282 lines
42 KiB
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282 lines
42 KiB
XML
<document id="F772A8A46A27EAFB60C00DC93A4D97D2" ID-CLB-Dataset="7981" ID-DOI="10.11646/zootaxa.4933.3.2" ID-GBIF-Dataset="92979f75-7e46-49c9-ad01-d17939694c30" ID-ISSN="1175-5326" ID-Zenodo-Dep="4554920" ID-ZooBank="2DAB754B-1029-4D55-A4F5-DA3205DF679F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613985881098" checkinUser="plazi" docAuthor="Bertani, Rogério & Almeida, Marlus Queiroz" docDate="2021" docId="039D87E67E04BC2BFF55FC8CA76DFE00" docLanguage="en" docName="zootaxa.4933.3.2.pdf" docOrigin="Zootaxa 4933 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Yanomamius Bertani & Almeida 2021, n. gen." docType="treatment" docVersion="8" lastPageNumber="328" masterDocId="FFA4FF9E7E05BC2FFFC2FFF7A555FFD2" masterDocTitle="Yanomamius n. gen., a new genus of tarantula from Brazilian and Venezuelan Amazon (Araneae, Theraphosidae), with description of three new species" masterLastPageNumber="340" masterPageNumber="324" pageNumber="325" updateTime="1698930701653" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="819786324EB0F3D8DD733A0D36D100B3">Yanomamius n. gen., a new genus of tarantula from Brazilian and Venezuelan Amazon (Araneae, Theraphosidae), with description of three new species</mods:title>
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<mods:name id="BC30BE19F542E99AA38F72B0EA5B9F05" type="personal">
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<mods:namePart id="2A2F73FB29E89AFA2A6410C0CAA16A4B">Bertani, Rogério</mods:namePart>
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<mods:affiliation id="55AAB740C3FBAD8DF2392C4DC9D26935">Instituto Butantan, Laboratório de Ecologia e Evolução, Av. Vital Brazil, 1500 CEP 05503 - 900, São Paulo, SP, Brazil. bertani. rogerio @ gmail. com; https: // orcid. org / 0000 - 0002 - 8186 - 8316</mods:affiliation>
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<mods:namePart id="42031F43DA4519AB3DA35E97130EBDAD">Almeida, Marlus Queiroz</mods:namePart>
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<mods:affiliation id="5EF4971EA57CB1D84DE537A8F303ABFA">Instituto Nacional de Pesquisas da Amazônia, Laboratório de Sistemática e Ecologia de Invertebrados do Solo, CEP 69060 - 001, Av. André Araújo, Manaus, AM, Brazil.</mods:affiliation>
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<mods:title id="5123BDADB648BA0B4FE6D3CE223F2AE9">Zootaxa</mods:title>
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<mods:date id="A9CE666772F0447953EF975403C5A7D3">2021</mods:date>
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<taxonomicName id="4C344D737E04BC2EFF55FC8CA461FC47" ID-CoL="98X8B" authority="Bertani & Almeida, 2021" authorityName="Bertani & Almeida" authorityYear="2021" box="[151,308,891,917]" class="Arachnida" family="Theraphosidae" genus="Yanomamius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B940EAE27E04BC2EFF55FC8CA461FC47" bold="true" box="[151,308,891,917]" italics="true" pageId="1" pageNumber="325">Yanomamius</emphasis>
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(
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<figureCitation id="130F2A757E04BC2EFF5DFC68A444FC68" box="[159,273,927,954]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–14" captionStart-2="FIGURES 15–20" captionStart-3="FIGURES 21–26" captionStart-4="FIGURES 27–34" captionStart-5="FIGURES 35–41" captionStart-6="FIGURES 42–49" captionStart-7="FIGURE 50" captionStartId-0="2.[151,264,1906,1930]" captionStartId-1="3.[151,264,1896,1920]" captionStartId-2="6.[151,264,1973,1997]" captionStartId-3="7.[151,264,1930,1954]" captionStartId-4="8.[151,264,1896,1920]" captionStartId-5="12.[151,264,1895,1919]" captionStartId-6="13.[151,264,1895,1919]" captionStartId-7="14.[151,250,1901,1925]" captionTargetBox-0="[215,1371,181,1881]" captionTargetBox-1="[260,1368,190,1872]" captionTargetBox-2="[158,1430,633,1950]" captionTargetBox-3="[207,1379,181,1905]" captionTargetBox-4="[228,1368,190,1872]" captionTargetBox-5="[219,1368,181,1871]" captionTargetBox-6="[249,1367,188,1871]" captionTargetBox-7="[151,1435,1234,1877]" captionTargetId-0="figure-19@2.[215,1371,181,1881]" captionTargetId-1="figure-26@3.[219,1368,181,1872]" captionTargetId-2="figure-206@6.[158,1430,633,1950]" captionTargetId-3="figure-26@7.[207,1379,181,1905]" captionTargetId-4="figure-19@8.[219,1368,181,1872]" captionTargetId-5="figure-19@12.[219,1368,181,1871]" captionTargetId-6="figure-26@13.[219,1368,181,1871]" captionTargetId-7="figure-353@14.[151,1435,1234,1877]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="12" captionTargetPageId-6="13" captionTargetPageId-7="14" captionText-0="FIGURES 1–6. Y. franciscoi n. sp. from Ducke Reserve, Manaus, state of Amazonas, Brazil. 1–3 paratype female (INPA 9085). 1. carapace and chelicerae. 2. eye tubercle. 3. sternum, labium, maxillae and coxae. 4–6 holotype male (INPA 9084). 4. carapace and chelicerae. 5. eye tubercle. 6. sternum, labium, maxillae and coxae. Scale bars = 1 mm." captionText-1="FIGURES 7–14. Y. franciscoi n. sp. from Ducke Reserve, Manaus, state of Amazonas, Brazil. 7–13 holotype male (INPA 9084). 7–9 left bulb. 7. retrolateral. 8. prolateral. 9. dorsal 10–12 left tibial apophysis. 10. ventral. 11. prolateral. 12. retrolateral. 13. left palp, retrolateral. 14. paratype female (INPA 9085), spermathecae. The arrows indicate the ridges behind the retrolateral process of tibial apophysis. Scale bars = 1 mm." captionText-2="FIGURES 15–20. Y. franciscoi n. sp. Living specimens from Ducke Reserve, Manaus, state of Amazonas, Brazil. 15–16 burrows in an embankment. 17. adult female, paratype (INPA 9085). 18–20 immatures. Photographs: R. Bertani." captionText-3="FIGURES 21–26. Y. raonii n. sp. from Presidente Figueiredo, state of Amazonas, Brazil. 21–23 paratype female (INPA 1045) 21. carapace and chelicerae. 22. eye tubercle. 23. sternum, labium, maxillae and coxae. 24–26 holotype male (INPA 9088). 24. carapace and chelicerae. 25. eye tubercle. 26. sternum, labium, maxillae and coxae. Scale bars = 1 mm." captionText-4="FIGURES 27–34. Y. raonii n. sp. from Presidente Figueiredo, state of Amazonas, Brazil. 27–33 holotype male (INPA 9088). 27–29 left bulb. 27. retrolateral. 28. prolateral. 29. dorsal 30–32 left tibial apophysis. 30. ventral. 31. prolateral. 32. retrolateral. 33. left palp, retrolateral. 34 paratype female (INPA 1045), spermathecae. The arrows indicate the protuberance behind the retrolateral process of tibial apophysis. Scale bars = 1 mm." captionText-5="FIGURES 35–41. Y. neblina n. sp. from Pico da Neblina, S„o Gabriel da Cachoeira, state of Amazonas, Brazil. 35–38 paratype female (INPA 9092). 35. carapace and chelicerae. 36. eye tubercle. 37. sternum, labium, maxillae and coxae. 38. detail of the sternum showing black spiniform setae. 39–41 holotype male (INPA 9091). 39. carapace and chelicerae. 40. eye tubercle. 41. sternum, labium, maxillae and coxae. Scale bars = 1 mm." captionText-6="FIGURES 42–49. Y. neblina n. sp. from Pico da Neblina, S„o Gabriel da Cachoeira, state of Amazonas, Brazil. 42–48 holotype male (INPA 9091) 42–44 left bulb. 42. prolateral. 43. retrolateral. 44. dorsal 45–48 left tibial apophysis. 45. ventral. 46. prolateral. 47. retrolateral. 48. left palp, retrolateral. 49. paratype female (INPA 9092), spermathecae. The arrows indicate the ridges behind the retrolateral process of tibial apophysis. Scale bars = 1 mm." captionText-7="FIGURE 50. Map showing records for Yanomamius n. gen. in Northern South America. Red star = Y. franciscoi n. sp. Blue circle = Y. raonii n. sp. Yellow triangle = Y. neblina n. sp. Orange square = Y. waikoshiemi. Purple hexagon = Yanomamius sp." figureDoi-0="http://doi.org/10.5281/zenodo.4554922" figureDoi-1="http://doi.org/10.5281/zenodo.4554924" figureDoi-2="http://doi.org/10.5281/zenodo.4554926" figureDoi-3="http://doi.org/10.5281/zenodo.4554928" figureDoi-4="http://doi.org/10.5281/zenodo.4554930" figureDoi-5="http://doi.org/10.5281/zenodo.4554932" figureDoi-6="http://doi.org/10.5281/zenodo.4554934" figureDoi-7="http://doi.org/10.5281/zenodo.4554936" httpUri-0="https://zenodo.org/record/4554922/files/figure.png" httpUri-1="https://zenodo.org/record/4554924/files/figure.png" httpUri-2="https://zenodo.org/record/4554926/files/figure.png" httpUri-3="https://zenodo.org/record/4554928/files/figure.png" httpUri-4="https://zenodo.org/record/4554930/files/figure.png" httpUri-5="https://zenodo.org/record/4554932/files/figure.png" httpUri-6="https://zenodo.org/record/4554934/files/figure.png" httpUri-7="https://zenodo.org/record/4554936/files/figure.png" pageId="1" pageNumber="325">
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Figs 1
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<emphasis id="B940EAE27E04BC2EFF24FC68A5A1FC6B" bold="true" box="[230,244,927,953]" pageId="1" pageNumber="325">–</emphasis>
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50
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)
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</paragraph>
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<treatmentCitationGroup id="AB2411DE7E04BC2EFF55FC29A0C8FBC7" pageId="1" pageNumber="325">
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<treatmentCitation id="0A9510E17E04BC2EFF55FC29A4E9FC24" author="Karsch, F." box="[151,444,990,1014]" page="544" pageId="1" pageNumber="325" year="1879">
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<taxonomicName id="4C344D737E04BC2EFF55FC29A4E9FC24" ID-CoL="4Y4C" authority="Karsch, 1879: 544" authorityName="Karsch" authorityPageNumber="544" authorityYear="1879" box="[151,444,990,1014]" class="Arachnida" family="Theraphosidae" genus="Holothele" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="genus">
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<emphasis id="B940EAE27E04BC2EFF55FC29A5AEFC24" box="[151,251,990,1014]" italics="true" pageId="1" pageNumber="325">Holothele</emphasis>
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<bibRefCitation id="EFA54B017E04BC2EFEC3FC29A4E9FC24" author="Karsch, F." box="[257,444,990,1014]" pageId="1" pageNumber="325" pagination="534 - 562" refId="ref11403" refString="Karsch, F. (1879) Arachnologische Beitrage. Zeitschrift f ʾ r die Gesammten Naturwissenschaften, 52, 534 - 562." type="journal article" year="1879">Karsch, 1879: 544</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(in part,
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<taxonomicName id="4C344D737E04BC2EFDDBFC28A6FBFC24" ID-CoL="3HN9N" authority="Bertani & Araujo, 2006" authorityName="Bertani & Araujo" authorityYear="2006" box="[537,942,990,1014]" class="Arachnida" family="Theraphosidae" genus="Guyruita" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="species" species="waikoshiemi">
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<emphasis id="B940EAE27E04BC2EFDDBFC28A7E2FC24" box="[537,695,990,1014]" italics="true" pageId="1" pageNumber="325">H. waikoshiemi</emphasis>
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<bibRefCitation id="EFA54B017E04BC2EFD7FFC29A6FBFC24" author="Bertani, R. & Araujo, Y." box="[701,942,990,1014]" pageId="1" pageNumber="325" pagination="13 - 16" refId="ref10816" refString="Bertani, R. & Araujo, Y. (2006) A new species of Holothele Karsch, 1879 (Theraphosidae, Ischnocolinae) from Venezuela. Revista Iberica de Aracnologia, 12, 13 - 16." type="journal article" year="2006">Bertani & Araújo, 2006</bibRefCitation>
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</taxonomicName>
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,
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<typeStatus id="548F88527E04BC2EFC7BFC29A147FC24" box="[953,1042,990,1014]" pageId="1" pageNumber="325" type="holotype">holotype</typeStatus>
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female deposited at UCV, examined).
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<treatmentCitation id="0A9510E17E04BC2EFF55FC0AA67AFBC7" author="Guadanucci, J. P. L. & Lucas, S. M. & Indicatti, R. P. & Yamamoto, F. U." box="[151,815,1021,1045]" page="992" pageId="1" pageNumber="325" year="2007">
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<taxonomicName id="4C344D737E04BC2EFF55FC0AA67AFBC7" ID-CoL="4RNM" authority="Guadanucci, Lucas, Indicatti & Yamamoto, 2007: 992" authorityName="Guadanucci, Lucas, Indicatti & Yamamoto" authorityPageNumber="992" authorityYear="2007" box="[151,815,1021,1045]" class="Arachnida" family="Theraphosidae" genus="Guyruita" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="genus">
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<emphasis id="B940EAE27E04BC2EFF55FC0AA5A7FBC6" box="[151,242,1021,1044]" italics="true" pageId="1" pageNumber="325">Guyruita</emphasis>
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<bibRefCitation id="EFA54B017E04BC2EFF3EFC0AA67AFBC7" author="Guadanucci, J. P. L. & Lucas, S. M. & Indicatti, R. P. & Yamamoto, F. U." box="[252,815,1021,1045]" pageId="1" pageNumber="325" pagination="991 - 996" refId="ref11096" refString="Guadanucci, J. P. L., Lucas, S. M., Indicatti, R. P. & Yamamoto, F. U. (2007) Description of Guyruita gen. nov. and two new spe- cies (Ischnocolinae, Theraphosidae). Revista Brasileira de Zoologia, 24 (4), 991 - 996. https: // doi. org / 10.1590 / S 0101 - 81752007000400015." type="journal article" year="2007">Guadanucci, Lucas, Indicatti & Yamamoto, 2007: 992</bibRefCitation>
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</taxonomicName>
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(in part,
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<taxonomicName id="4C344D737E04BC2EFC56FC0AA004FBC7" ID-CoL="3HN9N" authority="(Bertani & Araujo, 2006))" authorityName="Bertani & Almeida" authorityYear="2021" baseAuthorityName="Bertani & Araujo" baseAuthorityYear="2006" box="[916,1361,1021,1045]" class="Arachnida" family="Theraphosidae" genus="Guyruita" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="species" species="waikoshiemi">
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<emphasis id="B940EAE27E04BC2EFC56FC0AA160FBC7" box="[916,1077,1021,1045]" italics="true" pageId="1" pageNumber="325">G. waikoshiemi</emphasis>
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(
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<bibRefCitation id="EFA54B017E04BC2EFB86FC0AA015FBC7" author="Bertani, R. & Araujo, Y." box="[1092,1344,1021,1045]" pageId="1" pageNumber="325" pagination="13 - 16" refId="ref10816" refString="Bertani, R. & Araujo, Y. (2006) A new species of Holothele Karsch, 1879 (Theraphosidae, Ischnocolinae) from Venezuela. Revista Iberica de Aracnologia, 12, 13 - 16." type="journal article" year="2006">Bertani & Araújo, 2006</bibRefCitation>
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))
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</taxonomicName>
|
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;
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<bibRefCitation id="EFA54B017E04BC2EFA9CFC0AA0C8FBC7" author="World Spider Catalog" box="[1374,1437,1021,1045]" pageId="1" pageNumber="325" refId="ref11714" refString="World Spider Catalog (2020) World Spider Catalog. Version 21.5. Natural History Museum Bern, Bern. Available from: http: // wsc. nmbe. ch (accessed 21 July 2020)" type="url" year="2020">World</bibRefCitation>
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</treatmentCitationGroup>
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Spider Catalog, 2020.
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|
||
<paragraph id="8B8B36F07E04BC2EFF55FBAAA7D4FBA5" blockId="1.[151,1437,1116,2043]" box="[151,641,1116,1143]" pageId="1" pageNumber="325">
|
||
<emphasis id="B940EAE27E04BC2EFF55FBAAA467FBA4" bold="true" box="[151,306,1116,1143]" pageId="1" pageNumber="325">
|
||
<typeStatus id="548F88527E04BC2EFF55FBAAA587FBA5" box="[151,210,1117,1143]" pageId="1" pageNumber="325">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C344D737E04BC2EFEFBFBAAA76BFBA5" ID-CoL="99BC5" authority="Bertani & Almeida, 2021" authorityName="Bertani & Almeida" authorityYear="2021" box="[313,574,1117,1143]" class="Arachnida" family="Theraphosidae" genus="Yanomamius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="species" species="franciscoi" status="sp. nov.">
|
||
<emphasis id="B940EAE27E04BC2EFEFBFBAAA76BFBA5" box="[313,574,1117,1143]" italics="true" pageId="1" pageNumber="325">Yanomamius franciscoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27357997E04BC2EFD87FBAAA7D4FBA5" box="[581,641,1117,1143]" pageId="1" pageNumber="325" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E657B7E04BC2EFF05FB77A727FAF9" pageId="1" pageNumber="325" type="diagnosis">
|
||
<paragraph id="8B8B36F07E04BC2EFF05FB77A727FAF9" blockId="1.[151,1437,1116,2043]" pageId="1" pageNumber="325">
|
||
<emphasis id="B940EAE27E04BC2EFF05FB77A417FB48" bold="true" box="[199,322,1152,1178]" pageId="1" pageNumber="325">Diagnosis.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4C344D737E04BC2EFE70FB76A715FB48" authority="Bertani & Almeida, 2021" authorityName="Bertani & Almeida" authorityYear="2021" box="[434,576,1153,1178]" class="Arachnida" family="Theraphosidae" genus="Yanomamius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B940EAE27E04BC2EFE70FB76A715FB48" box="[434,576,1153,1178]" italics="true" pageId="1" pageNumber="325">Yanomamius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27357997E04BC2EFD85FB76A7C6FB49" box="[583,659,1153,1179]" pageId="1" pageNumber="325" rank="genus">n. gen.</taxonomicNameLabel>
|
||
resemble those of
|
||
<taxonomicName id="4C344D737E04BC2EFCA5FB77A144FB48" authorityName="Karsch" authorityYear="1879" box="[871,1041,1152,1178]" class="Arachnida" family="Theraphosidae" genus="Schismatothele" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940EAE27E04BC2EFCA5FB77A144FB48" box="[871,1041,1152,1178]" italics="true" pageId="1" pageNumber="325">Schismatothele</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C344D737E04BC2EFB85FB77A185FB48" authorityName="Simon" authorityYear="1889" box="[1095,1232,1152,1178]" class="Arachnida" family="Theraphosidae" genus="Euthycaelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940EAE27E04BC2EFB85FB77A185FB48" box="[1095,1232,1152,1178]" italics="true" pageId="1" pageNumber="325">Euthycaelus</emphasis>
|
||
</taxonomicName>
|
||
by having several short spines closely positioned at the distal retrolateral palp tibia (
|
||
<figureCitation id="130F2A757E04BC2EFC42FB53A682FB6D" box="[896,983,1188,1215]" captionStart="FIGURES 7–14" captionStartId="3.[151,264,1896,1920]" captionTargetBox="[260,1368,190,1872]" captionTargetId="figure-26@3.[219,1368,181,1872]" captionTargetPageId="3" captionText="FIGURES 7–14. Y. franciscoi n. sp. from Ducke Reserve, Manaus, state of Amazonas, Brazil. 7–13 holotype male (INPA 9084). 7–9 left bulb. 7. retrolateral. 8. prolateral. 9. dorsal 10–12 left tibial apophysis. 10. ventral. 11. prolateral. 12. retrolateral. 13. left palp, retrolateral. 14. paratype female (INPA 9085), spermathecae. The arrows indicate the ridges behind the retrolateral process of tibial apophysis. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4554924" httpUri="https://zenodo.org/record/4554924/files/figure.png" pageId="1" pageNumber="325">Figs 13</figureCitation>
|
||
,
|
||
<figureCitation id="130F2A757E04BC2EFC26FB52A157FB6D" box="[996,1026,1189,1215]" captionStart="FIGURES 27–34" captionStartId="8.[151,264,1896,1920]" captionTargetBox="[228,1368,190,1872]" captionTargetId="figure-19@8.[219,1368,181,1872]" captionTargetPageId="8" captionText="FIGURES 27–34. Y. raonii n. sp. from Presidente Figueiredo, state of Amazonas, Brazil. 27–33 holotype male (INPA 9088). 27–29 left bulb. 27. retrolateral. 28. prolateral. 29. dorsal 30–32 left tibial apophysis. 30. ventral. 31. prolateral. 32. retrolateral. 33. left palp, retrolateral. 34 paratype female (INPA 1045), spermathecae. The arrows indicate the protuberance behind the retrolateral process of tibial apophysis. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4554930" httpUri="https://zenodo.org/record/4554930/files/figure.png" pageId="1" pageNumber="325">33</figureCitation>
|
||
,
|
||
<figureCitation id="130F2A757E04BC2EFBCDFB52A17AFB6D" box="[1039,1071,1189,1215]" captionStart="FIGURES 42–49" captionStartId="13.[151,264,1895,1919]" captionTargetBox="[249,1367,188,1871]" captionTargetId="figure-26@13.[219,1368,181,1871]" captionTargetPageId="13" captionText="FIGURES 42–49. Y. neblina n. sp. from Pico da Neblina, S„o Gabriel da Cachoeira, state of Amazonas, Brazil. 42–48 holotype male (INPA 9091) 42–44 left bulb. 42. prolateral. 43. retrolateral. 44. dorsal 45–48 left tibial apophysis. 45. ventral. 46. prolateral. 47. retrolateral. 48. left palp, retrolateral. 49. paratype female (INPA 9092), spermathecae. The arrows indicate the ridges behind the retrolateral process of tibial apophysis. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4554934" httpUri="https://zenodo.org/record/4554934/files/figure.png" pageId="1" pageNumber="325">48</figureCitation>
|
||
). They differ by the embolus tapering to its tip (
|
||
<figureCitation id="130F2A757E04BC2EFE90FB3FA4EDFB31" box="[338,440,1224,1251]" captionStart="FIGURES 7–14" captionStartId="3.[151,264,1896,1920]" captionTargetBox="[260,1368,190,1872]" captionTargetId="figure-26@3.[219,1368,181,1872]" captionTargetPageId="3" captionText="FIGURES 7–14. Y. franciscoi n. sp. from Ducke Reserve, Manaus, state of Amazonas, Brazil. 7–13 holotype male (INPA 9084). 7–9 left bulb. 7. retrolateral. 8. prolateral. 9. dorsal 10–12 left tibial apophysis. 10. ventral. 11. prolateral. 12. retrolateral. 13. left palp, retrolateral. 14. paratype female (INPA 9085), spermathecae. The arrows indicate the ridges behind the retrolateral process of tibial apophysis. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4554924" httpUri="https://zenodo.org/record/4554924/files/figure.png" pageId="1" pageNumber="325">Figs 7–9</figureCitation>
|
||
,
|
||
<figureCitation id="130F2A757E04BC2EFE06FB3EA758FB31" box="[452,525,1225,1251]" captionStart="FIGURES 27–34" captionStartId="8.[151,264,1896,1920]" captionTargetBox="[228,1368,190,1872]" captionTargetId="figure-19@8.[219,1368,181,1872]" captionTargetPageId="8" captionText="FIGURES 27–34. Y. raonii n. sp. from Presidente Figueiredo, state of Amazonas, Brazil. 27–33 holotype male (INPA 9088). 27–29 left bulb. 27. retrolateral. 28. prolateral. 29. dorsal 30–32 left tibial apophysis. 30. ventral. 31. prolateral. 32. retrolateral. 33. left palp, retrolateral. 34 paratype female (INPA 1045), spermathecae. The arrows indicate the protuberance behind the retrolateral process of tibial apophysis. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4554930" httpUri="https://zenodo.org/record/4554930/files/figure.png" pageId="1" pageNumber="325">27–29</figureCitation>
|
||
,
|
||
<figureCitation id="130F2A757E04BC2EFDDBFB3EA735FB31" box="[537,608,1225,1251]" captionStart="FIGURES 42–49" captionStartId="13.[151,264,1895,1919]" captionTargetBox="[249,1367,188,1871]" captionTargetId="figure-26@13.[219,1368,181,1871]" captionTargetPageId="13" captionText="FIGURES 42–49. Y. neblina n. sp. from Pico da Neblina, S„o Gabriel da Cachoeira, state of Amazonas, Brazil. 42–48 holotype male (INPA 9091) 42–44 left bulb. 42. prolateral. 43. retrolateral. 44. dorsal 45–48 left tibial apophysis. 45. ventral. 46. prolateral. 47. retrolateral. 48. left palp, retrolateral. 49. paratype female (INPA 9092), spermathecae. The arrows indicate the ridges behind the retrolateral process of tibial apophysis. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4554934" httpUri="https://zenodo.org/record/4554934/files/figure.png" pageId="1" pageNumber="325">42–44</figureCitation>
|
||
) and by the spines in the distal retrolateral palp tibia disposed in a group, instead of in one (
|
||
<taxonomicName id="4C344D737E04BC2EFE9CFB1BA75DFAD4" authorityName="Karsch" authorityYear="1879" box="[350,520,1260,1286]" class="Arachnida" family="Theraphosidae" genus="Schismatothele" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940EAE27E04BC2EFE9CFB1BA75DFAD4" box="[350,520,1260,1286]" italics="true" pageId="1" pageNumber="325">Schismatothele</emphasis>
|
||
</taxonomicName>
|
||
) or two (
|
||
<taxonomicName id="4C344D737E04BC2EFDACFB1BA7A2FAD4" authorityName="Simon" authorityYear="1889" box="[622,759,1260,1286]" class="Arachnida" family="Theraphosidae" genus="Euthycaelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940EAE27E04BC2EFDACFB1BA7A2FAD4" box="[622,759,1260,1286]" italics="true" pageId="1" pageNumber="325">Euthycaelus</emphasis>
|
||
</taxonomicName>
|
||
) rows. Females differ from all other schismatotheline genera by the incrassate tibia I in adults (
|
||
<figureCitation id="130F2A757E04BC2EFDD3FAE7A731FAF9" box="[529,612,1296,1323]" captionStart="FIGURES 15–20" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[158,1430,633,1950]" captionTargetId="figure-206@6.[158,1430,633,1950]" captionTargetPageId="6" captionText="FIGURES 15–20. Y. franciscoi n. sp. Living specimens from Ducke Reserve, Manaus, state of Amazonas, Brazil. 15–16 burrows in an embankment. 17. adult female, paratype (INPA 9085). 18–20 immatures. Photographs: R. Bertani." figureDoi="http://doi.org/10.5281/zenodo.4554926" httpUri="https://zenodo.org/record/4554926/files/figure.png" pageId="1" pageNumber="325">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E657B7E04BC2EFF05FAC2A003FA44" pageId="1" pageNumber="325" type="etymology">
|
||
<paragraph id="8B8B36F07E04BC2EFF05FAC2A003FA44" blockId="1.[151,1437,1116,2043]" pageId="1" pageNumber="325">
|
||
<emphasis id="B940EAE27E04BC2EFF05FAC2A41DFA9D" bold="true" box="[199,328,1333,1359]" pageId="1" pageNumber="325">Etymology</emphasis>
|
||
. The generic name is a homage to the Yanomami Indians. The distribution of the species of the genus on the Guiana Shield largely overlaps the territory of the Yanomamis. Furthermore, the first known species was found in a study on the food resources used by Yanomamis and Yekuanas. The genus is masculine in gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E657B7E04BC2BFF05FA57A62DFECF" lastPageId="4" lastPageNumber="328" pageId="1" pageNumber="325" type="description">
|
||
<paragraph id="8B8B36F07E04BC2EFF05FA57A795F964" blockId="1.[151,1437,1116,2043]" pageId="1" pageNumber="325">
|
||
<emphasis id="B940EAE27E04BC2EFF05FA57A40CFA68" bold="true" box="[199,345,1440,1466]" pageId="1" pageNumber="325">Description.</emphasis>
|
||
Carapace longer than wide, caput raised, cephalic and thoracic striae conspicuous. Fovea procurved (females) or slightly procurved (males). Clypeus short (females) or absent (males). Eight eyes arranged on tubercle, anterior eye row straight, posterior slightly recurved. Intercheliceral intumescence absent. Rastellum absent. Labium: rectangular with 141–401 cuspules (females), 423–728 cuspules (male). Labiosternal groove shallow, flat, with pair of sigilla. Maxillae: 153–198 cuspules (females), 167–272 cuspules (males) on upper mound in inner angle. Heel distinct. Anterior lobe distinct, short. Lyra absent. Short, black, spiniform setae on parts of palp and legs trochanters, maxillae and coxae of females. Sternum: Three pairs of sigilla, ovals, all ca. one diameter from margin. Book lung combs absent. Urticating setae absent.
|
||
</paragraph>
|
||
<paragraph id="8B8B36F07E04BC2EFF05F936A582F8B9" blockId="1.[151,1437,1116,2043]" pageId="1" pageNumber="325">Legs: formula IV-I-II-III (males), I=IV-II-III or I-IV-II-III (females). Preening combs absent. Clavate trichobothria in two rows. Claws: ITC absent. STC bare on all legs (females), or with two very small teeth on all or only on the III-IV legs (males); 1 bare claw on female palp. Tarsi II–IV, III–IV with medial ventral weakness (cracked) (males) or integral (females). Scopulae of tarsi III–IV of males and females and palp of females divided by broad band of setae.</paragraph>
|
||
<paragraph id="8B8B36F07E04BC2EFF05F883A059F828" blockId="1.[151,1437,1116,2043]" pageId="1" pageNumber="325">
|
||
Spinnerets: Apical segment of PLS digitiform. Male tibial apophysis with two processes, a small protuberance or a series of ridges behind the retrolateral process. Male palp tibia with distal 16–28 spines on retrolateral aspect, in a compact group. Metatarsus I narrow, pallid, weakly sclerotized, when folded it touches the retrolateral process tip (
|
||
<taxonomicName id="4C344D737E04BC2EFF07F816A461F828" authority="Bertani & Almeida, 2021" authorityName="Bertani & Almeida" authorityYear="2021" box="[197,308,2016,2042]" class="Arachnida" family="Theraphosidae" genus="Yanomamius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="325" phylum="Arthropoda" rank="species" species="neblina" status="sp. nov.">
|
||
<emphasis id="B940EAE27E04BC2EFF07F816A461F828" box="[197,308,2016,2042]" italics="true" pageId="1" pageNumber="325">Y. neblina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27357997E04BC2EFEF9F816A42CF829" box="[315,377,2017,2043]" pageId="1" pageNumber="325" rank="species">n. sp.</taxonomicNameLabel>
|
||
), or passes without touching it (other species). Tibia I of adult females incrassate.
|
||
</paragraph>
|
||
<caption id="DF4B66787E07BC2DFF55F885A160F803" ID-DOI="http://doi.org/10.5281/zenodo.4554922" ID-Zenodo-Dep="4554922" httpUri="https://zenodo.org/record/4554922/files/figure.png" pageId="2" pageNumber="326" startId="2.[151,264,1906,1930]" targetBox="[215,1371,181,1881]" targetPageId="2">
|
||
<paragraph id="8B8B36F07E07BC2DFF55F885A160F803" blockId="2.[151,1437,1906,2002]" pageId="2" pageNumber="326">
|
||
<emphasis id="B940EAE27E07BC2DFF55F885A46FF858" bold="true" box="[151,314,1906,1930]" pageId="2" pageNumber="326">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="4C344D737E07BC2DFEFDF884A4E9F858" authority="Bertani & Almeida, 2021" authorityName="Bertani & Almeida" authorityYear="2021" box="[319,444,1906,1930]" class="Arachnida" family="Theraphosidae" genus="Yanomamius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="326" phylum="Arthropoda" rank="species" species="franciscoi" status="sp. nov.">
|
||
<emphasis id="B940EAE27E07BC2DFEFDF884A4E9F858" box="[319,444,1906,1930]" italics="true" pageId="2" pageNumber="326">Y. franciscoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27357997E07BC2DFE00F885A4ADF85B" box="[450,504,1906,1929]" pageId="2" pageNumber="326" rank="species">n. sp.</taxonomicNameLabel>
|
||
from Ducke Reserve, Manaus, state of Amazonas, Brazil.
|
||
<emphasis id="B940EAE27E07BC2DFB81F885A13CF858" bold="true" box="[1091,1129,1906,1930]" pageId="2" pageNumber="326">1–3</emphasis>
|
||
paratype female (INPA 9085).
|
||
<emphasis id="B940EAE27E07BC2DFF55F861A5FFF87C" bold="true" box="[151,170,1942,1966]" pageId="2" pageNumber="326">1.</emphasis>
|
||
carapace and chelicerae.
|
||
<emphasis id="B940EAE27E07BC2DFE68F861A4E8F87C" bold="true" box="[426,445,1942,1966]" pageId="2" pageNumber="326">2.</emphasis>
|
||
eye tubercle.
|
||
<emphasis id="B940EAE27E07BC2DFD8BF861A709F87C" bold="true" box="[585,604,1942,1966]" pageId="2" pageNumber="326">3.</emphasis>
|
||
sternum, labium, maxillae and coxae.
|
||
<emphasis id="B940EAE27E07BC2DFC1DF861A151F87C" bold="true" box="[991,1028,1942,1966]" pageId="2" pageNumber="326">4–6</emphasis>
|
||
holotype male (INPA 9084).
|
||
<emphasis id="B940EAE27E07BC2DFAE9F861A06BF87C" bold="true" box="[1323,1342,1942,1966]" pageId="2" pageNumber="326">4.</emphasis>
|
||
carapace and chelicerae.
|
||
<emphasis id="B940EAE27E07BC2DFEF7F84DA41DF800" bold="true" box="[309,328,1978,2002]" pageId="2" pageNumber="326">5.</emphasis>
|
||
eye tubercle.
|
||
<emphasis id="B940EAE27E07BC2DFE14F84DA4BCF800" bold="true" box="[470,489,1978,2002]" pageId="2" pageNumber="326">6.</emphasis>
|
||
sternum, labium, maxillae and coxae. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4B66787E06BC2CFF55F89FA776F83E" ID-DOI="http://doi.org/10.5281/zenodo.4554924" ID-Zenodo-Dep="4554924" httpUri="https://zenodo.org/record/4554924/files/figure.png" pageId="3" pageNumber="327" startId="3.[151,264,1896,1920]" targetBox="[260,1368,190,1872]" targetPageId="3">
|
||
<paragraph id="8B8B36F07E06BC2CFF55F89FA776F83E" blockId="3.[151,1437,1896,2028]" pageId="3" pageNumber="327">
|
||
<emphasis id="B940EAE27E06BC2CFF55F89FA413F852" bold="true" box="[151,326,1896,1920]" pageId="3" pageNumber="327">FIGURES 7–14.</emphasis>
|
||
<taxonomicName id="4C344D737E06BC2CFE89F89EA492F852" authority="Bertani & Almeida, 2021" authorityName="Bertani & Almeida" authorityYear="2021" box="[331,455,1896,1920]" class="Arachnida" family="Theraphosidae" genus="Yanomamius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="327" phylum="Arthropoda" rank="species" species="franciscoi" status="sp. nov.">
|
||
<emphasis id="B940EAE27E06BC2CFE89F89EA492F852" box="[331,455,1896,1920]" italics="true" pageId="3" pageNumber="327">Y. franciscoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27357997E06BC2CFE0FF89EA757F852" box="[461,514,1897,1920]" pageId="3" pageNumber="327" rank="species">n. sp.</taxonomicNameLabel>
|
||
from Ducke Reserve, Manaus, state of Amazonas, Brazil.
|
||
<emphasis id="B940EAE27E06BC2CFB88F89FA128F852" bold="true" box="[1098,1149,1896,1920]" pageId="3" pageNumber="327">7–13</emphasis>
|
||
holotype male (INPA 9084).
|
||
<emphasis id="B940EAE27E06BC2CFF55F87BA5E8F876" bold="true" box="[151,189,1932,1956]" pageId="3" pageNumber="327">7–9</emphasis>
|
||
left bulb.
|
||
<emphasis id="B940EAE27E06BC2CFEE1F87BA463F876" bold="true" box="[291,310,1932,1956]" pageId="3" pageNumber="327">7.</emphasis>
|
||
retrolateral.
|
||
<emphasis id="B940EAE27E06BC2CFE74F87BA49CF876" bold="true" box="[438,457,1932,1956]" pageId="3" pageNumber="327">8.</emphasis>
|
||
prolateral.
|
||
<emphasis id="B940EAE27E06BC2CFDF9F87BA71BF876" bold="true" box="[571,590,1932,1956]" pageId="3" pageNumber="327">9.</emphasis>
|
||
dorsal
|
||
<emphasis id="B940EAE27E06BC2CFD55F87BA783F876" bold="true" box="[663,726,1932,1956]" pageId="3" pageNumber="327">10–12</emphasis>
|
||
left tibial apophysis.
|
||
<emphasis id="B940EAE27E06BC2CFC6CF87BA698F876" bold="true" box="[942,973,1932,1956]" pageId="3" pageNumber="327">10.</emphasis>
|
||
ventral.
|
||
<emphasis id="B940EAE27E06BC2CFBE7F87BA116F876" bold="true" box="[1061,1091,1932,1956]" pageId="3" pageNumber="327">11.</emphasis>
|
||
prolateral.
|
||
<emphasis id="B940EAE27E06BC2CFB77F87BA180F876" bold="true" box="[1205,1237,1932,1956]" pageId="3" pageNumber="327">12.</emphasis>
|
||
retrolateral.
|
||
<emphasis id="B940EAE27E06BC2CFA97F87BA020F876" bold="true" box="[1365,1397,1932,1956]" pageId="3" pageNumber="327">13.</emphasis>
|
||
left palp, retrolateral.
|
||
<emphasis id="B940EAE27E06BC2CFE88F847A43CF81A" bold="true" box="[330,361,1968,1992]" pageId="3" pageNumber="327">14.</emphasis>
|
||
paratype female (INPA 9085), spermathecae. The arrows indicate the ridges behind the retrolateral process of tibial apophysis. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8B36F07E01BC2BFF05FF60A62DFECF" blockId="4.[151,1437,151,321]" pageId="4" pageNumber="328">
|
||
Male palp: bulb pyriform, embolus thick, short or slightly elongate with two parallel prolateral keels in all species and an additional ventral keel in
|
||
<taxonomicName id="4C344D737E01BC2BFDF9FF4CA7FEFF07" authority="Bertani & Almeida, 2021" authorityName="Bertani & Almeida" authorityYear="2021" box="[571,683,187,213]" class="Arachnida" family="Theraphosidae" genus="Yanomamius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="328" phylum="Arthropoda" rank="species" species="neblina" status="sp. nov.">
|
||
<emphasis id="B940EAE27E01BC2BFDF9FF4CA7FEFF07" box="[571,683,187,213]" italics="true" pageId="4" pageNumber="328">Y. neblina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27357997E01BC2BFD70FF4CA7BAFF07" box="[690,751,187,213]" pageId="4" pageNumber="328" rank="species">n. sp.</taxonomicNameLabel>
|
||
Cymbium triangular in retrolateral view, with short incision. Spermathecae: two, each weakly sclerotized, with distal constriction, apical well sclerotized cluster of lobules and a well sclerotized subdistal or median lobule or cluster of lobules.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E657B7E01BC2BFF05FED1A790FE93" box="[199,709,294,321]" pageId="4" pageNumber="328" type="distribution">
|
||
<paragraph id="8B8B36F07E01BC2BFF05FED1A790FE93" blockId="4.[151,1437,151,321]" box="[199,709,294,321]" pageId="4" pageNumber="328">
|
||
<emphasis id="B940EAE27E01BC2BFF05FED1A434FE92" bold="true" box="[199,353,294,320]" pageId="4" pageNumber="328">Distribution.</emphasis>
|
||
<collectingCountry id="F32376607E01BC2BFEA5FED0A4FEFE93" box="[359,427,295,321]" name="Brazil" pageId="4" pageNumber="328">Brazil</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32376607E01BC2BFE20FED0A700FE93" box="[482,597,295,321]" name="Venezuela" pageId="4" pageNumber="328">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="130F2A757E01BC2BFDA6FED0A7EDFE93" box="[612,696,294,321]" captionStart="FIGURE 50" captionStartId="14.[151,250,1901,1925]" captionTargetBox="[151,1435,1234,1877]" captionTargetId="figure-353@14.[151,1435,1234,1877]" captionTargetPageId="14" captionText="FIGURE 50. Map showing records for Yanomamius n. gen. in Northern South America. Red star = Y. franciscoi n. sp. Blue circle = Y. raonii n. sp. Yellow triangle = Y. neblina n. sp. Orange square = Y. waikoshiemi. Purple hexagon = Yanomamius sp." figureDoi="http://doi.org/10.5281/zenodo.4554936" httpUri="https://zenodo.org/record/4554936/files/figure.png" pageId="4" pageNumber="328">Fig. 50</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E657B7E01BC2BFF55FE64A76DFE00" pageId="4" pageNumber="328" type="key">
|
||
<paragraph id="8B8B36F07E01BC2BFF55FE64A73CFE7F" blockId="4.[151,617,402,466]" box="[151,617,402,429]" pageId="4" pageNumber="328">
|
||
<emphasis id="B940EAE27E01BC2BFF55FE64A426FE7E" bold="true" box="[151,371,402,429]" pageId="4" pageNumber="328">Key for species of</emphasis>
|
||
<taxonomicName id="4C344D737E01BC2BFEB9FE64A745FE7E" ID-CoL="98X8B" authority="Bertani & Almeida, 2021" authorityName="Bertani & Almeida" authorityYear="2021" box="[379,528,403,428]" class="Arachnida" family="Theraphosidae" genus="Yanomamius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="328" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B940EAE27E01BC2BFEB9FE64A745FE7E" box="[379,528,403,428]" italics="true" pageId="4" pageNumber="328">Yanomamius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940EAE27E01BC2BFDD5FE64A73CFE7F" bold="true" box="[535,617,403,429]" pageId="4" pageNumber="328">
|
||
<taxonomicNameLabel id="A27357997E01BC2BFDD5FE64A73CFE7F" box="[535,617,403,429]" pageId="4" pageNumber="328" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8B36F07E01BC2BFF55FE40A76DFE00" blockId="4.[151,617,402,466]" box="[151,568,439,466]" pageId="4" pageNumber="328">
|
||
(Male of
|
||
<taxonomicName id="4C344D737E01BC2BFF3DFE40A4F3FE03" ID-CoL="3HN9N" authorityName="Bertani & Almeida" authorityYear="2021" baseAuthorityName="Bertani & Araujo" baseAuthorityYear="2006" box="[255,422,439,465]" class="Arachnida" family="Theraphosidae" genus="Guyruita" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="328" phylum="Arthropoda" rank="species" species="waikoshiemi">
|
||
<emphasis id="B940EAE27E01BC2BFF3DFE40A4F3FE03" box="[255,422,439,465]" italics="true" pageId="4" pageNumber="328">Y. waikoshiemi</emphasis>
|
||
</taxonomicName>
|
||
is unknown)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |