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<document id="02BA2BA9C16FB10DB6E06ECFA6A12E8F" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="038776236221FF8A2DEFB6CBFBF2596F" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Parena) picipes Shi &amp; Liang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="106" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="104" updateTime="1699506634024" updateUser="plazi">
<mods:mods id="59563547109290B7FED77D038A1B1AAA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="33E1C00D9006762F4E321F19483D2E74">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="360D1B6203837610E9647DAF54574968">Shi, Hongliang</mods:namePart>
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<mods:namePart id="C3042BE7833C32D95EEF12CD8F3F2C02">Liang, Hongbin</mods:namePart>
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<mods:title id="573FE8A6A37BD12433D54D4EAD317BF7">Zootaxa</mods:title>
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<mods:date id="D9210749944F0F420A78C1BD57756FEE">2023</mods:date>
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<mods:number id="23B7BB21E9F9EA66EF9324E11650690C">2023-05-18</mods:number>
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<mods:number id="A61390F95F7166313AF34D90B349027C">5286</mods:number>
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<mods:classification id="E40D2340A40B79A93C4C59DF4507CC3B">journal article</mods:classification>
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<mods:identifier id="17E501606C832A8C7E42F3F8D26273B8" type="DOI">10.11646/zootaxa.5286.1.1</mods:identifier>
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<treatment id="038776236221FF8A2DEFB6CBFBF2596F" ID-DOI="http://doi.org/10.5281/zenodo.7963781" ID-GBIF-Taxon="209628965" ID-Zenodo-Dep="7963781" LSID="urn:lsid:plazi:treatment:038776236221FF8A2DEFB6CBFBF2596F" httpUri="http://treatment.plazi.org/id/038776236221FF8A2DEFB6CBFBF2596F" lastPageId="105" lastPageNumber="106" pageId="103" pageNumber="104">
<subSubSection id="C33494BE6221FF842DEFB6CBFDBD5990" box="[151,602,1199,1226]" pageId="103" pageNumber="104" type="nomenclature">
<paragraph id="8B91C7356221FF842DEFB6CBFDBD5990" blockId="103.[151,1113,1199,1262]" box="[151,602,1199,1226]" pageId="103" pageNumber="104">
<emphasis id="B95A1B276221FF842DEFB6CBFDBD5990" bold="true" box="[151,602,1199,1226]" pageId="103" pageNumber="104">
[30]
<taxonomicName id="4C2EBCB66221FF842DB7B6CBFE125990" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[207,501,1199,1226]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="picipes" status="sp. nov." subGenus="Parena">
<emphasis id="B95A1B276221FF842DB7B6CBFEC05990" bold="true" box="[207,295,1200,1226]" italics="true" pageId="103" pageNumber="104">Parena</emphasis>
(
<emphasis id="B95A1B276221FF842C40B6CBFE775990" bold="true" box="[312,400,1200,1226]" italics="true" pageId="103" pageNumber="104">Parena</emphasis>
)
<emphasis id="B95A1B276221FF842CDAB6CBFE125990" bold="true" box="[418,501,1200,1226]" italics="true" pageId="103" pageNumber="104">picipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269A65C6221FF842C84B6CBFDBD5990" box="[508,602,1200,1226]" pageId="103" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6221FF842DEFB6AFFBBE59B4" box="[151,1113,1236,1262]" pageId="103" pageNumber="104" type="description">
<paragraph id="8B91C7356221FF842DEFB6AFFBBE59B4" blockId="103.[151,1113,1199,1262]" box="[151,1113,1236,1262]" pageId="103" pageNumber="104">
Habitus:
<figureCitation id="1315DBB06221FF842D85B6AFFE4259B4" box="[253,421,1236,1262]" captionStart="FIGURE 62" captionStartId="104.[151,250,1585,1609]" captionTargetBox="[163,1434,168,1536]" captionTargetId="figure-28@104.[151,1436,166,1561]" captionTargetPageId="104" captionText="FIGURE 62. Habitus of Parena (Parena) nigrolineata species group. A Holotype of P. picipes sp. n. B Paratype of P. picipes sp. n., female. C Holotype of Parena andrewesi Jedlička. D P. andrewesi Jedlička, female from Philippine. E Lectotype of Parena politissima (Chaudoir). F P. politissima (Chaudoir), female from Vanuatu. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959066" httpUri="https://zenodo.org/record/7959066/files/figure.png" pageId="103" pageNumber="104">Figs 62A, 62B</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB06221FF842F27B6AFFD5659B4" box="[607,689,1236,1262]" captionStart="FIGURE 63" captionStartId="105.[151,250,1921,1945]" captionTargetBox="[160,1429,1096,1897]" captionTargetId="figure-458@105.[151,1436,1086,1898]" captionTargetPageId="105" captionText="FIGURE 63. Median lobe of male genitalia for Parena (Parena) picipes sp. n., right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959068" httpUri="https://zenodo.org/record/7959068/files/figure.png" pageId="103" pageNumber="104">Fig. 63</figureCitation>
. Gonocoxites of ovipositor:
<figureCitation id="1315DBB06221FF842E88B6AFFBB259B4" box="[1008,1109,1236,1262]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="103" pageNumber="104">Fig. 11O</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6221FF842DEFB767FE7658B0" pageId="103" pageNumber="104" type="materials_examined">
<paragraph id="8B91C7356221FF842DEFB767FB8B586C" blockId="103.[151,1437,1308,2018]" box="[151,1132,1308,1335]" pageId="103" pageNumber="104">
<materialsCitation id="3B46CD686221FF842DEFB767FB8F586C" ID-GBIF-Occurrence="4115871438" box="[151,1128,1308,1335]" county="Borneo" elevation="1182" latitude="5.443" location="Reserve" longLatPrecision="7" longitude="116.4512" municipality="Trus Madi forest" pageId="103" pageNumber="104" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B95A1B276221FF842DEFB767FED7586C" bold="true" box="[151,304,1308,1334]" pageId="103" pageNumber="104">
<typeStatus id="549579976221FF842DEFB767FF35586C" box="[151,210,1308,1334]" pageId="103" pageNumber="104">Type</typeStatus>
locality
</emphasis>
.
<collectingCounty id="62F0BFB96221FF842C46B767FE74586C" box="[318,403,1308,1334]" pageId="103" pageNumber="104">Borneo</collectingCounty>
,
<collectingRegion id="49EA09D76221FF842CE6B767FE02586C" box="[414,485,1308,1334]" country="Malaysia" name="Sabah" pageId="103" pageNumber="104">Sabah</collectingRegion>
,
<collectingMunicipality id="6BF55D4F6221FF842C88B767FD4B586C" box="[496,684,1308,1334]" pageId="103" pageNumber="104">Trus Madi forest</collectingMunicipality>
<location id="8EF191EE6221FF842FCAB767FCE8586C" LSID="urn:lsid:plazi:treatment:038776236221FF8A2DEFB6CBFBF2596F:8EF191EE6221FF842FCAB767FCE8586C" box="[690,783,1308,1334]" county="Borneo" latitude="5.443" longLatPrecision="7" longitude="116.4512" municipality="Trus Madi forest" name="Reserve" pageId="103" pageNumber="104" stateProvince="Sabah">Reserve</location>
,
<geoCoordinate id="EE1AA1F26221FF842E63B767FC98586D" box="[795,895,1308,1335]" degrees="5.4430" direction="north" orientation="latitude" pageId="103" pageNumber="104" precision="5" value="5.443">N5.4430</geoCoordinate>
,
<geoCoordinate id="EE1AA1F26221FF842EF2B767FBE1586D" box="[906,1030,1308,1335]" degrees="116.4512" direction="east" orientation="longitude" pageId="103" pageNumber="104" precision="5" value="116.4512">E116.4512</geoCoordinate>
,
<quantity id="4CD66AD06221FF842969B767FB8F586C" box="[1041,1128,1308,1334]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="103" pageNumber="104" unit="m" value="1182.0">
<elevation id="000320066221FF842969B767FB8F586C" box="[1041,1128,1308,1334]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="103" pageNumber="104" unit="m" value="1182.0">1182 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8B91C7356221FF842DBFB73BFE7658B0" blockId="103.[151,1437,1308,2018]" pageId="103" pageNumber="104">
<emphasis id="B95A1B276221FF842DBFB73BFE965800" bold="true" box="[199,369,1344,1370]" pageId="103" pageNumber="104">Type material</emphasis>
.
<materialsCitation id="3B46CD686221FF842CFAB73BFA6058F8" ID-GBIF-Occurrence="4115871492" collectingDate="2016-04-28" collectorName="Liu Y. Lgt." county="Borneo" elevation="1182" latitude="5.443" location="Jungle Girl Camp" longLatPrecision="7" longitude="116.4512" municipality="Keningau" pageId="103" pageNumber="104" specimenCount="2" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B95A1B276221FF842CFAB73BFE085800" bold="true" box="[386,495,1344,1370]" pageId="103" pageNumber="104">
<typeStatus id="549579976221FF842CFAB73BFE085800" box="[386,495,1344,1370]" pageId="103" pageNumber="104" type="holotype">Holotype</typeStatus>
</emphasis>
(IZAS,
<figureCitation id="1315DBB06221FF842F29B73BFD5A5800" box="[593,701,1344,1370]" captionStart="FIGURE 62" captionStartId="104.[151,250,1585,1609]" captionTargetBox="[163,1434,168,1536]" captionTargetId="figure-28@104.[151,1436,166,1561]" captionTargetPageId="104" captionText="FIGURE 62. Habitus of Parena (Parena) nigrolineata species group. A Holotype of P. picipes sp. n. B Paratype of P. picipes sp. n., female. C Holotype of Parena andrewesi Jedlička. D P. andrewesi Jedlička, female from Philippine. E Lectotype of Parena politissima (Chaudoir). F P. politissima (Chaudoir), female from Vanuatu. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959066" httpUri="https://zenodo.org/record/7959066/files/figure.png" pageId="103" pageNumber="104">Fig. 62A</figureCitation>
): male, body length=
<quantity id="4CD66AD06221FF842EC3B73BFBF15800" box="[955,1046,1344,1371]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="103" pageNumber="104" unit="mm" value="8.5">8.5 mm</quantity>
, board mounted, &quot;
<collectingCounty id="62F0BFB96221FF842991B73BFAA45800" box="[1257,1347,1344,1370]" pageId="103" pageNumber="104">Borneo</collectingCounty>
:
<collectingRegion id="49EA09D76221FF842829B73BFA7F5800" box="[1361,1432,1344,1370]" country="Malaysia" name="Sabah" pageId="103" pageNumber="104">Sabah</collectingRegion>
,
<collectingMunicipality id="6BF55D4F6221FF842DEFB71FFEE35824" box="[151,260,1380,1406]" pageId="103" pageNumber="104">Keningau</collectingMunicipality>
/ district,
<location id="8EF191EE6221FF842C0EB71FFDD95824" LSID="urn:lsid:plazi:treatment:038776236221FF8A2DEFB6CBFBF2596F:8EF191EE6221FF842C0EB71FFDD95824" box="[374,574,1380,1406]" county="Borneo" latitude="5.443" longLatPrecision="7" longitude="116.4512" municipality="Keningau" name="Jungle Girl Camp" pageId="103" pageNumber="104" stateProvince="Sabah">Jungle Girl Camp</location>
/
<geoCoordinate id="EE1AA1F26221FF842F2CB71FFD5E5825" box="[596,697,1380,1407]" degrees="5.4430" direction="north" orientation="latitude" pageId="103" pageNumber="104" precision="5" value="5.443">N5.4430</geoCoordinate>
,
<geoCoordinate id="EE1AA1F26221FF842FBCB71FFCA75825" box="[708,832,1380,1407]" degrees="116.4512" direction="east" orientation="longitude" pageId="103" pageNumber="104" precision="5" value="116.4512">E116.4512</geoCoordinate>
;
<quantity id="4CD66AD06221FF842E34B71FFC475824" box="[844,928,1380,1406]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="103" pageNumber="104" unit="m" value="1182.0">
<elevation id="000320066221FF842E34B71FFC475824" box="[844,928,1380,1406]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="103" pageNumber="104" unit="m" value="1182.0">1182 m</elevation>
</quantity>
/ Shi H.L. &amp;
<collectorName id="26DBA2E36221FF842942B71FFB525824" box="[1082,1205,1380,1406]" pageId="103" pageNumber="104">Liu Y. Lgt.</collectorName>
<collectingMethod id="526FBF226221FF8429C3B71FFAC55824" box="[1211,1314,1380,1406]" pageId="103" pageNumber="104">light trap</collectingMethod>
/ Ins. Zoo. CAS.
<date id="FF90E1F56221FF842DA4B7F3FEBE58F8" box="[220,345,1416,1442]" pageId="103" pageNumber="104" value="2016-04-28">
<collectingDate id="EFD4181D6221FF842DA4B7F3FEBE58F8" box="[220,345,1416,1442]" pageId="103" pageNumber="104" value="2016-04-28">2016.IV.28</collectingDate>
</date>
N&quot;, &quot;HOLOTYPE /
<taxonomicName id="4C2EBCB66221FF842F43B7F3FD0A58F8" box="[571,749,1416,1442]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="subGenus" subGenus="Parena">Parena (Parena)</taxonomicName>
/ picipes
<taxonomicNameLabel id="A269A65C6221FF842E22B7F3FC5758F8" box="[858,944,1416,1442]" pageId="103" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
/ des. Shi H.L. 2022&quot; [red label] &lt;
<figureCitation id="1315DBB06221FF84284EB7F3FA6058F8" box="[1334,1415,1416,1442]" captionStart="FIGURE 63" captionStartId="105.[151,250,1921,1945]" captionTargetBox="[160,1429,1096,1897]" captionTargetId="figure-458@105.[151,1436,1086,1898]" captionTargetPageId="105" captionText="FIGURE 63. Median lobe of male genitalia for Parena (Parena) picipes sp. n., right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959068" httpUri="https://zenodo.org/record/7959068/files/figure.png" pageId="103" pageNumber="104">Fig. 63</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD686221FF842DEFB7D7FE9A58B0" ID-GBIF-Occurrence="4115871454" collectingDate="2016-04-24" collectingDateMax="2016-05-02" collectingDateMin="2016-04-24" collectorName="Liu Y. Lgt." county="Borneo" elevation="1182" latitude="5.443" location="Jungle Girl Camp" longLatPrecision="7" longitude="116.4512" municipality="Keningau" pageId="103" pageNumber="104" specimenCount="8" specimenCount-female="5" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B95A1B276221FF842DEFB7D7FEE8589C" bold="true" box="[151,271,1452,1478]" pageId="103" pageNumber="104">
<typeStatus id="549579976221FF842DEFB7D7FEE8589C" box="[151,271,1452,1478]" pageId="103" pageNumber="104" type="paratype">Paratypes</typeStatus>
</emphasis>
(IZAS):
<specimenCount id="9D280CBC6221FF842C0EB7D7FE28589C" box="[374,463,1452,1478]" pageId="103" pageNumber="104" type="male">3 males</specimenCount>
,
<specimenCount id="9D280CBC6221FF842CA3B7D7FDAD589C" box="[475,586,1452,1479]" pageId="103" pageNumber="104" type="female">5 females</specimenCount>
, same data as the holotype, with different date from
<date id="FF90E1F56221FF8429EFB7D7FAF4589C" box="[1175,1299,1452,1478]" pageId="103" pageNumber="104" value="2016-04-24">
<collectingDate id="EFD4181D6221FF8429EFB7D7FAF4589C" box="[1175,1299,1452,1478]" pageId="103" pageNumber="104" value="2016-04-24">2016.IV.24</collectingDate>
</date>
to
<date id="FF90E1F56221FF842840B7D7FA7B589C" box="[1336,1436,1452,1478]" pageId="103" pageNumber="104" value="2016-05-02">
<collectingDate id="EFD4181D6221FF842840B7D7FA7B589C" box="[1336,1436,1452,1478]" pageId="103" pageNumber="104" value="2016-05-02">2016.V.2</collectingDate>
</date>
&lt;
<figureCitation id="1315DBB06221FF842DDEB7ABFEE558B0" box="[166,258,1488,1514]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. A–D females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. E–J males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="103" pageNumber="104">Figs 8H</figureCitation>
,
<figureCitation id="1315DBB06221FF842C76B7ABFED858B0" box="[270,319,1488,1514]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="103" pageNumber="104">11O</figureCitation>
,
<figureCitation id="1315DBB06221FF842C34B7ABFE9A58B0" box="[332,381,1488,1514]" captionStart="FIGURE 62" captionStartId="104.[151,250,1585,1609]" captionTargetBox="[163,1434,168,1536]" captionTargetId="figure-28@104.[151,1436,166,1561]" captionTargetPageId="104" captionText="FIGURE 62. Habitus of Parena (Parena) nigrolineata species group. A Holotype of P. picipes sp. n. B Paratype of P. picipes sp. n., female. C Holotype of Parena andrewesi Jedlička. D P. andrewesi Jedlička, female from Philippine. E Lectotype of Parena politissima (Chaudoir). F P. politissima (Chaudoir), female from Vanuatu. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959066" httpUri="https://zenodo.org/record/7959066/files/figure.png" pageId="103" pageNumber="104">62B</figureCitation>
</materialsCitation>
&gt;.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6221FF842DBFB78FFDBF5A2C" pageId="103" pageNumber="104" type="diagnosis">
<paragraph id="8B91C7356221FF842DBFB78FFE225B20" blockId="103.[151,1437,1308,2018]" pageId="103" pageNumber="104">
<emphasis id="B95A1B276221FF842DBFB78FFE2D5B54" bold="true" box="[199,458,1524,1550]" pageId="103" pageNumber="104">Diagnostic characters</emphasis>
. Dorsum reddish brown, elytra with black lateral stripes, without metallic lustre, elytra lateral margins black; legs with apical third of femora and entire tibiae black; postgenae without suborbital setae; median lobe of aedeagus stout, right margin weakly sinuate before apex, apical lamella narrow, LL 1.5 times LW, apex slightly bent dorsally.
</paragraph>
<paragraph id="8B91C7356221FF842DBFB4FFFDBF5A2C" blockId="103.[151,1437,1308,2018]" pageId="103" pageNumber="104">
<emphasis id="B95A1B276221FF842DBFB4FFFE815BC4" bold="true" box="[199,358,1668,1694]" pageId="103" pageNumber="104">Comparisons</emphasis>
.
<taxonomicName id="4C2EBCB66221FF842C0CB4FEFE385BC4" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[372,479,1669,1694]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="picipes" status="sp. nov.">
<emphasis id="B95A1B276221FF842C0CB4FEFE385BC4" box="[372,479,1669,1694]" italics="true" pageId="103" pageNumber="104">P. picipes</emphasis>
</taxonomicName>
<emphasis id="B95A1B276221FF842C9FB4FFFDC05BC4" bold="true" box="[487,551,1668,1694]" pageId="103" pageNumber="104">
<taxonomicNameLabel id="A269A65C6221FF842C9FB4FFFDC05BC4" box="[487,551,1668,1694]" pageId="103" pageNumber="104" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is distinguishable by its black tibiae among all
<taxonomicName id="4C2EBCB66221FF842946B4FEFB0F5BC4" box="[1086,1256,1668,1694]" pageId="103" pageNumber="104">
<emphasis id="B95A1B276221FF842946B4FEFB685BC4" box="[1086,1167,1669,1694]" italics="true" pageId="103" pageNumber="104">Parena</emphasis>
species
</taxonomicName>
with elytral lateral stripes. It is most similar to
<taxonomicName id="4C2EBCB66221FF842F79B4D2FD415B98" baseAuthorityName="Chaudoir" baseAuthorityYear="1852" box="[513,678,1704,1730]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="nigrolineata">
<emphasis id="B95A1B276221FF842F79B4D2FD415B98" box="[513,678,1704,1730]" italics="true" pageId="103" pageNumber="104">P. nigrolineata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2EBCB66221FF842FA5B4D2FC295B98" authorityName="Habu. Scale" authorityYear="1964" box="[733,974,1704,1730]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="amamiooshimensis">
<emphasis id="B95A1B276221FF842FA5B4D2FC295B98" box="[733,974,1704,1730]" italics="true" pageId="103" pageNumber="104">P. amamiooshimensis</emphasis>
</taxonomicName>
, but different in the black color of tibiae, femoral apices, and elytra lateral margins. The new species also has the male genitalia very different from the other two species. Compared to
<taxonomicName id="4C2EBCB66221FF842CB9B48AFD825A50" baseAuthorityName="Chaudoir" baseAuthorityYear="1852" box="[449,613,1776,1802]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="nigrolineata">
<emphasis id="B95A1B276221FF842CB9B48AFD825A50" box="[449,613,1776,1802]" italics="true" pageId="103" pageNumber="104">P. nigrolineata</emphasis>
</taxonomicName>
, the new species is different in the median lobe stouter (AL/AW 4.4–4.6, versus 5.3–6.0 in
<taxonomicName id="4C2EBCB66221FF842C19B56EFDE15A74" baseAuthorityName="Chaudoir" baseAuthorityYear="1852" box="[353,518,1812,1838]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="nigrolineata">
<emphasis id="B95A1B276221FF842C19B56EFDE15A74" box="[353,518,1812,1838]" italics="true" pageId="103" pageNumber="104">P. nigrolineata</emphasis>
</taxonomicName>
) and apical lamella much narrower (LL 1.5 times LW, versus LL slightly greater than LW in
<taxonomicName id="4C2EBCB66221FF842C59B542FE205A08" baseAuthorityName="Chaudoir" baseAuthorityYear="1852" box="[289,455,1848,1874]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="nigrolineata">
<emphasis id="B95A1B276221FF842C59B542FE205A08" box="[289,455,1848,1874]" italics="true" pageId="103" pageNumber="104">P. nigrolineata</emphasis>
</taxonomicName>
). Compared to
<taxonomicName id="4C2EBCB66221FF842F05B542FC885A08" authorityName="Habu. Scale" authorityYear="1964" box="[637,879,1848,1874]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="amamiooshimensis">
<emphasis id="B95A1B276221FF842F05B542FC885A08" box="[637,879,1848,1874]" italics="true" pageId="103" pageNumber="104">P. amamiooshimensis</emphasis>
</taxonomicName>
, the new species is different in median lobe less stout and apical lamella much narrower.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE6221FF8A2DBFB5FBFE6E5E93" lastPageId="105" lastPageNumber="106" pageId="103" pageNumber="104" type="description">
<paragraph id="8B91C7356221FF8A2DBFB5FBFE6E5E93" blockId="103.[151,1437,1308,2018]" lastBlockId="105.[151,1437,151,1077]" lastPageId="105" lastPageNumber="106" pageId="103" pageNumber="104">
<emphasis id="B95A1B276221FF842DBFB5FBFEB65AC0" bold="true" box="[199,337,1920,1946]" pageId="103" pageNumber="104">Description</emphasis>
. Body length
<quantity id="4CD66AD06221FF842C80B5FBFD625AC0" box="[504,645,1920,1947]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" metricValueMax="9.3" metricValueMin="8.5" pageId="103" pageNumber="104" unit="mm" value="8.9" valueMax="9.3" valueMin="8.5">8.5–9.3 mm</quantity>
; body median-sized in the genus, subconvex.
<emphasis id="B95A1B276221FF8429E2B5FBFB045AC0" bold="true" box="[1178,1251,1920,1946]" pageId="103" pageNumber="104">Color.</emphasis>
Dorsum mostly reddish brown, elytra with black lateral stripes, without trace of metallic lustre; mouthparts yellowish brown, inner margin of mandibles black; antennae bicolor with sharp contrast, antennomeres 1–2 brown, antennomere 3 brown to piceous; the rests black to pitchy brown; pronotum reddish brown, lateral margins slightly darkened; elytra with lateral black stripes, sixth to ninth intervals and elytra lateral margins completely black, median red patch usually widest to fifth striae, lateral stripes widely continuous at apex, reaching interval 2 or 3 at base; venter reddish brown, mesosternal process dark brown; legs with apical third of femora and entire tibiae black, tarsomeres dark brown.
<emphasis id="B95A1B27622EFF8B2DEFB534FF305A33" bold="true" box="[151,215,1871,1897]" pageId="104" pageNumber="105">Head</emphasis>
with sparse fine punctures on vertex; frons with V-shaped depression; eyes large and strongly prominent; tempora short, abruptly narrowed behind eyes, length of tempora plus neck-constriction approximately one-third of diameter of eye; postgenae without suborbital setae. Antennae barely reaching pronotal base. Labrum nearly quadrate, apex nearly straight; mandibles short and wide; mentum with a pair of minute median setae, barely visible in some specimens, lateral lobes short and wide, inner margins strongly oblique, outer margins completely rounded, epilobes wide.
<emphasis id="B95A1B27622FFF8A2C30B2ECFE265DEB" bold="true" box="[328,449,151,177]" pageId="105" pageNumber="106">Pronotum</emphasis>
rounded-hexagon, PW/PL = 1.34–1.40, usually slightly wider than head, PW/HW = 1.00–1.09, widest at anterior third, lateral explanations slightly wide; lateral margins fully rounded at anterior half and then strongly narrowed to base with shallow sinuation before posterior angles; posterior angles obtuse, rounded at apex; anterior margin nearly straight at middle; posterior margin gradually oblique at sides; disc convex, usually with very fine transverse wrinkles.
<emphasis id="B95A1B27622FFF8A2F52B35CFD915C1B" bold="true" box="[554,630,295,321]" pageId="105" pageNumber="106">Elytra</emphasis>
weakly convex, slightly dilated to apex. Striae shallowly incised, with rows of fine punctures; intervals slightly convex, with sparse fine punctures about half dense as interval punctures. Disc with very faint depressions near basal third of intervals 3 to 6, lateral sides slightly depressed near anterior third. Elytral basal pore present on base of first stria; interval 3 with three discal setigerous pores: first one on level of scutellar apex, adjacent to stria 3; second one slightly before middle, adjacent to stria 3; third one on apical eighth, adjacent to stria 2; interval 9 with 21–23 umbilicular pores. Apical truncation indistinct, evenly rounded, sutural angles indistinct.
<emphasis id="B95A1B27622FFF8A2C26B384FE495F43" bold="true" box="[350,430,511,537]" pageId="105" pageNumber="106">Venter</emphasis>
. Apex of abdominal sternite VII very shallowly emarginate in males, straight in females, with two setae on each side in both sexes. Males with biseriate adhesive setae on apical half of mesotarsomere 1 ventrally, and almost full length of mesotarsomeres 2–3.
<emphasis id="B95A1B27622FFF8A2E76B03CFC5F5F3A" bold="true" box="[782,952,582,609]" pageId="105" pageNumber="106">Male genitalia</emphasis>
(
<figureCitation id="1315DBB0622FFF8A2EBEB03CFBFF5F3B" box="[966,1048,583,609]" captionStart="FIGURE 63" captionStartId="105.[151,250,1921,1945]" captionTargetBox="[160,1429,1096,1897]" captionTargetId="figure-458@105.[151,1436,1086,1898]" captionTargetPageId="105" captionText="FIGURE 63. Median lobe of male genitalia for Parena (Parena) picipes sp. n., right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959068" httpUri="https://zenodo.org/record/7959068/files/figure.png" pageId="105" pageNumber="106">Fig. 63</figureCitation>
). Median lobe of aedeagus slightly stout (AL/AW = 4.4–4.6); in lateral view, ventral margin almost straight near middle, apical lamella slightly bent to dorsum; in dorsal view, right margin slightly sinuate before apex, apical lamella narrow, LL 1.5 times LW, slightly narrowed to apex, apex rounded. Endophallus with distinct flared basal expansion of primary sclerite; flagellum extending to the base of apical orifice; apical sclerite narrowly V-shaped, well chitinized, basal core distinct, ovate, heavily chitinized; squamate sheath heavily scaled, basal sheath larger than apical sheath; squamate sac heavily scaled and well divided, near middle of median lobe, dorsal-right to squamate sheath; proximal and distal sacs laterally compressed, forming a pair of file-like pieces in dorsal view, distal sac slightly smaller and closer to apex than proximal one.
<emphasis id="B95A1B27622FFF8A2C41B11CFE1A5EDA" bold="true" box="[313,509,870,897]" pageId="105" pageNumber="106">Female genitalia</emphasis>
. Gonocoxite II (
<figureCitation id="1315DBB0622FFF8A2FCDB11CFCFE5EDB" box="[693,793,871,897]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="105" pageNumber="106">Fig. 11O</figureCitation>
) of ovipositor nearly quadrate, slightly wider than length, apex slightly concave, slightly pointed near inner apical angle, with three ensiform setae on inner apical angle, two on outer apical angle.
</paragraph>
</subSubSection>
<caption id="DF5197BD622EFF8B2DEFB44AFB455BCB" ID-DOI="http://doi.org/10.5281/zenodo.7959066" ID-Zenodo-Dep="7959066" httpUri="https://zenodo.org/record/7959066/files/figure.png" pageId="104" pageNumber="105" startId="104.[151,250,1585,1609]" targetBox="[163,1434,168,1536]" targetPageId="104" targetType="figure">
<paragraph id="8B91C735622EFF8B2DEFB44AFB455BCB" blockId="104.[151,1436,1585,1681]" pageId="104" pageNumber="105">
<emphasis id="B95A1B27622EFF8B2DEFB44AFEFD5B13" bold="true" box="[151,282,1585,1609]" pageId="104" pageNumber="105">FIGURE 62</emphasis>
. Habitus of
<taxonomicName id="4C2EBCB6622EFF8B2CE1B449FD215B13" baseAuthorityName="Chaudoir" baseAuthorityYear="1852" box="[409,710,1585,1609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="nigrolineata" subGenus="Parena">
<emphasis id="B95A1B27622EFF8B2CE1B449FE045B13" box="[409,483,1586,1609]" italics="true" pageId="104" pageNumber="105">Parena</emphasis>
(
<emphasis id="B95A1B27622EFF8B2C89B449FDDC5B13" box="[497,571,1586,1609]" italics="true" pageId="104" pageNumber="105">Parena</emphasis>
)
<emphasis id="B95A1B27622EFF8B2F32B44AFD215B13" box="[586,710,1585,1609]" italics="true" pageId="104" pageNumber="105">nigrolineata</emphasis>
</taxonomicName>
species group.
<emphasis id="B95A1B27622EFF8B2E1EB44AFC9F5B13" bold="true" box="[870,888,1585,1609]" pageId="104" pageNumber="105">A</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6622EFF8B2978B449FB865B13" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[1024,1121,1586,1609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="picipes" status="sp. nov.">
<emphasis id="B95A1B27622EFF8B2978B449FB865B13" box="[1024,1121,1586,1609]" italics="true" pageId="104" pageNumber="105">P. picipes</emphasis>
</taxonomicName>
<emphasis id="B95A1B27622EFF8B2910B44AFB5E5B13" bold="true" box="[1128,1209,1585,1609]" pageId="104" pageNumber="105">
<taxonomicNameLabel id="A269A65C622EFF8B2910B44AFB465B13" box="[1128,1185,1585,1609]" pageId="104" pageNumber="105" rank="species">sp. n.</taxonomicNameLabel>
B
</emphasis>
Paratype of
<taxonomicName id="4C2EBCB6622EFF8B2843B449FA7B5B13" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[1339,1436,1586,1609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="picipes" status="sp. nov.">
<emphasis id="B95A1B27622EFF8B2843B449FA7B5B13" box="[1339,1436,1586,1609]" italics="true" pageId="104" pageNumber="105">P. picipes</emphasis>
</taxonomicName>
<emphasis id="B95A1B27622EFF8B2DEFB42EFF345B37" bold="true" box="[151,211,1621,1645]" pageId="104" pageNumber="105">
<taxonomicNameLabel id="A269A65C622EFF8B2DEFB42EFF345B37" box="[151,211,1621,1645]" pageId="104" pageNumber="105" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, female.
<emphasis id="B95A1B27622EFF8B2C4DB42EFEA05B37" bold="true" box="[309,327,1621,1645]" pageId="104" pageNumber="105">C</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6622EFF8B2CADB42DFCEC5B37" authority="Jedlicka. D" authorityName="Jedlicka. D" box="[469,779,1621,1645]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B95A1B27622EFF8B2CADB42DFD695B37" box="[469,654,1621,1645]" italics="true" pageId="104" pageNumber="105">Parena andrewesi</emphasis>
Jedlička.
<emphasis id="B95A1B27622EFF8B2F81B42EFCEC5B37" bold="true" box="[761,779,1621,1645]" pageId="104" pageNumber="105">D</emphasis>
</taxonomicName>
<taxonomicName id="4C2EBCB6622EFF8B2E6CB42DFC125B37" authority="Jedlicka" authorityName="Jedlicka, Philippine. Scale" authorityYear="1934" box="[788,1013,1621,1645]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B95A1B27622EFF8B2E6CB42DFC715B37" box="[788,918,1621,1645]" italics="true" pageId="104" pageNumber="105">P. andrewesi</emphasis>
Jedlička
</taxonomicName>
, female from Philippine.
<emphasis id="B95A1B27622EFF8B2878B42EFAF65B37" bold="true" box="[1280,1297,1621,1645]" pageId="104" pageNumber="105">E</emphasis>
Lectotype of
<taxonomicName id="4C2EBCB6622EFF8B2DEFB401FE285BCB" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1883" box="[151,463,1657,1681]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="politissima">
<emphasis id="B95A1B27622EFF8B2DEFB401FEB15BCB" box="[151,342,1657,1681]" italics="true" pageId="104" pageNumber="105">Parena politissima</emphasis>
(Chaudoir)
</taxonomicName>
.
<emphasis id="B95A1B27622EFF8B2CA1B402FE0F5BCB" bold="true" box="[473,488,1657,1681]" pageId="104" pageNumber="105">F</emphasis>
<taxonomicName id="4C2EBCB6622EFF8B2C96B401FD155BCB" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1883" box="[494,754,1657,1681]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="politissima">
<emphasis id="B95A1B27622EFF8B2C96B401FE1A5BCB" box="[494,509,1658,1681]" italics="true" pageId="104" pageNumber="105">P</emphasis>
.
<emphasis id="B95A1B27622EFF8B2F72B402FD9E5BCB" box="[522,633,1657,1681]" italics="true" pageId="104" pageNumber="105">politissima</emphasis>
(Chaudoir)
</taxonomicName>
, female from Vanuatu. Scale bars = 2 mm.
</paragraph>
</caption>
<subSubSection id="C33494BE622FFF8A2DBFB1A9FBAF5EB7" box="[199,1096,978,1005]" pageId="105" pageNumber="106" type="distribution">
<paragraph id="8B91C735622FFF8A2DBFB1A9FBAF5EB7" blockId="105.[151,1437,151,1077]" box="[199,1096,978,1005]" pageId="105" pageNumber="106">
<emphasis id="B95A1B27622FFF8A2DBFB1A9FEBE5EB6" bold="true" box="[199,345,978,1004]" pageId="105" pageNumber="106">Distribution</emphasis>
(
<figureCitation id="1315DBB0622FFF8A2C10B1A8FE255EB7" box="[360,450,979,1005]" captionStart="MAP 10" captionStartId="95.[152,211,1171,1195]" captionTargetBox="[164,1423,181,1147]" captionTargetId="figure-19@95.[164,1423,181,1147]" captionText="MAP 10. Distributions for species in the Parena (Parena) nigrolineata group." figureDoi="http://doi.org/10.5281/zenodo.7963687" httpUri="https://zenodo.org/record/7963687/files/figure.png" pageId="105" pageNumber="106">Map 10</figureCitation>
, green). Only known from the
<typeStatus id="54957997622FFF8A2E64B1A8FCAB5EB7" box="[796,844,979,1005]" pageId="105" pageNumber="106">type</typeStatus>
locality in N. Boreno.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE622FFF8A2DBFB18CFBF2596F" pageId="105" pageNumber="106" type="etymology">
<paragraph id="8B91C735622FFF8A2DBFB18CFBF2596F" blockId="105.[151,1437,151,1077]" pageId="105" pageNumber="106">
<emphasis id="B95A1B27622FFF8A2DBFB18CFEAF594B" bold="true" box="[199,328,1015,1041]" pageId="105" pageNumber="106">Etymology</emphasis>
. The scientific name of the new species is derived from Latin meaning black legs. The black-colored tibiae is one of the most important diagnostic character of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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