630.0">1630 m</quantity>
elevation along the Río Albarregas (misspelled “Abbaregas” by 
<bibRefCitation id="EF9E45959835FFF006EF74A1D35FFA52" author="Thomas, O." box="[144,300,1469,1493]" pageId="34" pageNumber="35" pagination="301 - 314" refId="ref39327" refString="Thomas, O. 1896. On new small mammals from the Neotropical region. Annals and Magazine of Natural History (ser. 6) 18: 301 - 314." type="journal article" year="1896">Thomas, 1896</bibRefCitation>
) near the city of 
<collectingRegion id="49CBF6869835FFF0078C74A1D031FA52" box="[499,578,1469,1493]" country="Venezuela" name="Merida" pageId="34" pageNumber="35">Mérida</collectingRegion>
(fig. 17: locality 114) in the Venezuelan state of 
<collectingRegion id="49CBF6869835FFF0042E74C2D0D3FA71" box="[593,672,1502,1526]" country="Venezuela" name="Merida" pageId="34" pageNumber="35">Mérida</collectingRegion>
on 
<date id="FFB11EA49835FFF006CB74E3D34CF990" box="[180,319,1535,1559]" pageId="34" pageNumber="35" value="1896-04-06">6 April 1896</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3156BEF9835FFF006CF773CD6D5FDDA" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BB038649835FFF006CF773CD0D3F95A" blockId="34.[144,673,350,1757]" pageId="34" pageNumber="35">
DISTRIBUTION, HABITATS, AND SYMPATRY: Examined specimens of 
<taxonomicName id="4C0F43E79835FFF007E2775ED004F9DE" authorityName=": Gardner and Creighton" authorityYear="1989" box="[413,631,1601,1625]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769835FFF007E2775ED004F9DE" box="[413,631,1601,1625]" italics="true" pageId="34" pageNumber="35">Marmosops fuscatus</emphasis>
</taxonomicName>
are from four Venezuelan localities, two in the 
<collectingRegion id="49CBF6869835FFF006EF779FD2ACF91C" box="[144,223,1667,1691]" country="Venezuela" name="Merida" pageId="34" pageNumber="35">Mérida</collectingRegion>
Andes (Cordillera de 
<collectingRegion id="49CBF6869835FFF007B0779FD051F91C" box="[463,546,1667,1691]" country="Venezuela" name="Merida" pageId="34" pageNumber="35">Mérida</collectingRegion>
), one in the Serranía de San Luis of western 
<collectingRegion id="49CBF6869835FFF0078E77B8D04FF93B" box="[497,572,1700,1724]" country="Venezuela" name="Falcon" pageId="34" pageNumber="35">Falcón</collectingRegion>
, and one in the Caribbean coastal lowlands of eastern Fal-
</paragraph>
<paragraph id="8BB038649835FFF004AF71E6D6D5FDDA" blockId="34.[720,1248,250,1761]" pageId="34" pageNumber="35">
cón (fig. 17). Although recorded elevations range from 
<quantity id="4CF795819835FFF0056C7007D1B2FEB5" box="[787,961,283,307]" metricMagnitude="3" metricUnit="m" metricValue="1.2375" metricValueMax="2.35" metricValueMin="0.125" pageId="34" pageNumber="35" unit="m" value="1237.5" valueMax="2350.0" valueMin="125.0">125 to 2350 m</quantity>
above sea level, capture habitats seem to be either premontane or montane rain forest. 
<superScript id="7C7A952C9835FFF005037046D1F6FEEC" attach="left" box="[892,901,346,363]" fontSize="7" pageId="34" pageNumber="35">6</superScript>
Previous reports of this species from 
<collectingCountry id="F31878F49835FFF0056D7062D10CFE11" box="[786,895,382,406]" name="Colombia" pageId="34" pageNumber="35">Colombia</collectingCountry>
(e.g., 
<bibRefCitation id="EF9E45959835FFF005B37062D174FE30" author="Gardner, A. L. &amp; G. K. Creighton" pageId="34" pageNumber="35" pagination="61 - 74" refId="ref36627" refString="Gardner, A. L., and G. K. Creighton. 2008 (&quot; 2007 &quot;). Genus Marmosops Matschie, 1916. In A. L. Gardner (editor), Mammals of South America, vol. 1: marsupials, xenarthrans, shrews, and bats: 61 - 74. Chicago: University of Chicago Press." type="book chapter" year="2008">Gardner and Creighton, 2008</bibRefCitation>
: map 28) were based on misidentified material (see Remarks, below). 
<taxonomicName id="4C0F43E79835FFF0058F70DDD6BBFE5F" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1008,1224,448,472]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769835FFF0058F70DDD6BBFE5F" box="[1008,1224,448,472]" italics="true" pageId="34" pageNumber="35">Marmosops fuscatus</emphasis>
</taxonomicName>
is not known to occur sympatrically with any congeneric species, although it could be expected to do so with 
<taxonomicName id="4C0F43E79835FFF0052D7338D1DEFDBC" baseAuthorityName="Allen and Chapman" baseAuthorityYear="1897" box="[850,941,548,571]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="carri">
<emphasis id="B97BE4769835FFF0052D7338D1DEFDBC" box="[850,941,548,571]" italics="true" pageId="34" pageNumber="35">M. carri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0F43E79835FFF005967338D621FDBC" box="[1001,1106,548,571]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="ojastii">
<emphasis id="B97BE4769835FFF005967338D621FDBC" box="[1001,1106,548,571]" italics="true" pageId="34" pageNumber="35">M. ojastii</emphasis>
</taxonomicName>
, which have adjacent or overlapping geographic ranges.
</paragraph>
</subSubSection>
<subSubSection id="C3156BEF9835FFF6048F7379D3D8FEF2" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
<paragraph id="8BB038649835FFF0048F7379D603FB57" blockId="34.[720,1248,250,1761]" pageId="34" pageNumber="35">
DESCRIPTION: Body pelage dark grayish brown (near Sepia) middorsally, indistinctly paler laterally, and about 
<quantity id="4CF795819835FFF0059273B4D63FFD38" box="[1005,1100,680,704]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="34" pageNumber="35" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
long at midback (shorter in specimens from lower elevations than in those from the 
<collectingRegion id="49CBF6869835FFF005A473F6D658FC85" box="[987,1067,746,770]" country="Venezuela" name="Merida" pageId="34" pageNumber="35">Mérida</collectingRegion>
Andes); ventral pelage superficially whitish but uniformly gray based except for the apex of the chin, oral margins, and scrotum (which are covered with selfwhitish hairs). (One examined specimen, AMNH 276509, has small patches of self-white hairs along the inner thighs.) Manus covered dorsally with uniformly pale hairs (the metacarpals not contrasting in color with the digits); lateral carpal tubercles bladelike in all examined adult males. Mammary formula unknown. Tail longer than combined length of head and body (mean LT/HBL × 100 = 110%); dorsal caudal surface dark, but indistinctly paler distally in some specimens; ventral surface distinctly paler.
</paragraph>
<paragraph id="8BB038649835FFF0048F75C6D679F966" blockId="34.[720,1248,250,1761]" pageId="34" pageNumber="35">
Nasal bones long (extending well behind the lacrimals) and uniformly narrow (without distinct lateral expansion at the maxillary-frontal suture). Interorbital region broad, with rounded supraorbital margins in females and young adult males, but relatively narrower and tending to develop squared supraorbital margins in large adult males; postorbital processes absent or 
<superScript id="7C7A952C9835FFF004AF74E9D0A4F986" attach="left" box="[720,727,1525,1537]" fontSize="5" pageId="34" pageNumber="35">6</superScript>
<bibRefCitation id="EF9E45959835FFF004A074EAD1A0F98E" author="Handley, C. O., Jr. &amp; L. K Gordon" box="[735,979,1526,1545]" pageId="34" pageNumber="35" pagination="65 - 72" refId="ref36974" refString="Handley, C. O., Jr., and L. K Gordon. 1979. New species of mammals from northern South America: mouse opossums, genus Marmosa Gray. In J. F. Eisenberg (editor), Vertebrate ecology in the northern Neotropics: 65 - 72. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1979">Handley and Gordon (1979)</bibRefCitation>
reported that the 
<typeStatus id="54B486C69835FFF0021074EAD6E7F98E" box="[1135,1172,1526,1545]" pageId="34" pageNumber="35">type</typeStatus>
series of 
<taxonomicName id="4C0F43E79835FFF004AF7712D110F9A7" box="[720,867,1550,1568]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769835FFF004AF7712D110F9A7" box="[720,867,1550,1568]" italics="true" pageId="34" pageNumber="35">Marmosa cracens</emphasis>
</taxonomicName>
was collected between 125 and 
<quantity id="4CF795819835FFF0020B7712D6DAF9A6" box="[1140,1193,1550,1569]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="34" pageNumber="35" unit="m" value="170.0">170 m</quantity>
above sea level on “the steep, moist, north slope of an isolated, low mountain” near the Caribbean coast. Vegetation of distinctly montane character can extend downslope to within 
<quantity id="4CF795819835FFF002D8774AD6ACF9EE" box="[1191,1247,1622,1641]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="35" unit="m" value="100.0">100 m</quantity>
above sea level on such isolated peaks, which intercept moisture-laden sea breezes and are often shrouded in low-lying clouds. The cloud forest (montane rain forest) vegetation at capture localities in the Serranía de San Luis was briefly described by 
<bibRefCitation id="EF9E45959835FFF0053D77D2D675F966" author="Anderson, R. P. &amp; E. E. Gutierrez &amp; J. Ochoa-G. &amp; F. J. Garcia &amp; M. Aguilera" box="[834,1030,1742,1761]" pageId="34" pageNumber="35" pagination="157 - 170" refId="ref35112" refString="Anderson, R. P., E. E. Gutierrez, J. Ochoa-G., F. J. Garcia, and M. Aguilera. 2012. Faunal nestedness and species-area relationship for small non-volant mammals in &quot; sky islands &quot; of northern Venezuela. Studies on Neotropical Fauna and Environment 47: 157 - 170." type="journal article" year="2012">Anderson et al. (2012)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB038649834FFF106EF71E0D3A4FDD8" blockId="35.[144,673,252,1566]" pageId="35" pageNumber="36">
indistinct. Lacrimal foramina variable (the upper foramen is usually not visible in lateral view, but the lower foramen can be either laterally visible or concealed inside the anterior orbital margin); 
<superScript id="7C7A952C9834FFF104E87041D0D3FEEA" attach="none" box="[663,672,349,365]" fontSize="7" pageId="35" pageNumber="36">7</superScript>
zygomatic process of squamosal broadly overlapped dorsally by the jugal. Palatine fenestrae present or absent (when present these can appear as a single small perforation on each side or as several small irregular perforations on both sides). Dorsolateral margin of ethmoid foramen usually formed by the frontal.
</paragraph>
<paragraph id="8BB038649834FFF106CF7375D0F2FBAA" blockId="35.[144,673,252,1566]" pageId="35" pageNumber="36">
Upper canine (C1) sexually dimorphic (longer and unicuspid in males, shorter and with posterior accessory cusp in young female specimens with unworn teeth). Upper third molar (M3) anterolabial cingulum discontinuous with preprotocrista (anterior cingulum incomplete). Lower canine (c1) sexually dimorphic (erect, without a posterior accessory cusp, and taller than p 
<quantity id="4CF795819834FFF106A6726DD28CFC0E" box="[217,255,881,905]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="1.0">1 in</quantity>
males versus procumbent, with a posterior accessory cusp, and subequal in height to p 
<quantity id="4CF795819834FFF106DE72AFD2B8FC4C" box="[161,203,947,971]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="1.0">1 in</quantity>
females); c1 anterolingual accessory cusp usually absent in both sexes. Entoconid of m1 subequal in height to adjacent m2 paraconid; unworn m4 talonid with three distinct cusps.
</paragraph>
<paragraph id="8BB038649834FFF106CF752BD3F1FB16" blockId="35.[144,673,252,1566]" pageId="35" pageNumber="36">
COMPARISONS: Comparisons between 
<taxonomicName id="4C0F43E79834FFF104167524D34EFBF7" authorityName=": Gardner and Creighton" authorityYear="1989" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF104167524D34EFBF7" italics="true" pageId="35" pageNumber="36">Marmosops fuscatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0F43E79834FFF107087545D3A2FBF7" baseAuthorityName="Allen and Chapman" baseAuthorityYear="1897" box="[375,465,1113,1136]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="carri">
<emphasis id="B97BE4769834FFF107087545D3A2FBF7" box="[375,465,1113,1136]" italics="true" pageId="35" pageNumber="36">M. carri</emphasis>
</taxonomicName>
have already been described (see above).
</paragraph>
<paragraph id="8BB038649834FFF106CF7587D0D3F999" blockId="35.[144,673,252,1566]" pageId="35" pageNumber="36">
<taxonomicName id="4C0F43E79834FFF106CF7587D3E5FB35" authorityName=": Gardner and Creighton" authorityYear="1989" box="[176,406,1178,1202]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF106CF7587D3E5FB35" box="[176,406,1178,1202]" italics="true" pageId="35" pageNumber="36">Marmosops fuscatus</emphasis>
</taxonomicName>
is substantially larger than 
<taxonomicName id="4C0F43E79834FFF106B475A0D33CFB54" baseAuthorityName="Pine" baseAuthorityYear="1981" box="[203,335,1211,1235]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF106B475A0D33CFB54" box="[203,335,1211,1235]" italics="true" pageId="35" pageNumber="36">M. handleyi</emphasis>
</taxonomicName>
in head-and-body length and has a proportionately shorter tail (tables 5, 6). Externally the two species differ in the same characters that distinguish 
<taxonomicName id="4C0F43E79834FFF107AF7403D043FAB1" baseAuthorityName="Allen and Chapman" baseAuthorityYear="1897" box="[464,560,1311,1334]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="carri">
<emphasis id="B97BE4769834FFF107AF7403D043FAB1" box="[464,560,1311,1334]" italics="true" pageId="35" pageNumber="36">M. carri</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C0F43E79834FFF104FE7403D29FFAD0" baseAuthorityName="Pine" baseAuthorityYear="1981" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF104FE7403D29FFAD0" italics="true" pageId="35" pageNumber="36">M. handleyi</emphasis>
</taxonomicName>
, namely dorsal pelage coloration (dark grayish brown in 
<taxonomicName id="4C0F43E79834FFF1072B747DD3D9FAFE" authorityName=": Gardner and Creighton" authorityYear="1989" box="[340,426,1377,1401]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1072B747DD3D9FAFE" box="[340,426,1377,1401]" italics="true" pageId="35" pageNumber="36">fuscatus</emphasis>
</taxonomicName>
, dark brown in 
<taxonomicName id="4C0F43E79834FFF1041F747DD2C4FA1D" baseAuthorityName="Pine" baseAuthorityYear="1981" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF1041F747DD2C4FA1D" italics="true" pageId="35" pageNumber="36">handleyi</emphasis>
</taxonomicName>
) and forefoot coloration (the manus is entirely whitish in 
<taxonomicName id="4C0F43E79834FFF1072674BFD3DCFA3C" authorityName=": Gardner and Creighton" authorityYear="1989" box="[345,431,1443,1467]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1072674BFD3DCFA3C" box="[345,431,1443,1467]" italics="true" pageId="35" pageNumber="36">fuscatus</emphasis>
</taxonomicName>
, whereas the pale digits contrast with the dark metacarpals in 
<taxonomicName id="4C0F43E79834FFF1041F74D8D2C5FA7A" baseAuthorityName="Pine" baseAuthorityYear="1981" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF1041F74D8D2C5FA7A" italics="true" pageId="35" pageNumber="36">handleyi</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C0F43E79834FFF106AD74FAD3C7FA7A" authorityName=": Gardner and Creighton" authorityYear="1989" box="[210,436,1509,1533]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF106AD74FAD3C7FA7A" box="[210,436,1509,1533]" italics="true" pageId="35" pageNumber="36">Marmosops fuscatus</emphasis>
</taxonomicName>
also averages larger than 
<taxonomicName id="4C0F43E79834FFF106B3771BD322F999" baseAuthorityName="Pine" baseAuthorityYear="1981" box="[204,337,1542,1566]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF106B3771BD322F999" box="[204,337,1542,1566]" italics="true" pageId="35" pageNumber="36">M. handleyi</emphasis>
</taxonomicName>
in most craniodental dimen-
</paragraph>
<paragraph id="8BB038649834FFF106EF7721D352F966" blockId="35.[144,672,1597,1761]" pageId="35" pageNumber="36">
<superScript id="7C7A952C9834FFF106EF7721D2E4F9CE" attach="left" box="[144,151,1597,1609]" fontSize="5" pageId="35" pageNumber="36">7</superScript>
The lower lacrimal foramen is laterally visible in some specimens (e.g., BMNH 3.1.5.2, 3.1.5.3, FMNH 22174, USNM 442719) but not in others (e.g., AMNH 276580, 276509). The specimens in which this foramen is visible are the largest in overall size and have the most heavily ossified skulls, but we lack sufficient material to determine whether or not this trait is age correlated.
</paragraph>
<paragraph id="8BB038649834FFF104AF71E6D60FFCE3" blockId="35.[720,1249,250,1762]" pageId="35" pageNumber="36">
sions, including four measurements (CBL, ZB, PL, MTR, and WM3) that exhibit nonoverlapping variation in same-sex comparisons. These species also differ in some of the same qualitative craniodental characters that distinguish 
<taxonomicName id="4C0F43E79834FFF102F77063D693FE11" baseAuthorityName="Allen and Chapman" baseAuthorityYear="1897" box="[1160,1248,383,406]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="carri">
<emphasis id="B97BE4769834FFF102F77063D693FE11" box="[1160,1248,383,406]" italics="true" pageId="35" pageNumber="36">M. carri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0F43E79834FFF1057C70BCD1FBFE30" baseAuthorityName="Pine" baseAuthorityYear="1981" box="[771,904,415,439]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF1057C70BCD1FBFE30" box="[771,904,415,439]" italics="true" pageId="35" pageNumber="36">M. handleyi</emphasis>
</taxonomicName>
; in particular, 
<taxonomicName id="4C0F43E79834FFF1024C70BCD6C1FE30" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1075,1202,415,439]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1024C70BCD6C1FE30" box="[1075,1202,415,439]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
has much narrower nasals than 
<taxonomicName id="4C0F43E79834FFF1025A70DDD6C2FE5F" baseAuthorityName="Pine" baseAuthorityYear="1981" box="[1061,1201,448,472]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF1025A70DDD6C2FE5F" box="[1061,1201,448,472]" italics="true" pageId="35" pageNumber="36">M. handleyi</emphasis>
</taxonomicName>
(in which these bones are conspicuously expanded laterally at the maxillary-frontal suture), and males of 
<taxonomicName id="4C0F43E79834FFF1054A7338D1C1FDBC" authorityName=": Gardner and Creighton" authorityYear="1989" box="[821,946,547,571]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1054A7338D1C1FDBC" box="[821,946,547,571]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
have unicuspid C1 and c1, whereas these teeth consistently have a small posterior accessory cusp in males of 
<taxonomicName id="4C0F43E79834FFF10225737AD6AAFDFA" baseAuthorityName="Pine" baseAuthorityYear="1981" box="[1114,1241,613,637]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF10225737AD6AAFDFA" box="[1114,1241,613,637]" italics="true" pageId="35" pageNumber="36">M. handleyi</emphasis>
</taxonomicName>
. Additionally, the dorsolateral margin of the ethmoid foramen is usually formed by the frontal in 
<taxonomicName id="4C0F43E79834FFF104AF73D5D13EFD66" authorityName=": Gardner and Creighton" authorityYear="1989" box="[720,845,713,737]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF104AF73D5D13EFD66" box="[720,845,713,737]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
, whereas the dorsolateral margin of this foramen is formed by the orbitosphenoid in 
<taxonomicName id="4C0F43E79834FFF104AF7217D123FCA4" baseAuthorityName="Pine" baseAuthorityYear="1981" box="[720,848,779,803]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF104AF7217D123FCA4" box="[720,848,779,803]" italics="true" pageId="35" pageNumber="36">M. handleyi</emphasis>
</taxonomicName>
. Lastly, the distinct lingual accessory cusp of c1 that appears to be consistently present in 
<taxonomicName id="4C0F43E79834FFF104917252D11CFCE2" baseAuthorityName="Pine" baseAuthorityYear="1981" box="[750,879,845,869]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="handleyi">
<emphasis id="B97BE4769834FFF104917252D11CFCE2" box="[750,879,845,869]" italics="true" pageId="35" pageNumber="36">M. handleyi</emphasis>
</taxonomicName>
is absent in 
<taxonomicName id="4C0F43E79834FFF105857252D605FCE2" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1018,1142,845,869]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF105857252D605FCE2" box="[1018,1142,845,869]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB038649834FFF6048F7273D3D8FEF2" blockId="35.[720,1249,250,1762]" lastBlockId="36.[144,673,250,1497]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<taxonomicName id="4C0F43E79834FFF1048F7273D1A5FC01" authorityName=": Gardner and Creighton" authorityYear="1989" box="[752,982,878,902]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1048F7273D1A5FC01" box="[752,982,878,902]" italics="true" pageId="35" pageNumber="36">Marmosops fuscatus</emphasis>
</taxonomicName>
is substantially larger than 
<taxonomicName id="4C0F43E79834FFF10573728CD1FBFC20" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[780,904,912,935]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF10573728CD1FBFC20" box="[780,904,912,935]" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
in all external dimensions. In qualitative external characters, 
<taxonomicName id="4C0F43E79834FFF1024172ADD6B1FC4F" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1086,1218,944,968]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1024172ADD6B1FC4F" box="[1086,1218,944,968]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
is most readily distinguished from 
<taxonomicName id="4C0F43E79834FFF1023E72CED6CEFC6E" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[1089,1213,978,1001]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF1023E72CED6CEFC6E" box="[1089,1213,978,1001]" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
by its paler dorsal coloration (the dorsal pelage of 
<taxonomicName id="4C0F43E79834FFF104AF7508D155FBAC" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[720,806,1044,1067]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF104AF7508D155FBAC" box="[720,806,1044,1067]" italics="true" pageId="35" pageNumber="36">invictus</emphasis>
</taxonomicName>
is consistently much darker), by its entirely pale forefeet (the metacarpals are dark in 
<taxonomicName id="4C0F43E79834FFF104AF754AD13BFBEA" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[720,840,1110,1133]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF104AF754AD13BFBEA" box="[720,840,1110,1133]" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
), and by its more distinctly bicolored tail (the tail is only indistinctly paler ventrally than dorsally in 
<taxonomicName id="4C0F43E79834FFF105017584D187FB28" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[894,1012,1176,1199]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF105017584D187FB28" box="[894,1012,1176,1199]" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C0F43E79834FFF102757584D693FB37" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1034,1248,1176,1200]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF102757584D693FB37" box="[1034,1248,1176,1200]" italics="true" pageId="35" pageNumber="36">Marmosops fuscatus</emphasis>
</taxonomicName>
averages larger than 
<taxonomicName id="4C0F43E79834FFF105CC75A5D65EFB57" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[947,1069,1209,1232]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF105CC75A5D65EFB57" box="[947,1069,1209,1232]" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
in most craniodental dimensions, including five measurements (CBL, NL, ZB, PL, MTR) that exhibit nonoverlapping variation in same-sex comparisons. Craniodental qualitative differences between these quite dissimilar species include the shape of the nasals (uniformly narrow in 
<taxonomicName id="4C0F43E79834FFF1027B749CD60DFA10" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1028,1150,1407,1431]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1027B749CD60DFA10" box="[1028,1150,1407,1431]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
, laterally expanded in 
<taxonomicName id="4C0F43E79834FFF1052174BDD1A5FA3F" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[862,982,1441,1464]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF1052174BDD1A5FA3F" box="[862,982,1441,1464]" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
), dorsolateral margin of the ethmoid foramen (usually formed by the frontal in 
<taxonomicName id="4C0F43E79834FFF1053E74FFD1CDFA7D" authorityName=": Gardner and Creighton" authorityYear="1989" box="[833,958,1506,1530]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1053E74FFD1CDFA7D" box="[833,958,1506,1530]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
, by the orbitosphenoid in 
<taxonomicName id="4C0F43E79834FFF104AF7718D134F99C" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[720,839,1540,1563]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF104AF7718D134F99C" box="[720,839,1540,1563]" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
), C1 posterior accessory cusp (absent in males of 
<taxonomicName id="4C0F43E79834FFF1052E7739D1BEF9BB" authorityName=": Gardner and Creighton" authorityYear="1989" box="[849,973,1572,1596]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1052E7739D1BEF9BB" box="[849,973,1572,1596]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
, present in both sexes of 
<taxonomicName id="4C0F43E79834FFF104AF775AD13AF9DA" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[720,841,1606,1629]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF104AF775AD13AF9DA" box="[720,841,1606,1629]" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
), M3 anterior cingulum (incomplete in 
<taxonomicName id="4C0F43E79834FFF1048F777BD11CF9F8" authorityName=": Gardner and Creighton" authorityYear="1989" box="[752,879,1639,1663]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1048F777BD11CF9F8" box="[752,879,1639,1663]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
, narrowly complete in 
<taxonomicName id="4C0F43E79834FFF10204777BD082F918" baseAuthorityName="Goldman" baseAuthorityYear="1912" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF10204777BD082F918" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
), c1 morphology (erect and unicuspid in male 
<taxonomicName id="4C0F43E79834FFF1056F77B5D1E2F946" authorityName=": Gardner and Creighton" authorityYear="1989" box="[784,913,1705,1729]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769834FFF1056F77B5D1E2F946" box="[784,913,1705,1729]" italics="true" pageId="35" pageNumber="36">M. fuscatus</emphasis>
</taxonomicName>
, procumbent and premolariform in 
<taxonomicName id="4C0F43E79834FFF1055077D6D1D8F966" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[815,939,1738,1761]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769834FFF1055077D6D1D8F966" box="[815,939,1738,1761]" italics="true" pageId="35" pageNumber="36">M. invictus</emphasis>
</taxonomicName>
), c1 lingual accessory cusp (absent in 
<taxonomicName id="4C0F43E79833FFF6076F71E6D3E0FE95" authorityName=": Gardner and Creighton" authorityYear="1989" box="[272,403,250,274]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769833FFF6076F71E6D3E0FE95" box="[272,403,250,274]" italics="true" pageId="36" pageNumber="37">M. fuscatus</emphasis>
</taxonomicName>
, usually present in 
<taxonomicName id="4C0F43E79833FFF604FD71E6D297FEB5" baseAuthorityName="Goldman" baseAuthorityYear="1912" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769833FFF604FD71E6D297FEB5" italics="true" pageId="36" pageNumber="37">M. invictus</emphasis>
</taxonomicName>
), and m1 entoconid (subequal to adjacent m2 paraconid in 
<taxonomicName id="4C0F43E79833FFF607F87020D3AEFED3" authorityName=": Gardner and Creighton" authorityYear="1989" box="[391,477,316,340]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769833FFF607F87020D3AEFED3" box="[391,477,316,340]" italics="true" pageId="36" pageNumber="37">fuscatus</emphasis>
</taxonomicName>
, shorter than m2 paraconid in 
<taxonomicName id="4C0F43E79833FFF6075C7042D3E8FEF2" baseAuthorityName="Goldman" baseAuthorityYear="1912" box="[291,411,350,373]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="invictus">
<emphasis id="B97BE4769833FFF6075C7042D3E8FEF2" box="[291,411,350,373]" italics="true" pageId="36" pageNumber="37">M. invictus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3156BEF9833FFF606CF7062D639FB76" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8BB038649833FFF606CF7062D3EEFBCB" blockId="36.[144,673,250,1497]" pageId="36" pageNumber="37">
REMARKS: 
<taxonomicName id="4C0F43E79833FFF607587063D39CFE11" authorityName="Thomas" authorityYear="1924" box="[295,495,382,406]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="perfusca">
<emphasis id="B97BE4769833FFF607587063D39CFE11" box="[295,495,382,406]" italics="true" pageId="36" pageNumber="37">Marmosa perfusca</emphasis>
</taxonomicName>
, a nominal species that 
<bibRefCitation id="EF9E45959833FFF606907083D3E0FE30" author="Thomas, O." box="[239,403,415,439]" pageId="36" pageNumber="37" pagination="234 - 237" refId="ref39388" refString="Thomas, O. 1924. New South American mammals. Annals and Magazine of Natural History (ser. 9) 13: 234 - 237." type="journal article" year="1924">Thomas (1924)</bibRefCitation>
described from the Cordillera Oriental of 
<collectingCountry id="F31878F49833FFF6071D70DCD3A3FE5F" box="[354,464,448,472]" name="Colombia" pageId="36" pageNumber="37">Colombia</collectingCountry>
, was synonymized with 
<taxonomicName id="4C0F43E79833FFF606B670FED334FE7E" authorityName=": Gardner and Creighton" authorityYear="1989" box="[201,327,481,505]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769833FFF606B670FED334FE7E" box="[201,327,481,505]" italics="true" pageId="36" pageNumber="37">M. fuscatus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF9E45959833FFF6070D70FED054FE7E" author="Tate, G. H. H." box="[370,551,481,505]" pageId="36" pageNumber="37" pagination="1 - 250" refId="ref39277" refString="Tate, G. H. H. 1933. A systematic revision of the marsupial genus Marmosa with a discussion of the adaptive radiation of the murine opossums (Marmosa). Bulletin of the American Museum of Natural History 66: 1 - 250 + 26 pls." type="book chapter" year="1933">Tate (1933: 173)</bibRefCitation>
, and some authors (e.g., 
<bibRefCitation id="EF9E45959833FFF60758731ED0EBFD9D" author="Gardner, A. L. &amp; G. K. Creighton" box="[295,664,514,538]" pageId="36" pageNumber="37" pagination="61 - 74" refId="ref36627" refString="Gardner, A. L., and G. K. Creighton. 2008 (&quot; 2007 &quot;). Genus Marmosops Matschie, 1916. In A. L. Gardner (editor), Mammals of South America, vol. 1: marsupials, xenarthrans, shrews, and bats: 61 - 74. Chicago: University of Chicago Press." type="book chapter" year="2008">Gardner and Creighton, 2008: 66</bibRefCitation>
) have treated it as a valid subspecies (“ 
<taxonomicName id="4C0F43E79833FFF6045B7338D32BFDDB" baseAuthorityName="Thomas" baseAuthorityYear="1924" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="subSpecies" species="fuscatus" subSpecies="perfuscus">
<emphasis id="B97BE4769833FFF6045B7338D32BFDDB" italics="true" pageId="36" pageNumber="37">Marmosops fuscatus perfuscus</emphasis>
</taxonomicName>
”). However, the 
<typeStatus id="54B486C69833FFF604647359D039FDDA" box="[539,586,581,605]" pageId="36" pageNumber="37">type</typeStatus>
of 
<taxonomicName id="4C0F43E79833FFF6040A7359D2BBFDFA" authorityName="Thomas" authorityYear="1924" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="perfusca">
<emphasis id="B97BE4769833FFF6040A7359D2BBFDFA" italics="true" pageId="36" pageNumber="37">perfusca</emphasis>
</taxonomicName>
(BMNH 1923.11.13.18, consisting of the skin and skull of an old adult female) has posteriorly widened nasals, a short subsquamosal foramen (not exposing the petrosal behind the sulcus of the prootic sinus), and only one antebrachial vibrissa. In these and other characters it closely resembles 
<taxonomicName id="4C0F43E79833FFF607237230D2A3FCE2" authority="(Thomas, 1900)" baseAuthorityName="Thomas" baseAuthorityYear="1900" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="caucae">
<emphasis id="B97BE4769833FFF607237230D05DFCC4" box="[348,558,812,835]" italics="true" pageId="36" pageNumber="37">Marmosops caucae</emphasis>
(
<bibRefCitation id="EF9E45959833FFF6043F7230D2BBFCE2" author="Thomas, O." pageId="36" pageNumber="37" pagination="217 - 222" refId="ref39359" refString="Thomas, O. 1900. Descriptions of new Neotropical mammals. Annals and Magazine of Natural History (ser. 7) 5: 217 - 222." type="journal article" year="1900">Thomas, 1900</bibRefCitation>
)
</taxonomicName>
, a member of the nominotypical subgenus of 
<taxonomicName id="4C0F43E79833FFF606CF7273D35EFC01" authorityName="Matschie" authorityYear="1916" box="[176,301,879,902]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B97BE4769833FFF606CF7273D35EFC01" box="[176,301,879,902]" italics="true" pageId="36" pageNumber="37">Marmosops</emphasis>
</taxonomicName>
. A second Colombian specimen that 
<bibRefCitation id="EF9E45959833FFF606BB7293D331FC20" author="Tate, G. H. H." box="[196,322,911,935]" pageId="36" pageNumber="37" pagination="1 - 250" refId="ref39277" refString="Tate, G. H. H. 1933. A systematic revision of the marsupial genus Marmosa with a discussion of the adaptive radiation of the murine opossums (Marmosa). Bulletin of the American Museum of Natural History 66: 1 - 250 + 26 pls." type="book chapter" year="1933">Tate (1933)</bibRefCitation>
referred to 
<taxonomicName id="4C0F43E79833FFF607B5728CD0F5FC20" box="[458,646,911,935]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscata">
<emphasis id="B97BE4769833FFF607B5728CD0F5FC20" box="[458,646,911,935]" italics="true" pageId="36" pageNumber="37">Marmosa fuscata</emphasis>
</taxonomicName>
is AMNH 34641, also from the Cordillera Oriental; 
<superScript id="7C7A952C9833FFF604E872B1D0D3FC39" attach="none" box="[663,672,941,958]" fontSize="7" pageId="36" pageNumber="37">8</superScript>
like BMNH 1923.11.13.18, this specimen appears to represent 
<taxonomicName id="4C0F43E79833FFF6076872EFD3F1FB8D" baseAuthorityName="Thomas" baseAuthorityYear="1900" box="[279,386,1011,1034]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="caucae">
<emphasis id="B97BE4769833FFF6076872EFD3F1FB8D" box="[279,386,1011,1034]" italics="true" pageId="36" pageNumber="37">M. caucae</emphasis>
</taxonomicName>
. To the best of our knowledge, there are no valid records of 
<taxonomicName id="4C0F43E79833FFF6045C7508D295FBCB" authorityName=": Gardner and Creighton" authorityYear="1989" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769833FFF6045C7508D295FBCB" italics="true" pageId="36" pageNumber="37">Marmosops fuscatus</emphasis>
</taxonomicName>
from 
<collectingCountry id="F31878F49833FFF607547528D3EAFBCB" box="[299,409,1076,1100]" name="Colombia" pageId="36" pageNumber="37">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="8BB038649833FFF606CF7549D0D3FA5E" blockId="36.[144,673,250,1497]" pageId="36" pageNumber="37">
<bibRefCitation id="EF9E45959833FFF606CF7549D05BFBEA" author="Handley, C. O., Jr. &amp; L. K Gordon" box="[176,552,1109,1133]" pageId="36" pageNumber="37" pagination="65 - 72" refId="ref36974" refString="Handley, C. O., Jr., and L. K Gordon. 1979. New species of mammals from northern South America: mouse opossums, genus Marmosa Gray. In J. F. Eisenberg (editor), Vertebrate ecology in the northern Neotropics: 65 - 72. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1979">Handley and Gordon (1979: 67)</bibRefCitation>
described 
<taxonomicName id="4C0F43E79833FFF606EF756BD33EFB09" box="[144,333,1143,1166]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769833FFF606EF756BD33EFB09" box="[144,333,1143,1166]" italics="true" pageId="36" pageNumber="37">Marmosa cracens</emphasis>
</taxonomicName>
as “in almost every respect of pelage and skull character ... a miniature replica of 
<taxonomicName id="4C0F43E79833FFF606D275A5D36CFB56" box="[173,287,1209,1233]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscata">
<emphasis id="B97BE4769833FFF606D275A5D36CFB56" box="[173,287,1209,1233]" italics="true" pageId="36" pageNumber="37">M. fuscata</emphasis>
</taxonomicName>
, distinguishable from it only by its small size and the short, broad shape of its palate.” Indeed, the male 
<typeStatus id="54B486C69833FFF6070675E7D3A4FA94" box="[377,471,1275,1299]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C0F43E79833FFF6078775E7D036FA95" box="[504,581,1275,1298]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769833FFF6078775E7D036FA95" box="[504,581,1275,1298]" italics="true" pageId="36" pageNumber="37">cracens</emphasis>
</taxonomicName>
(USNM 418503) is substantially smaller than any adult male specimen of 
<taxonomicName id="4C0F43E79833FFF6071C7422D032FAD2" authorityName=": Gardner and Creighton" authorityYear="1989" box="[355,577,1341,1365]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769833FFF6071C7422D032FAD2" box="[355,577,1341,1365]" italics="true" pageId="36" pageNumber="37">Marmosops fuscatus</emphasis>
</taxonomicName>
that we measured. For example, the condylobasal length of the skull of USNM 418503 is 28.4 mm, whereas the range of condylobasal lengths of five adult male 
<taxonomicName id="4C0F43E79833FFF6076F74DED3FDFA5E" authorityName=": Gardner and Creighton" authorityYear="1989" box="[272,398,1473,1497]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769833FFF6076F74DED3FDFA5E" box="[272,398,1473,1497]" italics="true" pageId="36" pageNumber="37">M. fuscatus</emphasis>
</taxonomicName>
is 31.8–35.4 mm. How-
</paragraph>
<paragraph id="8BB038649833FFF606EF74E9D0F7F966" blockId="36.[144,674,1525,1761]" pageId="36" pageNumber="37">
<superScript id="7C7A952C9833FFF606EF74E9D2E4F986" attach="left" box="[144,151,1525,1537]" fontSize="5" pageId="36" pageNumber="37">8</superScript>
This specimen was collected by G. O’Connell on 
<date id="FFB11EA49833FFF6044774EAD0D2F98E" box="[568,673,1526,1545]" pageId="36" pageNumber="37" value="1913-04-04">4 April 1913</date>
at “El Roble,” a locality that 
<bibRefCitation id="EF9E45959833FFF607087712D38EF9A6" author="Tate, G. H. H." box="[375,509,1550,1569]" pageId="36" pageNumber="37" pagination="1 - 250" refId="ref39277" refString="Tate, G. H. H. 1933. A systematic revision of the marsupial genus Marmosa with a discussion of the adaptive radiation of the murine opossums (Marmosa). Bulletin of the American Museum of Natural History 66: 1 - 250 + 26 pls." type="book chapter" year="1933">Tate (1933: 173)</bibRefCitation>
erroneously placed in Departamento 
<collectingRegion id="49CBF6869833FFF6075C773AD324F9BE" box="[291,343,1574,1593]" country="Colombia" name="Cauca" pageId="36" pageNumber="37">Cauca</collectingRegion>
and that 
<bibRefCitation id="EF9E45959833FFF607DA773AD2DFF9D6" author="Gardner, A. L. &amp; G. K. Creighton" pageId="36" pageNumber="37" pagination="61 - 74" refId="ref36627" refString="Gardner, A. L., and G. K. Creighton. 2008 (&quot; 2007 &quot;). Genus Marmosops Matschie, 1916. In A. L. Gardner (editor), Mammals of South America, vol. 1: marsupials, xenarthrans, shrews, and bats: 61 - 74. Chicago: University of Chicago Press." type="book chapter" year="2008">Gardner and Creighton (2008: 66)</bibRefCitation>
located in Departamento 
<collectingRegion id="49CBF6869833FFF607F77722D3BCF9D6" box="[392,463,1598,1617]" country="Colombia" name="Quindio" pageId="36" pageNumber="37">Quindío</collectingRegion>
(on the western slope of the Cordillera Central; 
<bibRefCitation id="EF9E45959833FFF60716774AD396F9EE" author="Paynter, R. A., Jr." box="[361,485,1622,1641]" pageId="36" pageNumber="37" refId="ref38517" refString="Paynter, R. A., Jr. 1997. Ornithological gazetteer of Colombia, 2 nd ed. Cambridge, MA: Museum of Comparative Zoology (Harvard University)." type="book" year="1997">Paynter, 1997</bibRefCitation>
). However, Geoffroy O’Connell was a member of the 1913 AMNH expedition that worked at a different place called El Roble, “a posada on the trail from 
<collectingRegion id="49CBF6869833FFF606BD7782D28DF936" box="[194,254,1694,1713]" country="Colombia" name="Bogota" pageId="36" pageNumber="37">Bogotá</collectingRegion>
to Fusugasugá [= Fusagasugá]” on the western slope of the Cordillera Oriental (
<bibRefCitation id="EF9E45959833FFF607DD77AAD021F94E" author="Chapman, F. M." box="[418,594,1718,1737]" pageId="36" pageNumber="37" pagination="1 - 729" refId="ref35772" refString="Chapman, F. M. 1917. The distribution of birdlife in Colombia; a contribution to a biological survey of South America. Bulletin of the American Museum of Natural History 36: 1 - 729." type="journal article" year="1917">Chapman, 1917: 645</bibRefCitation>
); according to 
<bibRefCitation id="EF9E45959833FFF606B877D2D337F966" author="Paynter, R. A., Jr." box="[199,324,1742,1761]" pageId="36" pageNumber="37" refId="ref38517" refString="Paynter, R. A., Jr. 1997. Ornithological gazetteer of Colombia, 2 nd ed. Cambridge, MA: Museum of Comparative Zoology (Harvard University)." type="book" year="1997">Paynter (1997)</bibRefCitation>
, this locality is near 
<geoCoordinate id="EE3B5EA39833FFF6078F77D2D05EF966" box="[496,557,1741,1761]" degrees="4" direction="north" minutes="23" orientation="latitude" pageId="36" pageNumber="37" precision="925" value="4.383333">4°23′N</geoCoordinate>
, 
<geoCoordinate id="EE3B5EA39833FFF6044977D2D0F3F966" box="[566,640,1741,1761]" degrees="74" direction="west" minutes="19" orientation="longitude" pageId="36" pageNumber="37" precision="925" value="-74.316666">74°19′W</geoCoordinate>
.
</paragraph>
<paragraph id="8BB038649833FFF604AF71E6D6ACFBAB" blockId="36.[720,1249,250,1266]" pageId="36" pageNumber="37">
ever, this size difference is plausibly explained by ontogeny: the type series of 
<taxonomicName id="4C0F43E79833FFF6026C7007D612FEB5" box="[1043,1121,283,306]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769833FFF6026C7007D612FEB5" box="[1043,1121,283,306]" italics="true" pageId="36" pageNumber="37">cracens</emphasis>
</taxonomicName>
consists of subadults (with incompletely erupted P3s), not young adults (contra 
<bibRefCitation id="EF9E45959833FFF605A57041D17AFE11" author="Handley, C. O., Jr. &amp; L. K Gordon" pageId="36" pageNumber="37" pagination="65 - 72" refId="ref36974" refString="Handley, C. O., Jr., and L. K Gordon. 1979. New species of mammals from northern South America: mouse opossums, genus Marmosa Gray. In J. F. Eisenberg (editor), Vertebrate ecology in the northern Neotropics: 65 - 72. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1979">Handley and Gordon, 1979</bibRefCitation>
). The subadult 
<typeStatus id="54B486C69833FFF605C17062D66CFE11" box="[958,1055,382,406]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54B486C69833FFF602207062D6CCFE11" box="[1119,1215,382,406]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C0F43E79833FFF604AF70BCD16DFE30" box="[720,798,416,439]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769833FFF604AF70BCD16DFE30" box="[720,798,416,439]" italics="true" pageId="36" pageNumber="37">cracens</emphasis>
</taxonomicName>
do have relatively wider palates (PB/PL = 0.57–0.59) than topotypical adult 
<taxonomicName id="4C0F43E79833FFF6024070DCD6E6FE5F" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1087,1173,448,472]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769833FFF6024070DCD6E6FE5F" box="[1087,1173,448,472]" italics="true" pageId="36" pageNumber="37">fuscatus</emphasis>
</taxonomicName>
(0.51– 0.53), but relative width of the palate is another ontogenetically variable trait because didelphid palates become proportionately longer and narrower with advancing age (
<bibRefCitation id="EF9E45959833FFF605817358D6A6FDDB" author="Abdala, F. &amp; D. A. Flores &amp; N. P. Giannini" box="[1022,1237,580,604]" pageId="36" pageNumber="37" pagination="190 - 200" refId="ref34829" refString="Abdala, F., D. A. Flores, and N. P. Giannini. 2001. Postweaning ontogeny of the skull of Didelphis albiventris. Journal of Mammalogy 82: 190 - 200." type="journal article" year="2001">Abdala et al., 2001</bibRefCitation>
). Lastly, we (
<bibRefCitation id="EF9E45959833FFF6052E7379D615FDFA" author="Diaz-Nieto, J. F. &amp; S. A. Jansa &amp; R. S. Voss" box="[849,1126,613,637]" pageId="36" pageNumber="37" pagination="914 - 940" refId="ref36051" refString="Diaz-Nieto, J. F., S. A. Jansa, and R. S. Voss. 2016 a. DNA sequencing reveals unexpected Recent diversity and an ancient dichotomy in the American marsupial genus Marmosops (Didelphidae: Thylamyini). Zoological Journal of the Linnean Society 176: 914 - 940." type="journal article" year="2016">Díaz-Nieto et al., 2016a</bibRefCitation>
) amplified partial (579–619 bp) cytochrome- 
<emphasis id="B97BE4769833FFF60227739AD616FD19" box="[1112,1125,646,670]" italics="true" pageId="36" pageNumber="37">b</emphasis>
sequences from the 
<typeStatus id="54B486C69833FFF6053D73BBD1D0FD38" box="[834,931,679,703]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
and a 
<typeStatus id="54B486C69833FFF6058373B4D62FFD47" box="[1020,1116,680,704]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C0F43E79833FFF602F573B4D6AAFD38" box="[1162,1241,680,703]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769833FFF602F573B4D6AAFD38" box="[1162,1241,680,703]" italics="true" pageId="36" pageNumber="37">cracens</emphasis>
</taxonomicName>
, which differ by only 2.2% (uncorrected) from a partial (1008 bp) cytochrome- 
<emphasis id="B97BE4769833FFF6026973F6D650FC85" box="[1046,1059,746,770]" italics="true" pageId="36" pageNumber="37">b</emphasis>
sequence that we amplified from a topotype of 
<taxonomicName id="4C0F43E79833FFF602787217D6F3FCA4" authorityName=": Gardner and Creighton" authorityYear="1989" box="[1031,1152,779,803]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769833FFF602787217D6F3FCA4" box="[1031,1152,779,803]" italics="true" pageId="36" pageNumber="37">M. fuscatus</emphasis>
</taxonomicName>
(BMNH 1903.1.5.2). Because this genetic distance is well within the range of values commonly obtained from comparisons of conspecific sequences of other mammals, and because the phenotypic differences that 
<bibRefCitation id="EF9E45959833FFF6052172ACD6FCFC4F" author="Handley, C. O., Jr. &amp; L. K Gordon" box="[862,1167,944,968]" pageId="36" pageNumber="37" pagination="65 - 72" refId="ref36974" refString="Handley, C. O., Jr., and L. K Gordon. 1979. New species of mammals from northern South America: mouse opossums, genus Marmosa Gray. In J. F. Eisenberg (editor), Vertebrate ecology in the northern Neotropics: 65 - 72. Washington, DC: Smithsonian Institution Press." type="book chapter" year="1979">Handley and Gordon (1979)</bibRefCitation>
alleged between 
<taxonomicName id="4C0F43E79833FFF6054F72CED10EFC6E" box="[816,893,978,1001]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769833FFF6054F72CED10EFC6E" box="[816,893,978,1001]" italics="true" pageId="36" pageNumber="37">cracens</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0F43E79833FFF605CD72CDD67BFC6E" authorityName=": Gardner and Creighton" authorityYear="1989" box="[946,1032,977,1001]" class="Mammalia" family="Didelphidae" genus="Marmosops" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="fuscatus">
<emphasis id="B97BE4769833FFF605CD72CDD67BFC6E" box="[946,1032,977,1001]" italics="true" pageId="36" pageNumber="37">fuscatus</emphasis>
</taxonomicName>
seem to be attributable to the immaturity of their type series, we treat the former as a subjective junior synonym.
</paragraph>
<paragraph id="8BB038649833FFF6048F7528D639FB76" blockId="36.[720,1249,250,1266]" pageId="36" pageNumber="37">
<materialsCitation id="3B6732399833FFF6048F7528D184FB28" ID-GBIF-Occurrence="3059923310" collectionCode="USNM, AMNH" country="Venezuela" location="Serrania de San Luis" municipality="Near La Pastora" pageId="36" pageNumber="37" specimenCode="USNM 418503, AMNH 276509, 276580" specimenCount="1" stateProvince="Falcon" typeStatus="holotype">
SPECIMENS EXAMINED (
<emphasis id="B97BE4769833FFF605877529D678FBCB" box="[1016,1035,1077,1100]" italics="true" pageId="36" pageNumber="37">N</emphasis>
= 8): 
<emphasis id="B97BE4769833FFF6022C7528D6B4FBCB" bold="true" box="[1107,1223,1076,1100]" pageId="36" pageNumber="37">
<collectingCountry id="F31878F49833FFF6022C7528D6B4FBCB" box="[1107,1223,1076,1100]" name="Venezuela" pageId="36" pageNumber="37">Venezuela</collectingCountry>
</emphasis>
— 
<emphasis id="B97BE4769833FFF604AF7549D16EFBEA" box="[720,797,1109,1133]" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="49CBF6869833FFF604AF7549D16AFBEA" box="[720,793,1109,1133]" country="Venezuela" name="Falcon" pageId="36" pageNumber="37">Falcón</collectingRegion>
,
</emphasis>
<collectingMunicipality id="6BD4A21E9833FFF60558754AD1A8FBEA" box="[807,987,1110,1133]" pageId="36" pageNumber="37">Near La Pastora</collectingMunicipality>
(
<specimenCode id="DBA9901F9833FFF605947549D6E0FBEA" box="[1003,1171,1109,1133]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="36" pageNumber="37" type="Museum">USNM 418503</specimenCode>
[
<typeStatus id="54B486C69833FFF602DD7549D08CFB08" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C0F43E79833FFF60557756BD105FB09" box="[808,886,1143,1166]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="cracens">
<emphasis id="B97BE4769833FFF60557756BD105FB09" box="[808,886,1143,1166]" italics="true" pageId="36" pageNumber="37">cracens</emphasis>
</taxonomicName>
], 442719), 
<location id="8ED06EBF9833FFF60587756AD693FB09" LSID="urn:lsid:plazi:treatment:03A689729835FFF606AD71E6D639FB76:8ED06EBF9833FFF60587756AD693FB09" box="[1016,1248,1142,1167]" country="Venezuela" municipality="Near La Pastora" name="Serrania de San Luis" pageId="36" pageNumber="37" stateProvince="Falcon">Serranía de San Luís</location>
(
<specimenCode id="DBA9901F9833FFF604A87584D1FFFB28" box="[727,908,1175,1200]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="36" pageNumber="37">AMNH 276509</specimenCode>
, 
<specimenCode id="DBA9901F9833FFF605E3758BD183FB28" box="[924,1008,1175,1199]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="36" pageNumber="37">276580</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B6732399833FFF60278758BD635FB76" ID-GBIF-Occurrence="3059923312" collectionCode="FMNH, BMNH" collectorName="La Azulita" country="Venezuela" location="Merida" pageId="36" pageNumber="37" specimenCode="FMNH 22174, BMNH 1896.11" specimenCount="1" stateProvince="Merida" typeStatus="holotype">
<emphasis id="B97BE4769833FFF60278758BD62EFB28" box="[1031,1117,1175,1199]" italics="true" pageId="36" pageNumber="37">
<collectingRegion id="49CBF6869833FFF60278758BD62AFB28" box="[1031,1113,1175,1199]" country="Venezuela" name="Merida" pageId="36" pageNumber="37">Mérida</collectingRegion>
,
</emphasis>
<collectorName id="26FA5DB29833FFF602167584D6ACFB28" box="[1129,1247,1175,1199]" pageId="36" pageNumber="37">La Azulita</collectorName>
(
<specimenCode id="DBA9901F9833FFF604A875A5D1F3FB56" box="[727,896,1209,1233]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="36" pageNumber="37" type="Museum">FMNH 22174</specimenCode>
), 
<collectingRegion id="49CBF6869833FFF605E675A5D199FB56" box="[921,1002,1209,1233]" country="Venezuela" name="Merida" pageId="36" pageNumber="37">Mérida</collectingRegion>
(
<specimenCode id="DBA9901F9833FFF6058075A5D6C8FB57" box="[1023,1211,1208,1233]" collectionCode="BMNH" pageId="36" pageNumber="37">BMNH 1896.11</specimenCode>
.1.6 [
<typeStatus id="54B486C69833FFF604A975C6D14EFB75" box="[726,829,1242,1266]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
], 1903.1.5.2, 1903.1.5.3)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>