treatments-xml/data/03/87/76/03877623623DFF9E2DEFB695FCFB5F3B.xml

<document id="FB608F5A2DE18E8CBB9A6C65F58858C5" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="03877623623DFF9E2DEFB695FCFB5F3B" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Parena) fulva Shi &amp; Liang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="126" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="124" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="008282C22DD945E1EE546C1AE555294B">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="A010B1AAC4C26989AC6EA5CC735CD0E9">Shi, Hongliang</mods:namePart>
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<mods:namePart id="FBC1C6617106EC381881B952E96D1FC2">Liang, Hongbin</mods:namePart>
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<subSubSection id="C33494BE623DFF982DEFB695FDA35852" box="[151,580,1261,1288]" pageId="123" pageNumber="124" type="nomenclature">
<paragraph id="8B91C735623DFF982DEFB695FDA35852" blockId="123.[151,1113,1261,1325]" box="[151,580,1261,1288]" pageId="123" pageNumber="124">
<emphasis id="B95A1B27623DFF982DEFB695FDA35852" bold="true" box="[151,580,1261,1288]" pageId="123" pageNumber="124">
[39] 
<taxonomicName id="4C2EBCB6623DFF982DB7B695FE075852" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[207,480,1261,1288]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="fulva" status="sp. nov." subGenus="Parena">
<emphasis id="B95A1B27623DFF982DB7B695FEC05852" bold="true" box="[207,295,1262,1288]" italics="true" pageId="123" pageNumber="124">Parena</emphasis>
(
<emphasis id="B95A1B27623DFF982C40B695FE775852" bold="true" box="[312,400,1262,1288]" italics="true" pageId="123" pageNumber="124">Parena</emphasis>
) 
<emphasis id="B95A1B27623DFF982CDAB696FE075852" bold="true" box="[418,480,1261,1288]" italics="true" pageId="123" pageNumber="124">fulva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269A65C623DFF982C9FB695FDA35852" box="[487,580,1262,1288]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE623DFF982DEFB769FBBE5876" box="[151,1113,1298,1325]" pageId="123" pageNumber="124" type="description">
<paragraph id="8B91C735623DFF982DEFB769FBBE5876" blockId="123.[151,1113,1261,1325]" box="[151,1113,1298,1325]" pageId="123" pageNumber="124">
Habitus: 
<figureCitation id="1315DBB0623DFF982D85B769FE425876" box="[253,421,1298,1324]" captionStart="FIGURE 73" captionStartId="121.[151,250,1902,1926]" captionTargetBox="[167,1409,298,1862]" captionTargetId="figure-87@121.[151,1436,294,1878]" captionTargetPageId="121" captionText="FIGURE 73. Habitus of Parena (Parena) stigmatica species group. A Lectotype of Parena stigmatica (Fairmaire). B Holotype of Parena dorae Basilewsky. C Holotype of Parena fulva sp. n. D Paratype of Parena fulva sp. n., female from Zimbabwe. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959090" httpUri="https://zenodo.org/record/7959090/files/figure.png" pageId="123" pageNumber="124">Figs 73C, 73D</figureCitation>
. Male genitalia: 
<figureCitation id="1315DBB0623DFF982F27B769FD565877" box="[607,689,1298,1325]" captionStart="FIGURE 75" captionStartId="124.[151,250,777,801]" captionTargetBox="[164,1425,172,747]" captionTargetId="figure-28@124.[151,1436,166,753]" captionTargetPageId="124" captionText="FIGURE 75. Median lobe of male genitalia for Parena (Parena) fulva sp. n., right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Holotype. B Paratype from Zimbabwe. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959094" httpUri="https://zenodo.org/record/7959094/files/figure.png" pageId="123" pageNumber="124">Fig. 75</figureCitation>
. Gonocoxites of ovipositor: 
<figureCitation id="1315DBB0623DFF982E88B769FBB15876" box="[1008,1110,1298,1324]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="123" pageNumber="124">Fig. 11U</figureCitation>
.
</paragraph>
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<paragraph id="8B91C735623DFF982DEFB721FCC0582E" blockId="123.[151,1437,1369,2008]" box="[151,807,1369,1397]" pageId="123" pageNumber="124">
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<typeStatus id="54957997623DFF982DEFB721FF35582E" box="[151,210,1370,1396]" pageId="123" pageNumber="124">Type</typeStatus>
locality
</emphasis>
. 
<collectingCountry id="F33987A5623DFF982C46B721FE41582E" box="[318,422,1370,1396]" name="Tanzania" pageId="123" pageNumber="124">Tanzania</collectingCountry>
, 
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, 
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, 
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, 
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<elevation id="00032006623DFF982FB4B721FCC4582E" box="[716,803,1370,1396]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="123" pageNumber="124" unit="m" value="1120.0">1120 m</elevation>
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.
</paragraph>
<paragraph id="8B91C735623DFF982DBFB705FE335A13" blockId="123.[151,1437,1369,2008]" pageId="123" pageNumber="124">
<emphasis id="B95A1B27623DFF982DBFB705FE8858CD" bold="true" box="[199,367,1405,1432]" pageId="123" pageNumber="124">Type material</emphasis>
. 
<materialsCitation id="3B46CD68623DFF982C05B705FF2D5B5F" ID-GBIF-Occurrence="4115871316" collectorName="Coll. Ch.ALLUAUD" determinerName="Alluaud" location="Dodoma" pageId="123" pageNumber="124" specimenCount="2" stateProvince="Dodoma" typeStatus="holotype">
<emphasis id="B95A1B27623DFF982C05B705FE0D58C2" bold="true" box="[381,490,1406,1432]" pageId="123" pageNumber="124">
<typeStatus id="54957997623DFF982C05B705FE0D58C2" box="[381,490,1406,1432]" pageId="123" pageNumber="124" type="holotype">Holotype</typeStatus>
</emphasis>
(MNHN, 
<figureCitation id="1315DBB0623DFF982F27B705FD2158C2" box="[607,710,1406,1432]" captionStart="FIGURE 73" captionStartId="121.[151,250,1902,1926]" captionTargetBox="[167,1409,298,1862]" captionTargetId="figure-87@121.[151,1436,294,1878]" captionTargetPageId="121" captionText="FIGURE 73. Habitus of Parena (Parena) stigmatica species group. A Lectotype of Parena stigmatica (Fairmaire). B Holotype of Parena dorae Basilewsky. C Holotype of Parena fulva sp. n. D Paratype of Parena fulva sp. n., female from Zimbabwe. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959090" httpUri="https://zenodo.org/record/7959090/files/figure.png" pageId="123" pageNumber="124">Fig. 73C</figureCitation>
): male, body length= 
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, board mounted, &quot; 
<emphasis id="B95A1B27623DFF98299EB705FA7A58C2" box="[1254,1437,1406,1432]" italics="true" pageId="123" pageNumber="124">
<collectingRegion id="49EA09D7623DFF98299EB705FAAE58C2" box="[1254,1353,1406,1432]" country="Tanzania" name="Dodoma" pageId="123" pageNumber="124">Dodoma</collectingRegion>
, 
<collectorName id="26DBA2E3623DFF98282DB704FA7A58C2" box="[1365,1437,1406,1432]" pageId="123" pageNumber="124">E. Afr.</collectorName>
</emphasis>
/ 
<emphasis id="B95A1B27623DFF982DDEB7D8FE7458E6" box="[166,403,1442,1468]" italics="true" pageId="123" pageNumber="124">
<collectorName id="26DBA2E3623DFF982DDEB7D8FEA858E6" box="[166,335,1442,1468]" pageId="123" pageNumber="124">H.L. Andrewes</collectorName>
, 1918
</emphasis>
&quot;, &quot; 
<taxonomicName id="4C2EBCB6623DFF982CC0B7D9FDA958E6" authority="Chaudoir, 1873" box="[440,590,1442,1468]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27623DFF982CC0B7D9FDA958E6" box="[440,590,1442,1468]" italics="true" pageId="123" pageNumber="124">Crossoglossa</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C2EBCB6623DFF982F1DB7D9FD3C58E6" baseAuthorityName="Chaudoir" baseAuthorityYear="1878" box="[613,731,1442,1468]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B95A1B27623DFF982F1DB7D9FD3C58E6" box="[613,731,1442,1468]" italics="true" pageId="123" pageNumber="124">ferruginea</emphasis>
</taxonomicName>
/ 
<emphasis id="B95A1B27623DFF982F8AB7D9FCDA58E6" box="[754,829,1442,1468]" italics="true" pageId="123" pageNumber="124">Chaud</emphasis>
. / 
<determinerName id="7E4D2BF1623DFF982E21B7D9FC5158E6" box="[857,950,1442,1468]" pageId="123" pageNumber="124">Alluaud</determinerName>
, det. 19 
<emphasis id="B95A1B27623DFF982976B7D9FBCD58E6" box="[1038,1066,1442,1468]" italics="true" pageId="123" pageNumber="124">30</emphasis>
&quot;, &quot;MUSEUM PARIS / 
<collectorName id="26DBA2E3623DFF98284EB7D9FA7A58E6" box="[1334,1437,1442,1468]" pageId="123" pageNumber="124">Coll. Ch.</collectorName>
ALLUAUD&quot; [blue label], &quot;HOLOTYPE / 
<taxonomicName id="4C2EBCB6623DFF982F03B7BDFCC858BA" box="[635,815,1478,1504]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="subGenus" subGenus="Parena">Parena (Parena)</taxonomicName>
/ fulva 
<taxonomicNameLabel id="A269A65C623DFF982EF1B7BDFC0658BA" box="[905,993,1478,1504]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
/ des. Shi H.L. 2022&quot; [red label] &lt;
<figureCitation id="1315DBB0623DFF982808B7BDFF2D5B5F" captionStart="FIGURE 75" captionStartId="124.[151,250,777,801]" captionTargetBox="[164,1425,172,747]" captionTargetId="figure-28@124.[151,1436,166,753]" captionTargetPageId="124" captionText="FIGURE 75. Median lobe of male genitalia for Parena (Parena) fulva sp. n., right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Holotype. B Paratype from Zimbabwe. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959094" httpUri="https://zenodo.org/record/7959094/files/figure.png" pageId="123" pageNumber="124">Fig. 75A</figureCitation>
</materialsCitation>
&gt;. 
<materialsCitation id="3B46CD68623DFF982D9DB791FCD65B59" ID-GBIF-Occurrence="4115871407" box="[229,817,1513,1540]" country="Tanzania" location="Tanzania" pageId="123" pageNumber="124" specimenCount="7" specimenCount-female="3" specimenCount-male="4" typeStatus="paratype">
<emphasis id="B95A1B27623DFF982D9DB791FEBA5B5E" bold="true" box="[229,349,1514,1540]" pageId="123" pageNumber="124">
<typeStatus id="54957997623DFF982D9DB791FEBA5B5E" box="[229,349,1514,1540]" pageId="123" pageNumber="124" type="paratype">Paratypes</typeStatus>
</emphasis>
(a total of 
<specimenCount id="9D280CBC623DFF982CAFB791FDD75B5E" box="[471,560,1514,1540]" pageId="123" pageNumber="124" type="male">4 males</specimenCount>
, 
<specimenCount id="9D280CBC623DFF982F43B791FD4A5B5E" box="[571,685,1514,1540]" pageId="123" pageNumber="124" type="female">3 females</specimenCount>
): 
<collectingCountry id="F33987A5623DFF982FC7B792FCD65B59" box="[703,817,1513,1539]" name="Tanzania" pageId="123" pageNumber="124">
<emphasis id="B95A1B27623DFF982FC7B792FCD65B59" bold="true" box="[703,817,1513,1539]" pageId="123" pageNumber="124">Tanzania</emphasis>
</collectingCountry>
</materialsCitation>
<emphasis id="B95A1B27623DFF982E49B792FCD15B59" bold="true" box="[817,822,1513,1539]" pageId="123" pageNumber="124">:</emphasis>
<specimenCount id="9D280CBC623DFF982E44B791FC7A5B5E" box="[828,925,1514,1540]" pageId="123" pageNumber="124" type="female">1 female</specimenCount>
(MNHN), 
<materialsCitation id="3B46CD68623DFF982962B791FC9C5B72" ID-GBIF-Occurrence="4115871471" collectorName="H. L. Andrewes &amp; Coll. Ch." country="Tanzania" location="Afr." pageId="123" pageNumber="124" specimenCount="1" stateProvince="Dodoma" typeStatus="paratype">
&quot; 
<emphasis id="B95A1B27623DFF98295EB791FB3C5B5E" box="[1062,1243,1514,1540]" italics="true" pageId="123" pageNumber="124">
<collectingRegion id="49EA09D7623DFF98295EB791FB6E5B5E" box="[1062,1161,1514,1540]" country="Tanzania" name="Dodoma" pageId="123" pageNumber="124">Dodoma</collectingRegion>
, E. 
<location id="8EF191EE623DFF9829CBB791FB3C5B5E" LSID="urn:lsid:plazi:treatment:03877623623DFF9E2DEFB695FCFB5F3B:8EF191EE623DFF9829CBB791FB3C5B5E" box="[1203,1243,1514,1540]" country="Tanzania" name="Afr." pageId="123" pageNumber="124" stateProvince="Dodoma">Afr.</location>
</emphasis>
/ 
<emphasis id="B95A1B27623DFF982988B790FF285B72" italics="true" pageId="123" pageNumber="124">
<collectorName id="26DBA2E3623DFF982988B790FA7F5B5E" box="[1264,1432,1514,1540]" pageId="123" pageNumber="124">H.L. Andrewes</collectorName>
, 1918
</emphasis>
&quot;; &quot;MUSEUM PARIS / 
<collectorName id="26DBA2E3623DFF982C9DB475FDA85B72" box="[485,591,1550,1576]" pageId="123" pageNumber="124">Coll. Ch.</collectorName>
ALLUAUD&quot; [blue label]
</materialsCitation>
. 
<specimenCount id="9D280CBC623DFF982EF0B475FC315B72" box="[904,982,1550,1576]" pageId="123" pageNumber="124" type="male">1 male</specimenCount>
(NHML), 
<materialsCitation id="3B46CD68623DFF98292EB475FA7E5B17" ID-GBIF-Occurrence="4115871565" collectorName="H. L. Andrewes" country="Tanzania" determinerName="Alluaud." location="Afr." pageId="123" pageNumber="124" specimenCount="1" stateProvince="Dodoma" typeStatus="paratype">
&quot; 
<emphasis id="B95A1B27623DFF98291AB475FAF95B72" box="[1122,1310,1550,1576]" italics="true" pageId="123" pageNumber="124">
<collectingRegion id="49EA09D7623DFF98291AB475FB225B72" box="[1122,1221,1550,1576]" country="Tanzania" name="Dodoma" pageId="123" pageNumber="124">Dodoma</collectingRegion>
, E. 
<location id="8EF191EE623DFF98298DB475FAF95B72" LSID="urn:lsid:plazi:treatment:03877623623DFF9E2DEFB695FCFB5F3B:8EF191EE623DFF98298DB475FAF95B72" box="[1269,1310,1550,1576]" country="Tanzania" name="Afr." pageId="123" pageNumber="124" stateProvince="Dodoma">Afr.</location>
</emphasis>
/ 
<emphasis id="B95A1B27623DFF982840B474FECB5B16" italics="true" pageId="123" pageNumber="124">
<collectorName id="26DBA2E3623DFF982840B474FF0F5B16" pageId="123" pageNumber="124">H.L. Andrewes</collectorName>
, 1918
</emphasis>
&quot;, &quot; 
<emphasis id="B95A1B27623DFF982C29B449FD0E5B16" box="[337,745,1586,1612]" italics="true" pageId="123" pageNumber="124">
<taxonomicName id="4C2EBCB6623DFF982C29B449FE005B16" authority="Chaudoir, 1873" box="[337,487,1586,1612]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">Crossoglossa</taxonomicName>
/ 
<taxonomicName id="4C2EBCB6623DFF982C84B449FD955B16" baseAuthorityName="Chaudoir" baseAuthorityYear="1878" box="[508,626,1586,1612]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="ferruginea">ferruginea</taxonomicName>
/ Chaud. /
</emphasis>
<determinerName id="7E4D2BF1623DFF982F88B449FCB55B16" box="[752,850,1586,1612]" pageId="123" pageNumber="124">Alluaud.</determinerName>
det. 1924&quot;, &quot;H.E. Andrewes Coll. / B.M. 1945-97.&quot;
</materialsCitation>
. 
<materialsCitation id="3B46CD68623DFF982DEFB42DFA725B35" ID-GBIF-Occurrence="4115871391" box="[151,1429,1621,1649]" collectorName="H. L. Andrewes &amp; H. E. Andrewes Coll." country="Zimbabwe" location="Afr." pageId="123" pageNumber="124" specimenCount="2" specimenCount-male="2" stateProvince="Dodoma" typeStatus="paratype">
<specimenCount id="9D280CBC623DFF982DEFB42DFF095B2A" box="[151,238,1622,1648]" pageId="123" pageNumber="124" type="male">2 males</specimenCount>
(NHML), &quot; 
<emphasis id="B95A1B27623DFF982C0DB42DFDCB5B2A" box="[373,556,1622,1648]" italics="true" pageId="123" pageNumber="124">
<collectingRegion id="49EA09D7623DFF982C0DB42DFE3F5B2A" box="[373,472,1622,1648]" country="Tanzania" name="Dodoma" pageId="123" pageNumber="124">Dodoma</collectingRegion>
, E. 
<location id="8EF191EE623DFF982F7BB42DFDCB5B2A" LSID="urn:lsid:plazi:treatment:03877623623DFF9E2DEFB695FCFB5F3B:8EF191EE623DFF982F7BB42DFDCB5B2A" box="[515,556,1622,1648]" country="Zimbabwe" name="Afr." pageId="123" pageNumber="124" stateProvince="Dodoma">Afr.</location>
</emphasis>
/ 
<emphasis id="B95A1B27623DFF982F3AB42CFCC85B2A" box="[578,815,1622,1648]" italics="true" pageId="123" pageNumber="124">
<collectorName id="26DBA2E3623DFF982F3AB42CFD0C5B2A" box="[578,747,1622,1648]" pageId="123" pageNumber="124">H.L. Andrewes</collectorName>
, 1918
</emphasis>
&quot;, &quot; 
<collectorName id="26DBA2E3623DFF982E2AB42DFBD85B2A" box="[850,1087,1622,1648]" pageId="123" pageNumber="124">H.E. Andrewes Coll.</collectorName>
/ B.M. 1945-97.&quot;. 
<emphasis id="B95A1B27623DFF98286DB42EFA725B35" bold="true" box="[1301,1429,1621,1647]" pageId="123" pageNumber="124">
<collectingCountry id="F33987A5623DFF98286DB42EFA725B35" box="[1301,1429,1621,1647]" name="Zimbabwe" pageId="123" pageNumber="124">Zimbabwe</collectingCountry>
</emphasis>
</materialsCitation>
: 
<specimenCount id="9D280CBC623DFF982DEFB401FF1B5BCE" box="[151,252,1658,1684]" pageId="123" pageNumber="124" type="female">1 female</specimenCount>
(CSF), 
<materialsCitation id="3B46CD68623DFF982C24B401FDF55BE2" ID-GBIF-Occurrence="4115871511" collectingDate="1998-12-08" collectorName="S. Becvar" country="Zimbabwe" determinerName="Det. Facchini" location="Bubi river" pageId="123" pageNumber="124" specimenCount="1" typeStatus="paratype">
&quot;ZIMBABWE-S / BUBI env. / 
<location id="8EF191EE623DFF982FB6B401FCA35BCE" LSID="urn:lsid:plazi:treatment:03877623623DFF9E2DEFB695FCFB5F3B:8EF191EE623DFF982FB6B401FCA35BCE" box="[718,836,1658,1684]" country="Zimbabwe" name="Bubi river" pageId="123" pageNumber="124">Bubi river</location>
/ lgt. 
<collectorName id="26DBA2E3623DFF982EE8B401FBE75BCE" box="[912,1024,1658,1684]" pageId="123" pageNumber="124">S. Bečvář</collectorName>
<date id="FF90E1F5623DFF982973B401FB665BCE" box="[1035,1153,1658,1684]" pageId="123" pageNumber="124" value="1998-12-08">
<collectingDate id="EFD4181D623DFF982973B401FB665BCE" box="[1035,1153,1658,1684]" pageId="123" pageNumber="124" value="1998-12-08">8.xii.1998</collectingDate>
</date>
&quot;; &quot; 
<taxonomicName id="4C2EBCB6623DFF9829DDB400FB115BCE" box="[1189,1270,1659,1684]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27623DFF9829DDB400FB115BCE" box="[1189,1270,1659,1684]" italics="true" pageId="123" pageNumber="124">Parena</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C2EBCB6623DFF98286CB401FA6D5BCE" baseAuthorityName="Chaudoir" baseAuthorityYear="1878" box="[1300,1418,1658,1684]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B95A1B27623DFF98286CB401FA6D5BCE" box="[1300,1418,1658,1684]" italics="true" pageId="123" pageNumber="124">ferruginea</emphasis>
</taxonomicName>
/ 
<emphasis id="B95A1B27623DFF982DEFB4E5FF3D5BE2" box="[151,218,1694,1720]" italics="true" pageId="123" pageNumber="124">CHD.</emphasis>
/ 
<determinerName id="7E4D2BF1623DFF982D88B4E5FE6B5BE2" box="[240,396,1694,1720]" pageId="123" pageNumber="124">Det. Facchini</determinerName>
&quot; &lt;
<figureCitation id="1315DBB0623DFF982CD3B4E5FDF55BE2" box="[427,530,1694,1720]" captionStart="FIGURE 73" captionStartId="121.[151,250,1902,1926]" captionTargetBox="[167,1409,298,1862]" captionTargetId="figure-87@121.[151,1436,294,1878]" captionTargetPageId="121" captionText="FIGURE 73. Habitus of Parena (Parena) stigmatica species group. A Lectotype of Parena stigmatica (Fairmaire). B Holotype of Parena dorae Basilewsky. C Holotype of Parena fulva sp. n. D Paratype of Parena fulva sp. n., female from Zimbabwe. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959090" httpUri="https://zenodo.org/record/7959090/files/figure.png" pageId="123" pageNumber="124">Fig. 73D</figureCitation>
</materialsCitation>
&gt;. 
<specimenCount id="9D280CBC623DFF982F56B4E5FD685BE2" box="[558,655,1694,1720]" pageId="123" pageNumber="124" type="female">1 female</specimenCount>
(NHML), 
<materialsCitation id="3B46CD68623DFF982E73B4E5FD515A5A" ID-GBIF-Occurrence="4115871355" collectingDate="1915-12" collectorName="R. Lowe Thompson" country="Zimbabwe" location="Victoria Falls" municipality="Victoria Falls." pageId="123" pageNumber="124" specimenCount="1" typeStatus="paratype">
&quot; 
<collectorName id="26DBA2E3623DFF982E6DB4E5FC465BE2" box="[789,929,1694,1720]" pageId="123" pageNumber="124">S. Rhodesia</collectorName>
: / 
<collectingMunicipality id="6BF55D4F623DFF982EC3B4E5FBBC5BE2" box="[955,1115,1694,1720]" pageId="123" pageNumber="124">
<location id="8EF191EE623DFF982EC3B4E5FBB05BE2" LSID="urn:lsid:plazi:treatment:03877623623DFF9E2DEFB695FCFB5F3B:8EF191EE623DFF982EC3B4E5FBB05BE2" box="[955,1111,1694,1720]" country="Zimbabwe" municipality="Victoria Falls." name="Victoria Falls" pageId="123" pageNumber="124">Victoria Falls</location>
.
</collectingMunicipality>
/ 
<date id="FF90E1F5623DFF982909B4E5FAC35BE3" box="[1137,1316,1694,1721]" pageId="123" pageNumber="124" value="1915-12">
<collectingDate id="EFD4181D623DFF982909B4E5FAC35BE3" box="[1137,1316,1694,1721]" pageId="123" pageNumber="124" value="1915-12">
<emphasis id="B95A1B27623DFF982909B4E5FB035BE2" box="[1137,1252,1694,1720]" italics="true" pageId="123" pageNumber="124">December</emphasis>
1915
</collectingDate>
</date>
/ 
<collectorName id="26DBA2E3623DFF982843B4E5FEF25B86" pageId="123" pageNumber="124">R. Lowe Thompson</collectorName>
&quot;, &quot;Pres. By / Imp. Bur. Ent. / Brit. Mus. / 1925-184.&quot;, &quot; 
<taxonomicName id="4C2EBCB6623DFF982EE5B4B9FBEF5B86" authorityName="Chaudoir" authorityYear="1873" box="[925,1032,1730,1756]" class="Insecta" family="Carabidae" genus="Metallica" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27623DFF982EE5B4B9FBEF5B86" box="[925,1032,1730,1756]" italics="true" pageId="123" pageNumber="124">Metallica</emphasis>
</taxonomicName>
/
<emphasis id="B95A1B27623DFF98295BB4B9FB745B86" box="[1059,1171,1730,1756]" italics="true" pageId="123" pageNumber="124">? sp. nov.</emphasis>
&quot;, &quot; 
<emphasis id="B95A1B27623DFF9829C2B4B8FF165A5A" italics="true" pageId="123" pageNumber="124">
<taxonomicName id="4C2EBCB6623DFF9829C2B4B8FAEC5B86" box="[1210,1291,1731,1756]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/ 
<taxonomicName id="4C2EBCB6623DFF98285EB4B9FF055A5A" authority="Chaud" authorityName="Chaud" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="ferruginea">ferruginea Chaud</taxonomicName>
/
</emphasis>
<collectorName id="26DBA2E3623DFF982D80B49DFE605A5A" box="[248,391,1766,1792]" pageId="123" pageNumber="124">E.B. Britton.</collectorName>
/ det. 
<emphasis id="B95A1B27623DFF982CB5B49DFE105A5A" box="[461,503,1766,1792]" italics="true" pageId="123" pageNumber="124">6.3.</emphasis>
193 
<emphasis id="B95A1B27623DFF982F59B49DFDC85A5A" box="[545,559,1766,1792]" italics="true" pageId="123" pageNumber="124">6</emphasis>
&quot; &lt;
<figureCitation id="1315DBB0623DFF982F29B49DFD515A5A" box="[593,694,1766,1792]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="123" pageNumber="124">Fig. 11U</figureCitation>
</materialsCitation>
&gt;. 
<specimenCount id="9D280CBC623DFF982FA9B49DFCFB5A5A" box="[721,796,1766,1792]" pageId="123" pageNumber="124" type="male">1 male</specimenCount>
(NHML), 
<materialsCitation id="3B46CD68623DFF982EEFB49DFE585A13" ID-GBIF-Occurrence="4115871480" country="Zimbabwe" county="Rhodesia" location="Empanduri" municipality="Salisbury" pageId="123" pageNumber="124" specimenCount="1" typeStatus="paratype">
&quot; 
<emphasis id="B95A1B27623DFF982EDAB49DFBED5A5A" box="[930,1034,1766,1792]" italics="true" pageId="123" pageNumber="124">
<collectingCounty id="62F0BFB9623DFF982EDAB49DFBED5A5A" box="[930,1034,1766,1792]" pageId="123" pageNumber="124">Rhodesia</collectingCounty>
</emphasis>
/ 
<emphasis id="B95A1B27623DFF982958B49DFB6D5A5A" box="[1056,1162,1766,1792]" italics="true" pageId="123" pageNumber="124">
<collectingMunicipality id="6BF55D4F623DFF982958B49DFB6D5A5A" box="[1056,1162,1766,1792]" pageId="123" pageNumber="124">Salisbury</collectingMunicipality>
</emphasis>
/ 
<emphasis id="B95A1B27623DFF9829D8B49CFAEC5A5A" box="[1184,1291,1766,1792]" italics="true" pageId="123" pageNumber="124">ja. O'herl</emphasis>
/ 
<emphasis id="B95A1B27623DFF982859B49DFA6D5A5A" box="[1313,1418,1766,1792]" italics="true" pageId="123" pageNumber="124">1920.143</emphasis>
&quot;, &quot; 
<emphasis id="B95A1B27623DFF982DDBB571FEC65A7E" box="[163,289,1802,1828]" italics="true" pageId="123" pageNumber="124">
<location id="8EF191EE623DFF982DDBB571FEC65A7E" LSID="urn:lsid:plazi:treatment:03877623623DFF9E2DEFB695FCFB5F3B:8EF191EE623DFF982DDBB571FEC65A7E" box="[163,289,1802,1828]" country="Zimbabwe" county="Rhodesia" municipality="Salisbury" name="Empanduri" pageId="123" pageNumber="124">Empanduri</location>
</emphasis>
/13.1.19&quot;, &quot;Pres. by / Imp. Bur. Ent.&quot;, &quot; 
<taxonomicName id="4C2EBCB6623DFF982F9DB571FC9C5A7E" authority="Chaudoir, 1873" box="[741,891,1802,1828]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27623DFF982F9DB571FC9C5A7E" box="[741,891,1802,1828]" italics="true" pageId="123" pageNumber="124">Crossoglossa</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C2EBCB6623DFF982EE8B571FBAC5A7E" authority=", Chd." authorityName="Chd." box="[912,1099,1802,1828]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B95A1B27623DFF982EE8B571FBAC5A7E" box="[912,1099,1802,1828]" italics="true" pageId="123" pageNumber="124">ferruginea, Chd.</emphasis>
</taxonomicName>
/ DET. FR. DESCR. / G.A.K. MARSHALL.&quot; &lt;
<figureCitation id="1315DBB0623DFF982C23B555FE585A13" box="[347,447,1838,1865]" captionStart="FIGURE 75" captionStartId="124.[151,250,777,801]" captionTargetBox="[164,1425,172,747]" captionTargetId="figure-28@124.[151,1436,166,753]" captionTargetPageId="124" captionText="FIGURE 75. Median lobe of male genitalia for Parena (Parena) fulva sp. n., right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. A Holotype. B Paratype from Zimbabwe. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959094" httpUri="https://zenodo.org/record/7959094/files/figure.png" pageId="123" pageNumber="124">Fig. 75B</figureCitation>
</materialsCitation>
&gt;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE623DFF9F2DBFB52AFB465988" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124" type="diagnosis">
<paragraph id="8B91C735623DFF982DBFB52AFC555A82" blockId="123.[151,1437,1369,2008]" pageId="123" pageNumber="124">
<emphasis id="B95A1B27623DFF982DBFB52AFE2B5A36" bold="true" box="[199,460,1873,1900]" pageId="123" pageNumber="124">Diagnostic characters</emphasis>
. Dorsum yellowish brown, elytra without pattern; antennae with basal three antennomeres yellow, apical half of fourth and remaining tarsomeres black; tarsi black; postgenae with a pair of suborbital setae; pronotum with lateral margins nearly straight before posterior angles; elytra discal depression indistinct; median lobe of aedeagus very stout; apical lamella thick and coniform.
</paragraph>
<caption id="DF5197BD623AFF9F2DEFB172FC585E1F" ID-DOI="http://doi.org/10.5281/zenodo.7959094" ID-Zenodo-Dep="7959094" httpUri="https://zenodo.org/record/7959094/files/figure.png" pageId="124" pageNumber="125" startId="124.[151,250,777,801]" targetBox="[164,1425,172,747]" targetPageId="124" targetType="figure">
<paragraph id="8B91C735623AFF9F2DEFB172FC585E1F" blockId="124.[151,1437,777,838]" pageId="124" pageNumber="125">
<emphasis id="B95A1B27623AFF9F2DEFB172FEFD5E7B" bold="true" box="[151,282,777,801]" pageId="124" pageNumber="125">FIGURE 75</emphasis>
. Median lobe of male genitalia for 
<taxonomicName id="4C2EBCB6623AFF9F2FF3B171FC965E7B" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[651,881,777,801]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="fulva" status="sp. nov." subGenus="Parena">
<emphasis id="B95A1B27623AFF9F2FF3B171FD325E7B" box="[651,725,778,801]" italics="true" pageId="124" pageNumber="125">Parena</emphasis>
(
<emphasis id="B95A1B27623AFF9F2F9CB171FCC95E7B" box="[740,814,778,801]" italics="true" pageId="124" pageNumber="125">Parena</emphasis>
) 
<emphasis id="B95A1B27623AFF9F2E46B172FC965E7B" box="[830,881,777,801]" italics="true" pageId="124" pageNumber="125">fulva</emphasis>
</taxonomicName>
<emphasis id="B95A1B27623AFF9F2E00B172FC545E7B" bold="true" box="[888,947,777,801]" pageId="124" pageNumber="125">
<taxonomicNameLabel id="A269A65C623AFF9F2E00B172FC545E7B" box="[888,947,777,801]" pageId="124" pageNumber="125" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views. 
<emphasis id="B95A1B27623AFF9F2C3FB156FEBE5E1F" bold="true" box="[327,345,813,837]" pageId="124" pageNumber="125">A</emphasis>
Holotype. 
<emphasis id="B95A1B27623AFF9F2CB3B156FE3B5E1F" bold="true" box="[459,476,813,837]" pageId="124" pageNumber="125">B</emphasis>
Paratype from Zimbabwe. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B91C735623AFF9F2DBFB108FB465988" blockId="124.[151,1437,883,2026]" pageId="124" pageNumber="125">
<emphasis id="B95A1B27623AFF9F2DBFB108FE815ED7" bold="true" box="[199,358,883,909]" pageId="124" pageNumber="125">Comparisons</emphasis>
. 
<taxonomicName id="4C2EBCB6623AFF9F2C0CB10FFE205ED7" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[372,455,883,909]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="fulva" status="sp. nov.">
<emphasis id="B95A1B27623AFF9F2C0CB10FFE205ED7" box="[372,455,883,909]" italics="true" pageId="124" pageNumber="125">P. fulva</emphasis>
</taxonomicName>
<emphasis id="B95A1B27623AFF9F2CB6B108FDE95ED7" bold="true" box="[462,526,883,909]" pageId="124" pageNumber="125">
<taxonomicNameLabel id="A269A65C623AFF9F2CB6B108FDE95ED7" box="[462,526,883,909]" pageId="124" pageNumber="125" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is different from most African species in having suborbital setae. Compared with the two other African species with suborbital setae, the new species can be readily distinguished in having the dorsum with paler color, antennae distinctly bicolored, elytra without discal depressions, and pronotum with lateral margins nearly straight before posterior angles. From the original labels, we found that several specimens of this new species were misidentified as 
<taxonomicName id="4C2EBCB6623AFF9F2F63B67FFD4B5947" box="[539,684,1027,1053]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B95A1B27623AFF9F2F63B67FFD4B5947" box="[539,684,1027,1053]" italics="true" pageId="124" pageNumber="125">P. ferruginea</emphasis>
</taxonomicName>
. However, 
<taxonomicName id="4C2EBCB6623AFF9F2E56B67FFC655947" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[814,898,1027,1053]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="fulva" status="sp. nov.">
<emphasis id="B95A1B27623AFF9F2E56B67FFC655947" box="[814,898,1027,1053]" italics="true" pageId="124" pageNumber="125">P. fulva</emphasis>
</taxonomicName>
<emphasis id="B95A1B27623AFF9F2EF0B678FC2F5947" bold="true" box="[904,968,1027,1053]" pageId="124" pageNumber="125">
<taxonomicNameLabel id="A269A65C623AFF9F2EF0B678FC2F5947" box="[904,968,1027,1053]" pageId="124" pageNumber="125" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is different from the latter species in having suborbital setae present and the pronotum wider and with lateral margins only faintly sinuate before posterior angles. In several important features, the &quot;unicolor&quot; form of 
<taxonomicName id="4C2EBCB6623AFF9F2E34B637FC25593F" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[844,962,1099,1125]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27623AFF9F2E34B637FC25593F" box="[844,962,1099,1125]" italics="true" pageId="124" pageNumber="125">P. fasciata</emphasis>
</taxonomicName>
distributed in the 
<collectingRegion id="49EA09D7623AFF9F29ECB630FB17593F" box="[1172,1264,1099,1125]" country="Democratic Republic of the Congo" name="Orientale" pageId="124" pageNumber="125">Oriental</collectingRegion>
Realm is most similar to 
<taxonomicName id="4C2EBCB6623AFF9F2C74B60BFE8659D3" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[268,353,1135,1161]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="fulva" status="sp. nov.">
<emphasis id="B95A1B27623AFF9F2C74B60BFE8659D3" box="[268,353,1135,1161]" italics="true" pageId="124" pageNumber="125">P. fulva</emphasis>
</taxonomicName>
<emphasis id="B95A1B27623AFF9F2C11B614FE4D59D3" bold="true" box="[361,426,1135,1161]" pageId="124" pageNumber="125">
<taxonomicNameLabel id="A269A65C623AFF9F2C11B614FE4D59D3" box="[361,426,1135,1161]" pageId="124" pageNumber="125" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but the 
<collectingRegion id="49EA09D7623AFF9F2F69B614FD8A59D3" box="[529,621,1135,1161]" country="Democratic Republic of the Congo" name="Orientale" pageId="124" pageNumber="125">Oriental</collectingRegion>
species differs in having antennae uniformly yellow and male genitalia much slenderer. The new species also differs from all other African species in having its male genitalia very stout (AL/AW = 3.7–3.9), resembling some species from the 
<collectingRegion id="49EA09D7623AFF9F2E7FB6CCFC84598B" box="[775,867,1207,1233]" country="Democratic Republic of the Congo" name="Orientale" pageId="124" pageNumber="125">Oriental</collectingRegion>
realm, such as 
<taxonomicName id="4C2EBCB6623AFF9F2968B6C3FB7E598B" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[1040,1177,1207,1233]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="latecincta">
<emphasis id="B95A1B27623AFF9F2968B6C3FB7E598B" box="[1040,1177,1207,1233]" italics="true" pageId="124" pageNumber="125">P. latecincta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE623AFF9E2DBFB6A0FEBD5CAF" lastPageId="125" lastPageNumber="126" pageId="124" pageNumber="125" type="description">
<paragraph id="8B91C735623AFF9E2DBFB6A0FEBD5CAF" blockId="124.[151,1437,883,2026]" lastBlockId="125.[151,1437,151,609]" lastPageId="125" lastPageNumber="126" pageId="124" pageNumber="125">
<emphasis id="B95A1B27623AFF9F2DBFB6A0FEB659AF" bold="true" box="[199,337,1243,1269]" pageId="124" pageNumber="125">Description</emphasis>
. Body length 
<quantity id="4CD66AD0623AFF9F2C95B6A0FD6359AC" box="[493,644,1243,1270]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="10.4" metricValueMin="9.2" pageId="124" pageNumber="125" unit="mm" value="9.8" valueMax="10.4" valueMin="9.2">9.2–10.4 mm</quantity>
; body median-sized in the genus, subconvex. 
<emphasis id="B95A1B27623AFF9F2905B6A0FB2159AF" bold="true" box="[1149,1222,1243,1269]" pageId="124" pageNumber="125">Color.</emphasis>
Dorsum uniformly yellowish brown, head and pronotal disc slightly darker than elytra; apices of mandibles dark brown; antennae distinctly bicolor: antennomeres 1–3, and basal half of antennomere 4 yellowish brown, remainder of antennae nearly black; venter uniformly reddish brown; legs yellowish brown, tarsomeres dark brown. 
<emphasis id="B95A1B27623AFF9F29A9B73CFAF6583B" bold="true" box="[1233,1297,1351,1377]" pageId="124" pageNumber="125">Head</emphasis>
with sparse fine punctures on vertex; frons with shallow V-shaped depression; eyes large and strongly prominent; tempora very short, abruptly narrowed behind eyes, length of tempora plus neck-constriction approximately one-third of diameter of eye; postgenae with a pair of suborbital setae, slightly shorter than supraorbital ones. Antennae nearly extended to pronotal base. Labrum quadrate, apex weakly convex; mandibles short and wide; mentum with a pair of very short median setae, lateral lobes short and wide, inner margins strongly oblique, outer margins completely rounded, epilobes wide. 
<emphasis id="B95A1B27623AFF9F2CCEB464FDC85B63" bold="true" box="[438,559,1567,1593]" pageId="124" pageNumber="125">Pronotum</emphasis>
nearly quadrate, PW/PL = 1.46–1.56, slightly wider than head, PW/HW = 1.01–1.08, widest at anterior third, lateral explanations slightly wide; lateral margins rounded at anterior half and then gradually narrowed to base, straight before posterior angles; posterior angles obtuse, rounded at apex; anterior margin nearly straight at middle; posterior margin gradually oblique at sides; disc convex, usually with very fine punctures and transverse wrinkles. 
<emphasis id="B95A1B27623AFF9F2F59B4D4FD8A5B93" bold="true" box="[545,621,1711,1737]" pageId="124" pageNumber="125">Elytra</emphasis>
weakly convex, slightly dilated to apex, surface without microsculpture. Striae very shallowly incised, with rows of fine punctures; intervals faintly convex, with very sparse fine punctures. Disc without depressions, lateral sides weakly depressed near anterior third. Elytral basal pore present on base of stria 1; interval 3 with three discal setigerous pores: first one on level of scutellar apex, adjacent to stria 3; second one slightly before middle, adjacent to stria 3; third one on apical eighth, adjacent to stria 2; interval 9 with 24–26 umbilicular pores. Apical truncation indistinct, evenly rounded, outer angles completely rounded, sutural angles indistinct. 
<emphasis id="B95A1B27623AFF9F2C77B5FCFEB85AFB" bold="true" box="[271,351,1927,1953]" pageId="124" pageNumber="125">Venter</emphasis>
. Apex of abdominal sternite VII with two setae on each side in both sexes, straight in females, very weakly emarginate in males. Males with biseriate adhesive setae on full length of mesotarsomeres 2–3. 
<emphasis id="B95A1B27623AFF9F2818B5D0FF195AB3" bold="true" pageId="124" pageNumber="125">Male genitalia</emphasis>
. Median lobe of aedeagus very stout (AL/AW = 3.7–3.9); in lateral view, ventral margin slightly swollen near middle, dorsal margin strongly expanded, apical lamella thick, coniform, very weakly bent to dorsum; in dorsal view, right margin weakly sinuate before apex, apical lamella narrow, LL equal to LW, slightly narrowed to apex, apex rounded. Endophallus with very large and thick flared basal expansion of primary sclerite; flagellum thick and short, extending slightly beyond middle of median lobe; apical sclerite widely V-shaped, weakly defined, basal core distinct, strongly extended forming a well-defined transverse scaled belt; basal sheath coarsely scaled, apical sheath finely scaled, smaller than basal sheath; squamate sac well divided, near middle of median lobe, dorsal to squamate sheath; proximal sac small and dorsal-ventrally compressed, distal sac larger than proximal sac. 
<emphasis id="B95A1B27623BFF9E29A8B314FA715CD2" bold="true" box="[1232,1430,366,393]" pageId="125" pageNumber="126">Female genitalia</emphasis>
. Gonocoxite II of ovipositor nearly quadrate, slightly wider than length, apex nearly straight, weakly pointed near inner apical angle, with six ensiform setae on apical margin, two of them grouped on inner apical angle, other four equally arranged.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE623BFF9E2DBFB385FD1C5F43" box="[199,763,510,537]" pageId="125" pageNumber="126" type="distribution">
<paragraph id="8B91C735623BFF9E2DBFB385FD1C5F43" blockId="125.[151,1437,151,609]" box="[199,763,510,537]" pageId="125" pageNumber="126">
<emphasis id="B95A1B27623BFF9E2DBFB385FEBE5F42" bold="true" box="[199,345,510,536]" pageId="125" pageNumber="126">Distribution</emphasis>
(
<figureCitation id="1315DBB0623BFF9E2C10B384FE255F43" box="[360,450,511,537]" captionStart="MAP 12" captionStartId="120.[152,211,1374,1398]" captionTargetBox="[151,1436,700,1351]" captionTargetId="figure-240@120.[151,1436,700,1351]" captionText="MAP 12. Distributions for species in the Parena (Parena) stigmatica group" figureDoi="http://doi.org/10.5281/zenodo.7963697" httpUri="https://zenodo.org/record/7963697/files/figure.png" pageId="125" pageNumber="126">Map 12</figureCitation>
, red). 
<collectingCountry id="F33987A5623BFF9E2F71B384FD965F43" box="[521,625,511,537]" name="Tanzania" pageId="125" pageNumber="126">Tanzania</collectingCountry>
, 
<collectingCountry id="F33987A5623BFF9E2F03B384FD115F43" box="[635,758,511,537]" name="Zimbabwe" pageId="125" pageNumber="126">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE623BFF9E2DBFB058FCFB5F3B" pageId="125" pageNumber="126" type="etymology">
<paragraph id="8B91C735623BFF9E2DBFB058FCFB5F3B" blockId="125.[151,1437,151,609]" pageId="125" pageNumber="126">
<emphasis id="B95A1B27623BFF9E2DBFB058FEAF5F67" bold="true" box="[199,328,547,573]" pageId="125" pageNumber="126">Etymology</emphasis>
. The scientific name &quot; 
<taxonomicName id="4C2EBCB6623BFF9E2F30B058FD675F67" authorityName="Shi &amp; Liang" authorityYear="2023" box="[584,640,547,573]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="fulva">
<emphasis id="B95A1B27623BFF9E2F30B058FD675F67" box="[584,640,547,573]" italics="true" pageId="125" pageNumber="126">fulva</emphasis>
</taxonomicName>
&quot; is derived from the Latin root &quot; 
<emphasis id="B95A1B27623BFF9E2E8CB058FBC05F67" box="[1012,1063,547,573]" italics="true" pageId="125" pageNumber="126">fulv-</emphasis>
&quot; meaning yellow. It refers to the uniformly reddish yellow dorsal color of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>