treatments-xml/data/03/94/D6/0394D665BE6CFFFCFF3C5506272EE8AC.xml
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<document id="CFD66D7ABAEB051F439BDA76DA4034D3" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629612136760" checkinUser="felipe" docAuthor="Ballantyne, Lesley A. &amp; Lambkin, Christine" docDate="2009" docId="0394D665BE6CFFFCFF3C5506272EE8AC" docLanguage="en" docName="zt01997p188.pdf" docOrigin="Zootaxa 1997" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Lloydiella wareo Ballantyne &amp; Lambkin 2009, sp. n." docType="treatment" docVersion="3" lastPageNumber="65" masterDocId="FFADAE1DBE2CFFBCFFAB57492227EE30" masterDocTitle="Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae)" masterLastPageNumber="188" masterPageNumber="1" pageNumber="65" updateTime="1699083011144" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F0D7D8FAC23DFC92145757DB5D528C1C">Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="393FA9D585145D2D312148EE7A58D3EF">Ballantyne, Lesley A.</mods:namePart>
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<mods:namePart id="0C85979B3B22D8B0F5E694A037FEAAD2">Lambkin, Christine</mods:namePart>
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<mods:date id="0391490A24FF6E2248EA903BD466AF55">2009</mods:date>
<mods:detail id="F5A36DB0D0666F334AEB10E7D663AF35" type="pubDate">
<mods:number id="351F668E7546FEF6F4CEFEEAFA7E89DD">2009-02-04</mods:number>
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<mods:detail id="C7E254AC1BDB1476822E0A44A71D1EC9" type="volume">
<mods:number id="EF851609DB0F74892D2F844B63385EB2">1997</mods:number>
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<treatment id="0394D665BE6CFFFCFF3C5506272EE8AC" ID-DOI="http://doi.org/10.5281/zenodo.5324390" ID-Zenodo-Dep="5324390" LSID="urn:lsid:plazi:treatment:0394D665BE6CFFFCFF3C5506272EE8AC" httpUri="http://treatment.plazi.org/id/0394D665BE6CFFFCFF3C5506272EE8AC" lastPageNumber="65" pageId="64" pageNumber="65">
<subSubSection id="C32734F8BE6CFFFCFF3C5506238EEC59" box="[151,425,591,617]" pageId="64" pageNumber="65" type="nomenclature">
<paragraph id="8B826773BE6CFFFCFF3C5506238EEC59" blockId="64.[151,506,591,652]" box="[151,425,591,617]" pageId="64" pageNumber="65">
<heading id="D0CAD01FBE6CFFFCFF3C5506238EEC59" bold="true" box="[151,425,591,617]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
<emphasis id="B949BB61BE6CFFFCFF3C5506238EEC59" bold="true" box="[151,425,591,617]" pageId="64" pageNumber="65">
<taxonomicName id="4C3D1CF0BE6CFFFCFF3C55062378EC59" ID-CoL="3VQSQ" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[151,351,591,617]" class="Insecta" family="Lampyridae" genus="Lloydiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="wareo" status="sp. nov.">
<emphasis id="B949BB61BE6CFFFCFF3C55062378EC59" bold="true" box="[151,351,591,617]" italics="true" pageId="64" pageNumber="65">Lloydiella wareo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A061ABE6CFFFCFECC5506238EEC59" box="[359,425,591,617]" pageId="64" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE6CFFFCFF3C553B23DDECBC" box="[151,506,626,652]" pageId="64" pageNumber="65" type="description">
<paragraph id="8B826773BE6CFFFCFF3C553B23DDECBC" blockId="64.[151,506,591,652]" box="[151,506,626,652]" pageId="64" pageNumber="65">
(
<figureCitation id="13067BF6BE6CFFFCFF34553B235AECBC" box="[159,381,626,652]" captionStart="FIGURES 204211" captionStartId="162.[151,269,1874,1898]" captionTargetBox="[151,1435,204,1847]" captionTargetId="figure-76@162.[151,1436,194,1850]" captionTargetPageId="162" captionText="FIGURES 204211. Lloydiella spp. paratype males. 204, 208, 210 Ll. uberia; 205, 209 Ll. japenensis; 206, 207, 211 Ll wareo (BPBM). 204206 dorsal whole body (204, 206 have abdomen dissected); 207210 dorsal pronota; 211 four specimens on one card, L to R male ventral, female ventral (no head or pronotum), male dorsal, male ventral." pageId="64" pageNumber="65">Figs 206, 207, 211</figureCitation>
,
<figureCitation id="13067BF6BE6CFFFCFE22553B23D7ECBC" box="[393,496,626,652]" captionStart="FIGURES 212224" captionStartId="163.[151,269,1698,1722]" captionTargetBox="[151,1436,194,1656]" captionTargetId="figure-75@163.[151,1436,194,1674]" captionTargetPageId="163" captionText="FIGURES 212224. Lloydiella spp. paratype males. Ll. japenensis 212214, 217219, 222. Ll. majuscula holotype 223. Ll. uberia 215, 220, 224. Ll. wareo 216, 221, 224. 212, 215, 216 aedeagi ventral; 213 aedeagus left laterodorsal; 214 aedeagus slightly laterodorsal; 217, 218, 220, 221 aedeagal sheath, ventral; 219 aedeagal sheath dorsal; 222224 abdominal ventrites 6, 7 and tergite 8 ventral." pageId="64" pageNumber="65">216, 221</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE6CFFFCFF3C558B23C2ED64" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="8B826773BE6CFFFCFF3C558B22D6ED34" blockId="64.[151,1436,706,1692]" pageId="64" pageNumber="65">
<materialsCitation id="3B556D2EBE6CFFFCFF3C558B22D6ED34" collectionCode="SAM" collectorName="Finschaven &amp; Wareo &amp; L Wagner" country="Papua New Guinea" latitude="-6.6" location="Madang" longLatPrecision="5582" longitude="147.85" pageId="64" pageNumber="65" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<emphasis id="B949BB61BE6CFFFCFF3C558B232EECEC" bold="true" box="[151,265,706,732]" pageId="64" pageNumber="65">
<typeStatus id="5486D9D1BE6CFFFCFF3C558B232EECEC" box="[151,265,706,732]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
</emphasis>
. Male.
<emphasis id="B949BB61BE6CFFFCFECD558B20B3ECEC" bold="true" box="[358,660,706,732]" pageId="64" pageNumber="65">
<collectingCountry id="F32A27E3BE6CFFFCFECD558B20B3ECEC" box="[358,660,706,732]" name="Papua New Guinea" pageId="64" pageNumber="65">PAPUA NEW GUINEA</collectingCountry>
</emphasis>
:
<collectingRegion id="49F9A991BE6CFFFCFD0E558B2122ECEC" box="[677,773,706,732]" country="Papua New Guinea" name="Madang" pageId="64" pageNumber="65">Madang</collectingRegion>
Pr.,
<geoCoordinate id="EE0901B4BE6CFFFCFC94558B2151ECEC" box="[831,886,706,732]" degrees="6.6" direction="south" orientation="latitude" pageId="64" pageNumber="65" precision="5555" value="-6.6">6.6S</geoCoordinate>
,
<geoCoordinate id="EE0901B4BE6CFFFCFC2F558B21CDECEC" box="[900,1002,706,732]" degrees="147.85" direction="east" orientation="longitude" pageId="64" pageNumber="65" precision="555" value="147.85">147.85E</geoCoordinate>
,
<collectorName id="26C802A5BE6CFFFCFC5C558B265BECEC" box="[1015,1148,706,732]" pageId="64" pageNumber="65">Finschaven</collectorName>
(sic),
<collectorName id="26C802A5BE6CFFFCFB61558B273EECEC" box="[1226,1305,706,732]" pageId="64" pageNumber="65">Wareo</collectorName>
,
<collectorName id="26C802A5BE6CFFFCFA8D558B27BCECEC" box="[1318,1435,706,732]" pageId="64" pageNumber="65">L Wagner</collectorName>
(
<collectionCode id="ED2CFFB6BE6CFFFCFF3455A322C3ED34" box="[159,228,746,772]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="64" pageNumber="65">SAM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B826773BE6CFFFCFF6D545B23C2ED64" blockId="64.[151,1436,706,1692]" pageId="64" pageNumber="65">
<materialsCitation id="3B556D2EBE6CFFFCFF6D545B23C6ED64" collectionCode="SAM" collectorName="L Wagner" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="64" pageNumber="65" specimenCount="11" specimenCount-male="11" typeStatus="paratype">
<emphasis id="B949BB61BE6CFFFCFF6D545B2364ED1C" bold="true" box="[198,323,786,812]" pageId="64" pageNumber="65">
<typeStatus id="5486D9D1BE6CFFFCFF6D545B2364ED1C" box="[198,323,786,812]" pageId="64" pageNumber="65" type="paratype">Paratypes</typeStatus>
</emphasis>
(12)
<emphasis id="B949BB61BE6CFFFCFE2F545B2097ED1C" bold="true" box="[388,688,786,812]" pageId="64" pageNumber="65">
<collectingCountry id="F32A27E3BE6CFFFCFE2F545B2097ED1C" box="[388,688,786,812]" name="Papua New Guinea" pageId="64" pageNumber="65">PAPUA NEW GUINEA</collectingCountry>
</emphasis>
: Finschaven (sic), Wareo,
<collectorName id="26C802A5BE6CFFFCFC47545B2647ED1C" box="[1004,1120,786,812]" pageId="64" pageNumber="65">L Wagner</collectorName>
<specimenCount id="9D3BACFABE6CFFFCFBC3545B26F4ED1C" box="[1128,1235,786,812]" pageId="64" pageNumber="65" type="male">11 males</specimenCount>
, female (missing head and pronotum) (
<collectionCode id="ED2CFFB6BE6CFFFCFE39547323FFED64" box="[402,472,826,852]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="64" pageNumber="65">SAM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE6CFFFCFF6D542B23FBE844" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="8B826773BE6CFFFCFF6D542B21DCED94" blockId="64.[151,1436,706,1692]" pageId="64" pageNumber="65">
<emphasis id="B949BB61BE6CFFFCFF6D542B2319ED4C" bold="true" box="[198,318,866,892]" pageId="64" pageNumber="65">Diagnosis</emphasis>
. Very similar to
<taxonomicName id="4C3D1CF0BE6CFFFCFDA6542B2094ED4B" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[525,691,866,891]" class="Insecta" family="Lampyridae" genus="Lloydiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="japenensis">
<emphasis id="B949BB61BE6CFFFCFDA6542B2094ED4B" box="[525,691,866,891]" italics="true" pageId="64" pageNumber="65">Ll. japenensis</emphasis>
</taxonomicName>
, distinguished by its locality, the narrowly separated antennal sockets, and the very narrow posterior half of the aedeagal sheath sternite.
</paragraph>
<paragraph id="8B826773BE6CFFFCFF6D54FB274AEB9C" blockId="64.[151,1436,706,1692]" pageId="64" pageNumber="65">
<emphasis id="B949BB61BE6CFFFCFF6D54FB2322EDFC" bold="true" box="[198,261,946,972]" pageId="64" pageNumber="65">Male</emphasis>
. 11.0
<quantity id="4CC5CA96BE6CFFFCFEF354FB2398EDFC" box="[344,447,946,972]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="64" pageNumber="65" unit="mm" value="12.3">12.3 mm</quantity>
long; 4.55.0 mm wide; W/L 0.4. Colour (
<figureCitation id="13067BF6BE6CFFFCFC6954FB2681EDFC" box="[962,1190,946,972]" captionStart="FIGURES 204211" captionStartId="162.[151,269,1874,1898]" captionTargetBox="[151,1435,204,1847]" captionTargetId="figure-76@162.[151,1436,194,1850]" captionTargetPageId="162" captionText="FIGURES 204211. Lloydiella spp. paratype males. 204, 208, 210 Ll. uberia; 205, 209 Ll. japenensis; 206, 207, 211 Ll wareo (BPBM). 204206 dorsal whole body (204, 206 have abdomen dissected); 207210 dorsal pronota; 211 four specimens on one card, L to R male ventral, female ventral (no head or pronotum), male dorsal, male ventral." pageId="64" pageNumber="65">Figs 206, 207, 211</figureCitation>
): Pronotum, MS and MN orange yellow; pronotum with very faint traces of median paired brown marks in 4/12; elytra very dark brown almost black; head and maxillary palpi dark brown, antennae and labial palpi slightly lighter brown; ventral abdomen very dark brown except for white LO; abdominal tergites dark brown, T7, 8 pale, semitransparent, T7 may be faintly mottled in middle. Pronotum: 2.0
<quantity id="4CC5CA96BE6CFFFCFCF4531B219FEA5C" box="[863,952,1106,1132]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="64" pageNumber="65" unit="mm" value="2.3">2.3 mm</quantity>
long;
<quantity id="4CC5CA96BE6CFFFCFBAF531B26B6EA5C" box="[1028,1169,1106,1132]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="64" pageNumber="65" unit="mm" value="3.65" valueMax="3.9" valueMin="3.4">3.43.9 mm</quantity>
wide; W/L 1.7; lateral margins diverge posteriorly with some slight convergence in posterior area (8/12) or diverge along their length (3/12) with
<specimenCount id="9D3BACFABE6CFFFCFE8C53EB23ECEA8C" box="[295,459,1186,1212]" pageId="64" pageNumber="65" type="generic">one specimen</specimenCount>
having divergent left margin and convergent right margin; posterolateral corners angulate, &lt;90°, and inclined obliquely to the median line. Elytron: 9.010.0 mm long; with 2 welldefined interstitial lines (1, 2). Head: GHW 2.12.6; SIW 0.30.4; SIW/GHW 0.1. Apical segment of labial palpi with 2 teeth (broad basal and more slender apical). Abdomen, ventrites (
<figureCitation id="13067BF6BE6CFFFCFC055253262AEB04" box="[942,1037,1306,1332]" captionStart="FIGURES 204211" captionStartId="162.[151,269,1874,1898]" captionTargetBox="[151,1435,204,1847]" captionTargetId="figure-76@162.[151,1436,194,1850]" captionTargetPageId="162" captionText="FIGURES 204211. Lloydiella spp. paratype males. 204, 208, 210 Ll. uberia; 205, 209 Ll. japenensis; 206, 207, 211 Ll wareo (BPBM). 204206 dorsal whole body (204, 206 have abdomen dissected); 207210 dorsal pronota; 211 four specimens on one card, L to R male ventral, female ventral (no head or pronotum), male dorsal, male ventral." pageId="64" pageNumber="65">Fig.211</figureCitation>
): MPP relatively broad, similar to that of
<taxonomicName id="4C3D1CF0BE6CFFFCFF47520A23B2EB6C" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[236,405,1347,1372]" class="Insecta" family="Lampyridae" genus="Lloydiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="japenensis">
<emphasis id="B949BB61BE6CFFFCFF47520A23B2EB6C" box="[236,405,1347,1372]" italics="true" pageId="64" pageNumber="65">Ll. Japenensis</emphasis>
</taxonomicName>
, with apex inclining slightly dorsally. Tergites: T8 not heavily sclerotised, lateral margins converge slightly posteriorly, posterior margin broadly rounded, ventral surface lacking median longitudinal groove. Aedeagus (
<figureCitation id="13067BF6BE6CFFFCFE1752DB2004EB9C" box="[444,547,1426,1452]" captionStart="FIGURES 212224" captionStartId="163.[151,269,1698,1722]" captionTargetBox="[151,1436,194,1656]" captionTargetId="figure-75@163.[151,1436,194,1674]" captionTargetPageId="163" captionText="FIGURES 212224. Lloydiella spp. paratype males. Ll. japenensis 212214, 217219, 222. Ll. majuscula holotype 223. Ll. uberia 215, 220, 224. Ll. wareo 216, 221, 224. 212, 215, 216 aedeagi ventral; 213 aedeagus left laterodorsal; 214 aedeagus slightly laterodorsal; 217, 218, 220, 221 aedeagal sheath, ventral; 219 aedeagal sheath dorsal; 222224 abdominal ventrites 6, 7 and tergite 8 ventral." pageId="64" pageNumber="65">Fig. 216</figureCitation>
). Aedeagal sheath (
<figureCitation id="13067BF6BE6CFFFCFCA352DB214BEB9C" box="[776,876,1426,1452]" captionStart="FIGURES 212224" captionStartId="163.[151,269,1698,1722]" captionTargetBox="[151,1436,194,1656]" captionTargetId="figure-75@163.[151,1436,194,1674]" captionTargetPageId="163" captionText="FIGURES 212224. Lloydiella spp. paratype males. Ll. japenensis 212214, 217219, 222. Ll. majuscula holotype 223. Ll. uberia 215, 220, 224. Ll. wareo 216, 221, 224. 212, 215, 216 aedeagi ventral; 213 aedeagus left laterodorsal; 214 aedeagus slightly laterodorsal; 217, 218, 220, 221 aedeagal sheath, ventral; 219 aedeagal sheath dorsal; 222224 abdominal ventrites 6, 7 and tergite 8 ventral." pageId="64" pageNumber="65">Fig. 221</figureCitation>
) with posterior half of sternite very narrow.
</paragraph>
<paragraph id="8B826773BE6CFFFCFF6D52F3212DE87C" blockId="64.[151,1436,706,1692]" pageId="64" pageNumber="65">
<emphasis id="B949BB61BE6CFFFCFF6D52F32306EBE4" bold="true" box="[198,289,1466,1492]" pageId="64" pageNumber="65">Female</emphasis>
. Assumed capable of flight. Missing head and pronotum, otherwise coloured as for male except for very pale, whitish and semitransparent V7 and 8. Posterior margin of V7 medially shallowly emarginated, lateral areas broadly rounded; median posterior margin of V8 narrowly emarginated. T8 with lateral margins converging posteriorly and posterior margin rounded.
</paragraph>
<paragraph id="8B826773BE6CFFFCFF6D511323FBE844" blockId="64.[151,1436,706,1692]" box="[198,476,1626,1652]" pageId="64" pageNumber="65">
<emphasis id="B949BB61BE6CFFFCFF6D51132335E844" bold="true" box="[198,274,1626,1652]" pageId="64" pageNumber="65">Larva</emphasis>
.
<emphasis id="B949BB61BE6CFFFCFE8A511323FBE844" bold="true" box="[289,476,1626,1652]" pageId="64" pageNumber="65">Not associated.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE6CFFFCFF6D51CB272EE8AC" box="[198,1289,1666,1692]" pageId="64" pageNumber="65" type="etymology">
<paragraph id="8B826773BE6CFFFCFF6D51CB272EE8AC" blockId="64.[151,1436,706,1692]" box="[198,1289,1666,1692]" pageId="64" pageNumber="65">
<emphasis id="B949BB61BE6CFFFCFF6D51CB236CE8AC" bold="true" box="[198,331,1666,1692]" pageId="64" pageNumber="65">Etymology</emphasis>
. The specific name is considered a noun in apposition reflecting the
<typeStatus id="5486D9D1BE6CFFFCFBD951CB2683E8AC" box="[1138,1188,1666,1692]" pageId="64" pageNumber="65">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>