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<document id="CE26F652B132D1BD53D2083AA994AD4A" ID-DOI="10.11646/zootaxa.5403.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10561558" ID-ZooBank="7458460D-2367-48C5-9ACE-D28457C27463" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1706103452799" checkinUser="plazi" docAuthor="Dondorp, Esther &amp; Denzer, Wolfgang" docDate="2024" docId="0387E3417C37FFC774C8CD61CBCEEB3F" docLanguage="en" docName="zootaxa.5403.1.4.pdf" docOrigin="Zootaxa 5403 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lophyrus sumatranus Schlegel 1851" docType="treatment" docVersion="3" lastPageNumber="75" masterDocId="FFBE9B397C3EFFCD745FCA6FC845E926" masterDocTitle="Annotated catalogue of acrodont lizard types in the collection of the Naturalis Biodiverstity Center (Squamata: Agamidae, Chamaeleonidae)" masterLastPageNumber="79" masterPageNumber="65" pageNumber="74" updateTime="1706278432058" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="BD70ACD955477171CEC99770DB07F607">Annotated catalogue of acrodont lizard types in the collection of the Naturalis Biodiverstity Center (Squamata: Agamidae, Chamaeleonidae)</mods:title>
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<mods:namePart id="798E8EF675F89CFD9B21FA57752EBF1E">Denzer, Wolfgang</mods:namePart>
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<emphasis id="B95A8E457C37FFC474C8CD61CCAAEE0E" bold="true" box="[151,1263,1806,1833]" pageId="9" pageNumber="74">
<subSubSection id="C33401DC7C37FFC474C8CD61CA3CEE0E" box="[151,633,1806,1833]" pageId="9" pageNumber="74" type="nomenclature">
<paragraph id="8B9152577C37FFC474C8CD61CA3CEE0E" blockId="9.[151,1263,1806,1833]" box="[151,633,1806,1833]" pageId="9" pageNumber="74">
<heading id="D0D9E53B7C37FFC474C8CD61CA3CEE0E" bold="true" box="[151,633,1806,1833]" fontSize="11" level="1" pageId="9" pageNumber="74" reason="1">
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<treatmentCitation id="0A8F74467C37FFC474C8CD61CA3CEE0E" author="Schlegel, H." box="[151,633,1806,1833]" page="5" pageId="9" pageNumber="74" year="1851">
<taxonomicName id="4C2E29D47C37FFC474C8CD61CA3CEE0E" ID-CoL="3W563" authority="Schlegel, 1851: 5" authorityName="Schlegel" authorityPageNumber="5" authorityYear="1851" box="[151,633,1806,1833]" class="Squamata" family="Agamidae" genus="Lophyrus" kingdom="Animalia" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="sumatranus">
<emphasis id="B95A8E457C37FFC474C8CD61C9E4EE0F" bold="true" box="[151,417,1806,1833]" italics="true" pageId="9" pageNumber="74">Lophyrus sumatranus</emphasis>
<bibRefCitation id="EFBF2FA67C37FFC475F6CD61CA3CEE0E" author="Schlegel, H." box="[425,633,1806,1832]" pageId="9" pageNumber="74" pagination="4 - 6" refId="ref11831" refString="Schlegel, H. (1851) Descriptions de plusieurs expeces nouvelles du genre Lophyrus. Bijdragen tot de Dierkunde, Amsterdam, 3 (1), 4 - 6. https: // doi. org / 10.1163 / 26660644 - 00301002" type="journal article" year="1851">Schlegel, 1851: 5</bibRefCitation>
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</paragraph>
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<subSubSection id="C33401DC7C37FFC476DFCD61CCAAEE0E" box="[640,1263,1806,1833]" pageId="9" pageNumber="74" type="reference_group">
<paragraph id="8B9152577C37FFC476DFCD61CCAAEE0E" blockId="9.[151,1263,1806,1833]" box="[640,1263,1806,1833]" pageId="9" pageNumber="74">
=
<taxonomicName id="4C2E29D47C37FFC476C7CD61CCAAEE0E" authority="(Laurenti, 1768)" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[664,1263,1806,1833]" class="Squamata" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="chamaeleontinus">
<emphasis id="B95A8E457C37FFC476C7CD61CC58EE0F" bold="true" box="[664,1053,1806,1833]" italics="true" pageId="9" pageNumber="74">Gonocephalus chamaeleontinus</emphasis>
(
<bibRefCitation id="EFBF2FA67C37FFC47073CD61CCA2EE0E" author="Laurenti, J. N." box="[1068,1255,1806,1832]" pageId="9" pageNumber="74" refId="ref10764" refString="Laurenti, J. N. (1768) Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium austracorum, quod authoritate et consensu. Joan. Thomae, Vienna, 217 pp. https: // doi. org / 10.5962 / bhl. title. 5108" type="book" year="1768">Laurenti, 1768</bibRefCitation>
)
</taxonomicName>
</paragraph>
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</emphasis>
<subSubSection id="C33401DC7C37FFC474C8CD39CA02EEB2" pageId="9" pageNumber="74" type="materials_examined">
<paragraph id="8B9152577C37FFC474C8CD39CBD8EE56" blockId="9.[151,1437,1878,2013]" box="[151,925,1878,1905]" pageId="9" pageNumber="74">
<materialsCitation id="3B46580A7C37FFC474C8CD39CBD8EE56" box="[151,925,1878,1905]" collectionCode="RMNH, RENA" location="Sumatra" pageId="9" pageNumber="74" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="B95A8E457C37FFC474C8CD39C941EE56" bold="true" box="[151,260,1878,1904]" pageId="9" pageNumber="74">
<typeStatus id="5495ECF57C37FFC474C8CD39C941EE56" box="[151,260,1878,1904]" pageId="9" pageNumber="74" type="holotype">Holotype</typeStatus>
</emphasis>
(by monotypy):
<collectionCode id="ED3FCA927C37FFC4759ECD39CA5FEE57" box="[449,538,1878,1905]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="74" type="Museum">RMNH</collectionCode>
.
<collectionCode id="ED3FCA927C37FFC47641CD39CA2AEE57" box="[542,623,1878,1905]" pageId="9" pageNumber="74">RENA</collectionCode>
.3051, male,
<collectingRegion id="49EA9CB57C37FFC476A5CD39CB1FEE56" box="[762,858,1878,1904]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="74">Sumatra</collectingRegion>
, 1848
</materialsCitation>
</paragraph>
<paragraph id="8B9152577C37FFC47498CD15CA02EEB2" blockId="9.[151,1437,1878,2013]" box="[199,583,1914,1941]" pageId="9" pageNumber="74">
<emphasis id="B95A8E457C37FFC47498CD15C925EEB2" bold="true" box="[199,352,1914,1940]" pageId="9" pageNumber="74">Type locality</emphasis>
:
<collectingRegion id="49EA9CB57C37FFC47530CD15C98AEEB2" box="[367,463,1914,1940]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="74">Sumatra</collectingRegion>
,
<collectingCountry id="F33912C77C37FFC47585CD15CA02EEB2" box="[474,583,1914,1940]" name="Indonesia" pageId="9" pageNumber="74">Indonesia</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C33401DC7C37FFC77498CDF1CBCEEB3F" lastPageId="10" lastPageNumber="75" pageId="9" pageNumber="74" type="discussion">
<paragraph id="8B9152577C37FFC47498CDF1CDD8EEFB" blockId="9.[151,1437,1878,2013]" pageId="9" pageNumber="74">
<emphasis id="B95A8E457C37FFC47498CDF1C971EE9E" bold="true" box="[199,308,1950,1976]" pageId="9" pageNumber="74">Remarks</emphasis>
:
<bibRefCitation id="EFBF2FA67C37FFC4751CCDF1C9B0EE9F" author="Schlegel, H." box="[323,501,1950,1977]" pageId="9" pageNumber="74" pagination="4 - 6" refId="ref11831" refString="Schlegel, H. (1851) Descriptions de plusieurs expeces nouvelles du genre Lophyrus. Bijdragen tot de Dierkunde, Amsterdam, 3 (1), 4 - 6. https: // doi. org / 10.1163 / 26660644 - 00301002" type="journal article" year="1851">Schlegel (1851)</bibRefCitation>
did not indicate how many specimens he had to hand and the only measurement he gave was a total length of 17 pouces (1 pouce approx.
<quantity id="4CD6FFB27C37FFC4775BCDADCB11EEFB" box="[772,852,1986,2013]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="74" unit="mm" value="25.0">25 mm</quantity>
in 1851). This agrees well with the size of RMNH.
</paragraph>
<paragraph id="8B9152577C34FFC774C8CAF9CBCEEB3F" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="75">
RENA.3051 (
<quantity id="4CD6FFB27C34FFC7756CCAF8C9DCE997" box="[307,409,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="4.3" pageId="10" pageNumber="75" unit="mm" value="430.0">430 mm</quantity>
). There exists another specimen in the Naturalis collection that may constitute a
<typeStatus id="5495ECF57C34FFC77162CAF8CDDDE997" box="[1341,1432,151,177]" pageId="10" pageNumber="75" type="syntype">syntype</typeStatus>
: RMNH.RENA.3055, male,
<collectingRegion id="49EA9CB57C34FFC77590CAD4CA6AE9F3" box="[463,559,187,213]" country="Indonesia" name="Sumatera Utara" pageId="10" pageNumber="75">Sumatra</collectingRegion>
, coll. S. Müller 1826. However, the year of collection given in the handwritten catalogue does not coincide with Müllers visit to
<collectingRegion id="49EA9CB57C34FFC776BECAB0CB7BE9DF" box="[737,830,223,249]" country="Indonesia" name="Sumatera Utara" pageId="10" pageNumber="75">Sumatra</collectingRegion>
(18331835). As we could not ascertain the precise collection data for RMNH.RENA.3055 we currently consider RMNH.RENA.3051 as the
<typeStatus id="5495ECF57C34FFC770C9CB6CCCBDE83B" box="[1174,1272,259,285]" pageId="10" pageNumber="75" type="holotype">holotype</typeStatus>
by monotypy.
<bibRefCitation id="EFBF2FA67C34FFC774C8CB48C900E867" author="Barbour, T." box="[151,325,295,321]" pageId="10" pageNumber="75" pagination="1 - 203" refId="ref8618" refString="Barbour, T. (1912) A contribution to the zoogeography of the East Indian Islands. Memoirs of the Museum of Comparative Zoology Harvard, 44, 1 - 203. https: // doi. org / 10.5962 / bhl. title. 52042" type="journal article" year="1912">Barbour (1912)</bibRefCitation>
stated that „Schlegels
<emphasis id="B95A8E457C34FFC77613CB48CA25E867" box="[588,608,295,321]" italics="true" pageId="10" pageNumber="75">G</emphasis>
. [
<emphasis id="B95A8E457C34FFC77627CB48CB43E867" box="[632,774,295,321]" italics="true" pageId="10" pageNumber="75">onocephalus</emphasis>
]
<taxonomicName id="4C2E29D47C34FFC77749CB48CC58E866" authorityName="Schlegel" authorityYear="1851" box="[790,1053,295,321]" class="Squamata" family="Agamidae" genus="Lophyrus" kingdom="Animalia" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="sumatranus" subGenus="Lophyrus">
(
<emphasis id="B95A8E457C34FFC77740CB48CBCCE867" box="[799,905,295,321]" italics="true" pageId="10" pageNumber="75">Lophyrus</emphasis>
)
<emphasis id="B95A8E457C34FFC777C5CB48CC58E866" box="[922,1053,295,320]" italics="true" pageId="10" pageNumber="75">sumatranus</emphasis>
</taxonomicName>
should be added to the synonymy [of
<taxonomicName id="4C2E29D47C34FFC7749ECB24C9E1E843" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[193,420,331,357]" class="Squamata" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="chamaeleontinus">
<emphasis id="B95A8E457C34FFC7749ECB24C9E1E843" box="[193,420,331,357]" italics="true" pageId="10" pageNumber="75">G. chamaeleontinus</emphasis>
</taxonomicName>
]“ and the species has been treated as such ever since with one major exception, namely
<bibRefCitation id="EFBF2FA67C34FFC774C8CB00C9CDE8AF" author="de Rooij, N." box="[151,392,366,393]" pageId="10" pageNumber="75" refId="ref9589" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, xiv + 384 pp. https: // doi. org / 10.5962 / bhl. title. 10610" type="book" year="1915">de Rooij (1915: 103)</bibRefCitation>
. With regard to Sumatran populations currently considered to represent
<taxonomicName id="4C2E29D47C34FFC770E4CB00CDD9E8AF" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[1211,1436,367,393]" class="Squamata" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="chamaeleontinus">
<emphasis id="B95A8E457C34FFC770E4CB00CDD9E8AF" box="[1211,1436,367,393]" italics="true" pageId="10" pageNumber="75">G. chamaeleontinus</emphasis>
</taxonomicName>
the name
<taxonomicName id="4C2E29D47C34FFC77557CBFCCA73E88A" box="[264,566,403,429]" class="Squamata" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="sumatranus">
<emphasis id="B95A8E457C34FFC77557CBFCCA73E88A" box="[264,566,403,429]" italics="true" pageId="10" pageNumber="75">Gonocephalus sumatranus</emphasis>
</taxonomicName>
is available for nomenclatural purposes. The specimen depicted in
<bibRefCitation id="EFBF2FA67C34FFC77163CBFCC89EE8F7" author="Schlegel, H." pageId="10" pageNumber="75" pagination="4 - 6" refId="ref11831" refString="Schlegel, H. (1851) Descriptions de plusieurs expeces nouvelles du genre Lophyrus. Bijdragen tot de Dierkunde, Amsterdam, 3 (1), 4 - 6. https: // doi. org / 10.1163 / 26660644 - 00301002" type="journal article" year="1851">Schlegel (1851</bibRefCitation>
, plate III) and the
<typeStatus id="5495ECF57C34FFC775F4CBD8CA48E8F7" box="[427,525,439,465]" pageId="10" pageNumber="75" type="holotype">holotype</typeStatus>
exhibit morphological traits such as well a developed dorsal crest (not present in several other
<taxonomicName id="4C2E29D47C34FFC77572CBB4CA49E8D3" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[301,524,475,501]" class="Squamata" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="chamaeleontinus">
<emphasis id="B95A8E457C34FFC77572CBB4CA49E8D3" box="[301,524,475,501]" italics="true" pageId="10" pageNumber="75">G. chamaeleontinus</emphasis>
</taxonomicName>
populations) that may warrant species status. Regarding the date of publication of Schlegels original description see comment under
<taxonomicName id="4C2E29D47C34FFC7768CCB90CBC1EB3F" box="[723,900,511,537]" class="Squamata" family="Agamidae" genus="Lophyrus" kingdom="Animalia" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="B95A8E457C34FFC7768CCB90CBC1EB3F" box="[723,900,511,537]" italics="true" pageId="10" pageNumber="75">Lophyrus kuhlii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>