treatments-xml/data/03/87/76/038776236259FFC12DEFB3A0FF175A8D.xml

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<mods:title id="34FD4E9ACE5067FD6D53B9F3C964F2AD">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="20887397D07D1EEC1BA6ED22AC4727E7">Shi, Hongliang</mods:namePart>
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<subSubSection id="C33494BE6259FFFC2DEFB3A0FCEE5CAF" box="[151,777,475,502]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8B91C7356259FFFC2DEFB3A0FCEE5CAF" blockId="31.[151,1130,475,537]" box="[151,777,475,502]" pageId="31" pageNumber="32">
<emphasis id="B95A1B276259FFFC2DEFB3A0FCEE5CAF" bold="true" box="[151,777,475,502]" pageId="31" pageNumber="32">
[2] 
<taxonomicName id="4C2EBCB66259FFFC2DB9B3A0FCEE5CAF" ID-CoL="4DKPN" authority="(Bates, 1873)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[193,777,475,502]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="laesipennis" subGenus="Crossoglossa">
<emphasis id="B95A1B276259FFFC2DB9B3A0FEFE5CAF" bold="true" box="[193,281,475,501]" italics="true" pageId="31" pageNumber="32">Parena</emphasis>
(
<emphasis id="B95A1B276259FFFC2C52B3A0FE2F5CAC" bold="true" box="[298,456,475,502]" italics="true" pageId="31" pageNumber="32">Crossoglossa</emphasis>
) 
<emphasis id="B95A1B276259FFFC2CA1B3A0FD875CAC" bold="true" box="[473,608,475,502]" italics="true" pageId="31" pageNumber="32">laesipennis</emphasis>
(
<bibRefCitation id="EFBFBAC46259FFFC2F08B3A0FCE65CAF" author="Bates, H. W." box="[624,769,475,501]" pageId="31" pageNumber="32" pagination="229 - 322" refId="ref109990" refString="Bates, H. W. (1873) On the Geodephagous Coleoptera of Japan. Fam. Carabidae. The Transactions of the Entomological Society of London, 1873 (2), 229 - 322." type="journal article" year="1873">Bates, 1873</bibRefCitation>
)
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6259FFFC2DEFB384FB8D5F43" box="[151,1130,511,537]" pageId="31" pageNumber="32" type="description">
<paragraph id="8B91C7356259FFFC2DEFB384FB8D5F43" blockId="31.[151,1130,475,537]" box="[151,1130,511,537]" pageId="31" pageNumber="32">
Habitus: 
<figureCitation id="1315DBB06259FFFC2D85B384FE6D5F43" box="[253,394,511,537]" captionStart="FIGURE 17" captionStartId="31.[151,250,1926,1950]" captionTargetBox="[154,1424,569,1887]" captionTargetId="figure-130@31.[151,1436,561,1903]" captionTargetPageId="31" captionText="FIGURE 17. Habitus of Parena (Crossoglossa) laesipennis species group. A Lectotype of Parena laesipennis (Bates). B P. laesipennis (Bates), male from Taiwan. C P. laesipennis (Bates), female from Yunnan. D P. laesipennis (Bates), male from Nepal. E Holotype of Parena levata Andrewes. F P. levata Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958971" httpUri="https://zenodo.org/record/7958971/files/figure.png" pageId="31" pageNumber="32">Figs 17A–D</figureCitation>
. Male genitalia: 
<figureCitation id="1315DBB06259FFFC2F3DB384FD7D5F43" box="[581,666,511,537]" captionStart="FIGURE 18" captionStartId="32.[151,250,1101,1125]" captionTargetBox="[164,1427,365,1072]" captionTargetId="figure-113@32.[151,1436,352,1078]" captionTargetPageId="32" captionText="FIGURE 18. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Yunnan. B Yunnan. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958973" httpUri="https://zenodo.org/record/7958973/files/figure.png" pageId="31" pageNumber="32">Figs 18</figureCitation>
, 
<figureCitation id="1315DBB06259FFFC2FDEB384FD225F43" box="[678,709,511,537]" captionStart="FIGURE 19" captionStartId="32.[151,250,1946,1970]" captionTargetBox="[161,1425,1199,1917]" captionTargetId="figure-151@32.[151,1436,1186,1923]" captionTargetPageId="32" captionText="FIGURE 19. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates) from different localities, left-lateral (l) and dorsal (d) views: A Laos. B Taiwan. C Nepal. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958975" httpUri="https://zenodo.org/record/7958975/files/figure.png" pageId="31" pageNumber="32">19</figureCitation>
. Gonocoxites of ovipositor: 
<figureCitation id="1315DBB06259FFFC297BB384FB825F43" box="[1027,1125,511,537]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="31" pageNumber="32">Fig. 11Z</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5197BD6259FFFC2DEFB5FDFB5E5ABC" ID-DOI="http://doi.org/10.5281/zenodo.7958971" ID-Zenodo-Dep="7958971" httpUri="https://zenodo.org/record/7958971/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1926,1950]" targetBox="[154,1424,569,1887]" targetPageId="31" targetType="figure">
<paragraph id="8B91C7356259FFFC2DEFB5FDFB5E5ABC" blockId="31.[151,1437,1926,2023]" pageId="31" pageNumber="32">
<emphasis id="B95A1B276259FFFC2DEFB5FDFEFF5AC4" bold="true" box="[151,280,1926,1950]" pageId="31" pageNumber="32">FIGURE 17</emphasis>
. Habitus of 
<taxonomicName id="4C2EBCB66259FFFC2CEBB5FCFD165AC5" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[403,753,1927,1951]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="laesipennis" subGenus="Crossoglossa">
<emphasis id="B95A1B276259FFFC2CEBB5FCFE3A5AC4" box="[403,477,1927,1950]" italics="true" pageId="31" pageNumber="32">Parena</emphasis>
(
<emphasis id="B95A1B276259FFFC2C92B5FCFD955AC5" box="[490,626,1927,1951]" italics="true" pageId="31" pageNumber="32">Crossoglossa</emphasis>
) 
<emphasis id="B95A1B276259FFFC2F07B5FCFD165AC5" box="[639,753,1927,1951]" italics="true" pageId="31" pageNumber="32">laesipennis</emphasis>
</taxonomicName>
species group. 
<emphasis id="B95A1B276259FFFC2EF4B5FDFC795AC4" bold="true" box="[908,926,1926,1950]" pageId="31" pageNumber="32">A</emphasis>
Lectotype of 
<taxonomicName id="4C2EBCB66259FFFC2950B5FCFADC5AC5" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[1064,1339,1927,1951]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="laesipennis">
<emphasis id="B95A1B276259FFFC2950B5FCFB0E5AC5" box="[1064,1257,1927,1951]" italics="true" pageId="31" pageNumber="32">Parena laesipennis</emphasis>
(Bates)
</taxonomicName>
. 
<emphasis id="B95A1B276259FFFC283CB5FCFAB25AC5" bold="true" box="[1348,1365,1927,1951]" pageId="31" pageNumber="32">B</emphasis>
<taxonomicName id="4C2EBCB66259FFFC2822B5FCFEDC5A99" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="laesipennis">
<emphasis id="B95A1B276259FFFC2822B5FCFA8E5AC4" box="[1370,1385,1927,1950]" italics="true" pageId="31" pageNumber="32">P</emphasis>
. 
<emphasis id="B95A1B276259FFFC280DB5FCFF0D5A98" italics="true" pageId="31" pageNumber="32">laesipennis</emphasis>
(Bates)
</taxonomicName>
, male from Taiwan. 
<emphasis id="B95A1B276259FFFC2F73B5D1FDFA5A98" bold="true" box="[523,541,1962,1986]" pageId="31" pageNumber="32">C</emphasis>
<taxonomicName id="4C2EBCB66259FFFC2F5BB5D0FCE45A99" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[547,771,1963,1987]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="laesipennis">
<emphasis id="B95A1B276259FFFC2F5BB5D0FDD55A98" box="[547,562,1963,1986]" italics="true" pageId="31" pageNumber="32">P</emphasis>
. 
<emphasis id="B95A1B276259FFFC2F47B5D0FD565A99" box="[575,689,1963,1987]" italics="true" pageId="31" pageNumber="32">laesipennis</emphasis>
(Bates)
</taxonomicName>
, female from Yunnan. 
<emphasis id="B95A1B276259FFFC2E90B5D0FC1D5A99" bold="true" box="[1000,1018,1963,1987]" pageId="31" pageNumber="32">D</emphasis>
<taxonomicName id="4C2EBCB66259FFFC2979B5D0FB075A99" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[1025,1248,1963,1987]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="laesipennis">
<emphasis id="B95A1B276259FFFC2979B5D0FBF75A98" box="[1025,1040,1963,1986]" italics="true" pageId="31" pageNumber="32">P</emphasis>
. 
<emphasis id="B95A1B276259FFFC2965B5D0FB685A99" box="[1053,1167,1963,1987]" italics="true" pageId="31" pageNumber="32">laesipennis</emphasis>
(Bates)
</taxonomicName>
, male from Nepal. 
<emphasis id="B95A1B276259FFFC2DEFB5B4FF4F5ABD" bold="true" box="[151,168,1999,2023]" pageId="31" pageNumber="32">E</emphasis>
Holotype of 
<taxonomicName id="4C2EBCB66259FFFC2C56B5B4FDA25ABD" authority="Andrewes. F" authorityName="Andrewes. F" box="[302,581,1999,2023]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="levata">
<emphasis id="B95A1B276259FFFC2C56B5B4FE5B5ABD" box="[302,444,1999,2023]" italics="true" pageId="31" pageNumber="32">Parena levata</emphasis>
Andrewes. 
<emphasis id="B95A1B276259FFFC2F4EB5B4FDA25ABD" bold="true" box="[566,581,1999,2023]" pageId="31" pageNumber="32">F</emphasis>
</taxonomicName>
<taxonomicName id="4C2EBCB66259FFFC2F34B5B4FCF65ABD" authority="Andrewes" authorityName="Andrewes, Borneo. D" authorityYear="1931" box="[588,785,1999,2023]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="levata">
<emphasis id="B95A1B276259FFFC2F34B5B4FD445ABD" box="[588,675,1999,2023]" italics="true" pageId="31" pageNumber="32">P. levata</emphasis>
Andrewes
</taxonomicName>
, female from Borneo. Scale bars = 2 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE6266FFC32DEFB2ECFE065C12" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="8B91C7356266FFC32DEFB2ECFC405DB6" blockId="32.[151,1437,151,328]" pageId="32" pageNumber="33">
<bibRefCitation id="EFBFBAC46266FFC32DEFB2ECFEA35DF5" author="Bates, H. W." box="[151,324,151,175]" pageId="32" pageNumber="33" pagination="229 - 322" refId="ref109990" refString="Bates, H. W. (1873) On the Geodephagous Coleoptera of Japan. Fam. Carabidae. The Transactions of the Entomological Society of London, 1873 (2), 229 - 322." type="journal article" year="1873">Bates, 1873: 317</bibRefCitation>
(original: 
<taxonomicName id="4C2EBCB66266FFC32CC9B2ECFDDE5DF5" ID-CoL="8KWFM" authority="Chaudoir, 1873" box="[433,569,151,175]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B276266FFC32CC9B2ECFDDE5DF5" box="[433,569,151,175]" italics="true" pageId="32" pageNumber="33">Crossoglossa</emphasis>
</taxonomicName>
; type locality: Nagasaki; 
<typeStatus id="549579976266FFC32E39B2ECFC795DF5" box="[833,926,151,175]" pageId="32" pageNumber="33" type="lectotype">lectotype</typeStatus>
in MNHN); Jacobson, 1907: 403 (
<taxonomicName id="4C2EBCB66266FFC3287DB2ECFA6A5DF5" ID-CoL="8KWFM" authority="Chaudoir, 1873" box="[1285,1421,151,175]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B276266FFC3287DB2ECFA6A5DF5" box="[1285,1421,151,175]" italics="true" pageId="32" pageNumber="33">Crossoglossa</emphasis>
</taxonomicName>
); 
<bibRefCitation id="EFBFBAC46266FFC32DBFB2CEFE935D97" author="Jedlicka, A." box="[199,372,181,205]" pageId="32" pageNumber="33" refId="ref111358" refString="Jedlicka, A. (1946) Description des Carabides nouveaux de 1 ' Asie orient. Praha: A. Jedlicka, 16 pp." type="book" year="1946">Jedlička, 1946: 8</bibRefCitation>
; 
<bibRefCitation id="EFBFBAC46266FFC32C05B2CEFDA35D97" author="Jedlicka, A." box="[381,580,181,205]" pageId="32" pageNumber="33" pagination="269 - 579" refId="ref111456" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article" year="1963">Jedlička, 1963: 439</bibRefCitation>
; Habu, 1967: 168; 
<bibRefCitation id="EFBFBAC46266FFC32E78B2CEFC4D5D97" author="Habu, A." box="[768,938,181,205]" pageId="32" pageNumber="33" pagination="83 - 118" refId="ref110913" refString="Habu, A. (1982) Revised and supplementary notes on and descriptions of the Truncatipennes group of Japan (2) (Coleoptera, Carabidae). Entomological Review of Japan, 37 (2), 83 - 118." type="journal article" year="1982">Habu, 1982: 118</bibRefCitation>
; 
<bibRefCitation id="EFBFBAC46266FFC32ECBB2CEFBBB5D97" author="Lafer, G. S." box="[947,1116,181,205]" pageId="32" pageNumber="33" pagination="71 - 222" refId="ref111842" refString="Lafer, G. S. (1989) 4. Semeystvo Carabidae - Zhuzhelitsy. In: Ler, P. A. (ed.), Opredelitel nasekomykh Dalnego Vostoka SSSR. Chast 1. Zhestkokrylye, ili zhuki. Chast 1. Leningrad: Nauka, 576 pp. [pp. 71 - 222.]" type="book chapter" year="1989">Lafer, 1989: 213</bibRefCitation>
; 
<bibRefCitation id="EFBFBAC46266FFC3291DB2CEFAB45D97" author="Park, J. K. &amp; Kwon, Y. J." box="[1125,1363,181,205]" pageId="32" pageNumber="33" pagination="35 - 41" refId="ref112486" refString="Park, J. K. &amp; Kwon, Y. J. (1998) Ground-beetles of the genus Parena Motschulsky from Korea (Coleoptera: Harpalidae). Korean Journal of Soil Zoology, 3 (1), 35 - 41." type="journal article" year="1998">Park &amp; Kwon, 1998: 36</bibRefCitation>
(
<collectingCountry id="F33987A56266FFC32818B2CEFEEE5DB6" name="South Korea" pageId="32" pageNumber="33">South Korea</collectingCountry>
); 
<bibRefCitation id="EFBFBAC46266FFC32C63B2AFFDF45DB6" author="Kirschenhofer, E." box="[283,531,212,236]" pageId="32" pageNumber="33" pagination="61 - 75" refId="ref111662" refString="Kirschenhofer, E. (2011) Beschreibungen neuer truncatipenner Carabiden aus Ost- und Sudostasien; eine neue Art der Gattung Craspedophorus Hope, 1838 (Panageaeini) aus Sudostasien; Bemerkungen zu zwei japanischen Arten der Gattung Parena Motschulsky, 1859. Acta Coleopterologica, 27 (1), 61 - 75." type="journal article" year="2011">Kirschenhofer, 2011: 66</bibRefCitation>
. 
<bibRefCitation id="EFBFBAC46266FFC32F65B2AFFD1D5DB6" author="Karube, H. &amp; Matsumoto, K. &amp; Kishimoto, T. &amp; Ozono, A." box="[541,762,212,236]" pageId="32" pageNumber="33" pagination="1 - 16" refId="ref111529" refString="Karube, H., Matsumoto, K., Kishimoto, T. &amp; Ozono, A. (2012) Records of notable insect species from the Ogasawara Islands. Ogasawara Research, 38, 1 - 16." type="journal article" year="2012">
Karube 
<emphasis id="B95A1B276266FFC32F15B2AEFD7D5DB6" box="[621,666,212,236]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
., 2012: 3
</bibRefCitation>
(Ogasawara Is.).
</paragraph>
<paragraph id="8B91C7356266FFC32DEFB289FC975C51" blockId="32.[151,1437,151,328]" box="[151,880,242,267]" pageId="32" pageNumber="33">
<emphasis id="B95A1B276266FFC32DEFB289FEA85C50" bold="true" box="[151,335,242,266]" pageId="32" pageNumber="33">Misidentification</emphasis>
: 
<taxonomicName id="4C2EBCB66266FFC32C25B288FD955C51" ID-CoL="4DKP6" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[349,626,243,267]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cavipennis">
<emphasis id="B95A1B276266FFC32C25B288FDF95C51" bold="true" box="[349,542,243,267]" italics="true" pageId="32" pageNumber="33">Parena cavipennis</emphasis>
(Bates)
</taxonomicName>
: 
<bibRefCitation id="EFBFBAC46266FFC32F03B288FCFF5C51" author="Habu, A." box="[635,792,243,267]" pageId="32" pageNumber="33" pagination="76 - 79" refId="ref110652" refString="Habu, A. (1942) Notes on some Carabideous-beetles from Miyake Island. Kontyu, 16, 76 - 79." type="journal article" year="1942">Habu, 1942: 78</bibRefCitation>
(
<collectingCountry id="F33987A56266FFC32E5EB288FC825C51" box="[806,869,243,267]" name="Japan" pageId="32" pageNumber="33">Japan</collectingCountry>
).
</paragraph>
<paragraph id="8B91C7356266FFC32DEFB36AFE065C12" blockId="32.[151,1437,151,328]" pageId="32" pageNumber="33">
<emphasis id="B95A1B276266FFC32DEFB36AFEA85C73" bold="true" box="[151,335,273,297]" pageId="32" pageNumber="33">Misidentification</emphasis>
: 
<taxonomicName id="4C2EBCB66266FFC32C24B369FD6C5C73" ID-CoL="4DKQD" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1934" box="[348,651,273,297]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rufotestacea">
<emphasis id="B95A1B276266FFC32C24B369FDC95C73" bold="true" box="[348,558,273,297]" italics="true" pageId="32" pageNumber="33">Parena rufotestacea</emphasis>
Jedlička
</taxonomicName>
: 
<bibRefCitation id="EFBFBAC46266FFC32FECB36AFC825C73" author="Xie, W. P. &amp; Yu, P. Y." box="[660,869,273,297]" pageId="32" pageNumber="33" pagination="185 - 195" refId="ref113326" refString="Xie, W. P. &amp; Yu, P. Y. (1993) On the Chinese species of Parena Motschulsky (Coleoptera: Carabidae). Sinozoologia, 10, 185 - 195." type="journal article" year="1993">Xie &amp; Yu, 1993: 194</bibRefCitation>
(
<collectingCountry id="F33987A56266FFC32E0AB36AFC535C73" box="[882,948,273,297]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
: 
<collectingRegion id="49EA09D76266FFC32EC5B36AFBE75C73" box="[957,1024,273,297]" country="China" name="Fujian" pageId="32" pageNumber="33">Fujian</collectingRegion>
, 
<collectingRegion id="49EA09D76266FFC32971B36AFBBC5C73" box="[1033,1115,273,297]" country="China" name="Yunnan" pageId="32" pageNumber="33">Yunnan</collectingRegion>
); 
<bibRefCitation id="EFBFBAC46266FFC32914B36AFA845C73" author="Kirschenhofer, E." box="[1132,1379,273,297]" pageId="32" pageNumber="33" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 89</bibRefCitation>
; 
<bibRefCitation id="EFBFBAC46266FFC32816B36AFE8A5C12" author="Park, J. &amp; Trac, D. &amp; Will, K." pageId="32" pageNumber="33" pagination="85 - 105" refId="ref112590" refString="Park, J. K, Trac, D. H, &amp; Will, K. (2006) Carabidae from Vietnam (Coleoptera). Journal of Asia-Pacific Entomology, 9 (2), 85 - 105. https: // doi. org / 10.1016 / S 1226 - 8615 (08) 60280 - 0" type="journal article" year="2006">
Park 
<emphasis id="B95A1B276266FFC32DBFB34AFF135C12" box="[199,244,304,328]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
., 2006: 102
</bibRefCitation>
(
<collectingCountry id="F33987A56266FFC32C03B34BFE315C12" box="[379,470,304,328]" name="Vietnam" pageId="32" pageNumber="33">Vietnam</collectingCountry>
).
</paragraph>
</subSubSection>
<caption id="DF5197BD6266FFC32DEFB636FC6C59D3" ID-DOI="http://doi.org/10.5281/zenodo.7958973" ID-Zenodo-Dep="7958973" httpUri="https://zenodo.org/record/7958973/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1101,1125]" targetBox="[164,1427,365,1072]" targetPageId="32" targetType="figure">
<paragraph id="8B91C7356266FFC32DEFB636FC6C59D3" blockId="32.[151,1436,1101,1162]" pageId="32" pageNumber="33">
<emphasis id="B95A1B276266FFC32DEFB636FEFC593C" bold="true" box="[151,283,1101,1126]" pageId="32" pageNumber="33">FIGURE 18</emphasis>
. Median lobe of male genitalia for 
<taxonomicName id="4C2EBCB66266FFC32FF5B635FBA3593C" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[653,1092,1102,1126]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="laesipennis" subGenus="Crossoglossa">
<emphasis id="B95A1B276266FFC32FF5B635FD30593F" box="[653,727,1102,1125]" italics="true" pageId="32" pageNumber="33">Parena</emphasis>
(
<emphasis id="B95A1B276266FFC32F9FB635FC88593C" box="[743,879,1102,1126]" italics="true" pageId="32" pageNumber="33">Crossoglossa</emphasis>
) 
<emphasis id="B95A1B276266FFC32E07B635FC16593C" box="[895,1009,1102,1126]" italics="true" pageId="32" pageNumber="33">laesipennis</emphasis>
(Bates)
</taxonomicName>
, right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. 
<emphasis id="B95A1B276266FFC32CA4B60AFE0959D3" bold="true" box="[476,494,1137,1161]" pageId="32" pageNumber="33">A</emphasis>
Yunnan. 
<emphasis id="B95A1B276266FFC32F36B609FDB859D0" bold="true" box="[590,607,1138,1162]" pageId="32" pageNumber="33">B</emphasis>
Yunnan. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF5197BD6266FFC32DEFB5E1FC635A8C" ID-DOI="http://doi.org/10.5281/zenodo.7958975" ID-Zenodo-Dep="7958975" httpUri="https://zenodo.org/record/7958975/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1946,1970]" targetBox="[161,1425,1199,1917]" targetPageId="32" targetType="figure">
<paragraph id="8B91C7356266FFC32DEFB5E1FC635A8C" blockId="32.[151,1437,1946,2007]" pageId="32" pageNumber="33">
<emphasis id="B95A1B276266FFC32DEFB5E1FEFE5AE8" bold="true" box="[151,281,1946,1970]" pageId="32" pageNumber="33">FIGURE 19</emphasis>
. Median lobe of male genitalia for 
<taxonomicName id="4C2EBCB66266FFC32FF8B5E0FBD75AE9" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[640,1072,1947,1971]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="laesipennis" subGenus="Crossoglossa">
<emphasis id="B95A1B276266FFC32FF8B5E0FD2D5AE8" box="[640,714,1947,1970]" italics="true" pageId="32" pageNumber="33">Parena</emphasis>
(
<emphasis id="B95A1B276266FFC32FA0B5E0FC875AE9" box="[728,864,1947,1971]" italics="true" pageId="32" pageNumber="33">Crossoglossa</emphasis>
) 
<emphasis id="B95A1B276266FFC32E16B5E0FC075AE9" box="[878,992,1947,1971]" italics="true" pageId="32" pageNumber="33">laesipennis</emphasis>
(Bates)
</taxonomicName>
from different localities, left-lateral (l) and dorsal (d) views: 
<emphasis id="B95A1B276266FFC32CE9B5C5FE445A8C" bold="true" box="[401,419,1982,2006]" pageId="32" pageNumber="33">A</emphasis>
Laos. 
<emphasis id="B95A1B276266FFC32C9FB5C4FE1F5A8D" bold="true" box="[487,504,1983,2007]" pageId="32" pageNumber="33">B</emphasis>
Taiwan. 
<emphasis id="B95A1B276266FFC32F2DB5C5FD805A8C" bold="true" box="[597,615,1982,2006]" pageId="32" pageNumber="33">C</emphasis>
Nepal. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE6267FFC22DEFB2ECFAF75C1A" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8B91C7356267FFC22DEFB2ECFAF75C1A" blockId="33.[151,1437,150,2044]" pageId="33" pageNumber="34">
<emphasis id="B95A1B276267FFC22DEFB2ECFE5E5DEA" bold="true" box="[151,441,150,177]" pageId="33" pageNumber="34">Type material examined</emphasis>
. 
<emphasis id="B95A1B276267FFC22CB0B2EDFCDE5DEB" bold="true" box="[456,825,150,177]" pageId="33" pageNumber="34">
<taxonomicName id="4C2EBCB66267FFC22CB0B2EDFCD25DEB" authority="Bates" authorityName="Bates" authorityYear="1873" box="[456,821,150,177]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="laesipennis">
<emphasis id="B95A1B276267FFC22CB0B2EDFD005DEB" bold="true" box="[456,743,150,177]" italics="true" pageId="33" pageNumber="34">Crossoglossa laesipennis</emphasis>
Bates
</taxonomicName>
:
</emphasis>
<materialsCitation id="3B46CD686267FFC22E39B2ECFB0B5DA2" ID-GBIF-Occurrence="4115871364" collectorName="Ex Musaeo H. W. Bates" location="NAGASAKI" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Nagasaki" typeStatus="lectotype">
<emphasis id="B95A1B276267FFC22E39B2ECFC505DEB" bold="true" box="[833,951,151,177]" pageId="33" pageNumber="34">
<typeStatus id="549579976267FFC22E39B2ECFC505DEB" box="[833,951,151,177]" pageId="33" pageNumber="34" type="lectotype">Lectotype</typeStatus>
</emphasis>
(MNHN, 
<figureCitation id="1315DBB06267FFC22954B2ECFB745DEB" box="[1068,1171,151,177]" captionStart="FIGURE 17" captionStartId="31.[151,250,1926,1950]" captionTargetBox="[154,1424,569,1887]" captionTargetId="figure-130@31.[151,1436,561,1903]" captionTargetPageId="31" captionText="FIGURE 17. Habitus of Parena (Crossoglossa) laesipennis species group. A Lectotype of Parena laesipennis (Bates). B P. laesipennis (Bates), male from Taiwan. C P. laesipennis (Bates), female from Yunnan. D P. laesipennis (Bates), male from Nepal. E Holotype of Parena levata Andrewes. F P. levata Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958971" httpUri="https://zenodo.org/record/7958971/files/figure.png" pageId="33" pageNumber="34">Fig. 17A</figureCitation>
), designated herein: female, body length = 
<quantity id="4CD66AD06267FFC22CFAB2C0FE005D8F" box="[386,487,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="33" pageNumber="34" unit="mm" value="11.9">11.9 mm</quantity>
, pin mounted, &quot; 
<collectingRegion id="49EA09D76267FFC22FE0B2C0FCD35D8F" box="[664,820,187,213]" country="Japan" name="Nagasaki" pageId="33" pageNumber="34">NAGASAKI</collectingRegion>
&quot;, &quot;TYPE&quot; [red label], &quot; 
<collectorName id="26DBA2E36267FFC22940B2C0FAA15D8F" box="[1080,1350,187,213]" pageId="33" pageNumber="34">Ex Musaeo H.W. Bates</collectorName>
, 1892&quot;, &quot;MUSEUM PARIS 1952, Coll. R. OBERTHUR&quot;, &quot; 
<taxonomicName id="4C2EBCB66267FFC22FAEB2A5FBDD5DA3" authority="Bates" authorityName="Bates" authorityYear="1873" box="[726,1082,222,249]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="laesipennis">Crossoglossa laesipennis Bates</taxonomicName>
&quot; [hand written]
</materialsCitation>
. 
<materialsCitation id="3B46CD686267FFC2298FB2A4FAEB5C1A" ID-GBIF-Occurrence="4115871578" collectorName="Ex Musaeo H. W. Bates" location="NAGASAKI" pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" stateProvince="Nagasaki" typeStatus="paralectotype">
<emphasis id="B95A1B276267FFC2298FB2A4FA7B5DA3" bold="true" box="[1271,1436,223,249]" pageId="33" pageNumber="34">
<typeStatus id="549579976267FFC2298FB2A4FA7B5DA3" box="[1271,1436,223,249]" pageId="33" pageNumber="34" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
(MNHN): 
<specimenCount id="9D280CBC6267FFC22C68B379FE905C46" box="[272,375,258,284]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
, teneral, without abdomen, labia glued on board and solely pinned, &quot; 
<collectingRegion id="49EA09D76267FFC229F6B379FACD5C46" box="[1166,1322,258,284]" country="Japan" name="Nagasaki" pageId="33" pageNumber="34">NAGASAKI</collectingRegion>
&quot;, &quot;PARATYPE&quot; [red label], &quot; 
<collectorName id="26DBA2E36267FFC22C02B35DFD625C1B" box="[378,645,294,321]" pageId="33" pageNumber="34">Ex Musaeo H.W. Bates</collectorName>
, 1892&quot;, &quot;MUSEUM PARIS 1952, Coll. R. OBERTHUR&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6267FFC22DBFB331FB8C5F4D" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8B91C7356267FFC22DBFB331FB8C5F4D" blockId="33.[151,1437,150,2044]" pageId="33" pageNumber="34">
<emphasis id="B95A1B276267FFC22DBFB331FE925C3E" bold="true" box="[199,373,330,356]" pageId="33" pageNumber="34">Notes on types</emphasis>
. This species was originally described from an unspecified number of specimens from &quot; 
<collectingRegion id="49EA09D76267FFC2282CB331FF2D5CD2" country="Japan" name="Nagasaki" pageId="33" pageNumber="34">Nagasaki</collectingRegion>
&quot;, but more than 
<specimenCount id="9D280CBC6267FFC22CF9B314FDFC5CD2" box="[385,539,366,393]" pageId="33" pageNumber="34" type="generic">one specimen</specimenCount>
was implied. In the collection of MNHN, a total of 
<specimenCount id="9D280CBC6267FFC2291BB314FAED5CD2" box="[1123,1290,366,393]" pageId="33" pageNumber="34" type="generic">two specimens</specimenCount>
from Bates's collection accord with the original description. We herein designate the one bearing Bates's handwriting label (
<figureCitation id="1315DBB06267FFC22817B3E9FF2F5C8A" captionStart="FIGURE 17" captionStartId="31.[151,250,1926,1950]" captionTargetBox="[154,1424,569,1887]" captionTargetId="figure-130@31.[151,1436,561,1903]" captionTargetPageId="31" captionText="FIGURE 17. Habitus of Parena (Crossoglossa) laesipennis species group. A Lectotype of Parena laesipennis (Bates). B P. laesipennis (Bates), male from Taiwan. C P. laesipennis (Bates), female from Yunnan. D P. laesipennis (Bates), male from Nepal. E Holotype of Parena levata Andrewes. F P. levata Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958971" httpUri="https://zenodo.org/record/7958971/files/figure.png" pageId="33" pageNumber="34">Fig. 17A</figureCitation>
) as the 
<typeStatus id="549579976267FFC22C66B3CDFE6F5C8A" box="[286,392,438,464]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
. We also examined a female in NHML labeled as &quot;type&quot;, with a determination label not in accord with Bates's handwriting, and the locality &quot;Hiogo&quot; different from the type locality in the original description. Thus, this one is considered to be mislabeled as type and excluded from the type series.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6267FFC12DBFB059FD995954" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8B91C7356267FFC12DBFB059FD995954" blockId="33.[151,1437,150,2044]" lastBlockId="34.[151,1437,151,2007]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<emphasis id="B95A1B276267FFC22DBFB059FDFB5F61" bold="true" box="[199,540,545,572]" pageId="33" pageNumber="34">Non-type material examined</emphasis>
. 
<emphasis id="B95A1B276267FFC22F52B059FD945F66" bold="true" box="[554,627,546,572]" pageId="33" pageNumber="34">
<collectingCountry id="F33987A56267FFC22F52B059FD945F66" box="[554,627,546,572]" name="Japan" pageId="33" pageNumber="34">Japan</collectingCountry>
</emphasis>
: 
<specimenCount id="9D280CBC6267FFC22FFAB059FD045F61" box="[642,739,545,572]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(NHML), &quot;HIOGO&quot;, &quot;Japan. / G. Lewis. / 1910-320&quot;, &quot; 
<typeStatus id="549579976267FFC2282EB059FA735F66" box="[1366,1428,546,572]" pageId="33" pageNumber="34">Type</typeStatus>
&quot; [round label with red circle], &quot; 
<emphasis id="B95A1B276267FFC22C97B03EFC975F05" box="[495,880,581,608]" italics="true" pageId="33" pageNumber="34">
<taxonomicName id="4C2EBCB66267FFC22C97B03EFD625F05" authority="Chaudoir, 1873" box="[495,645,581,607]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Crossoglossa</taxonomicName>
/ 
<taxonomicName id="4C2EBCB66267FFC22FE4B03EFCFD5F05" authorityName="Bates" authorityYear="1873" box="[668,794,581,607]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="laesipennis">laesipennis</taxonomicName>
/ Bates
</emphasis>
&quot;. 
<specimenCount id="9D280CBC6267FFC22EF2B03EFC315F05" box="[906,982,581,607]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(IZAS), &quot;Mt. Tatsurayama Tsushima Is. 
<collectingRegion id="49EA09D76267FFC22DEFB012FEA65FD9" box="[151,321,617,643]" country="Japan" name="Nagasaki" pageId="33" pageNumber="34">Nagasaki Pref.</collectingRegion>
, 
<date id="FF90E1F56267FFC22C36B012FE335FD9" box="[334,468,617,643]" pageId="33" pageNumber="34" value="1989-08-02">1989.VIII.2</date>
, M. Ito leg.&quot;. 
<specimenCount id="9D280CBC6267FFC22F0EB012FD245FD9" box="[630,707,617,643]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(OMNH), &quot;Iwawaki 
<collectingRegion id="49EA09D76267FFC22EC1B012FBE55FD9" box="[953,1026,617,643]" country="Japan" name="Osaka" pageId="33" pageNumber="34">Osaka</collectingRegion>
, 
<date id="FF90E1F56267FFC22977B012FB775FD9" box="[1039,1168,617,644]" pageId="33" pageNumber="34" value="1959-06-27">1959.VI.27</date>
, T. Kawatsu&quot;. 
<specimenCount id="9D280CBC6267FFC22841B012FA7B5FD9" box="[1337,1436,617,643]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(OMNH), &quot;Izumi, Osaka-Pref., 
<date id="FF90E1F56267FFC22C80B0F6FD645FFD" box="[504,643,653,679]" pageId="33" pageNumber="34" value="1986-07-20">1986.VII.20</date>
, leg. O. Tohinaga&quot;. 
<specimenCount id="9D280CBC6267FFC22E26B0F6FC4F5FFD" box="[862,936,653,679]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(OMNH), &quot;Mt. Karasuga-dake, nr. Mikumo. Kouga 
<collectingRegion id="49EA09D76267FFC22D91B0CAFECB5F91" box="[233,300,689,715]" country="Japan" name="Shiga" pageId="33" pageNumber="34">Shiga</collectingRegion>
, 
<date id="FF90E1F56267FFC22C4FB0CAFE5E5F91" box="[311,441,689,715]" pageId="33" pageNumber="34" value="1980-11-30">1980.XI.30</date>
, Mitsuo Goto leg.&quot;. 
<specimenCount id="9D280CBC6267FFC22FE4B0CAFD1A5F91" box="[668,765,689,715]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(OMNH), &quot;Mt. Oikedake Mie-Pref., 
<date id="FF90E1F56267FFC229D9B0CAFAF85F91" box="[1185,1311,689,716]" pageId="33" pageNumber="34" value="1956-04-22">1956.IV.22</date>
, Coll. Zen. Naruse&quot;. 
<emphasis id="B95A1B276267FFC22C78B0AEFEAE5FB5" bold="true" box="[256,329,725,751]" pageId="33" pageNumber="34">
<collectingCountry id="F33987A56267FFC22C78B0AEFEAE5FB5" box="[256,329,725,751]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
</emphasis>
: 
<specimenCount id="9D280CBC6267FFC22C2FB0AEFE5F5FB5" box="[343,440,725,751]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(SNU), &quot; 
<collectingRegion id="49EA09D76267FFC22F67B0AEFD6E5FB5" box="[543,649,725,751]" country="China" name="Zhejiang" pageId="33" pageNumber="34">Zhejiang</collectingRegion>
, Lin’an, W. Tianmushan, 2008.VI., Huang Hao lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC229A5B0AEFAD95FB5" box="[1245,1342,725,751]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22DE7B082FEEE5E49" box="[159,265,761,787]" country="China" name="Zhejiang" pageId="33" pageNumber="34">Zhejiang</collectingRegion>
, W. Tianmushan, 
<quantity id="4CD66AD06267FFC22CA0B082FDD55E49" box="[472,562,761,787]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="m" value="1200.0">1200 m</quantity>
, Bi Wenxuan lgt., 
<date id="FF90E1F56267FFC22E74B082FC915E49" box="[780,886,761,787]" pageId="33" pageNumber="34" value="2010-05-02">2010.V.2</date>
&quot;. 
<specimenCount id="9D280CBC6267FFC22EF4B082FC175E49" box="[908,1008,761,787]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22908B082FB305E49" box="[1136,1239,761,787]" country="China" name="Zhejiang" pageId="33" pageNumber="34">Zhejiang</collectingRegion>
, W. Tianmushan, Song Haitian lgt., 
<date id="FF90E1F56267FFC22C0AB166FE305E6D" box="[370,471,797,823]" pageId="33" pageNumber="34" value="2010-05-01">2010.V.1</date>
&quot;. 1 ex (IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22FD9B166FD155E6D" box="[673,754,797,823]" country="China" name="Hunan" pageId="33" pageNumber="34">Hunan</collectingRegion>
, Ningxiang, Huaminglou, Shili vill. 
<date id="FF90E1F56267FFC229DBB166FAC35E62" box="[1187,1316,797,824]" pageId="33" pageNumber="34" value="2015-09-29">2015.IX.29</date>
D, 
<quantity id="4CD66AD06267FFC22832B166FA705E6D" box="[1354,1431,797,824]" metricMagnitude="2" metricUnit="m" metricValue="1.52" pageId="33" pageNumber="34" unit="m" value="152.0">152 m</quantity>
, 
<geoCoordinate id="EE1AA1F26267FFC22DEFB13AFEE15E01" box="[151,262,833,859]" degrees="28.0974" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="5" value="28.0974">N28.0974</geoCoordinate>
<geoCoordinate id="EE1AA1F26267FFC22C75B13BFE6D5E00" box="[269,394,832,858]" degrees="112.6638" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="5" value="112.6638">E112.6638</geoCoordinate>
, Zhao KD. lgt.&quot;. 1 ex (IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22F9FB13AFCDF5E01" box="[743,824,833,859]" country="China" name="Hunan" pageId="33" pageNumber="34">Hunan</collectingRegion>
, Ningxiang, Huaminglou, Shili vill. 
<date id="FF90E1F56267FFC229A8B13BFAB65E01" box="[1232,1361,832,859]" pageId="33" pageNumber="34" value="2015-09-27">2015.IX.27</date>
D, 
<quantity id="4CD66AD06267FFC2280AB13AFF485E25" metricMagnitude="2" metricUnit="m" metricValue="1.48" pageId="33" pageNumber="34" unit="m" value="148.0">148 m</quantity>
, 
<geoCoordinate id="EE1AA1F26267FFC22DC3B11EFECE5E25" box="[187,297,869,895]" degrees="28.0939" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="5" value="28.0939">N28.0939</geoCoordinate>
<geoCoordinate id="EE1AA1F26267FFC22C48B11FFE4A5E24" box="[304,429,868,894]" degrees="112.6640" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="5" value="112.664">E112.6640</geoCoordinate>
, Yao J. lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22F46B11EFD785E24" box="[574,671,868,895]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22E74B11EFCBA5E25" box="[780,861,869,895]" country="China" name="Hunan" pageId="33" pageNumber="34">Hunan</collectingRegion>
, Mangshan, 
<date id="FF90E1F56267FFC22E92B11EFB985E25" box="[1002,1151,869,895]" pageId="33" pageNumber="34" value="2007-08-24">2007.VIII.24</date>
, Tang Guo lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22844B11EFA7B5E24" box="[1340,1436,868,895]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22D85B1F3FEAC5EF8" box="[253,331,904,930]" country="China" name="Fujian" pageId="33" pageNumber="34">Fujian</collectingRegion>
, Jianyang, Huangkeng, Guilin, 
<quantity id="4CD66AD06267FFC22FDFB1F2FCC05EF9" box="[679,807,905,931]" metricMagnitude="2" metricUnit="m" metricValue="3.05" metricValueMax="3.4" metricValueMin="2.7" pageId="33" pageNumber="34" unit="m" value="305.0" valueMax="340.0" valueMin="270.0">270–340 m</quantity>
, 
<date id="FF90E1F56267FFC22E4AB1F3FC455EF8" box="[818,930,904,930]" pageId="33" pageNumber="34" value="1960-04-08">1960.IV.8</date>
, Zhang Yiran lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22901B1F2FB3E5EF8" box="[1145,1241,904,931]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC2283DB1F2FF355E9C" country="China" name="Guangdong" pageId="33" pageNumber="34">Guangdong</collectingRegion>
, Ruyuan, Nanling, Ruyang station, 
<geoCoordinate id="EE1AA1F26267FFC22F17B1D7FD085E9D" box="[623,751,940,967]" degrees="24.93450" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="24.9345">N24.93450</geoCoordinate>
, 
<geoCoordinate id="EE1AA1F26267FFC22F85B1D7FC605E9D" box="[765,903,940,967]" degrees="113.01588" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="113.01588">E113.01588</geoCoordinate>
; 
<quantity id="4CD66AD06267FFC22EEDB1D7FC085E9D" box="[917,1007,940,967]" metricMagnitude="3" metricUnit="m" metricValue="1.266" pageId="33" pageNumber="34" unit="m" value="1266.0">1266 m</quantity>
, 
<date id="FF90E1F56267FFC22E85B1D7FB6F5E9C" box="[1021,1160,940,966]" pageId="33" pageNumber="34" value="2008-07-20">2008.VII.20</date>
, Liang Hongbin lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC228F6B1D7FF045EB0" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22C2DB1ABFE4B5EB0" box="[341,428,976,1002]" country="China" name="Hainan" pageId="33" pageNumber="34">Hainan</collectingRegion>
, Bawangling, 
<quantity id="4CD66AD06267FFC22F29B1ABFD4D5EB1" box="[593,682,976,1003]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="34" unit="m" value="1000.0">1000 m</quantity>
, 
<date id="FF90E1F56267FFC22FC0B1ABFCCA5EB0" box="[696,813,976,1003]" pageId="33" pageNumber="34" value="1997-05-22">1997.V.22</date>
, Yu Peiyu lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22E93B1ABFBA95EB0" box="[1003,1102,976,1002]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC229B9B1ABFAFF5EB0" box="[1217,1304,976,1002]" country="China" name="Hainan" pageId="33" pageNumber="34">Hainan</collectingRegion>
, Limushan, Sanquling, 
<date id="FF90E1F56267FFC22C6EB18FFE715954" box="[278,406,1012,1038]" pageId="33" pageNumber="34" value="2007-11-30">2007.11.30</date>
, Zhu Xiaoyu lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22F1FB18FFD205954" box="[615,711,1012,1038]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(CAU), &quot; 
<collectingRegion id="49EA09D76267FFC22E57B18FFC615954" box="[815,902,1012,1038]" country="China" name="Hainan" pageId="33" pageNumber="34">Hainan</collectingRegion>
, Yinggeling, Baisha, Hongmao vill., 
<date id="FF90E1F56267FFC22867B18FFA7F5954" box="[1311,1432,1012,1038]" pageId="33" pageNumber="34" value="2007-10-19">2007.X.19</date>
, Wang Fang lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22C25B663FE4F5968" box="[349,424,1048,1074]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(ZWWC), &quot; 
<collectingRegion id="49EA09D76267FFC22F4AB663FD525968" box="[562,693,1048,1074]" country="China" name="Chongqing" pageId="33" pageNumber="34">Chongqing</collectingRegion>
, Jiangjin, Simianshan, 
<date id="FF90E1F56267FFC22EC1B663FBBC5968" box="[953,1115,1048,1074]" pageId="33" pageNumber="34" value="2007-10-22" valueMax="2007-10-23" valueMin="2007-10-22">2007.X.22–23</date>
, Zhang Weiwei lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22828B663FA7B5968" box="[1360,1436,1048,1074]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22D86B647FEBD590C" box="[254,346,1084,1110]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Xishuangbanna, Mengzhe, 
<quantity id="4CD66AD06267FFC22FE1B647FD05590C" box="[665,738,1084,1110]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="33" pageNumber="34" unit="m" value="870.0">870 m</quantity>
, 
<date id="FF90E1F56267FFC22F96B647FC97590C" box="[750,880,1084,1111]" pageId="33" pageNumber="34" value="1958-06-30">1958.VI.30</date>
, Wang Shuyong lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22914B647FB29590C" box="[1132,1230,1084,1110]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22844B647FA7F590C" box="[1340,1432,1084,1110]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Xishuangbanna, Xiaomengyang, 
<quantity id="4CD66AD06267FFC22F69B61BFDBA5920" box="[529,605,1120,1147]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="33" pageNumber="34" unit="m" value="850.0">850 m</quantity>
, 
<date id="FF90E1F56267FFC22F13B61BFD035921" box="[619,740,1120,1147]" pageId="33" pageNumber="34" value="1957-10-24">1957.X.24</date>
, Wang Shuyong lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22E90B61BFBAC5920" box="[1000,1099,1120,1146]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC229C6B61BFAFD5920" box="[1214,1306,1120,1146]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Xishuangbanna, Meng’a, 
<quantity id="4CD66AD06267FFC22C28B6F8FE0859C4" box="[336,495,1155,1182]" metricMagnitude="3" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" pageId="33" pageNumber="34" unit="m" value="1065.0" valueMax="1080.0" valueMin="1050.0">1050–1080 m</quantity>
, 
<date id="FF90E1F56267FFC22C84B6F8FD8859C4" box="[508,623,1155,1182]" pageId="33" pageNumber="34" value="1958-05-11">1958.V.11</date>
, Wang Shuyong lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22E09B6FFFC3359C4" box="[881,980,1156,1182]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC2293DB6FFFB4659C4" box="[1093,1185,1156,1182]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Mengla, Menglun 
<quantity id="4CD66AD06267FFC228F8B6F8FF59599B" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="33" pageNumber="34" unit="km" value="55.0">55 km</quantity>
, 
<quantity id="4CD66AD06267FFC22DB3B6D3FEF05998" box="[203,279,1192,1218]" metricMagnitude="2" metricUnit="m" metricValue="7.029999999999999" pageId="33" pageNumber="34" unit="m" value="703.0">703 m</quantity>
, 
<date id="FF90E1F56267FFC22C5DB6D3FE775998" box="[293,400,1192,1218]" pageId="33" pageNumber="34" value="2013-10-03">2013.X.3</date>
, Yang Xiaodong lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22FE0B6D3FD01599B" box="[664,742,1191,1218]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22E20B6D3FC535998" box="[856,948,1192,1218]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Fugong county, Pihe, Bajiao, 
<geoCoordinate id="EE1AA1F26267FFC22860B6DCFA7F5998" box="[1304,1432,1191,1218]" degrees="26.54816" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="26.54816">N26.54816</geoCoordinate>
, 
<geoCoordinate id="EE1AA1F26267FFC22DEFB6B0FEF359BC" box="[151,276,1227,1254]" degrees="98.89576" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="98.89576">E98.89576</geoCoordinate>
, 
<quantity id="4CD66AD06267FFC22C58B6B7FE9159BC" box="[288,374,1228,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="33" pageNumber="34" unit="m" value="1120.0">1120 m</quantity>
, 
<date id="FF90E1F56267FFC22CFAB6B0FDF059BC" box="[386,535,1227,1254]" pageId="33" pageNumber="34" value="2005-08-23">2005.VIII.23</date>
, Liang Hongbin lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22E6BB6B7FC9359BF" box="[787,884,1227,1254]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(CAS), &quot; 
<collectingRegion id="49EA09D76267FFC22EA2B6B7FBD159BC" box="[986,1078,1228,1254]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Tengchong county, Mangbang, Longwenqiao, 
<geoCoordinate id="EE1AA1F26267FFC22C46B694FE585850" box="[318,447,1263,1290]" degrees="25.02329" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="25.02329">N25.02329</geoCoordinate>
, 
<geoCoordinate id="EE1AA1F26267FFC22CB0B694FDA15853" box="[456,582,1263,1289]" degrees="98.67710" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="98.6771">E98.67710</geoCoordinate>
, 
<quantity id="4CD66AD06267FFC22F28B694FD415850" box="[592,678,1263,1290]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="33" pageNumber="34" unit="m" value="1290.0">1290 m</quantity>
, 
<date id="FF90E1F56267FFC22FC9B694FCC35850" box="[689,804,1263,1290]" pageId="33" pageNumber="34" value="2006-06-05">2006.VI.5</date>
, Liang Hongbin &amp; Hu Peng lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC229E6B694FB195853" box="[1182,1278,1263,1289]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(CAS), &quot; 
<collectingRegion id="49EA09D76267FFC2281AB694FF255874" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Tengchong coungty, Mangbang, Longwenqiao, 
<geoCoordinate id="EE1AA1F26267FFC22F8BB768FC945874" box="[755,883,1299,1326]" degrees="25.02396" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="25.02396">N25.02396</geoCoordinate>
, 
<geoCoordinate id="EE1AA1F26267FFC22EF9B768FC195874" box="[897,1022,1299,1326]" degrees="98.67675" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="98.67675">E98.67675</geoCoordinate>
, 
<quantity id="4CD66AD06267FFC22974B768FB815874" box="[1036,1126,1299,1326]" metricMagnitude="3" metricUnit="m" metricValue="1.285" pageId="33" pageNumber="34" unit="m" value="1285.0">1285 m</quantity>
, 
<date id="FF90E1F56267FFC2290DB768FB0E5874" box="[1141,1257,1299,1326]" pageId="33" pageNumber="34" value="2006-06-05">2006.VI.5</date>
, Kavanaugh D. &amp; Brett R. lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22C19B74CFE21580B" box="[353,454,1335,1361]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22F44B74CFD7F580B" box="[572,664,1335,1361]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Gongshan county, btw. Pulahekou and Gazutian, 
<date id="FF90E1F56267FFC2299DB74CFA805808" box="[1253,1383,1335,1362]" pageId="33" pageNumber="34" value="2007-09-25">2007.IX.25</date>
, Shi Hongliang lgt.&quot; &lt;
<figureCitation id="1315DBB06267FFC22C24B720FE24582F" box="[348,451,1371,1397]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="33" pageNumber="34">Figs 11Z</figureCitation>
, 
<figureCitation id="1315DBB06267FFC22CA8B720FDE7582F" box="[464,512,1371,1397]" captionStart="FIGURE 17" captionStartId="31.[151,250,1926,1950]" captionTargetBox="[154,1424,569,1887]" captionTargetId="figure-130@31.[151,1436,561,1903]" captionTargetPageId="31" captionText="FIGURE 17. Habitus of Parena (Crossoglossa) laesipennis species group. A Lectotype of Parena laesipennis (Bates). B P. laesipennis (Bates), male from Taiwan. C P. laesipennis (Bates), female from Yunnan. D P. laesipennis (Bates), male from Nepal. E Holotype of Parena levata Andrewes. F P. levata Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958971" httpUri="https://zenodo.org/record/7958971/files/figure.png" pageId="33" pageNumber="34">17C</figureCitation>
&gt;. 
<specimenCount id="9D280CBC6267FFC22F65B720FD8E582F" box="[541,617,1371,1397]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(IZAS), &quot;Tengchong, Qushi, Heiyuhe, 
<quantity id="4CD66AD06267FFC2295EB720FB99582C" box="[1062,1150,1371,1398]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="33" pageNumber="34" unit="m" value="1490.0">1490 m</quantity>
, 
<date id="FF90E1F56267FFC229F3B720FB11582F" box="[1163,1270,1371,1397]" pageId="33" pageNumber="34" value="2002-10-01">2002.X.1</date>
, Situ Yingxian lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22DACB704FEF958C3" box="[212,286,1407,1433]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(IZAS), &quot;Yongde county, Yinchang, 
<quantity id="4CD66AD06267FFC22FC1B704FCF758C3" box="[697,784,1407,1433]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="33" pageNumber="34" unit="m" value="2300.0">2300 m</quantity>
, 
<date id="FF90E1F56267FFC22E63B704FC7B58C3" box="[795,924,1407,1433]" pageId="33" pageNumber="34" value="2002-06-29">2002.VI.29</date>
, Song Jinxin &amp; Huang Kunyun lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22844B704FA7B58C3" box="[1340,1436,1407,1433]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22C79B7D8FEBA58E7" box="[257,349,1443,1469]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Ruili, Nongdao town, way btw Dengga and Sepengqiao, 
<geoCoordinate id="EE1AA1F26267FFC2297AB7D8FB6558E4" box="[1026,1154,1443,1470]" degrees="23.95285" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="23.95285">N23.95285</geoCoordinate>
, 
<geoCoordinate id="EE1AA1F26267FFC229E8B7D8FAEC58E4" box="[1168,1291,1443,1470]" degrees="97.59808" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="97.59808">E97.59808</geoCoordinate>
– 
<geoCoordinate id="EE1AA1F26267FFC22862B7D8FA7F58E4" box="[1306,1432,1443,1470]" degrees="23.97518" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="23.97518">N23.97518</geoCoordinate>
, 
<geoCoordinate id="EE1AA1F26267FFC22DEFB7BDFEF358BB" box="[151,276,1478,1505]" degrees="97.56944" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="97.56944">E97.56944</geoCoordinate>
, 927– 
<quantity id="4CD66AD06267FFC22C21B7BCFE4558BB" box="[345,418,1479,1505]" metricMagnitude="2" metricUnit="m" metricValue="8.07" pageId="33" pageNumber="34" unit="m" value="807.0">807 m</quantity>
, 
<date id="FF90E1F56267FFC22CD6B7BCFDA558BB" box="[430,578,1479,1505]" pageId="33" pageNumber="34" value="2009-08-11">2009.VIII.11</date>
, on vegetation, Shi Hongliang lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22EA6B7BCFBCD58BB" box="[990,1066,1479,1505]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC229E1B7BCFB1258BB" box="[1177,1269,1479,1505]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Jingmai, Huimin, Lancang, 100°01’34.42’’E, 22°12’04.79’’N, 
<quantity id="4CD66AD06267FFC22FB6B790FCC05B5F" box="[718,807,1515,1541]" metricMagnitude="3" metricUnit="m" metricValue="1.488" pageId="33" pageNumber="34" unit="m" value="1488.0">1488 m</quantity>
, Jianfeng Liang, Chengcong Yu, 
<date id="FF90E1F56267FFC229DBB790FACA5B5F" box="[1187,1325,1515,1541]" pageId="33" pageNumber="34" value="2017-07-11">2017.VII.11</date>
, Net capture&quot;. 
<specimenCount id="9D280CBC6267FFC22DA4B474FECF5B72" box="[220,296,1550,1577]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22CEEB474FE155B73" box="[406,498,1551,1577]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Ruili, Bangda vill, 
<quantity id="4CD66AD06267FFC22FADB474FCCB5B73" box="[725,812,1551,1577]" metricMagnitude="3" metricUnit="m" metricValue="1.432" pageId="33" pageNumber="34" unit="m" value="1432.0">1432 m</quantity>
, 
<date id="FF90E1F56267FFC22E41B475FC565B73" box="[825,945,1550,1577]" pageId="33" pageNumber="34" value="2015-09-03">2015-IX-3</date>
, Yang Xiaodong lgt.&quot; &lt;
<figureCitation id="1315DBB06267FFC229C2B475FAF95B73" box="[1210,1310,1550,1577]" captionStart="FIGURE 18" captionStartId="32.[151,250,1101,1125]" captionTargetBox="[164,1427,365,1072]" captionTargetId="figure-113@32.[151,1436,352,1078]" captionTargetPageId="32" captionText="FIGURE 18. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Yunnan. B Yunnan. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958973" httpUri="https://zenodo.org/record/7958973/files/figure.png" pageId="33" pageNumber="34">Fig. 18B</figureCitation>
&gt;. 
<specimenCount id="9D280CBC6267FFC22843B474FA7B5B72" box="[1339,1436,1550,1577]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC22D87B449FEBC5B16" box="[255,347,1586,1612]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Ruili, Bangda vill, 
<quantity id="4CD66AD06267FFC22F38B449FD7F5B17" box="[576,664,1586,1613]" metricMagnitude="3" metricUnit="m" metricValue="1.432" pageId="33" pageNumber="34" unit="m" value="1432.0">1432 m</quantity>
, 
<date id="FF90E1F56267FFC22FDDB449FCCB5B17" box="[677,812,1586,1613]" pageId="33" pageNumber="34" value="2015-09-10">2015-IX-10</date>
, Yang Xiaodong lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22957B449FB9C5B16" box="[1071,1147,1586,1612]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(IZAS), &quot;China, 
<collectingRegion id="49EA09D76267FFC22838B448FA7E5B17" box="[1344,1433,1587,1613]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Wenshan, Zhuilijie, 
<quantity id="4CD66AD06267FFC22C01B42DFE375B2B" box="[377,464,1622,1649]" metricMagnitude="3" metricUnit="m" metricValue="1.734" pageId="33" pageNumber="34" unit="m" value="1734.0">1734 m</quantity>
, 
<date id="FF90E1F56267FFC22CA4B42DFDB25B2A" box="[476,597,1622,1648]" pageId="33" pageNumber="34" value="2019-05-23">2019.V-23</date>
, Leg. Y.-Q. LU&quot;&lt;
<figureCitation id="1315DBB06267FFC22E65B42DFC655B2A" box="[797,898,1622,1648]" captionStart="FIGURE 18" captionStartId="32.[151,250,1101,1125]" captionTargetBox="[164,1427,365,1072]" captionTargetId="figure-113@32.[151,1436,352,1078]" captionTargetPageId="32" captionText="FIGURE 18. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Yunnan. B Yunnan. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958973" httpUri="https://zenodo.org/record/7958973/files/figure.png" pageId="33" pageNumber="34">Fig. 18A</figureCitation>
&gt;. 
<specimenCount id="9D280CBC6267FFC22EE5B42DFC0F5B2A" box="[925,1000,1622,1648]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(IZAS), &quot; 
<collectingRegion id="49EA09D76267FFC2292EB42DFB4A5B2A" box="[1110,1197,1622,1648]" country="China" name="Xizang" pageId="33" pageNumber="34">Xizang</collectingRegion>
, Mêdog county, 
<quantity id="4CD66AD06267FFC2281FB42DFF485BCF" metricMagnitude="3" metricUnit="m" metricValue="1.111" pageId="33" pageNumber="34" unit="m" value="1111.0">1111 m</quantity>
, 
<date id="FF90E1F56267FFC22DC5B401FEA95BCE" box="[189,334,1658,1684]" pageId="33" pageNumber="34" value="2014-07-30">2014-VII-30</date>
, Yang Xiaodong lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22F2CB401FD435BCE" box="[596,676,1658,1684]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
, 
<specimenCount id="9D280CBC6267FFC22FCAB401FCF25BCE" box="[690,789,1658,1684]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(CCCC), &quot;Taiwan, Taitung county, Orchid Island, 
<date id="FF90E1F56267FFC22825B401FF595BE2" pageId="33" pageNumber="34" value="2006-05-09">2006. V.9</date>
, Chen Changchin lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC22CA2B4E5FDCE5BE2" box="[474,553,1694,1720]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(CCCC), &quot;Taiwan, Taipei, Yingchiling, 
<date id="FF90E1F56267FFC22E85B4E5FB675BE2" box="[1021,1152,1694,1720]" pageId="33" pageNumber="34" value="1999-06-30">1999.VI.30</date>
, Chou Wen-I lgt.&quot; &lt;
<figureCitation id="1315DBB06267FFC22808B4E5FF2F5B86" captionStart="FIGURE 19" captionStartId="32.[151,250,1946,1970]" captionTargetBox="[161,1425,1199,1917]" captionTargetId="figure-151@32.[151,1436,1186,1923]" captionTargetPageId="32" captionText="FIGURE 19. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates) from different localities, left-lateral (l) and dorsal (d) views: A Laos. B Taiwan. C Nepal. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958975" httpUri="https://zenodo.org/record/7958975/files/figure.png" pageId="33" pageNumber="34">Fig. 19B</figureCitation>
&gt;. 
<specimenCount id="9D280CBC6267FFC22D9DB4B9FEA05B86" box="[229,327,1730,1756]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(CCCC), &quot;Taiwan, Nantou county, Puli town, Shitsaitou, 
<date id="FF90E1F56267FFC22EA0B4B9FBB25B87" box="[984,1109,1730,1757]" pageId="33" pageNumber="34" value="1994-04-15">1994.IV.15</date>
, Chen Changchin lgt.&quot;. 
<specimenCount id="9D280CBC6267FFC2281FB4B9FF2A5A5A" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(CCCC), &quot;Taiwan, Pintung county, Hengchun town, Nanrenshan, 
<date id="FF90E1F56267FFC22EBEB49DFBB65A5A" box="[966,1105,1766,1792]" pageId="33" pageNumber="34" value="1992-12-17">1992.XII.17</date>
, Chou Wen-I lgt.&quot; &lt;
<figureCitation id="1315DBB06267FFC22843B49DFA7F5A5A" box="[1339,1432,1766,1792]" captionStart="FIGURE 4" captionStartId="11.[151,250,1555,1579]" captionTargetBox="[155,1431,887,1528]" captionTargetId="figure-419@11.[151,1436,885,1532]" captionTargetPageId="11" captionText="FIGURE 4. Elytral base (left) of Parena spp. A P. (Parena) bicolor Motschulsky, Sumatra. B P. (Parena) africana (Alluaud), Cameroon. C P. (Crossoglossa) laesipennis (Bates), Taiwan. D P. (Bothynoptera) tripunctata (Bates), Beijing. E P. (Bothynoptera) dorsigera (Schaum), Guangxi. F P. (Bothynoptera) heteronycha sp. n., Laos, paratype. Scale bars = 0.5mm (a for fig. D, b for figs A B E F, c for fig. C). Abbreviations: bp elytral basal pore; sa the apex of scutellum; d1 the first discal setigerous pore." figureDoi="http://doi.org/10.5281/zenodo.7958945" httpUri="https://zenodo.org/record/7958945/files/figure.png" pageId="33" pageNumber="34">Figs 4C</figureCitation>
, 
<figureCitation id="1315DBB06267FFC22DEFB571FF275A7E" box="[151,192,1802,1828]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="33" pageNumber="34">14J</figureCitation>
&gt;. 
<specimenCount id="9D280CBC6267FFC22DA7B571FEA45A7E" box="[223,323,1802,1828]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(CCCC), &quot;Taiwan, Taipei county, Sanhsia town, 
<date id="FF90E1F56267FFC22E05B571FC045A7E" box="[893,995,1802,1828]" pageId="33" pageNumber="34" value="1992-05-02">1992.V.2</date>
, Chen Changchin lgt.&quot;. 2 ex (CCCC), &quot;Taiwan, Taoyuan county, Fuhsing, Sankuang, 
<date id="FF90E1F56267FFC22FDBB555FCD55A12" box="[675,818,1838,1864]" pageId="33" pageNumber="34" value="1994-07-14">1994.VII.14</date>
&quot;. 1 ex (MTMB), &quot;Taiwan, Nantou county, Puli, singled, 
<date id="FF90E1F56267FFC22DEFB529FEFE5A36" box="[151,281,1874,1900]" pageId="33" pageNumber="34" value="2002-11-19">19.XI.2002</date>
, leg. H.R. Tzuoo&quot;. 
<emphasis id="B95A1B276267FFC22C96B52AFDB35A31" bold="true" box="[494,596,1873,1899]" pageId="33" pageNumber="34">
<collectingCountry id="F33987A56267FFC22C96B52AFDB35A31" box="[494,596,1873,1899]" name="Vietnam" pageId="33" pageNumber="34">Vietnam</collectingCountry>
</emphasis>
: 1 ex (MNHN), &quot;Tonkin, Reg de 
<collectingRegion id="49EA09D76267FFC22EBFB529FBD35A31" box="[967,1076,1873,1900]" country="Vietnam" name="Hoa Binh" pageId="33" pageNumber="34">Hoa Binh</collectingRegion>
, A. de Cooman, 1927&quot;. 
<specimenCount id="9D280CBC6267FFC22844B529FA7B5A31" box="[1340,1436,1873,1900]" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
(CRS), &quot;Vietnam, 
<collectingRegion id="49EA09D76267FFC22C1BB50DFE195ACA" box="[355,510,1909,1936]" country="Vietnam" name="Lao Cai" pageId="33" pageNumber="34">Lao Cai prov.</collectingRegion>
, near Sa Pa, Cat Cat vill. env. H= 
<quantity id="4CD66AD06267FFC22E15B50DFC245ACA" box="[877,963,1910,1936]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="m" value="1200.0">1200 m</quantity>
, 
<date id="FF90E1F56267FFC22EB6B50DFB895ACA" box="[974,1134,1910,1936]" pageId="33" pageNumber="34" value="2018-05-10" valueMax="2018-05-20" valueMin="2018-05-10">10–20.V.2018</date>
&quot; &lt;
<figureCitation id="1315DBB06267FFC229F4B50EFB085ACA" box="[1164,1263,1909,1936]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="33" pageNumber="34">Fig. 14B</figureCitation>
&gt;. 
<emphasis id="B95A1B276267FFC22872B50DFAA35ACA" bold="true" box="[1290,1348,1910,1936]" pageId="33" pageNumber="34">
<collectingCountry id="F33987A56267FFC22872B50DFAA35ACA" box="[1290,1348,1910,1936]" name="Laos" pageId="33" pageNumber="34">Laos</collectingCountry>
</emphasis>
: 
<specimenCount id="9D280CBC6267FFC2282AB50DFA7B5AD5" box="[1362,1436,1909,1936]" pageId="33" pageNumber="34" type="male">1 male</specimenCount>
(NHMB), &quot;LAO, 
<collectingRegion id="49EA09D76267FFC22C1CB5E2FDFB5AEE" box="[356,540,1945,1972]" country="Laos" name="Phongsali" pageId="33" pageNumber="34">Phongsaly prov.</collectingRegion>
21°41-2' 
<geoCoordinate id="EE1AA1F26267FFC22FFAB5E2FD0E5AE9" box="[642,745,1945,1971]" degrees="102" direction="north" minutes="6" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="102.1">N 102°6</geoCoordinate>
-8'E, 
<date id="FF90E1F56267FFC22E5CB5E2FC395AE9" box="[804,990,1945,1971]" pageId="33" pageNumber="34" value="2003-05-28" valueMax="2003-06-20" valueMin="2003-05-28">28.v.-20.vi.2003</date>
, 
<collectingRegion id="49EA09D76267FFC22E94B5E2FB865AE9" box="[1004,1121,1945,1971]" country="Laos" name="Phongsali" pageId="33" pageNumber="34">Phongsaly</collectingRegion>
env., ~ 
<quantity id="4CD66AD06267FFC229CFB5E2FAF65AEE" box="[1207,1297,1945,1972]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="34" unit="m" value="1500.0">1500 m</quantity>
, Pacholatko leg.&quot;. 2 ex (NHMB), &quot;LAO, 
<collectingRegion id="49EA09D76267FFC22CA0B5C6FD6A5A82" box="[472,653,1981,2008]" country="Laos" name="Phongsali" pageId="33" pageNumber="34">Phongsaly prov.</collectingRegion>
21°41-2' 
<geoCoordinate id="EE1AA1F26267FFC22F88B5C6FCB25A8D" box="[752,853,1981,2007]" degrees="102" direction="north" minutes="6" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="102.1">N 102°6</geoCoordinate>
-8'E, 
<date id="FF90E1F56267FFC22EF4B5C6FBA15A8D" box="[908,1094,1981,2007]" pageId="33" pageNumber="34" value="2003-05-28" valueMax="2003-06-20" valueMin="2003-05-28">28.v.-20.vi.2003</date>
, 
<collectingRegion id="49EA09D76267FFC2292AB5C6FB205A8D" box="[1106,1223,1981,2007]" country="Laos" name="Phongsali" pageId="33" pageNumber="34">Phongsaly</collectingRegion>
env., ~ 
<quantity id="4CD66AD06267FFC2286FB5C6FA885A82" box="[1303,1391,1981,2008]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="34" unit="m" value="1500.0">1500 m</quantity>
, Vit Kuban leg.&quot;. 
<specimenCount id="9D280CBC6267FFC22C53B59AFE855AA6" box="[299,354,2017,2044]" pageId="33" pageNumber="34" type="generic">2 ex.</specimenCount>
(NHMB), &quot;LAOS-NE, 
<collectingRegion id="49EA09D76267FFC22F16B59AFC875AA6" box="[622,864,2017,2044]" country="Laos" name="Xiangkhouang" pageId="33" pageNumber="34">Xieng Khouang prov.</collectingRegion>
19°37-8' 
<geoCoordinate id="EE1AA1F26267FFC22EBBB59AFBDE5AA1" box="[963,1081,2017,2043]" degrees="103" direction="north" minutes="20" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="103.333336">N 103°20</geoCoordinate>
-1'E, 
<quantity id="4CD66AD06267FFC22915B59AFB535AA1" box="[1133,1204,2017,2043]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="34" unit="km" value="30.0">30 km</quantity>
NE Phonsavan: Ban Ba Lam- Phou Sane Mt., 
<quantity id="4CD66AD06264FFC12CC8B2ECFDAB5DEB" box="[432,588,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="34" pageNumber="35" unit="m" value="1500.0" valueMax="1700.0" valueMin="1300.0">1300–1700 m</quantity>
, 
<date id="FF90E1F56264FFC12F2FB2ECFD155DEB" box="[599,754,151,177]" pageId="34" pageNumber="35" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">10–30.v.2009</date>
, M. Geiser leg.&quot;. 3 ex (NHMB), &quot;LAOS-NE, 
<collectingRegion id="49EA09D76264FFC12993B2ECFF375D8F" country="Laos" name="Xiangkhouang" pageId="34" pageNumber="35">Xieng Khouang prov.</collectingRegion>
19°37-8' 
<geoCoordinate id="EE1AA1F26264FFC12C4CB2C0FE4C5D8F" box="[308,427,187,213]" degrees="103" direction="north" minutes="20" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="103.333336">N 103°20</geoCoordinate>
-1'E, Phonsavan (
<quantity id="4CD66AD06264FFC12F11B2C1FD4C5D8E" box="[617,683,186,212]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="35" unit="km" value="30.0">30km</quantity>
NE): Phou Sane Mt., 
<quantity id="4CD66AD06264FFC12EDFB2C0FBA25D8F" box="[935,1093,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="34" pageNumber="35" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">1400–1700 m</quantity>
, 
<date id="FF90E1F56264FFC12929B2C0FB005D8F" box="[1105,1255,187,213]" pageId="34" pageNumber="35" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">10-30.v.2009</date>
, D. Hauck leg.&quot; &lt;
<figureCitation id="1315DBB06264FFC12DDFB2A5FEE95DA3" box="[167,270,222,249]" captionStart="FIGURE 19" captionStartId="32.[151,250,1946,1970]" captionTargetBox="[161,1425,1199,1917]" captionTargetId="figure-151@32.[151,1436,1186,1923]" captionTargetPageId="32" captionText="FIGURE 19. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates) from different localities, left-lateral (l) and dorsal (d) views: A Laos. B Taiwan. C Nepal. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958975" httpUri="https://zenodo.org/record/7958975/files/figure.png" pageId="34" pageNumber="35">Fig. 19A</figureCitation>
&gt;. 1 ex (NHMB), &quot;LAOS-NE, 
<collectingRegion id="49EA09D76264FFC12F10B2A5FCBC5DA3" box="[616,859,222,249]" country="Laos" name="Xiangkhouang" pageId="34" pageNumber="35">Xieng Khouang prov.</collectingRegion>
19°37-8' 
<geoCoordinate id="EE1AA1F26264FFC12EC7B2A4FBD25DA3" box="[959,1077,223,249]" degrees="103" direction="north" minutes="20" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="103.333336">N 103°20'</geoCoordinate>
E, Phonsavan (
<quantity id="4CD66AD06264FFC129A4B2A5FAFA5DA2" box="[1244,1309,222,248]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="35" unit="km" value="30.0">30km</quantity>
NE): Phou Sane Mt., 
<quantity id="4CD66AD06264FFC12C76B379FE4B5C47" box="[270,428,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="34" pageNumber="35" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400–1500 m</quantity>
, 
<date id="FF90E1F56264FFC12CC2B379FDA85C46" box="[442,591,258,284]" pageId="34" pageNumber="35" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">10-30.v.2009</date>
, Z. Kraus leg.&quot;. 1 ex (NHMB), &quot;LAOS-NE, Houa Phan prov. 20°11-13’ 
<geoCoordinate id="EE1AA1F26264FFC128FCB379FF125C1B" degrees="103" direction="north" minutes="59" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="103.98333">N 103°59’</geoCoordinate>
-104°1’E, Ban Saluei-Phou Pane Mt, 
<date id="FF90E1F56264FFC12FE5B35DFCD25C1A" box="[669,821,294,320]" pageId="34" pageNumber="35" value="2009-06-09" valueMax="2009-06-17" valueMin="2009-06-09">9.-17.vi.2009</date>
, 
<quantity id="4CD66AD06264FFC12E3AB35DFC075C1B" box="[834,992,294,321]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="34" pageNumber="35" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">1300–1900 m</quantity>
, M. Geiser leg.&quot;. 
<emphasis id="B95A1B276264FFC129D2B35DFACD5C1A" bold="true" box="[1194,1322,294,320]" pageId="34" pageNumber="35">Myammar</emphasis>
: 
<specimenCount id="9D280CBC6264FFC12842B35DFA7B5C1A" box="[1338,1436,294,320]" pageId="34" pageNumber="35" type="female">1 female</specimenCount>
(NHML), &quot;Birmah, Rubymes, Dohelly&quot;, &quot;60863&quot;, &quot;Fry coll. 1905.100&quot;. 
<emphasis id="B95A1B276264FFC12EB5B331FBDD5C3E" bold="true" box="[973,1082,330,356]" pageId="34" pageNumber="35">
<collectingCountry id="F33987A56264FFC12EB5B331FBDD5C3E" box="[973,1082,330,356]" name="Thailand" pageId="34" pageNumber="35">Thailand</collectingCountry>
</emphasis>
: 1 ex (CMB), &quot;N.W. Thailand, 
<collectingRegion id="49EA09D76264FFC12DEFB314FEA35CD2" box="[151,324,366,393]" country="Thailand" name="Mae Hong Son" pageId="34" pageNumber="35">Mae Hong Son</collectingRegion>
, 17– 
<date id="FF90E1F56264FFC12C01B315FE0A5CD3" box="[377,493,366,393]" pageId="34" pageNumber="35" value="1999-05-18">18.5.1999</date>
, R. Grimm&quot;. 1 ex (CSF), &quot;Thailand, 
<collectingRegion id="49EA09D76264FFC12EF5B315FBB35CD3" box="[909,1108,366,393]" country="Thailand" name="Chiang Mai" pageId="34" pageNumber="35">Chiang Mai prov.</collectingRegion>
<quantity id="4CD66AD06264FFC12922B315FB455CD2" box="[1114,1186,366,393]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="34" pageNumber="35" unit="km" value="25.0">25 km</quantity>
N of CHIANG DAO, 28- 
<date id="FF90E1F56264FFC12DC2B3E9FEC85CF6" box="[186,303,402,428]" pageId="34" pageNumber="35" value="2002-06-30">30.6.2002</date>
, WGS 84: 19°40'N, 098°50'E, lgt. Fouque R.+H.&quot;. 1 ex (CSF), &quot;Thailand, 7.-12.5. 
<collectingRegion id="49EA09D76264FFC1287AB3E9FEF65C8A" country="Thailand" name="Mae Hong Son" pageId="34" pageNumber="35">MAE HONG SON prov.</collectingRegion>
SOPPONG, 
<quantity id="4CD66AD06264FFC12CD0B3CDFDE65C8A" box="[424,513,438,465]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="m" value="1500.0">1500 m</quantity>
, 19°27' 
<geoCoordinate id="EE1AA1F26264FFC12F2DB3CDFD595C8A" box="[597,702,438,464]" degrees="98" direction="north" minutes="20" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="98.333336">N 98°20'</geoCoordinate>
E, lgt. S. Becvar, 1996&quot;. 
<emphasis id="B95A1B276264FFC12EA3B3CEFBFC5C95" bold="true" box="[987,1051,437,463]" pageId="34" pageNumber="35">
<collectingCountry id="F33987A56264FFC12EA3B3CEFBFC5C95" box="[987,1051,437,463]" name="India" pageId="34" pageNumber="35">India</collectingCountry>
</emphasis>
: 1 ex (NHMB), &quot;Durpin 
<quantity id="4CD66AD06264FFC12847B3CDFA705C8A" box="[1343,1431,438,464]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="34" pageNumber="35" unit="m" value="1200.0">1200 m</quantity>
, 
<date id="FF90E1F56264FFC12DEFB3A1FEFC5CAE" box="[151,283,474,500]" pageId="34" pageNumber="35" value="1986-03-26">26.III.1986</date>
&quot;; &quot;Indian Darjeeling D. Bhakta B.&quot;. 1 ex (NHMB), &quot;Chuba Busty, 
<quantity id="4CD66AD06264FFC12964B3A1FB925CAE" box="[1052,1141,474,500]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="34" pageNumber="35" unit="m" value="1080.0">1080 m</quantity>
, 
<date id="FF90E1F56264FFC129FBB3A1FAE25CAE" box="[1155,1285,474,500]" pageId="34" pageNumber="35" value="1984-04-18">18.IV.1984</date>
&quot;; &quot;Darjeeling D. India Bhakta B.&quot;. 1 ex (NHMB), &quot;India VII–VIII, Darjeeling D. Bhakta B. 1985&quot;. 
<specimenCount id="9D280CBC6264FFC12909B385FB595F4D" box="[1137,1214,509,536]" pageId="34" pageNumber="35" type="male">1 male</specimenCount>
(NHMB), &quot;&quot;Indien Darjeeling D. Bhakta B.&quot;, &quot;Reenak, 9-13.V.90&quot;. 1 ex (NHMB), &quot;Kalimp. 
<quantity id="4CD66AD06264FFC12EACB059FBCB5F66" box="[980,1068,546,572]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="35" unit="m" value="1000.0">1000 m</quantity>
, Rinkingpong, 
<date id="FF90E1F56264FFC129A1B059FABB5F66" box="[1241,1372,546,572]" pageId="34" pageNumber="35" value="1987-04-22">22.IV.1987</date>
&quot;; &quot;Indien Darjeeling D. Ch.J. Rai&quot;. 1 ex (NHMB), &quot;Kalimpong, 
<date id="FF90E1F56264FFC12E42B03EFC5D5F3A" box="[826,954,581,608]" pageId="34" pageNumber="35" value="1981-07-05">5.VII.1981</date>
&quot;; &quot;Darjeeling Distr. India Bhakta B.&quot;. 1 ex (MNHN), &quot;Rhenok, 3000p., VII. 
<date id="FF90E1F56264FFC12F70B012FD9E5FD9" box="[520,633,617,643]" pageId="34" pageNumber="35" value="1900-08">VIII.1900</date>
, 
<collectingRegion id="49EA09D76264FFC12FFDB012FD395FD9" box="[645,734,617,643]" country="India" name="Sikkim" pageId="34" pageNumber="35">Sikkim</collectingRegion>
?&quot;. 
<specimenCount id="9D280CBC6264FFC12F85B012FCB95FD9" box="[765,862,617,643]" pageId="34" pageNumber="35" type="female">1 female</specimenCount>
(NHMB), &quot;NE India, 
<collectingRegion id="49EA09D76264FFC12923B012FB3D5FD9" box="[1115,1242,617,643]" country="India" name="Meghalaya" pageId="34" pageNumber="35">Meghalaya</collectingRegion>
, Tura peak, 
<quantity id="4CD66AD06264FFC1281CB012FF085FF2" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="34" pageNumber="35" unit="m" value="800.0" valueMax="1000.0" valueMin="600.0">600– 1000 m</quantity>
, 25°30’ 
<geoCoordinate id="EE1AA1F26264FFC12C3DB0F6FE4E5FFD" box="[325,425,653,680]" degrees="90" direction="north" minutes="14" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="90.23333">N 90°14</geoCoordinate>
&quot;E, L. Dembicky leg., 
<date id="FF90E1F56264FFC12FDDB0F6FCA85FFD" box="[677,847,653,679]" pageId="34" pageNumber="35" value="2007-06-12" valueMax="2007-06-22" valueMin="2007-06-12">12.-22.vi.2007</date>
&quot;. 
<emphasis id="B95A1B276264FFC12E1AB0F6FC4F5FFD" bold="true" box="[866,936,653,679]" pageId="34" pageNumber="35">
<collectingCountry id="F33987A56264FFC12E1AB0F6FC4F5FFD" box="[866,936,653,679]" name="Nepal" pageId="34" pageNumber="35">Nepal</collectingCountry>
</emphasis>
: 1 ex (NHMB), &quot;Burhanilkanth, 
<quantity id="4CD66AD06264FFC12866B0F6FF485F96" metricMagnitude="3" metricUnit="m" metricValue="1.545" metricValueMax="1.65" metricValueMin="1.44" pageId="34" pageNumber="35" unit="m" value="1545.0" valueMax="1650.0" valueMin="1440.0">1440–1650 m</quantity>
, 
<date id="FF90E1F56264FFC12DC4B0CAFEA55F91" box="[188,322,689,715]" pageId="34" pageNumber="35" value="1983-06-16">16.VI.1983</date>
&quot;; &quot;Nepal, Kathmandu v., M. Brancucci&quot;. 1 ex (NHMB), &quot;Nepal Kathmandu V., Godavari 
<quantity id="4CD66AD06264FFC12847B0CAFA705F96" box="[1343,1431,689,716]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="m" value="1500.0">1500 m</quantity>
, M. Brancucci&quot;; &quot;21.-27.VI, 1989&quot;. 1 ex (NHMB), &quot;Nepal Kathmandu V. Burhanikanth&quot;; &quot; 
<date id="FF90E1F56264FFC129E1B0AEFAEA5FB5" box="[1177,1293,725,751]" pageId="34" pageNumber="35" value="1989-05-19">19.V.1989</date>
<quantity id="4CD66AD06264FFC1286EB0AEFA885FB5" box="[1302,1391,725,752]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="34" pageNumber="35" unit="m" value="1450.0">1450 m</quantity>
, M. Brancucci&quot;. 1 ex (NHMB), &quot;Nepal Kathmandu V. Godavari 
<quantity id="4CD66AD06264FFC12E36B082FC4C5E49" box="[846,939,761,788]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="m" value="1500.0">1500 m</quantity>
&quot;; &quot; 
<date id="FF90E1F56264FFC12EB3B082FB825E49" box="[971,1125,761,787]" pageId="34" pageNumber="35" value="1989-05-21" valueMax="1989-05-27" valueMin="1989-05-21">21-27.V.1989</date>
<quantity id="4CD66AD06264FFC12917B082FB2E5E49" box="[1135,1225,761,788]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="34" pageNumber="35" unit="m" value="1450.0">1450 m</quantity>
, M. Brancucci&quot;. 
<specimenCount id="9D280CBC6264FFC128F6B082FF2A5E6D" pageId="34" pageNumber="35" type="male">1 male</specimenCount>
(NHMB), &quot;Godavari 
<quantity id="4CD66AD06264FFC12CBAB166FDB85E6D" box="[450,607,797,824]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="34" pageNumber="35" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500–1700 m</quantity>
, 21.5&quot;, &quot;Nepal, 1977 Wittmer, Brancucci&quot;. 
<specimenCount id="9D280CBC6264FFC12939B166FB6C5E6D" box="[1089,1163,797,823]" pageId="34" pageNumber="35" type="male">1 male</specimenCount>
(NHMB), &quot;Balaju 
<date id="FF90E1F56264FFC1281AB166FF285E01" pageId="34" pageNumber="35" value="1350-05-23">
23.V. 
<quantity id="4CD66AD06264FFC12DEFB13BFF285E01" box="[151,207,832,859]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="34" pageNumber="35" unit="m" value="1350.0">1350</quantity>
</date>
m&quot;, &quot;Nepal, 1977, Wittmer, Brancucci&quot; &lt;
<figureCitation id="1315DBB06264FFC12FD3B13AFCF15E01" box="[683,790,833,859]" captionStart="FIGURE 17" captionStartId="31.[151,250,1926,1950]" captionTargetBox="[154,1424,569,1887]" captionTargetId="figure-130@31.[151,1436,561,1903]" captionTargetPageId="31" captionText="FIGURE 17. Habitus of Parena (Crossoglossa) laesipennis species group. A Lectotype of Parena laesipennis (Bates). B P. laesipennis (Bates), male from Taiwan. C P. laesipennis (Bates), female from Yunnan. D P. laesipennis (Bates), male from Nepal. E Holotype of Parena levata Andrewes. F P. levata Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958971" httpUri="https://zenodo.org/record/7958971/files/figure.png" pageId="34" pageNumber="35">Figs 17D</figureCitation>
, 
<figureCitation id="1315DBB06264FFC12E5CB13AFCB35E01" box="[804,852,833,859]" captionStart="FIGURE 19" captionStartId="32.[151,250,1946,1970]" captionTargetBox="[161,1425,1199,1917]" captionTargetId="figure-151@32.[151,1436,1186,1923]" captionTargetPageId="32" captionText="FIGURE 19. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates) from different localities, left-lateral (l) and dorsal (d) views: A Laos. B Taiwan. C Nepal. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958975" httpUri="https://zenodo.org/record/7958975/files/figure.png" pageId="34" pageNumber="35">19C</figureCitation>
&gt;. 1 ex (NHMB), &quot;Nepal Bagm ati, Sindhupalchok, Parahang-Dapkakharka&quot;, &quot; 
<quantity id="4CD66AD06264FFC12CB8B11EFDFC5E25" box="[448,539,869,895]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="34" pageNumber="35" unit="m" value="2100.0">2100 m</quantity>
, 11.VI.99, M. Brancucci&quot;. 1 ex (NHMB), &quot;Nepal: 
<collectingRegion id="49EA09D76264FFC12920B11FFB6A5E24" box="[1112,1165,868,894]" country="Nepal" name="Kosi" pageId="34" pageNumber="35">Kosi</collectingRegion>
- #13 Depitar 27°27’N/ 87°17E to Barabishe 27°26N/87°18’E, 1250– 
<quantity id="4CD66AD06264FFC12FE1B1F3FD045EF9" box="[665,739,904,931]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="34" pageNumber="35" unit="m" value="560.0">560 m</quantity>
, 12.vi.01&quot;. 1 ex (NHMB), &quot;Nepal: 
<collectingRegion id="49EA09D76264FFC12908B1F3FB425EF8" box="[1136,1189,904,930]" country="Nepal" name="Kosi" pageId="34" pageNumber="35">Kosi</collectingRegion>
- #10 Mure 27°30’N/ 87°16E 
<quantity id="4CD66AD06264FFC12D8BB1D7FE775E9D" box="[243,400,940,967]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="34" pageNumber="35" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">2000–2100 m</quantity>
, 6–8/11–12.vi.01&quot;. 1 ex (CSF), &quot;NEPAL Annapurna Himal, Lumle, 17. – 
<date id="FF90E1F56264FFC129BCB1D7FAA35E9C" box="[1220,1348,940,966]" pageId="34" pageNumber="35" value="1999-06-22">22.06.1999</date>
, A. Kudrna JR. Lgt.&quot;. 
<emphasis id="B95A1B276264FFC12C3DB1ABFE555EB0" bold="true" box="[325,434,976,1002]" pageId="34" pageNumber="35">
<collectingCountry id="F33987A56264FFC12C3DB1ABFE555EB0" box="[325,434,976,1002]" name="Malaysia" pageId="34" pageNumber="35">Malaysia</collectingCountry>
</emphasis>
: 
<specimenCount id="9D280CBC6264FFC12CB9B1ABFDEA5EB0" box="[449,525,976,1002]" pageId="34" pageNumber="35" type="male">1 male</specimenCount>
(NHMB), &quot;MALAYSIA, 
<collectingRegion id="49EA09D76264FFC12E4EB1ABFC4F5EB0" box="[822,936,976,1002]" country="Malaysia" name="Pahang" pageId="34" pageNumber="35">PAHANG</collectingRegion>
, 2003 Cameron Highlands; TANAH RATA, 
<quantity id="4CD66AD06264FFC12DEFB18FFED35955" box="[151,308,1012,1039]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="34" pageNumber="35" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500–1700 m</quantity>
, P. Pacholatko leg. 24.-31.i.&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6264FFC12DBFB663FC6A590C" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8B91C7356264FFC12DBFB663FC6A590C" blockId="34.[151,1437,151,2007]" pageId="34" pageNumber="35">
<emphasis id="B95A1B276264FFC12DBFB663FE815968" bold="true" box="[199,358,1048,1074]" pageId="34" pageNumber="35">Comparisons</emphasis>
. 
<taxonomicName id="4C2EBCB66264FFC12C0CB662FDEA5968" authorityName="Bates" authorityYear="1873" box="[372,525,1048,1074]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="laesipennis">
<emphasis id="B95A1B276264FFC12C0CB662FDEA5968" box="[372,525,1048,1074]" italics="true" pageId="34" pageNumber="35">P. laesipennis</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C2EBCB66264FFC12FA3B662FC815968" box="[731,870,1048,1074]" class="Insecta" family="Arctiidae" genus="Paratype" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="obenbergi">
<emphasis id="B95A1B276264FFC12FA3B662FC815968" box="[731,870,1048,1074]" italics="true" pageId="34" pageNumber="35">P. obenbergi</emphasis>
</taxonomicName>
, and also might be confused with a sympatric species, 
<taxonomicName id="4C2EBCB66264FFC12DA9B646FE83590C" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[209,356,1085,1110]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cavipennis">
<emphasis id="B95A1B276264FFC12DA9B646FE83590C" box="[209,356,1085,1110]" italics="true" pageId="34" pageNumber="35">P. cavipennis</emphasis>
</taxonomicName>
. See &quot;Comparisons&quot; under the latter two species.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6264FFC12DBFB624FDD55B16" pageId="34" pageNumber="35" type="description">
<paragraph id="8B91C7356264FFC12DBFB624FDD55B16" blockId="34.[151,1437,151,2007]" pageId="34" pageNumber="35">
<emphasis id="B95A1B276264FFC12DBFB624FEB65923" bold="true" box="[199,337,1119,1145]" pageId="34" pageNumber="35">Description</emphasis>
. Body length 
<quantity id="4CD66AD06264FFC12C96B61BFD615920" box="[494,646,1120,1147]" metricMagnitude="-2" metricUnit="m" metricValue="1.105" metricValueMax="1.26" metricValueMin="0.95" pageId="34" pageNumber="35" unit="mm" value="11.05" valueMax="12.6" valueMin="9.5">9.5–12.6 mm</quantity>
. Dorsum yellowish brown to reddish brown; basal three antennomeres, and basal half of antennomere 4 brown, remainder of antennae black; venter brown. Pronotum strongly transverse, PW/PL = 1.50–1.65, usually much wider than head, PW/HW = 1.05–1.14; widest at anterior third, lateral explanations very wide. Elytra without microsculpture, or with very weak isodiametric microsculpture near apices; striae not incised, replaced by rows of fine punctures; intervals completely flat, very finely and sparsely punctate; disc well depressed near middle of intervals 3 to 6, depressions subtriangular; apical truncation distinct, straight or slightly curved, outer apical angles weakly angulate, forming very blunt obtuse angles; sutural angles sharply pointed, forming short denticles. Median lobe of aedeagus gradually narrowed to apex, ventral margin straight or gently curved; apical lamella more or less bent toward dorsum, length 1.50–1.65 folds as its basal width, apex rounded. Endophallus very densely scaled on basal sheath, base of apical sheath discoid in dorsal view; squamate sac equally divided, situated on basal fifth of median lobe, right to squamate sheath, distal sac slightly closer to base than proximal sac (
<figureCitation id="1315DBB06264FFC12C16B790FE305B5F" box="[366,471,1515,1541]" captionStart="FIGURE 18" captionStartId="32.[151,250,1101,1125]" captionTargetBox="[164,1427,365,1072]" captionTargetId="figure-113@32.[151,1436,352,1078]" captionTargetPageId="32" captionText="FIGURE 18. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Yunnan. B Yunnan. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958973" httpUri="https://zenodo.org/record/7958973/files/figure.png" pageId="34" pageNumber="35">Fig. 18A</figureCitation>
). Gonocoxite II of ovipositor dichotomous, inner branch with two long ensiform setae apically, outer branch slightly longer than the inner one, apex with two or three ensiform setae, much shorter than those on the inner branch (
<figureCitation id="1315DBB06264FFC12CBAB449FDC25B16" box="[450,549,1586,1612]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="34" pageNumber="35">Fig. 11Z</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6264FFC12DBFB42DFF175A8D" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8B91C7356264FFC12DBFB42DFAFC5BCE" blockId="34.[151,1437,151,2007]" pageId="34" pageNumber="35">
<emphasis id="B95A1B276264FFC12DBFB42DFEBE5B2A" bold="true" box="[199,345,1622,1648]" pageId="34" pageNumber="35">Distribution</emphasis>
(
<figureCitation id="1315DBB06264FFC12C10B42DFE555B2A" box="[360,434,1622,1648]" captionStart="MAP 3" captionStartId="30.[152,211,2011,2035]" captionTargetBox="[170,1417,1025,1987]" captionTargetId="figure-424@30.[170,1417,1025,1987]" captionText="MAP 3. Distributions for species in the Parena (Crossoglossa) laesipennis group." figureDoi="http://doi.org/10.5281/zenodo.7963958" httpUri="https://zenodo.org/record/7963958/files/figure.png" pageId="34" pageNumber="35">Map 3</figureCitation>
, red). 
<collectingCountry id="F33987A56264FFC12C81B42DFDDB5B2A" box="[505,572,1622,1648]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
(
<collectingRegion id="49EA09D76264FFC12F32B42DFD555B2A" box="[586,690,1622,1648]" country="China" name="Zhejiang" pageId="34" pageNumber="35">Zhejiang</collectingRegion>
, 
<collectingRegion id="49EA09D76264FFC12FC5B42CFCED5B2B" box="[701,778,1623,1649]" country="China" name="Hunan" pageId="34" pageNumber="35">Hunan</collectingRegion>
, 
<collectingRegion id="49EA09D76264FFC12E6DB42DFC875B2A" box="[789,864,1622,1648]" country="China" name="Fujian" pageId="34" pageNumber="35">Fujian</collectingRegion>
, 
<collectingRegion id="49EA09D76264FFC12E12B42DFC175B2A" box="[874,1008,1622,1648]" country="China" name="Guangdong" pageId="34" pageNumber="35">Guangdong</collectingRegion>
, 
<collectingRegion id="49EA09D76264FFC12E82B42DFBA95B2A" box="[1018,1102,1622,1648]" country="China" name="Hainan" pageId="34" pageNumber="35">Hainan</collectingRegion>
, 
<collectingRegion id="49EA09D76264FFC12920B42DFB305B2A" box="[1112,1239,1622,1648]" country="China" name="Chongqing" pageId="34" pageNumber="35">Chongqing</collectingRegion>
, 
<collectingRegion id="49EA09D76264FFC12999B42CFADD5B2B" box="[1249,1338,1623,1649]" country="China" name="Yunnan" pageId="34" pageNumber="35">Yunnan</collectingRegion>
, 
<collectingRegion id="49EA09D76264FFC1283CB42DFA7F5B2A" box="[1348,1432,1622,1648]" country="China" name="Xizang" pageId="34" pageNumber="35">Xizang</collectingRegion>
, 
<collectingRegion id="49EA09D76264FFC12DEFB401FF115BCE" box="[151,246,1658,1684]" country="China" name="Taiwan)" pageId="34" pageNumber="35">Taiwan)</collectingRegion>
, 
<collectingCountry id="F33987A56264FFC12C79B400FEA35BCF" box="[257,324,1659,1685]" name="Japan" pageId="34" pageNumber="35">Japan</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC12C36B401FE395BCF" box="[334,478,1658,1685]" name="South Korea" pageId="34" pageNumber="35">South Korea</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC12C90B401FDAD5BCE" box="[488,586,1658,1684]" name="Vietnam" pageId="34" pageNumber="35">Vietnam</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC12F2DB400FD695BCF" box="[597,654,1659,1685]" name="Laos" pageId="34" pageNumber="35">Laos</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC12FE1B401FD185BCE" box="[665,767,1658,1684]" name="Thailand" pageId="34" pageNumber="35">Thailand</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC12E73B400FC9D5BCF" box="[779,890,1659,1685]" name="Myanmar" pageId="34" pageNumber="35">Myanmar</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC12EFCB401FC0A5BCE" box="[900,1005,1658,1684]" name="Malaysia" pageId="34" pageNumber="35">Malaysia</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC12E81B401FBD45BCE" box="[1017,1075,1658,1684]" name="India" pageId="34" pageNumber="35">India</collectingCountry>
(Darjeeling), 
<collectingCountry id="F33987A56264FFC129A8B401FAF05BCE" box="[1232,1303,1658,1684]" name="Nepal" pageId="34" pageNumber="35">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="8B91C7356264FFC12DBFB4E5FF175A8D" blockId="34.[151,1437,151,2007]" pageId="34" pageNumber="35">
<emphasis id="B95A1B276264FFC12DBFB4E5FE065BE2" bold="true" box="[199,481,1694,1720]" pageId="34" pageNumber="35">Geographical variation</emphasis>
. The southwestern specimens (from southwest 
<collectingCountry id="F33987A56264FFC12975B4E5FBB45BE2" box="[1037,1107,1694,1720]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC1291AB4E4FB7C5BE3" box="[1122,1179,1695,1721]" name="Laos" pageId="34" pageNumber="35">Laos</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC129D2B4E5FAF75BE2" box="[1194,1296,1694,1720]" name="Thailand" pageId="34" pageNumber="35">Thailand</collectingCountry>
, 
<collectingCountry id="F33987A56264FFC12867B4E5FA825BE2" box="[1311,1381,1694,1720]" name="Nepal" pageId="34" pageNumber="35">Nepal</collectingCountry>
, and 
<collectingCountry id="F33987A56264FFC12DEFB4B9FF345B86" box="[151,211,1730,1756]" name="India" pageId="34" pageNumber="35">India</collectingCountry>
, 
<figureCitation id="1315DBB06264FFC12D98B4B9FE8C5B86" box="[224,363,1730,1756]" captionStart="FIGURE 17" captionStartId="31.[151,250,1926,1950]" captionTargetBox="[154,1424,569,1887]" captionTargetId="figure-130@31.[151,1436,561,1903]" captionTargetPageId="31" captionText="FIGURE 17. Habitus of Parena (Crossoglossa) laesipennis species group. A Lectotype of Parena laesipennis (Bates). B P. laesipennis (Bates), male from Taiwan. C P. laesipennis (Bates), female from Yunnan. D P. laesipennis (Bates), male from Nepal. E Holotype of Parena levata Andrewes. F P. levata Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958971" httpUri="https://zenodo.org/record/7958971/files/figure.png" pageId="34" pageNumber="35">Figs 17C–D</figureCitation>
, 
<figureCitation id="1315DBB06264FFC12C00B4B9FE0F5B86" box="[376,488,1730,1756]" captionStart="FIGURE 18" captionStartId="32.[151,250,1101,1125]" captionTargetBox="[164,1427,365,1072]" captionTargetId="figure-113@32.[151,1436,352,1078]" captionTargetPageId="32" captionText="FIGURE 18. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates), right-lateral (r), ventral (v), leftlateral (l), and dorsal (d) views. A Yunnan. B Yunnan. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958973" httpUri="https://zenodo.org/record/7958973/files/figure.png" pageId="34" pageNumber="35">18A, 18B</figureCitation>
, 
<figureCitation id="1315DBB06264FFC12C8DB4B9FD845B86" box="[501,611,1730,1756]" captionStart="FIGURE 19" captionStartId="32.[151,250,1946,1970]" captionTargetBox="[161,1425,1199,1917]" captionTargetId="figure-151@32.[151,1436,1186,1923]" captionTargetPageId="32" captionText="FIGURE 19. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates) from different localities, left-lateral (l) and dorsal (d) views: A Laos. B Taiwan. C Nepal. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958975" httpUri="https://zenodo.org/record/7958975/files/figure.png" pageId="34" pageNumber="35">19A, 19C</figureCitation>
) of 
<taxonomicName id="4C2EBCB66264FFC12FEBB4B8FCCA5B86" authorityName="Bates" authorityYear="1873" box="[659,813,1730,1756]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="laesipennis">
<emphasis id="B95A1B276264FFC12FEBB4B8FCCA5B86" box="[659,813,1730,1756]" italics="true" pageId="34" pageNumber="35">P. laesipennis</emphasis>
</taxonomicName>
are slightly different from the typical specimens (from 
<collectingCountry id="F33987A56264FFC12DEFB49DFF305A5A" box="[151,215,1766,1792]" name="Japan" pageId="34" pageNumber="35">Japan</collectingCountry>
and 
<collectingCountry id="F33987A56264FFC12C6BB49DFE8F5A5A" box="[275,360,1766,1792]" name="Taiwan" pageId="34" pageNumber="35">Taiwan</collectingCountry>
, 
<figureCitation id="1315DBB06264FFC12C0FB49DFDE35A5A" box="[375,516,1766,1792]" captionStart="FIGURE 17" captionStartId="31.[151,250,1926,1950]" captionTargetBox="[154,1424,569,1887]" captionTargetId="figure-130@31.[151,1436,561,1903]" captionTargetPageId="31" captionText="FIGURE 17. Habitus of Parena (Crossoglossa) laesipennis species group. A Lectotype of Parena laesipennis (Bates). B P. laesipennis (Bates), male from Taiwan. C P. laesipennis (Bates), female from Yunnan. D P. laesipennis (Bates), male from Nepal. E Holotype of Parena levata Andrewes. F P. levata Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958971" httpUri="https://zenodo.org/record/7958971/files/figure.png" pageId="34" pageNumber="35">Figs 17A–B</figureCitation>
, 
<figureCitation id="1315DBB06264FFC12F6BB49DFDA25A5A" box="[531,581,1766,1792]" captionStart="FIGURE 19" captionStartId="32.[151,250,1946,1970]" captionTargetBox="[161,1425,1199,1917]" captionTargetId="figure-151@32.[151,1436,1186,1923]" captionTargetPageId="32" captionText="FIGURE 19. Median lobe of male genitalia for Parena (Crossoglossa) laesipennis (Bates) from different localities, left-lateral (l) and dorsal (d) views: A Laos. B Taiwan. C Nepal. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7958975" httpUri="https://zenodo.org/record/7958975/files/figure.png" pageId="34" pageNumber="35">19B</figureCitation>
). They are usually slightly darker and smaller sized (9.5–11.0 mm versus 11.0– 
<quantity id="4CD66AD06264FFC12DAEB571FED95A7E" box="[214,318,1802,1828]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="34" pageNumber="35" unit="mm" value="12.6">12.6 mm</quantity>
), and with elytral microsculpture less obvious. The male genitalia also differ slightly as well: in the southwestern specimens, the apical lamella is generally slightly more bent to the dorsum, and the ventral margin is slightly more curved in lateral view. However, the above differences are subtle, not always stable, and intermediate status is present in some individuals from Eastern 
<collectingCountry id="F33987A56264FFC12FB2B50EFCF65AD5" box="[714,785,1909,1935]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
. Therefore, we believe that the above characters gradually vary by distance from northeast to southwest, and that all the above populations should be classified as the same species.
</paragraph>
</subSubSection>
</treatment>
</document>