124 lines
16 KiB
XML
124 lines
16 KiB
XML
<document id="B0EA949DCF0C9274549B00568ED2D234" ID-CLB-Dataset="30136" ID-DOI="10.11646/zootaxa.4416.1.1" ID-GBIF-Dataset="aa238a39-adf8-4b94-8808-c788670d0455" ID-ISSN="1175-5326" ID-Zenodo-Dep="1242263" ID-ZooBank="3341A80B-7E54-4C6B-8B46-1336094952E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525601053009" checkinUser="plazi" docAuthor="Barbosa, Diego N. & Azevedo, Celso O." docDate="2018" docId="039D87C45CABFF0F48F6FF48FCC6FB3E" docLanguage="en" docName="zootaxa.4416.1.1.pdf" docOrigin="Zootaxa 4416 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Anisepyris proteus Evans 1966" docType="treatment" docVersion="7" lastPageNumber="181" masterDocId="FFA4FFBC5C1FFFBB4861FFDFFFCEFFFD" masterDocTitle="Revision of Anisepyris Kieffer (Hymenoptera, Bethylidae), with description of 135 new species" masterLastPageNumber="258" masterPageNumber="1" pageNumber="181" updateTime="1698542699957" updateUser="ExternalLinkService">
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<mods:titleInfo id="3B35F2F7732FFE72775C7B9B36F9646F">
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<mods:title id="CEB76B72A990A2E0C4619B8E9C878A67">Revision of Anisepyris Kieffer (Hymenoptera, Bethylidae), with description of 135 new species</mods:title>
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<mods:roleTerm id="189BA1E71883A67768439371575C3797">Author</mods:roleTerm>
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<mods:namePart id="10859AC9BB5829A24706AAE64D3751F5">Barbosa, Diego N.</mods:namePart>
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<mods:name id="BE9C54C1485B3095293509DC081D74DF" type="personal">
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<mods:roleTerm id="102F0DD8E4429558C488F2C027078103">Author</mods:roleTerm>
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<mods:namePart id="63463B0ABCD4D4CD481C45B45AF9D06D">Azevedo, Celso O.</mods:namePart>
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<mods:typeOfResource id="876B97C2908214AA941026796CF20BF1">text</mods:typeOfResource>
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<mods:title id="A0633F256982EEB53038A5A8D51B0923">Zootaxa</mods:title>
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<mods:part id="2B40689ABFF4A8A30D501DB7AC263D84">
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<mods:date id="9F8667BE05D1DB104D3E6D0DC193CD7A">2018</mods:date>
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<mods:number id="9179EA2377820CFCEC917B69FFACEA61">4416</mods:number>
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<mods:identifier id="3AF37BAD3F6CF6E6C5B120CC12392E5C" type="DOI">10.11646/zootaxa.4416.1.1</mods:identifier>
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<treatment id="039D87C45CABFF0F48F6FF48FCC6FB3E" ID-DOI="http://doi.org/10.5281/zenodo.5961571" ID-GBIF-Taxon="143811596" ID-Zenodo-Dep="5961571" LSID="urn:lsid:plazi:treatment:039D87C45CABFF0F48F6FF48FCC6FB3E" httpUri="http://treatment.plazi.org/id/039D87C45CABFF0F48F6FF48FCC6FB3E" lastPageNumber="181" pageId="180" pageNumber="181">
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<subSubSection id="C32E65595CABFF0F48F6FF48FE7FFF2E" pageId="180" pageNumber="181" type="nomenclature">
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<paragraph id="8B8B36D25CABFF0F48F6FF48FE6DFF4F" blockId="180.[151,433,151,211]" box="[151,419,151,178]" pageId="180" pageNumber="181">
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<heading id="D0C381BE5CABFF0F48F6FF48FE6DFF4F" bold="true" box="[151,419,151,178]" fontSize="11" level="1" pageId="180" pageNumber="181" reason="1">
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<emphasis id="B940EAC05CABFF0F48F6FF48FE6DFF4F" bold="true" box="[151,419,151,178]" pageId="180" pageNumber="181">
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<taxonomicName id="4C344D515CABFF0F48F6FF48FF3EFF4C" authorityName="Evans" authorityYear="1966" box="[151,240,151,177]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="proteus">
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<emphasis id="B940EAC05CABFF0F48F6FF48FF3EFF4C" bold="true" box="[151,240,151,177]" italics="true" pageId="180" pageNumber="181">proteus</emphasis>
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</taxonomicName>
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species-group
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8B36D25CABFF0F48F6FF65FE7FFF2E" blockId="180.[151,433,151,211]" box="[151,433,186,211]" pageId="180" pageNumber="181">
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(
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<figureCitation id="130F2A575CABFF0F48FEFF65FEE4FF2E" box="[159,298,186,211]" captionStart="FIGURE 23" captionStartId="169.[151,250,1901,1923]" captionTargetBox="[233,1351,223,1843]" captionTargetId="figure@169.[192,1395,185,1886]" captionTargetPageId="169" captionText="FIGURE 23. Anisepyris: megacephalus species-group. Head in dorsal profile. A. A. werneri (Evans, 1965) (♀); B–E. Mandible infrontal profile;F–H.Mesosoma indorsal profile;I–K.Mesopleuronin frontalprofile;proteus species-group.L–Q. Head in dorsal profile. L. A. ancalagoni sp. nov.; M. A. daeroni sp. nov.; N. A. dairuini sp. nov.; O. A.dorlasi sp. nov.; P. A. draugluini sp.nov.; Q. A.eilineli sp. nov." httpUri="https://zenodo.org/record/1242309/files/figure.png" pageId="180" pageNumber="181">Figs 23L–Q</figureCitation>
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,
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<figureCitation id="130F2A575CABFF0F4954FF64FE9DFF2E" box="[309,339,187,211]" captionStart="FIGURE 24" captionStartId="200.[151,250,1942,1964]" captionTargetBox="[224,1359,232,1878]" captionTargetId="figure@200.[182,1404,193,1921]" captionTargetPageId="200" captionText="FIGURE 24. Anisepyris: proteus species-group. Head in dorsal profile.A.A. elemmirei sp. nov.; B. A. elenwei sp. nov.; C. A. emeldiri sp. nov.; D. A. haldani sp. nov.; E. A. haldari sp. nov.; F.A. haldiri sp. nov.; G. A. halmiri sp. nov.; H. A. harethi sp. nov.; I. A. isilduri sp. nov.; J. A.khimi sp. nov.; K. A. artus Santos & Azevedo, 2008; L. A. proteus Evans, 1966." httpUri="https://zenodo.org/record/1242311/files/figure.png" pageId="180" pageNumber="181">24</figureCitation>
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,
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<figureCitation id="130F2A575CABFF0F493EFF65FE67FF2E" box="[351,425,186,211]" captionStart="FIGURE25" captionStartId="201.[151,250,1899,1921]" captionTargetBox="[233,1351,223,1843]" captionTargetId="figure@201.[192,1395,185,1886]" captionTargetPageId="201" captionText="FIGURE25. Anisepyris: proteus species-group.Head in dorsal profile.A. A. rugosicollisBrues, 1908; B–D. Female mandible in frontal profile; E. Male mandible in frontal profile; F–H.Pronotal disc in dorsal profile; I–J. Mesopleuron in frontal profile; strictusspecies-group.K–O.Head in dorsalprofile.K.A.guilini sp. nov.; L.A.hadorisp.nov.; M.A.hathaldiri sp. nov.; N.A. hunthori sp. nov.; O.A.huori sp. nov." httpUri="https://zenodo.org/record/1242313/files/figure.png" pageId="180" pageNumber="181">25A–J</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E65595CABFF0F48F6FEDDFB38FCCA" pageId="180" pageNumber="181" type="description">
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<paragraph id="8B8B36D25CABFF0F48F6FEDDFE05FDC6" blockId="180.[151,1437,258,1219]" pageId="180" pageNumber="181">
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<emphasis id="B940EAC05CABFF0F48F6FEDDFF37FEE6" bold="true" box="[151,249,258,283]" pageId="180" pageNumber="181">Females</emphasis>
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. Mandible with five distal teeth, evenly developed; clypeus short; antennal scrobe ecarinate or present; frontal angle of ocellar triangle acute and right; vertex straight and convex. Pronotal disc with transverse carina complete; parapsidal signum with anterior half continuous; anterior margin of propodeal declivity not setose; mesopleuron with lower mesopleural fovea opened, mesopleural suture opened or closed, and posterior mesopleural sulcus present. Forewing with R1 vein indistinct. Mesotibia not spinose Metatrochanter with lateral expansion. Metasoma with distal spiracle of metasoma small. Posterior hypopygeal margin projected uniform or convex. Genitalia with 2vv shorter than 1vv, and apex slender; sting (1vv+2vv) very long [surpassing 3vv apex]; T9 apex very swollen; 1vf strongly curved downward; 1rv not curved or curved, with 1rv ventral process small, and posterior process short.
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</paragraph>
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<paragraph id="8B8B36D25CABFF0F48A6FD99FB38FCCA" blockId="180.[151,1437,258,1219]" pageId="180" pageNumber="181">
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<emphasis id="B940EAC05CABFF0F48A6FD99FEC1FDA2" bold="true" box="[199,271,582,607]" pageId="180" pageNumber="181">Males</emphasis>
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. Frons not or weakly
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<taxonomicName id="4C344D515CABFF0F4A69FD99FD4DFDA2" authorityName="Kieffer" authorityYear="1908" box="[520,643,582,607]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="coriaceous">coriaceous</taxonomicName>
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, without color reflect of body metallic. Clypeus short or long; median clypeal carina lower than epistomal suture, in lateral view; antenna not elongate; antennal scrobe ecarinate. Pronotal disc with transverse carina complete. Metapostnotum with first pair of metapostnotal carina and metapostnotal-propodeal suture convergence absent; propodeal declivity with sculpture present. Mesopleuron with lower mesopleural fovea and mesopleural suture opened. Forewing with Cu vein indistinct of cu-a and R-rs&Rs vein indistinct. Hind wing with two proximal and four distal hamuli. Posterior hypopygeal margin uniform or uniform. Genitalia with paramere longer than basiparamere; inner margin of basivolsella excavated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E65595CABFF0F48A6FC9DFCA3FCA6" box="[199,877,834,859]" pageId="180" pageNumber="181" type="distribution">
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<paragraph id="8B8B36D25CABFF0F48A6FC9DFCA3FCA6" blockId="180.[151,1437,258,1219]" box="[199,877,834,859]" pageId="180" pageNumber="181">
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<emphasis id="B940EAC05CABFF0F48A6FC9DFE97FCA6" bold="true" box="[199,345,834,859]" pageId="180" pageNumber="181">Distribution</emphasis>
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. This group is mostly from Neotropical region.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E65595CABFF0F48A6FCB9FCC6FB3E" pageId="180" pageNumber="181" type="discussion">
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<paragraph id="8B8B36D25CABFF0F48A6FCB9FCC6FB3E" blockId="180.[151,1437,258,1219]" pageId="180" pageNumber="181">
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<emphasis id="B940EAC05CABFF0F48A6FCB9FEFAFC82" bold="true" box="[199,308,870,895]" pageId="180" pageNumber="181">Remarks</emphasis>
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. This group was defined by
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<bibRefCitation id="EFA54B235CABFF0F4A12FCB8FCC7FC82" box="[627,777,870,895]" pageId="180" pageNumber="181" refId="ref280374" refString="Evans, H. E. (1966) A revision of the genus Anisepyris Kieffer (Hymenoptera. Bethylidae). Studia Entomologica, 9, 1 - 120." type="journal article">Evans (1966)</bibRefCitation>
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by having the following characteristics: the mandible with five distal teeth, the antennal scrobe ecarinate or present, flagellomere I ring-shaped, the transverse pronotal carina absent or present, the mesopleuron with the mesopleural and lower fovea opened or closed, and the posterior fovea absent or present, the mesotibia without or with distal spines. Using these variable diagnostic characteristics,
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<taxonomicName id="4C344D515CABFF0F48F6FC28FF24FBF3" authorityName="Evans" authorityYear="1966" box="[151,234,1015,1038]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="proteus">
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<emphasis id="B940EAC05CABFF0F48F6FC28FF24FBF3" box="[151,234,1015,1038]" italics="true" pageId="180" pageNumber="181">proteus</emphasis>
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</taxonomicName>
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covered many species, making it the richest within
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<taxonomicName id="4C344D515CABFF0F4B3FFC28FC1AFBF3" authorityName="Kieffer" authorityYear="1905" box="[862,980,1015,1038]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="genus">
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<emphasis id="B940EAC05CABFF0F4B3FFC28FC1AFBF3" box="[862,980,1015,1038]" italics="true" pageId="180" pageNumber="181">Anisepyris</emphasis>
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</taxonomicName>
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. It was not easily definable, however, making it a “taxonomic trash-basket”. that included more than 40% of all species in
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<taxonomicName id="4C344D515CABFF0F4C39FBC4FB00FBCF" authorityName="Kieffer" authorityYear="1905" box="[1112,1230,1051,1074]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="genus">
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<emphasis id="B940EAC05CABFF0F4C39FBC4FB00FBCF" box="[1112,1230,1051,1074]" italics="true" pageId="180" pageNumber="181">Anisepyris</emphasis>
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</taxonomicName>
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. Most of these 51 species could be reallocated to other species-groups. In fact, former members of the
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<taxonomicName id="4C344D515CABFF0F4C27FBE0FB57FBAB" authorityName="Evans" authorityYear="1966" box="[1094,1177,1087,1110]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="proteus">
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<emphasis id="B940EAC05CABFF0F4C27FBE0FB57FBAB" box="[1094,1177,1087,1110]" italics="true" pageId="180" pageNumber="181">proteus</emphasis>
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</taxonomicName>
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species-group are now distributed into 11 of the 13 recognized species-groups of
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<taxonomicName id="4C344D515CABFF0F4B40FBBCFC59FB87" authorityName="Kieffer" authorityYear="1905" box="[801,919,1123,1146]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="genus">
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<emphasis id="B940EAC05CABFF0F4B40FBBCFC59FB87" box="[801,919,1123,1146]" italics="true" pageId="180" pageNumber="181">Anisepyris</emphasis>
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</taxonomicName>
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. One of these species-groups contains species that are related to
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<taxonomicName id="4C344D515CABFF0F4900FB58FE1AFB63" authorityName="Evans" authorityYear="1966" box="[353,468,1159,1182]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="proteus">
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<emphasis id="B940EAC05CABFF0F4900FB58FE1AFB63" box="[353,468,1159,1182]" italics="true" pageId="180" pageNumber="181">A. proteus</emphasis>
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</taxonomicName>
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and match the diagnostic features listed above. The redescribed and redefined
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<taxonomicName id="4C344D515CABFF0F4D28FB58FA52FB63" authorityName="Evans" authorityYear="1966" box="[1353,1436,1159,1182]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="proteus">
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<emphasis id="B940EAC05CABFF0F4D28FB58FA52FB63" box="[1353,1436,1159,1182]" italics="true" pageId="180" pageNumber="181">proteus</emphasis>
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</taxonomicName>
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species-group is now composed of 23 species (
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<tableCitation id="C6B603695CABFF0F4AC2FB75FD39FB3E" box="[675,759,1194,1219]" captionStart="TABLE 1" captionStartId="248.[151,239,1352,1375]" captionText="TABLE 1. Check list of species of Anisepyris." pageId="180" pageNumber="181">Table 1</tableCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |