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<document id="529A7D42675650B0B0D408DC218A2A0D" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="03877623626EFFC92DEFB48BFEE75CF7" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Crossoglossa) mellea" docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="41" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="876449641A2FB62006C80501789FC196">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<treatment id="03877623626EFFC92DEFB48BFEE75CF7" ID-DOI="http://doi.org/10.5281/zenodo.7963729" ID-GBIF-Taxon="209628966" ID-Zenodo-Dep="7963729" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7" httpUri="http://treatment.plazi.org/id/03877623626EFFC92DEFB48BFEE75CF7" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection id="C33494BE626EFFCB2DEFB48BFCE45A53" box="[151,771,1775,1802]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8B91C735626EFFCB2DEFB48BFCE45A53" blockId="40.[151,771,1775,1838]" box="[151,771,1775,1802]" pageId="40" pageNumber="41">
<emphasis id="B95A1B27626EFFCB2DEFB48BFCE45A53" bold="true" box="[151,771,1775,1802]" pageId="40" pageNumber="41">
[7]
<taxonomicName id="4C2EBCB6626EFFCB2DB9B48BFCE45A53" ID-CoL="4DKPT" authority="(Chaudoir, 1872)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[193,771,1775,1802]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mellea" subGenus="Crossoglossa">
<emphasis id="B95A1B27626EFFCB2DB9B48BFEFE5A50" bold="true" box="[193,281,1776,1802]" italics="true" pageId="40" pageNumber="41">Parena</emphasis>
(
<emphasis id="B95A1B27626EFFCB2C52B494FE2F5A50" bold="true" box="[298,456,1775,1802]" italics="true" pageId="40" pageNumber="41">Crossoglossa</emphasis>
)
<emphasis id="B95A1B27626EFFCB2CA1B494FDCE5A50" bold="true" box="[473,553,1775,1802]" italics="true" pageId="40" pageNumber="41">mellea</emphasis>
(
<bibRefCitation id="EFBFBAC4626EFFCB2F40B494FD1C5A53" author="Chaudoir, B. M." box="[568,763,1775,1801]" pageId="40" pageNumber="41" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872</bibRefCitation>
)
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626EFFCB2DEFB56FFD0C5A74" box="[151,747,1812,1838]" pageId="40" pageNumber="41" type="description">
<paragraph id="8B91C735626EFFCB2DEFB56FFD0C5A74" blockId="40.[151,771,1775,1838]" box="[151,747,1812,1838]" pageId="40" pageNumber="41">
Habitus:
<figureCitation id="1315DBB0626EFFCB2D85B56FFE305A74" box="[253,471,1812,1838]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="40" pageNumber="41">Figs 21E, 21F, 21G</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB0626EFFCB2FEDB56FFD005A74" box="[661,743,1812,1838]" captionStart="FIGURE 24" captionStartId="42.[151,250,1142,1166]" captionTargetBox="[159,1425,461,1114]" captionTargetId="figure-146@42.[151,1436,453,1119]" captionTargetPageId="42" captionText="FIGURE 24. Median lobe of male genitalia for Parena (Crossoglossa) mellea (Chaudoir), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Philippine. B Maluku. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958986" httpUri="https://zenodo.org/record/7958986/files/figure.png" pageId="40" pageNumber="41">Fig. 24</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626EFFCB2DEFB523FA7E5AD5" pageId="40" pageNumber="41" type="reference_group">
<paragraph id="8B91C735626EFFCB2DEFB523FA7E5AD5" blockId="40.[151,1436,1880,1935]" pageId="40" pageNumber="41">
<materialsCitation id="3B46CD68626EFFCB2DEFB523FA725AD5" ID-GBIF-Occurrence="4115871417" collectorName="Andonare Is." country="Papua New Guinea" location="Sumatra" pageId="40" pageNumber="41" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<bibRefCitation id="EFBFBAC4626EFFCB2DEFB523FE845A2A" author="Chaudoir, B. M." box="[151,355,1880,1904]" pageId="40" pageNumber="41" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872: 179</bibRefCitation>
(original:
<taxonomicName id="4C2EBCB6626EFFCB2CB2B523FDB55A2A" ID-CoL="8KWFM" authority="Chaudoir, 1873" box="[458,594,1880,1904]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27626EFFCB2CB2B523FDB55A2A" box="[458,594,1880,1904]" italics="true" pageId="40" pageNumber="41">Crossoglossa</emphasis>
</taxonomicName>
; type locality: Moluques;
<typeStatus id="54957997626EFFCB2E2DB523FC495A2A" box="[853,942,1880,1904]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
in MNHN);
<bibRefCitation id="EFBFBAC4626EFFCB2952B523FAE75A2A" author="Andrewes, H. E." box="[1066,1280,1880,1904]" pageId="40" pageNumber="41" refId="ref109646" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects, pars 18 (carabidae), Carabidae. Calcutta: Government of India Central Publication Branch, xxii + 389 pp." type="book" year="1930">Andrewes, 1930: 257</bibRefCitation>
(
<collectorName id="26DBA2E3626EFFCB2873B523FA685A2A" box="[1291,1423,1880,1904]" pageId="40" pageNumber="41">Andonare Is.</collectorName>
).
<emphasis id="B95A1B27626EFFCB2DEFB50DFEA85AD4" bold="true" box="[151,335,1910,1934]" pageId="40" pageNumber="41">Misidentification</emphasis>
:
<taxonomicName id="4C2EBCB6626EFFCB2C25B50CFD9B5AD5" ID-CoL="4DKQR" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[349,636,1911,1935]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B27626EFFCB2C25B50CFDE55AD4" bold="true" box="[349,514,1911,1934]" italics="true" pageId="40" pageNumber="41">Parena testacea</emphasis>
(Chaudoir)
</taxonomicName>
:
<bibRefCitation id="EFBFBAC4626EFFCB2FFCB50CFCBA5AD5" author="Andrewes, H. E." box="[644,861,1911,1935]" pageId="40" pageNumber="41" pagination="303 - 340" refId="ref109622" refString="Andrewes, H. E. (1929) The Fauna of Sumatrensis. Tijdschrift voor Entomologie, 72, 303 - 340." type="journal article" year="1929">Andrewes, 1929: 314</bibRefCitation>
(
<collectingRegion id="49EA09D7626EFFCB2E13B50CFC225AD5" box="[875,965,1911,1935]" country="Indonesia" name="Sumatera Utara" pageId="40" pageNumber="41">Sumatra</collectingRegion>
);
<bibRefCitation id="EFBFBAC4626EFFCB2EAEB50CFB505AD5" author="Darlington, P. J." box="[982,1207,1911,1935]" pageId="40" pageNumber="41" pagination="1 - 253" refId="ref110388" refString="Darlington, P. J. (1968) The carabid beetles of New Guinea. Part 3. Harpalinae (Continued): Perigonine to Pseudomorphini. Bulletin of the Museum of Comparative Zoology at Harvard College, 137 (1), 1 - 253." type="journal article" year="1968">Darlington, 1968: 139</bibRefCitation>
(
<collectingCountry id="F33987A5626EFFCB29BCB50CFA695AD5" box="[1220,1422,1911,1935]" name="Papua New Guinea" pageId="40" pageNumber="41">Papua New Guinea</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626EFFCA2DEFB5C6FCF95D8F" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8B91C735626EFFCA2DEFB5C6FCF95D8F" blockId="40.[151,1436,1980,2043]" lastBlockId="41.[151,1437,150,1977]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis id="B95A1B27626EFFCB2DEFB5C6FE5C5A8D" bold="true" box="[151,443,1981,2007]" pageId="40" pageNumber="41">Type material examined</emphasis>
.
<emphasis id="B95A1B27626EFFCB2CB2B5C7FC575A8D" bold="true" box="[458,944,1980,2007]" pageId="40" pageNumber="41">
<taxonomicName id="4C2EBCB6626EFFCB2CB2B5C7FCD25A8D" authority="Chaudoir" authorityName="Chaudoir" authorityYear="1873" box="[458,821,1980,2007]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626EFFCB2CB2B5C7FD515A8D" bold="true" box="[458,694,1980,2007]" italics="true" pageId="40" pageNumber="41">Crossoglossa mellea</emphasis>
Chaudoir
</taxonomicName>
:
<typeStatus id="54957997626EFFCB2E3BB5C6FC575A8D" box="[835,944,1981,2007]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
(MNHN,
<figureCitation id="1315DBB0626EFFCB295FB5C6FB6A5A8D" box="[1063,1165,1981,2007]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="40" pageNumber="41">Fig. 21E</figureCitation>
): female, body length =
<quantity id="4CD66AD0626EFFCB2DEFB59AFF1B5AA1" box="[151,252,2017,2043]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="40" pageNumber="41" unit="mm" value="11.3">11.3 mm</quantity>
, pin mounted, &quot;Ex Musaeo / Chaudoir&quot;, &quot;
<emphasis id="B95A1B27626EFFCB2FB6B59AFB955AA1" box="[718,1138,2017,2043]" italics="true" pageId="40" pageNumber="41">
<taxonomicName id="4C2EBCB6626EFFCB2FB6B59AFCFE5AA1" authorityName="Chaudoir" authorityYear="1873" box="[718,793,2017,2043]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mellea">mellea</taxonomicName>
/ Chaud. /
<collectingCountry id="F33987A5626EFFCB2EEDB59AFBE45AA1" box="[917,1027,2017,2043]" name="Indonesia" pageId="40" pageNumber="41">Moluques</collectingCountry>
/ Lorquin
</emphasis>
&quot; [ex Chaudoir's box label, pinned under specimen now], &quot;MUSEUM PARIS / 1952 / Coll. R. OBERTHÜR&quot;, &quot;
<typeStatus id="54957997626FFFCA292EB2ECFB095DEB" box="[1110,1262,151,177]" pageId="41" pageNumber="42" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C2EBCB6626FFFCA2870B2ECFF055D8F" authorityName="Chaudoir" authorityYear="1873" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">Crossoglossa mellea</taxonomicName>
/
<bibRefCitation id="EFBFBAC4626FFFCA2D80B2C0FE425D8F" author="Chaudoir, B. M." box="[248,421,187,213]" pageId="41" pageNumber="42" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872</bibRefCitation>
/ det. SHI H.L. 2011&quot; [red label].
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626FFFCA2DBFB2A4FC415C3F" pageId="41" pageNumber="42" type="discussion">
<paragraph id="8B91C735626FFFCA2DBFB2A4FC415C3F" blockId="41.[151,1437,150,1977]" pageId="41" pageNumber="42">
<emphasis id="B95A1B27626FFFCA2DBFB2A4FE955DA3" bold="true" box="[199,370,223,249]" pageId="41" pageNumber="42">Notes on types</emphasis>
. This species was originally described from
<specimenCount id="9D280CBC626FFFCA2E1AB2A4FC3A5DA3" box="[866,989,223,249]" pageId="41" pageNumber="42" type="female">one female</specimenCount>
collected by
<collectorName id="26DBA2E3626FFFCA2909B2A4FB2C5DA3" box="[1137,1227,223,249]" pageId="41" pageNumber="42">Lorquin</collectorName>
from &quot;
<collectingCountry id="F33987A5626FFFCA286EB2A4FA775DA3" box="[1302,1424,223,249]" name="Indonesia" pageId="41" pageNumber="42">Moluques</collectingCountry>
&quot;, perfectly in accord with the examined
<typeStatus id="54957997626FFFCA2F33B378FD4A5C47" box="[587,685,259,285]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
in MNHN (
<figureCitation id="1315DBB0626FFFCA2E40B378FC795C47" box="[824,926,259,285]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="41" pageNumber="42">Fig. 21E</figureCitation>
). There is another male (
<figureCitation id="1315DBB0626FFFCA29CFB378FAFE5C47" box="[1207,1305,259,285]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="41" pageNumber="42">Fig. 21F</figureCitation>
) in MNHN labeled &quot;
<collectingCountry id="F33987A5626FFFCA2D8FB35CFE8C5C1B" box="[247,363,295,321]" name="Indonesia" pageId="41" pageNumber="42">Moluques</collectingCountry>
Lorquin&quot; from the ex-collection of Mniszech. This one seems to be collected together with the
<typeStatus id="54957997626FFFCA2DEFB330FF1B5C3F" box="[151,252,331,357]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
, but not known to
<location id="8EF191EE626FFFCA2CB5B330FDD25C3F" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2CB5B330FDD25C3F" box="[461,565,331,357]" country="Indonesia" name="Chaudoir" pageId="41" pageNumber="42">Chaudoir</location>
when the species was described.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626FFFCA2DBFB314FB395EDB" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="8B91C735626FFFCA2DBFB314FB395EDB" blockId="41.[151,1437,150,1977]" pageId="41" pageNumber="42">
<emphasis id="B95A1B27626FFFCA2DBFB314FDF85CD2" bold="true" box="[199,543,366,393]" pageId="41" pageNumber="42">Non-type material examined</emphasis>
.
<emphasis id="B95A1B27626FFFCA2F57B314FD695CD3" bold="true" box="[559,654,367,393]" pageId="41" pageNumber="42">Maluku</emphasis>
:
<materialsCitation id="3B46CD68626FFFCA2FE7B314FDBF5CAF" ID-GBIF-Occurrence="4115871540" collectorName="H. E. Andrewes Coll." location="Andonare" pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC626FFFCA2FE7B314FD0C5CD3" box="[671,747,367,393]" pageId="41" pageNumber="42" type="male">1 male</specimenCount>
(MNHN), &quot;Moluques Lorquin&quot; [hand written], &quot;Ex Musaeo Mniszech&quot;, &quot;MUSEUM PARIS 1952, Coll. R. OBERTHUR&quot; &lt;
<figureCitation id="1315DBB0626FFFCA2E20B3E8FC5C5CF7" box="[856,955,403,429]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="41" pageNumber="42">Figs 21F</figureCitation>
,
<figureCitation id="1315DBB0626FFFCA2EBEB3E8FC1F5CF7" box="[966,1016,403,429]" captionStart="FIGURE 24" captionStartId="42.[151,250,1142,1166]" captionTargetBox="[159,1425,461,1114]" captionTargetId="figure-146@42.[151,1436,453,1119]" captionTargetPageId="42" captionText="FIGURE 24. Median lobe of male genitalia for Parena (Crossoglossa) mellea (Chaudoir), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Philippine. B Maluku. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958986" httpUri="https://zenodo.org/record/7958986/files/figure.png" pageId="41" pageNumber="42">24B</figureCitation>
&gt;.
<emphasis id="B95A1B27626FFFCA296AB3E9FB745CF6" bold="true" box="[1042,1171,402,428]" pageId="41" pageNumber="42">
<location id="8EF191EE626FFFCA296AB3E9FB695CF6" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA296AB3E9FB695CF6" box="[1042,1166,402,428]" name="Andonare" pageId="41" pageNumber="42">Andonare</location>
:
</emphasis>
1 ex (NHML), &quot;Laboenarang / Andonare 2-4000' / Doherty XI&quot;; &quot;
<emphasis id="B95A1B27626FFFCA2F06B3CCFC155C8B" box="[638,1010,439,465]" italics="true" pageId="41" pageNumber="42">
Phloedromius /
<taxonomicName id="4C2EBCB6626FFFCA2E49B3CCFC9B5C8B" authorityName="Chaudoir" authorityYear="1873" box="[817,892,439,465]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">mellea</taxonomicName>
/ Chaud. /
</emphasis>
Compared with /
<typeStatus id="54957997626FFFCA29C5B3CCFB0A5C8B" box="[1213,1261,439,465]" pageId="41" pageNumber="42">type</typeStatus>
H.E.A.&quot;; &quot;
<collectorName id="26DBA2E3626FFFCA281FB3CCFEAD5CAF" pageId="41" pageNumber="42">H.E. Andrewes Coll.</collectorName>
/ B.M. 1945-97&quot;.
<emphasis id="B95A1B27626FFFCA2F58B3A0FDBF5CAF" bold="true" box="[544,600,475,501]" pageId="41" pageNumber="42">Java</emphasis>
</materialsCitation>
:
<materialsCitation id="3B46CD68626FFFCA2F11B3A0FF0E5F43" ID-GBIF-Occurrence="4115871560" location="Java" municipality="Banjoewangl" pageId="41" pageNumber="42" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626FFFCA2F11B3A0FD2B5CAF" box="[617,716,475,501]" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(NNML), &quot;
<collectingMunicipality id="6BF55D4F626FFFCA2E2AB3A0FC0B5CAF" box="[850,1004,475,501]" pageId="41" pageNumber="42">Banjoewangl</collectingMunicipality>
,
<location id="8EF191EE626FFFCA2E82B3A0FBC95CAF" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2E82B3A0FBC95CAF" box="[1018,1070,475,501]" municipality="Banjoewangl" name="Java" pageId="41" pageNumber="42">Java</location>
, 1911, Mac Gillavry&quot; [=Banyuwangi]
</materialsCitation>
.
<materialsCitation id="3B46CD68626FFFCA2D8AB384FBE35F43" ID-GBIF-Occurrence="4115871338" box="[242,1028,511,537]" pageId="41" pageNumber="42" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D280CBC626FFFCA2D8AB384FEAC5F43" box="[242,331,511,537]" pageId="41" pageNumber="42" type="male">2 males</specimenCount>
,
<specimenCount id="9D280CBC626FFFCA2C2DB384FE515F43" box="[341,438,511,537]" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(NNML), &quot;Banjoewangl, Java, 1910, Mac Gillavry&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68626FFFCA2976B384FEB05F67" ID-GBIF-Occurrence="4115871306" pageId="41" pageNumber="42" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626FFFCA2976B384FB895F43" box="[1038,1134,511,537]" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(NNML), &quot;Java, Drescher, 12.1913, Djoija&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68626FFFCA2C1AB058FBCE5F3A" ID-GBIF-Occurrence="4115871588" collectorName="P. H. V Doesburg &amp; C. J. Louwerens" location="Museum Leiden" municipality="Semarang-Java" pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC626FFFCA2C1AB058FE4A5F67" box="[354,429,547,573]" pageId="41" pageNumber="42" type="male">1 male</specimenCount>
(NNML), &quot;
<collectingMunicipality id="6BF55D4F626FFFCA2F4AB058FD035F67" box="[562,740,547,573]" pageId="41" pageNumber="42">Semarang-Java</collectingMunicipality>
,
<collectorName id="26DBA2E3626FFFCA2F97B058FC4A5F67" box="[751,941,547,573]" pageId="41" pageNumber="42">P.H.V Doesburg</collectorName>
&quot;, &quot;
<location id="8EF191EE626FFFCA2EB0B058FB605F67" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2EB0B058FB605F67" box="[968,1159,547,573]" municipality="Semarang-Java" name="Museum Leiden" pageId="41" pageNumber="42">Museum Leiden</location>
, ex. collection
<collectorName id="26DBA2E3626FFFCA284AB058FF0E5F3B" pageId="41" pageNumber="42">C.J. Louwerens</collectorName>
, rec. 1979&quot;, &quot;
<taxonomicName id="4C2EBCB6626FFFCA2CFEB03CFCC85F3B" authority="Chaud., E. B. Britton" authorityName="Chaud., E. B. Britton" box="[390,815,583,609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="testacea">Parena testacea Chaud., E.B. Britton</taxonomicName>
det. 1947&quot;.
<emphasis id="B95A1B27626FFFCA2EB9B03DFBCE5F3A" bold="true" box="[961,1065,582,608]" pageId="41" pageNumber="42">Sumatra</emphasis>
</materialsCitation>
:
<materialsCitation id="3B46CD68626FFFCA2942B03CFEDA5FF3" ID-GBIF-Occurrence="4115871543" collectingDate="1920-04-08" collectorName="B. H. Klynstra" location="Medan" municipality="Corporaal" pageId="41" pageNumber="42" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626FFFCA2942B03CFB795F3B" box="[1082,1182,583,609]" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(NNHL), &quot;
<collectingMunicipality id="6BF55D4F626FFFCA285AB03CFA7E5F3B" box="[1314,1433,583,609]" pageId="41" pageNumber="42">Corporaal</collectingMunicipality>
,
<location id="8EF191EE626FFFCA2DEFB010FF0F5FDF" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2DEFB010FF0F5FDF" box="[151,232,619,645]" municipality="Corporaal" name="Medan" pageId="41" pageNumber="42">Medan</location>
,
<date id="FF90E1F5626FFFCA2D8BB010FEB85FDF" box="[243,351,619,645]" pageId="41" pageNumber="42" value="1920-04-08">
<collectingDate id="EFD4181D626FFFCA2D8BB010FEB85FDF" box="[243,351,619,645]" pageId="41" pageNumber="42" value="1920-04-08">8-4-1920</collectingDate>
</date>
&quot;, &quot;Museum Leiden, Ex coll.
<collectorName id="26DBA2E3626FFFCA2FDCB010FCA05FDF" box="[676,839,619,645]" pageId="41" pageNumber="42">B.H. Klynstra</collectorName>
&quot;, &quot;
<taxonomicName id="4C2EBCB6626FFFCA2E1AB010FAFB5FDF" authority="Chaud. H. E. Andrewes" box="[866,1308,619,645]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="testacea">Parena testacea Chaud. H.E. Andrewes</taxonomicName>
det.&quot;, &quot;Par. Chd. testacea&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68626FFFCA2C31B0F4FACA5FF3" ID-GBIF-Occurrence="4115871518" box="[329,1325,655,681]" collectingDate="1921-12" collectorName="E. Jacobson." elevation="920" location="Sumatra" municipality="Fort de Kock." pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC626FFFCA2C31B0F4FE715FF3" box="[329,406,655,681]" pageId="41" pageNumber="42" type="male">1 male</specimenCount>
(NNHL), &quot;
<collectingMunicipality id="6BF55D4F626FFFCA2F6FB0F4FD5B5FF3" box="[535,700,655,681]" pageId="41" pageNumber="42">Fort de Kock.</collectingMunicipality>
, (
<location id="8EF191EE626FFFCA2FA9B0F4FCD55FF3" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2FA9B0F4FCD55FF3" box="[721,818,655,681]" municipality="Fort de Kock." name="Sumatra" pageId="41" pageNumber="42">Sumatra</location>
)
<quantity id="4CD66AD0626FFFCA2E3AB0F4FC6B5FF3" box="[834,908,655,681]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="41" pageNumber="42" unit="m" value="920.0">
<elevation id="00032006626FFFCA2E3AB0F4FC6B5FF3" box="[834,908,655,681]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="41" pageNumber="42" unit="m" value="920.0">920 m</elevation>
</quantity>
,
<date id="FF90E1F5626FFFCA2EE2B0F4FBB75FF3" box="[922,1104,655,681]" pageId="41" pageNumber="42" value="1921-12">
<collectingDate id="EFD4181D626FFFCA2EE2B0F4FBB75FF3" box="[922,1104,655,681]" pageId="41" pageNumber="42" value="1921-12">December 1921</collectingDate>
</date>
, leg.
<collectorName id="26DBA2E3626FFFCA29E8B0F4FAC25FF3" box="[1168,1317,655,681]" pageId="41" pageNumber="42">E. Jacobson.</collectorName>
&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68626FFFCA2841B0F4FABC5F97" ID-GBIF-Occurrence="4115871458" collectorName="E. Jacobson. &amp; H. E. Andrewes Coll." location="Sumatra" municipality="Fort de Kock" pageId="41" pageNumber="42" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626FFFCA2841B0F4FA7B5FF3" box="[1337,1436,655,681]" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(NHML), &quot;
<collectingMunicipality id="6BF55D4F626FFFCA2C6BB0C8FE575F97" box="[275,432,691,717]" pageId="41" pageNumber="42">Fort de Kock</collectingMunicipality>
, (
<location id="8EF191EE626FFFCA2CBCB0C8FDC25F97" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2CBCB0C8FDC25F97" box="[452,549,691,717]" municipality="Fort de Kock" name="Sumatra" pageId="41" pageNumber="42">Sumatra</location>
) 920 M, 1924, leg.
<collectorName id="26DBA2E3626FFFCA2E7FB0C8FC7C5F97" box="[775,923,691,717]" pageId="41" pageNumber="42">E. Jacobson.</collectorName>
&quot;, &quot;
<collectorName id="26DBA2E3626FFFCA2EC0B0C8FB415F97" box="[952,1190,691,717]" pageId="41" pageNumber="42">H.E. Andrewes Coll.</collectorName>
B.M. 1945-97&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68626FFFCA281FB0C8FEE45E4F" ID-GBIF-Occurrence="4115871361" collectorName="H. E. Andrewes Coll." location="Sumatra" pageId="41" pageNumber="42" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626FFFCA281FB0C8FF2A5FAB" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(NHML), &quot;
<location id="8EF191EE626FFFCA2C2BB0ACFE5D5FAB" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2C2BB0ACFE5D5FAB" box="[339,442,727,753]" name="Sumatra" pageId="41" pageNumber="42">Sumatra</location>
&quot;, &quot;J.T., 18.3.84&quot;, &quot;
<collectorName id="26DBA2E3626FFFCA2FF2B0ACFCF95FAB" box="[650,798,727,753]" pageId="41" pageNumber="42">Coll. Kraatz</collectorName>
&quot;, &quot;
<collectorName id="26DBA2E3626FFFCA2E43B0ACFBAB5FAB" box="[827,1100,727,753]" pageId="41" pageNumber="42">Ex. Deutsch. Ent. Mus.</collectorName>
&quot;, &quot;
<collectorName id="26DBA2E3626FFFCA2914B0ACFABD5FAB" box="[1132,1370,727,753]" pageId="41" pageNumber="42">H.E. Andrewes Coll.</collectorName>
B.M. 1945-97&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68626FFFCA2C69B080FA675E4F" ID-GBIF-Occurrence="4115871529" box="[273,1408,762,789]" collectorName="H. E. Andrewes Coll." location="Ex Mus. Stockholm" municipality="Medan Mjob." pageId="41" pageNumber="42" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626FFFCA2C69B080FE915E4F" box="[273,374,763,789]" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(NHML), &quot;
<collectingMunicipality id="6BF55D4F626FFFCA2C87B080FD425E4F" box="[511,677,763,789]" pageId="41" pageNumber="42">Medan Mjob.</collectingMunicipality>
&quot;, &quot;
<location id="8EF191EE626FFFCA2FBCB080FC535E4F" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2FBCB080FC535E4F" box="[708,948,763,789]" municipality="Medan Mjob." name="Ex Mus. Stockholm" pageId="41" pageNumber="42">Ex Mus. Stockholm</location>
&quot;, &quot;
<collectorName id="26DBA2E3626FFFCA2EABB080FB215E4F" box="[979,1222,763,789]" pageId="41" pageNumber="42">H.E. Andrewes Coll.</collectorName>
B.M. 1945-97&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68626FFFCA28F6B080FD9D5E63" ID-GBIF-Occurrence="4115871311" collectorName="Coll. Kraatz" pageId="41" pageNumber="42" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626FFFCA28F6B080FF045E63" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(SDEI), &quot;Sumatra&quot;, &quot;
<collectorName id="26DBA2E3626FFFCA2CA5B164FD945E63" box="[477,627,799,825]" pageId="41" pageNumber="42">Coll. Kraatz</collectorName>
&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68626FFFCA2FF0B164FDBF5E06" ID-GBIF-Occurrence="4115871507" collectingDate="1921-12" collectorName="The" country="Philippines" county="Wai Lima" location="Lampongs" municipality="Sum." pageId="41" pageNumber="42" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626FFFCA2FF0B164FD0B5E63" box="[648,748,799,825]" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(SDEI), &quot;
<collectingCounty id="62F0BFB9626FFFCA2E27B164FC325E63" box="[863,981,799,825]" pageId="41" pageNumber="42">Wai Lima</collectingCounty>
Z.
<collectingMunicipality id="6BF55D4F626FFFCA2979B164FBD95E63" box="[1025,1086,799,825]" pageId="41" pageNumber="42">Sum.</collectingMunicipality>
,
<location id="8EF191EE626FFFCA2934B164FB225E63" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2934B164FB225E63" box="[1100,1221,799,825]" country="Philippines" county="Wai Lima" municipality="Sum." name="Lampongs" pageId="41" pageNumber="42">Lampongs</location>
, Karny &amp; Siebers, XI.
<date id="FF90E1F5626FFFCA2DC4B138FEC25E07" box="[188,293,835,861]" pageId="41" pageNumber="42" value="1921-12">
<collectingDate id="EFD4181D626FFFCA2DC4B138FEC25E07" box="[188,293,835,861]" pageId="41" pageNumber="42" value="1921-12">XII.1921</collectingDate>
</date>
. No261&quot;.
<emphasis id="B95A1B27626FFFCA2CE2B138FDBF5E06" bold="true" box="[410,600,834,861]" pageId="41" pageNumber="42">
<collectorName id="26DBA2E3626FFFCA2CE2B138FE2E5E07" box="[410,457,835,861]" pageId="41" pageNumber="42">The</collectorName>
<collectingCountry id="F33987A5626FFFCA2CAAB139FDBF5E06" box="[466,600,834,860]" name="Philippines" pageId="41" pageNumber="42">Philippines</collectingCountry>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B46CD68626FFFCA2F10B138FEEE5EDB" ID-GBIF-Occurrence="4115871477" collectingDate="2014-08" country="Philippines" location="Pola" municipality="Filippine" pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1" stateProvince="Ifugao">
<specimenCount id="9D280CBC626FFFCA2F10B138FD515E07" box="[616,694,835,861]" pageId="41" pageNumber="42" type="male">1 male</specimenCount>
(CRS), &quot;
<collectingMunicipality id="6BF55D4F626FFFCA2E67B138FC605E07" box="[799,903,835,861]" pageId="41" pageNumber="42">Filippine</collectingMunicipality>
—N Luzon,
<collectingRegion id="49EA09D7626FFFCA2961B138FB825E07" box="[1049,1125,835,861]" country="Philippines" name="Ifugao" pageId="41" pageNumber="42">Ifugao</collectingRegion>
,
<location id="8EF191EE626FFFCA290AB138FB435E07" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA290AB138FB435E07" box="[1138,1188,835,861]" country="Philippines" municipality="Filippine" name="Pola" pageId="41" pageNumber="42" stateProvince="Ifugao">Pola</location>
<date id="FF90E1F5626FFFCA29D5B138FAF85E07" box="[1197,1311,835,861]" pageId="41" pageNumber="42" value="2014-08">
<collectingDate id="EFD4181D626FFFCA29D5B138FAF85E07" box="[1197,1311,835,861]" pageId="41" pageNumber="42" value="2014-08">VIII.2014</collectingDate>
</date>
&quot; &lt;
<figureCitation id="1315DBB0626FFFCA2838B138FA705E07" box="[1344,1431,835,861]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="41" pageNumber="42">Figs 8F</figureCitation>
,
<figureCitation id="1315DBB0626FFFCA2DEFB11CFF205EDB" box="[151,199,871,897]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. AH Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. IL Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="41" pageNumber="42">14G</figureCitation>
,
<figureCitation id="1315DBB0626FFFCA2DADB11CFEEE5EDB" box="[213,265,871,897]" captionStart="FIGURE 24" captionStartId="42.[151,250,1142,1166]" captionTargetBox="[159,1425,461,1114]" captionTargetId="figure-146@42.[151,1436,453,1119]" captionTargetPageId="42" captionText="FIGURE 24. Median lobe of male genitalia for Parena (Crossoglossa) mellea (Chaudoir), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Philippine. B Maluku. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958986" httpUri="https://zenodo.org/record/7958986/files/figure.png" pageId="41" pageNumber="42">24A</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD68626FFFCA2C5BB11CFB205EDB" ID-GBIF-Occurrence="4115871450" box="[291,1223,870,897]" collectingDate="2015-07" country="Philippines" county="Philippines" location="Subic" municipality="Luzon" pageId="41" pageNumber="42" specimenCount="1" specimenCount-female="1" stateProvince="Zambales">
<specimenCount id="9D280CBC626FFFCA2C5BB11CFE635EDB" box="[291,388,871,897]" pageId="41" pageNumber="42" type="female">1 female</specimenCount>
(CRS), &quot;
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W
<collectingMunicipality id="6BF55D4F626FFFCA2FEAB11CFD3B5EDB" box="[658,732,871,897]" pageId="41" pageNumber="42">Luzon</collectingMunicipality>
,
<collectingRegion id="49EA09D7626FFFCA2F90B11CFCBF5EDB" box="[744,856,871,897]" country="Philippines" name="Zambales" pageId="41" pageNumber="42">Zambales</collectingRegion>
,
<location id="8EF191EE626FFFCA2E1AB11CFC455EDB" LSID="urn:lsid:plazi:treatment:03877623626EFFC92DEFB48BFEE75CF7:8EF191EE626FFFCA2E1AB11CFC455EDB" box="[866,930,871,897]" country="Philippines" county="Philippines" municipality="Luzon" name="Subic" pageId="41" pageNumber="42" stateProvince="Zambales">Subic</location>
<date id="FF90E1F5626FFFCA2ED1B11DFBF55EDB" box="[937,1042,870,897]" pageId="41" pageNumber="42" value="2015-07">
<collectingDate id="EFD4181D626FFFCA2ED1B11DFBF55EDB" box="[937,1042,870,897]" pageId="41" pageNumber="42" value="2015-07">VII.2015</collectingDate>
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&quot; &lt;
<figureCitation id="1315DBB0626FFFCA2948B11CFB6C5EDB" box="[1072,1163,871,897]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="41" pageNumber="42">Figs 8C</figureCitation>
,
<figureCitation id="1315DBB0626FFFCA29EFB11CFB205EDB" box="[1175,1223,871,897]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="41" pageNumber="42">21G</figureCitation>
</materialsCitation>
&gt;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626FFFCA2DBFB1F1FC6F58C7" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="8B91C735626FFFCA2DBFB1F1FD6C5EB7" blockId="41.[151,1437,150,1977]" pageId="41" pageNumber="42">
<emphasis id="B95A1B27626FFFCA2DBFB1F1FE815EFE" bold="true" box="[199,358,906,932]" pageId="41" pageNumber="42">Comparisons</emphasis>
.
<taxonomicName id="4C2EBCB6626FFFCA2C0BB1F0FE3E5EFF" authorityName="Chaudoir" authorityYear="1873" box="[371,473,907,933]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2C0BB1F0FE3E5EFF" box="[371,473,907,933]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
is distinguishable from the other four species in this species group by the combination of: elytra with sutural angles shortly pointed, forming small denticles; outer apical angles almost rounded; elytra without black patch; tibiae yellowish brown.
</paragraph>
<paragraph id="8B91C735626FFFCA2DBFB18CFE9759B3" blockId="41.[151,1437,150,1977]" pageId="41" pageNumber="42">
This species is similar to
<taxonomicName id="4C2EBCB6626FFFCA2C9FB18CFDB8594A" box="[487,607,1015,1040]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B27626FFFCA2C9FB18CFDB8594A" box="[487,607,1015,1040]" italics="true" pageId="41" pageNumber="42">P. testacea</emphasis>
</taxonomicName>
, especially for those specimens from Sumatra and the
<collectingCountry id="F33987A5626FFFCA29B3B18CFAAB594B" box="[1227,1356,1015,1041]" name="Philippines" pageId="41" pageNumber="42">Philippines</collectingCountry>
, which have the same antennal color as
<taxonomicName id="4C2EBCB6626FFFCA2C86B660FD92596E" box="[510,629,1051,1076]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B27626FFFCA2C86B660FD92596E" box="[510,629,1051,1076]" italics="true" pageId="41" pageNumber="42">P. testacea</emphasis>
</taxonomicName>
. These two species are distinguishable by: (1) elytral sutural angles only slightly pointed, forming small denticles in
<taxonomicName id="4C2EBCB6626FFFCA2FF3B644FD145903" authorityName="Chaudoir" authorityYear="1873" box="[651,755,1087,1113]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2FF3B644FD145903" box="[651,755,1087,1113]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
, but sharply pointed forming long spines in
<taxonomicName id="4C2EBCB6626FFFCA298AB644FA8C5902" box="[1266,1387,1087,1112]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B27626FFFCA298AB644FA8C5902" box="[1266,1387,1087,1112]" italics="true" pageId="41" pageNumber="42">P. testacea</emphasis>
</taxonomicName>
; (2) elytral outer apical angles less distinct in
<taxonomicName id="4C2EBCB6626FFFCA2F16B618FD325927" authorityName="Chaudoir" authorityYear="1873" box="[622,725,1123,1149]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2F16B618FD325927" box="[622,725,1123,1149]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
; (3) in
<taxonomicName id="4C2EBCB6626FFFCA2E53B618FC745927" authorityName="Chaudoir" authorityYear="1873" box="[811,915,1123,1149]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2E53B618FC745927" box="[811,915,1123,1149]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
, elytra intervals with punctures much sparser, and stria with puncture rows more clearly defined; (4) in
<taxonomicName id="4C2EBCB6626FFFCA2E6AB6FCFC9F59FB" authorityName="Chaudoir" authorityYear="1873" box="[786,888,1159,1185]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2E6AB6FCFC9F59FB" box="[786,888,1159,1185]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
, apex of abdominal sternite VII notched in males (
<figureCitation id="1315DBB0626FFFCA2DE7B6D0FF13599F" box="[159,244,1195,1221]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="41" pageNumber="42">Fig. 8F</figureCitation>
), and projected in females (
<figureCitation id="1315DBB0626FFFCA2F5EB6D0FD9A599F" box="[550,637,1195,1221]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="41" pageNumber="42">Fig. 8C</figureCitation>
), but straight in both sexes of
<taxonomicName id="4C2EBCB6626FFFCA2EB3B6D0FBA6599E" box="[971,1089,1195,1220]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B27626FFFCA2EB3B6D0FBA6599E" box="[971,1089,1195,1220]" italics="true" pageId="41" pageNumber="42">P. testacea</emphasis>
</taxonomicName>
; (5) apical lamella of aedeagus longer in
<taxonomicName id="4C2EBCB6626FFFCA2C7AB6B4FE8E59B3" authorityName="Chaudoir" authorityYear="1873" box="[258,361,1231,1257]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2C7AB6B4FE8E59B3" box="[258,361,1231,1257]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B91C735626FFFCA2DBFB688FC6F58C7" blockId="41.[151,1437,150,1977]" pageId="41" pageNumber="42">
Some specimens of
<taxonomicName id="4C2EBCB6626FFFCA2CC8B688FDFE5857" authorityName="Chaudoir" authorityYear="1873" box="[432,537,1267,1293]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2CC8B688FDFE5857" box="[432,537,1267,1293]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
are similar in color to
<taxonomicName id="4C2EBCB6626FFFCA2E51B688FC385857" authorityName="Kirschenhofer. D" authorityYear="2006" box="[809,991,1267,1293]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="B95A1B27626FFFCA2E51B688FC385857" box="[809,991,1267,1293]" italics="true" pageId="41" pageNumber="42">P. sulawesiensis</emphasis>
</taxonomicName>
, especially for those from the
<collectingCountry id="F33987A5626FFFCA283DB688FF0D586B" name="Indonesia" pageId="41" pageNumber="42">Maluku Islands</collectingCountry>
. These two species are different in (1) elytral sutural angles only slightly pointed forming small denticles in
<taxonomicName id="4C2EBCB6626FFFCA2DEFB740FF19580F" authorityName="Chaudoir" authorityYear="1873" box="[151,254,1339,1365]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2DEFB740FF19580F" box="[151,254,1339,1365]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
, but sharply pointed forming long spines in
<taxonomicName id="4C2EBCB6626FFFCA2F8BB740FC40580F" authorityName="Kirschenhofer. D" authorityYear="2006" box="[755,935,1339,1365]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="B95A1B27626FFFCA2F8BB740FC40580F" box="[755,935,1339,1365]" italics="true" pageId="41" pageNumber="42">P. sulawesiensis</emphasis>
</taxonomicName>
; (2) in
<taxonomicName id="4C2EBCB6626FFFCA2E84B740FB84580F" authorityName="Chaudoir" authorityYear="1873" box="[1020,1123,1339,1365]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2E84B740FB84580F" box="[1020,1123,1339,1365]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
, apex of abdominal sternite VII more distinctly notched in males, and less projected in females than in
<taxonomicName id="4C2EBCB6626FFFCA2E89B724FB425823" authorityName="Kirschenhofer. D" authorityYear="2006" box="[1009,1189,1375,1401]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="B95A1B27626FFFCA2E89B724FB425823" box="[1009,1189,1375,1401]" italics="true" pageId="41" pageNumber="42">P. sulawesiensis</emphasis>
</taxonomicName>
; (3) apical lamella of aedeagus ovate in
<taxonomicName id="4C2EBCB6626FFFCA2C1DB7F8FE2B58C7" authorityName="Chaudoir" authorityYear="1873" box="[357,460,1411,1437]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626FFFCA2C1DB7F8FE2B58C7" box="[357,460,1411,1437]" italics="true" pageId="41" pageNumber="42">P. mellea</emphasis>
</taxonomicName>
, but subrectangular in
<taxonomicName id="4C2EBCB6626FFFCA2FB5B7F8FC6758C7" authorityName="Kirschenhofer. D" authorityYear="2006" box="[717,896,1411,1437]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="B95A1B27626FFFCA2FB5B7F8FC6758C7" box="[717,896,1411,1437]" italics="true" pageId="41" pageNumber="42">P. sulawesiensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626FFFCA2DBFB7DDFD655AE3" pageId="41" pageNumber="42" type="description">
<paragraph id="8B91C735626FFFCA2DBFB7DDFD655AE3" blockId="41.[151,1437,150,1977]" pageId="41" pageNumber="42">
<emphasis id="B95A1B27626FFFCA2DBFB7DDFEB6589A" bold="true" box="[199,337,1446,1472]" pageId="41" pageNumber="42">Description</emphasis>
. Body length
<quantity id="4CD66AD0626FFFCA2C8DB7DDFD68589B" box="[501,655,1446,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.005" metricValueMax="1.06" metricValueMin="0.95" pageId="41" pageNumber="42" unit="mm" value="10.05" valueMax="10.6" valueMin="9.5">9.510.6 mm</quantity>
. Dorsum pale yellow; antennomere 1 yellow, antennomeres 2, 3 and basal third of 4 reddish yellow to dark brown, remainder of antennae black; apices of mandibles black; terminal labial palpomere black, terminal and penultimate maxillary palpomeres black; tarsomeres black, in strong contrast with yellowish brown tibiae; venter pale yellow. Pronotum strongly transverse, PW/PL = 1.601.69, much wider than head, PW/HW = 1.101.23; widest at anterior third, lateral explanations very wide. Elytra without microsculpture or with very faint isodiametric microsculpture near apices; striae not incised, replaced by rows of very fine punctures; intervals completely flat, with punctures sparser than in striae; disc depressed near middle of intervals 3 to 6, depressions large and shallow; apical truncation distinct, straight or slightly arched; outer apical angles almost rounded, not forming distinct angles; sutural angles slightly pointed, forming short denticles. Apex of abdominal sternite VII notched in males (
<figureCitation id="1315DBB0626FFFCA2C95B490FDA45A5F" box="[493,579,1771,1797]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="41" pageNumber="42">Fig. 8F</figureCitation>
), weakly projected at middle in females (
<figureCitation id="1315DBB0626FFFCA296DB490FB895A5F" box="[1045,1134,1771,1797]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="41" pageNumber="42">Fig. 8C</figureCitation>
). Median lobe of aedeagus with dorsal margin curved before apical orifice, and then narrowed to apex; apical lamella ovate, weakly bent to dorsum, length slightly greater than its basal width, apex rounded. Endophallus densely and heavily scaled on basal sheath, base of apical sheath ovate in dorsal view; squamate sac hardly divided, on basal fifth of median lobe, right or right-ventral to squamate sheath (
<figureCitation id="1315DBB0626FFFCA2F4EB500FD6B5ACF" box="[566,652,1915,1941]" captionStart="FIGURE 24" captionStartId="42.[151,250,1142,1166]" captionTargetBox="[159,1425,461,1114]" captionTargetId="figure-146@42.[151,1436,453,1119]" captionTargetPageId="42" captionText="FIGURE 24. Median lobe of male genitalia for Parena (Crossoglossa) mellea (Chaudoir), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Philippine. B Maluku. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958986" httpUri="https://zenodo.org/record/7958986/files/figure.png" pageId="41" pageNumber="42">Fig. 24</figureCitation>
). Gonocoxite II of ovipositor dichotomous, inner and outer branches each with two short ensiform setae apically.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626CFFC92DBFB2EDFEE75CF7" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8B91C735626CFFC92DBFB2EDFF175D8F" blockId="42.[151,1437,150,429]" pageId="42" pageNumber="43">
<emphasis id="B95A1B27626CFFC92DBFB2EDFEBE5DEA" bold="true" box="[199,345,150,176]" pageId="42" pageNumber="43">Distribution</emphasis>
(
<figureCitation id="1315DBB0626CFFC92C10B2ECFE545DEB" box="[360,435,151,177]" captionStart="MAP 4" captionStartId="36.[152,211,1895,1919]" captionTargetBox="[158,1430,1002,1871]" captionTargetId="figure-71@36.[158,1430,1002,1871]" captionTargetPageId="36" captionText="MAP 4. Distributions for species in the Parena (Crossoglossa) testacea group." figureDoi="http://doi.org/10.5281/zenodo.7963663" httpUri="https://zenodo.org/record/7963663/files/figure.png" pageId="42" pageNumber="43">Map 4</figureCitation>
, blue).
<collectingCountry id="F33987A5626CFFC92F7EB2ECFD525DEB" box="[518,693,151,177]" name="Indonesia" pageId="42" pageNumber="43">Maluku Islands</collectingCountry>
, Adonara Island,
<collectingCountry id="F33987A5626CFFC92E00B2ECFBB55DEB" box="[888,1106,151,177]" name="Papua New Guinea" pageId="42" pageNumber="43">Papua New Guinea</collectingCountry>
, East Java, Sumatra, the
<collectingCountry id="F33987A5626CFFC9281EB2ECFF0B5D8F" name="Philippines" pageId="42" pageNumber="43">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8B91C735626CFFC92DBFB2A5FEE75CF7" blockId="42.[151,1437,150,429]" pageId="42" pageNumber="43">
<emphasis id="B95A1B27626CFFC92DBFB2A5FE3B5DA2" bold="true" box="[199,476,222,248]" pageId="42" pageNumber="43">Geographical variation</emphasis>
. The antennae color of
<taxonomicName id="4C2EBCB6626CFFC92F98B2A4FCA25DA3" authorityName="Chaudoir" authorityYear="1873" box="[736,837,223,249]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626CFFC92F98B2A4FCA25DA3" box="[736,837,223,249]" italics="true" pageId="42" pageNumber="43">P. mellea</emphasis>
</taxonomicName>
varies between different localities. In specimens from
<collectingCountry id="F33987A5626CFFC92DEFB378FEAE5C47" box="[151,329,259,285]" name="Indonesia" pageId="42" pageNumber="43">Maluku Islands</collectingCountry>
, Adonara Island, and
<collectingRegion id="49EA09D7626CFFC92F3AB378FD485C47" box="[578,687,259,285]" country="Indonesia" name="Jawa Timur" pageId="42" pageNumber="43">East Java</collectingRegion>
, antennomeres 2, 3 and the basal third of 4 are reddish yellow, in sharp contrast with the black apical antennomeres (
<figureCitation id="1315DBB0626CFFC92FA2B35CFCD85C1B" box="[730,831,295,321]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="42" pageNumber="43">Fig. 21F</figureCitation>
). In specimens from
<collectingRegion id="49EA09D7626CFFC92954B35CFB6E5C1B" box="[1068,1161,295,321]" country="Indonesia" name="Sumatera Utara" pageId="42" pageNumber="43">Sumatra</collectingRegion>
and the
<collectingCountry id="F33987A5626CFFC92994B35CFA8A5C1B" box="[1260,1389,295,321]" name="Philippines" pageId="42" pageNumber="43">Philippines</collectingCountry>
, antennomeres 2, 3, and the basal third of 4 are very dark brown, a color more similar to the black apical antennomeres (
<figureCitation id="1315DBB0626CFFC92DE7B314FEE45CD3" box="[159,259,367,393]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="42" pageNumber="43">Fig. 21G</figureCitation>
). We studied the male genitalia of specimens from these different localities and found them to be identical (
<figureCitation id="1315DBB0626CFFC92DE7B3E8FF135CF7" box="[159,244,403,429]" captionStart="FIGURE 24" captionStartId="42.[151,250,1142,1166]" captionTargetBox="[159,1425,461,1114]" captionTargetId="figure-146@42.[151,1436,453,1119]" captionTargetPageId="42" captionText="FIGURE 24. Median lobe of male genitalia for Parena (Crossoglossa) mellea (Chaudoir), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Philippine. B Maluku. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958986" httpUri="https://zenodo.org/record/7958986/files/figure.png" pageId="42" pageNumber="43">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>