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<document id="A4F4872B5468E9D1B3A6019CDFA9C4CE" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="038776236261FFCB2DEFB78EFB975BC4" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Crossoglossa) sulawesiensis Kirschenhofer. D 2006" docType="treatment" docVersion="8" lastPageNumber="41" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="40" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="8FA8AD89FB3207D44AAC396A39FA99FD">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<treatment id="038776236261FFCB2DEFB78EFB975BC4" ID-DOI="http://doi.org/10.5281/zenodo.7963727" ID-GBIF-Taxon="209629003" ID-Zenodo-Dep="7963727" LSID="urn:lsid:plazi:treatment:038776236261FFCB2DEFB78EFB975BC4" httpUri="http://treatment.plazi.org/id/038776236261FFCB2DEFB78EFB975BC4" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<subSubSection id="C33494BE6261FFC42DEFB78EFC985B55" box="[151,895,1525,1552]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8B91C7356261FFC42DEFB78EFC985B55" blockId="39.[151,1107,1525,1587]" box="[151,895,1525,1552]" pageId="39" pageNumber="40">
<emphasis id="B95A1B276261FFC42DEFB78EFC985B55" bold="true" box="[151,895,1525,1552]" pageId="39" pageNumber="40">
[6]
<taxonomicName id="4C2EBCB66261FFC42DB9B78DFC985B55" ID-CoL="4DKQN" authority="Kirschenhofer, 2006" authorityName="Kirschenhofer. D" authorityYear="2006" box="[193,895,1525,1552]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sulawesiensis" subGenus="Crossoglossa">
<emphasis id="B95A1B276261FFC42DB9B78DFEFE5B4A" bold="true" box="[193,281,1526,1552]" italics="true" pageId="39" pageNumber="40">Parena</emphasis>
(
<emphasis id="B95A1B276261FFC42C52B78EFE2F5B4A" bold="true" box="[298,456,1525,1552]" italics="true" pageId="39" pageNumber="40">Crossoglossa</emphasis>
)
<emphasis id="B95A1B276261FFC42CA1B78EFD9B5B4A" bold="true" box="[473,636,1525,1552]" italics="true" pageId="39" pageNumber="40">sulawesiensis</emphasis>
<bibRefCitation id="EFBFBAC46261FFC42FFBB78EFC985B55" author="Kirschenhofer, E." box="[643,895,1525,1551]" pageId="39" pageNumber="40" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6261FFC42DEFB462FBB45B69" box="[151,1107,1561,1587]" pageId="39" pageNumber="40" type="description">
<paragraph id="8B91C7356261FFC42DEFB462FBB45B69" blockId="39.[151,1107,1525,1587]" box="[151,1107,1561,1587]" pageId="39" pageNumber="40">
Habitus:
<figureCitation id="1315DBB06261FFC42D85B462FE435B69" box="[253,420,1561,1587]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="39" pageNumber="40">Figs 21C, 21D</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB06261FFC42F27B462FD565B69" box="[607,689,1561,1587]" captionStart="FIGURE 23" captionStartId="40.[151,250,877,901]" captionTargetBox="[158,1428,247,847]" captionTargetId="figure-52@40.[151,1436,237,854]" captionTargetPageId="40" captionText="FIGURE 23. Median lobe of male genitalia for Parena (Crossoglossa) sulawesiensis Kirschenhofer, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Sulawesi. B Holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958983" httpUri="https://zenodo.org/record/7958983/files/figure.png" pageId="39" pageNumber="40">Fig. 23</figureCitation>
. Gonocoxites of ovipositor:
<figureCitation id="1315DBB06261FFC42E88B462FBA75B69" box="[1008,1088,1561,1587]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="39" pageNumber="40">Fig. 11</figureCitation>
β.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6261FFC42DEFB423FCB35B2A" box="[151,852,1624,1648]" pageId="39" pageNumber="40" type="reference_group">
<paragraph id="8B91C7356261FFC42DEFB423FCB35B2A" blockId="39.[151,852,1624,1648]" box="[151,852,1624,1648]" pageId="39" pageNumber="40">
<bibRefCitation id="EFBFBAC46261FFC42DEFB423FE6A5B2A" author="Kirschenhofer, E." box="[151,397,1624,1648]" pageId="39" pageNumber="40" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 95</bibRefCitation>
(type locality:
<collectingRegion id="49EA09D76261FFC42F5FB423FD615B2A" box="[551,646,1624,1648]" country="Indonesia" name="Sulawesi Utara" pageId="39" pageNumber="40">Sulawesi</collectingRegion>
;
<typeStatus id="549579976261FFC42FE8B423FD0E5B2A" box="[656,745,1624,1648]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
in CDW).
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6261FFCB2DEFB4E4FE3D5D8F" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8B91C7356261FFC42DEFB4E4FCA65A13" blockId="39.[151,1437,1694,2009]" pageId="39" pageNumber="40">
<emphasis id="B95A1B276261FFC42DEFB4E4FE5D5BE2" bold="true" box="[151,442,1694,1721]" pageId="39" pageNumber="40">Type material examined</emphasis>
.
<emphasis id="B95A1B276261FFC42CB2B4E4FC125BE3" bold="true" box="[458,1013,1694,1721]" pageId="39" pageNumber="40">
<taxonomicName id="4C2EBCB66261FFC42CB2B4E4FC9D5BE2" authority="Kirschenhofer" authorityName="Kirschenhofer. D" authorityYear="2006" box="[458,890,1694,1721]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="B95A1B276261FFC42CB2B4E4FD255BE3" bold="true" box="[458,706,1694,1721]" italics="true" pageId="39" pageNumber="40">Parena sulawesiensis</emphasis>
Kirschenhofer
</taxonomicName>
:
<typeStatus id="549579976261FFC42EF0B4E4FC125BE3" box="[904,1013,1695,1721]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
</emphasis>
(CDW,
<figureCitation id="1315DBB06261FFC4292CB4E5FB5C5BE3" box="[1108,1211,1694,1721]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="39" pageNumber="40">Fig. 21C</figureCitation>
): male, body length =
<quantity id="4CD66AD06261FFC42DC8B4B8FEF05B87" box="[176,279,1731,1757]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="39" pageNumber="40" unit="mm" value="10.7">10.7 mm</quantity>
, board mounted, &quot;
<collectingCountry id="F33987A56261FFC42C9EB4B8FD665B87" box="[486,641,1731,1757]" name="Indonesia" pageId="39" pageNumber="40">INDONESIA</collectingCountry>
(C.
<collectingRegion id="49EA09D76261FFC42FCEB4B9FCFA5B86" box="[694,797,1730,1756]" country="Indonesia" name="Sulawesi Utara" pageId="39" pageNumber="40">Sulawesi</collectingRegion>
) / Kab. Donggala, village Toro / UTM51 S (WGS-84) / X 9833613, Y 170666. Alt
<quantity id="4CD66AD06261FFC42CB3B49DFDF45A5B" box="[459,531,1766,1793]" metricMagnitude="2" metricUnit="m" metricValue="8.15" pageId="39" pageNumber="40" unit="m" value="815.0">815 m</quantity>
/ From: T. cacao under diverse, / planted shade (canopy fogging) /
<date id="FF90E1F56261FFC42877B49CFA7B5A5B" box="[1295,1436,1767,1793]" pageId="39" pageNumber="40" value="2003-12-24">24-XII-2003</date>
M.M. Bos leg. / 05C241203E&quot;, &quot;
<emphasis id="B95A1B276261FFC42F7FB570FD8E5A7F" box="[519,617,1803,1829]" italics="true" pageId="39" pageNumber="40">
<collectingCountry id="F33987A56261FFC42F7FB570FDDB5A7F" box="[519,572,1803,1829]" name="Central African Republic" pageId="39" pageNumber="40">CAR</collectingCountry>
. 7B
</emphasis>
&quot; [green label], &quot;
<typeStatus id="549579976261FFC42E66B571FC715A7E" box="[798,918,1802,1828]" pageId="39" pageNumber="40" type="holotype">Holotypus</typeStatus>
/
<taxonomicName id="4C2EBCB66261FFC42ED4B570FC1F5A7F" box="[940,1016,1803,1829]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/ sulawesiensis
<taxonomicNameLabel id="A269A65C6261FFC429D5B570FB0E5A7F" box="[1197,1257,1803,1829]" pageId="39" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
/ det. Kirschenhofer&quot; [red label], &quot;COLL. WRASE / BERLIN&quot; &lt;
<figureCitation id="1315DBB06261FFC42FB0B555FCCB5A13" box="[712,812,1838,1865]" captionStart="FIGURE 23" captionStartId="40.[151,250,877,901]" captionTargetBox="[158,1428,247,847]" captionTargetId="figure-52@40.[151,1436,237,854]" captionTargetPageId="40" captionText="FIGURE 23. Median lobe of male genitalia for Parena (Crossoglossa) sulawesiensis Kirschenhofer, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Sulawesi. B Holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958983" httpUri="https://zenodo.org/record/7958983/files/figure.png" pageId="39" pageNumber="40">Fig. 23B</figureCitation>
&gt;.
</paragraph>
<paragraph id="8B91C7356261FFCB2DBFB528FE3D5D8F" blockId="39.[151,1437,1694,2009]" lastBlockId="40.[151,1437,151,213]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="B95A1B276261FFC42DBFB528FDF85A36" bold="true" box="[199,543,1874,1901]" pageId="39" pageNumber="40">Non-type material examined</emphasis>
.
<specimenCount id="9D280CBC6261FFC42F57B528FD755A36" box="[559,658,1874,1901]" pageId="39" pageNumber="40" type="female">1 female</specimenCount>
(NHMB), &quot;Drs. Sarasin, S. Cerebes, Lamontjong&quot;, &quot;Lebia sp.? Det. K.M. Heller 1916&quot; &lt;
<figureCitation id="1315DBB06261FFC42C05B50DFE305ACB" box="[381,471,1910,1937]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="39" pageNumber="40">Figs 8B</figureCitation>
,
<figureCitation id="1315DBB06261FFC42C9AB50CFE195ACB" box="[482,510,1911,1937]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="39" pageNumber="40">11</figureCitation>
β, 14E&gt;.
<specimenCount id="9D280CBC6261FFC42F1AB50CFD4B5ACA" box="[610,684,1910,1937]" pageId="39" pageNumber="40" type="male">1 male</specimenCount>
(CRS), &quot;Indon.—C.
<collectingRegion id="49EA09D76261FFC42EEEB50DFC195ACA" box="[918,1022,1910,1936]" country="Indonesia" name="Sulawesi Utara" pageId="39" pageNumber="40">Sulawesi</collectingRegion>
, Puncak Palopo,
<date id="FF90E1F56261FFC429C4B50CFAF65ACB" box="[1212,1297,1911,1937]" pageId="39" pageNumber="40" value="2013-02">II.2013</date>
&quot; &lt;
<figureCitation id="1315DBB06261FFC42856B50DFA7F5ACB" box="[1326,1432,1910,1937]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="39" pageNumber="40">Figs 21D</figureCitation>
,
<figureCitation id="1315DBB06261FFC42DEFB5E0FF2D5AEF" box="[151,202,1947,1973]" captionStart="FIGURE 23" captionStartId="40.[151,250,877,901]" captionTargetBox="[158,1428,247,847]" captionTargetId="figure-52@40.[151,1436,237,854]" captionTargetPageId="40" captionText="FIGURE 23. Median lobe of male genitalia for Parena (Crossoglossa) sulawesiensis Kirschenhofer, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Sulawesi. B Holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958983" httpUri="https://zenodo.org/record/7958983/files/figure.png" pageId="39" pageNumber="40">23A</figureCitation>
&gt;.
<specimenCount id="9D280CBC6261FFC42D9DB5E0FED75AEE" box="[229,304,1946,1973]" pageId="39" pageNumber="40" type="male">1 male</specimenCount>
(NNML), &quot;C.J. Louwerens, W.
<collectingRegion id="49EA09D76261FFC42FEFB5E1FD175AEE" box="[663,752,1946,1972]" country="Indonesia" name="Sulawesi Utara" pageId="39" pageNumber="40">Celebes</collectingRegion>
1800', G. Rangkoenau- Paloe,
<date id="FF90E1F56261FFC42932B5E0FB275AEF" box="[1098,1216,1947,1973]" pageId="39" pageNumber="40" value="1937-08">VIII.1937</date>
&quot;; &quot;Museum Leiden, ex. collection, C.J. Louwerens, rec. 1979&quot;; &quot;
<taxonomicName id="4C2EBCB66261FFC42FF3B5C4FBA45A82" authority="Chaud., H. E. Andrewes" authorityName="Chaud., H. E. Andrewes" box="[651,1091,1982,2009]" class="Insecta" family="Pyralidae" genus="Parnea" kingdom="Animalia" order="Lepidoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B276261FFC42FF3B5C4FC6D5A82" box="[651,906,1982,2008]" italics="true" pageId="39" pageNumber="40">Parnea mellea Chaud.</emphasis>
, H.E. Andrewes
</taxonomicName>
det.&quot;.
<specimenCount id="9D280CBC6261FFC429F6B5C4FB175A82" box="[1166,1264,1982,2009]" pageId="39" pageNumber="40" type="female">1 female</specimenCount>
(NNML), &quot;
<emphasis id="B95A1B276261FFCB280FB5C5FF0A5DEB" italics="true" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">Id.. Salayer</emphasis>
,
<emphasis id="B95A1B27626EFFCB2D83B2ECFEB85DEB" box="[251,351,151,177]" italics="true" pageId="40" pageNumber="41">Somarisi</emphasis>
&quot;; &quot;Museum Leiden, ex. collection, C.J. Louwerens, rec. 1979&quot;; &quot;
<emphasis id="B95A1B27626EFFCB2942B2ECFAD65DEB" box="[1082,1329,151,177]" italics="true" pageId="40" pageNumber="41">
<taxonomicName id="4C2EBCB6626EFFCB2942B2ECFB055DEB" box="[1082,1250,151,177]" class="Insecta" family="Pyralidae" genus="Parnea" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mellea">Parnea mellea</taxonomicName>
? Chd.,
</emphasis>
H.E. Andrewes det.&quot;[Selayar island].
</paragraph>
</subSubSection>
<caption id="DF5197BD626EFFCB2DEFB116FBE65EF3" ID-DOI="http://doi.org/10.5281/zenodo.7958983" ID-Zenodo-Dep="7958983" httpUri="https://zenodo.org/record/7958983/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,877,901]" targetBox="[158,1428,247,847]" targetPageId="40" targetType="figure">
<paragraph id="8B91C735626EFFCB2DEFB116FBE65EF3" blockId="40.[151,1436,877,938]" pageId="40" pageNumber="41">
<emphasis id="B95A1B27626EFFCB2DEFB116FEFC5EDF" bold="true" box="[151,283,877,901]" pageId="40" pageNumber="41">FIGURE 23</emphasis>
. Median lobe of male genitalia for
<taxonomicName id="4C2EBCB6626EFFCB2FF5B115FB435EDF" authority="Kirschenhofer" authorityName="Kirschenhofer. D" authorityYear="2006" box="[653,1188,877,902]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="sulawesiensis" subGenus="Crossoglossa">
<emphasis id="B95A1B27626EFFCB2FF5B115FD305EDF" box="[653,727,878,901]" italics="true" pageId="40" pageNumber="41">Parena</emphasis>
(
<emphasis id="B95A1B27626EFFCB2F9FB116FC885EDF" box="[743,879,877,901]" italics="true" pageId="40" pageNumber="41">Crossoglossa</emphasis>
)
<emphasis id="B95A1B27626EFFCB2E07B116FBEE5EDF" box="[895,1033,877,901]" italics="true" pageId="40" pageNumber="41">sulawesiensis</emphasis>
Kirschenhofer
</taxonomicName>
, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views.
<emphasis id="B95A1B27626EFFCB2F49B1EAFDA45EF3" bold="true" box="[561,579,913,937]" pageId="40" pageNumber="41">A</emphasis>
Sulawesi.
<emphasis id="B95A1B27626EFFCB2FC9B1E9FD255EF0" bold="true" box="[689,706,914,938]" pageId="40" pageNumber="41">B</emphasis>
Holotype. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE626EFFCB2DBFB1ACFC3D5904" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8B91C735626EFFCB2DBFB1ACFC3D5904" blockId="40.[151,1437,983,1694]" pageId="40" pageNumber="41">
<emphasis id="B95A1B27626EFFCB2DBFB1ACFE815EAB" bold="true" box="[199,358,983,1009]" pageId="40" pageNumber="41">Comparisons</emphasis>
.
<taxonomicName id="4C2EBCB6626EFFCB2C00B1A3FDC85EA8" authorityName="Kirschenhofer. D" authorityYear="2006" box="[376,559,984,1010]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="B95A1B27626EFFCB2C00B1A3FDC85EA8" box="[376,559,984,1010]" italics="true" pageId="40" pageNumber="41">P. sulawesiensis</emphasis>
</taxonomicName>
is distinguishable from the other four species in this species group by the combination of: basal three antennomeres yellow; elytra sutural angles sharply pointed, forming long spines; outer apical angles almost rounded; female sternite VII strongly projected at middle. This species is most similar to
<taxonomicName id="4C2EBCB6626EFFCB28F0B65BFF145907" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B27626EFFCB28F0B65BFF145907" italics="true" pageId="40" pageNumber="41">P. testacea</emphasis>
</taxonomicName>
. Comparison between them was provided under the latter species.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626EFFCB2DBFB61CFD425B20" pageId="40" pageNumber="41" type="description">
<paragraph id="8B91C735626EFFCB2DBFB61CFD425B20" blockId="40.[151,1437,983,1694]" pageId="40" pageNumber="41">
<emphasis id="B95A1B27626EFFCB2DBFB61CFEB659DB" bold="true" box="[199,337,1127,1153]" pageId="40" pageNumber="41">Description</emphasis>
. Body length
<quantity id="4CD66AD0626EFFCB2C97B613FD7359D8" box="[495,660,1128,1154]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" metricValueMax="1.08" metricValueMin="1.04" pageId="40" pageNumber="41" unit="mm" value="10.6" valueMax="10.8" valueMin="10.4">10.410.8 mm</quantity>
. Dorsum pale yellow; antennomeres 1 to 3 reddish yellow, basal third of antennomere 4 reddish yellow, remainder of antennae black; apices of mandibles black; terminal labial palpomere black, terminal and penultimate maxillary palpomeres black; tarsomeres black, in strong contrast with yellowish brown tibiae; venter pale yellow. Pronotum strongly transverse, PW/PL = 1.651.71, much wider than head, PW/ HW = 1.141.22; widest at anterior third, lateral explanations very wide. Elytra with faint isodiametric microsculpture on apical third; striae not incised, with rows of fine punctures; intervals completely flat, with punctures slightly finer and much sparser than in striae; disc depressed near middle of intervals 3 to 6, depressions large and shallow; apical truncation distinct, nearly straight, outer apical angles almost rounded, not forming distinct angles; sutural angles sharply pointed, forming long spines. Apex of abdominal sternite VII nearly straight in males, strongly projected at middle in females (
<figureCitation id="1315DBB0626EFFCB2CAAB7D7FDCE589C" box="[466,553,1452,1478]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="40" pageNumber="41">Fig. 8B</figureCitation>
). Median lobe of aedeagus with dorsal margin curved before apical orifice, and then narrowed to apex; apical lamella subrectangular, more or less bent to dorsum, length slightly greater than its basal width (L/W = 1.3), apex slightly truncate. Endophallus very coarsely scaled on basal sheath, base of apical sheath discoid in dorsal view; squamate sac hardly divided, on basal fifth of median lobe, right to squamate sheath (
<figureCitation id="1315DBB0626EFFCB2DE7B447FF145B0C" box="[159,243,1596,1622]" captionStart="FIGURE 23" captionStartId="40.[151,250,877,901]" captionTargetBox="[158,1428,247,847]" captionTargetId="figure-52@40.[151,1436,237,854]" captionTargetPageId="40" captionText="FIGURE 23. Median lobe of male genitalia for Parena (Crossoglossa) sulawesiensis Kirschenhofer, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Sulawesi. B Holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958983" httpUri="https://zenodo.org/record/7958983/files/figure.png" pageId="40" pageNumber="41">Fig. 23</figureCitation>
). Gonocoxite II of ovipositor dichotomous, inner branch with two ensiform setae apically, outer branch with two or three ensiform setae apically (
<figureCitation id="1315DBB0626EFFCB2F4FB41BFD6F5B20" box="[567,648,1632,1658]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="40" pageNumber="41">Fig. 11</figureCitation>
β).
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626EFFCB2DBFB4F8FB975BC4" box="[199,1136,1667,1694]" pageId="40" pageNumber="41" type="distribution">
<paragraph id="8B91C735626EFFCB2DBFB4F8FB975BC4" blockId="40.[151,1437,983,1694]" box="[199,1136,1667,1694]" pageId="40" pageNumber="41">
<emphasis id="B95A1B27626EFFCB2DBFB4F8FEBE5BC7" bold="true" box="[199,345,1667,1693]" pageId="40" pageNumber="41">Distribution</emphasis>
(
<figureCitation id="1315DBB0626EFFCB2C10B4FFFE535BC4" box="[360,436,1668,1694]" captionStart="MAP 4" captionStartId="36.[152,211,1895,1919]" captionTargetBox="[158,1430,1002,1871]" captionTargetId="figure-71@36.[158,1430,1002,1871]" captionTargetPageId="36" captionText="MAP 4. Distributions for species in the Parena (Crossoglossa) testacea group." figureDoi="http://doi.org/10.5281/zenodo.7963663" httpUri="https://zenodo.org/record/7963663/files/figure.png" pageId="40" pageNumber="41">Map 4</figureCitation>
, green). Only known from
<collectingRegion id="49EA09D7626EFFCB2F9DB4FFFCAA5BC4" box="[741,845,1668,1694]" country="Indonesia" name="Sulawesi Utara" pageId="40" pageNumber="41">Sulawesi</collectingRegion>
, including Selayar Island.
</paragraph>
</subSubSection>
</treatment>
</document>