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<mods:title id="FC580DF567F6FEF5E28D851E520B15AF">Novelties in Aspidistra (Asparagaceae, Convallarioideae) with erect stems in the flora of Vietnam: A. quanii, a new species, and A. erecta, a new national record</mods:title>
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<mods:namePart id="74C096F3F5EFF5962B525CC70F4544D0">Averyanov, Leonid V.</mods:namePart>
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<emphasis id="B959DB20FFE52C39FF6AF9D5FEBAF9E2" bold="true" box="[136,330,1553,1580]" italics="true" pageId="1" pageNumber="40">Aspidistra erecta</emphasis>
Yan Liu &amp; C.I Peng
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in 
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<bibRefCitation id="EFBC7AC3FFE52C39FDB1F9D6FCB2F9E2" author="Liu, Y. &amp; Kono, Y. &amp; Lin, C. - R. &amp; Xu, W. - B. &amp; Peng, C." box="[595,834,1554,1580]" pageId="1" pageNumber="40" pagination="367 - 373" refId="ref7475" refString="Liu, Y., Kono, Y., Lin, C. - R., Xu, W. - B. &amp; Peng, C. - I (2011) Aspidistra erecta (Asparagaceae), a new species from limestone areas in Guangxi, China. Botanical Studies 52: 367 - 373." type="journal article" year="2011">
Liu 
<emphasis id="B959DB20FFE52C39FD63F9D7FD49F9E2" box="[641,697,1554,1580]" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
(2011: 367)
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(
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&amp; 
<figureCitation id="13161BB7FFE52C39FF15F9F1FEF6F981" box="[247,262,1589,1615]" captionStart="FIGURE 2" captionStartId="3.[136,229,1866,1888]" captionTargetBox="[243,1344,190,1841]" captionTargetId="figure-18@3.[243,1344,190,1841]" captionTargetPageId="3" captionText="FIGURE 2. Aspidistra erecta. A. Flowering plant. B. Apical portion of stem. C. Leaves, adaxial and abaxial side. D, E. Flower bud and open flower, side view.F. Flower on shoot, front view. G. Flower, side and half-side views. H. Inflorescence bracts, abaxial side. I. Flower, front view (perianth lobes removed), showing stamens and stigma. J. Sagittal section of flower, front and side views. K. Perianth lobes, adaxial and abaxial surface. L. Peduncle and sagittally dissected flower (with perianth lobes removed), showing intact pistil. M. Stamen, front and side view. N. Pistil, side view. O. Pistil, view from above. P. Sagittal section of pistil. Bui Van Huong, Truong Ba Vuong, BVH s.n. Photos by Bui Van Huong, correction and design by L. Averyanov and T. Maisak." figureDoi="http://doi.org/10.5281/zenodo.13213476" httpUri="https://zenodo.org/record/13213476/files/figure.png" pageId="1" pageNumber="40">2</figureCitation>
)
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<emphasis id="B959DB20FFE52C39FF6AF9BFFF3CF95B" bold="true" box="[136,204,1659,1685]" pageId="1" pageNumber="40">Type:</emphasis>
— 
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. 
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Autonomous Region: Napo Xian (County), alt. 
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, in a valley on limestone hill, 
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, 
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0782
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<emphasis id="B959DB20FFE52C39FF5FF905FE1AF915" bold="true" box="[189,490,1729,1755]" pageId="1" pageNumber="40">Ecology and phenology:</emphasis>
—Primary and secondary evergreen broad-leaved forests on karstic limestone at elevations of 
<quantity id="4CD5AAD7FFE52C39FEFDF920FE5CF930" box="[287,428,1764,1790]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="1" pageNumber="40" unit="m" value="1000.0" valueMax="1300.0" valueMin="700.0">700–1300 m</quantity>
a.s.l. Occasionally common. Flowers in October–December.
</paragraph>
</subSubSection>
<subSubSection id="C33754B9FFE52C3CFF5FF8C2FBCBFD1A" lastPageId="4" lastPageNumber="43" pageId="1" pageNumber="40" type="distribution">
<paragraph id="8B920732FFE52C39FF5FF8C2FDF0F8A8" blockId="1.[136,1451,1659,2070]" pageId="1" pageNumber="40">
<emphasis id="B959DB20FFE52C39FF5FF8C2FEA9F8EE" bold="true" box="[189,345,1798,1824]" pageId="1" pageNumber="40">Distribution:</emphasis>
— 
<collectingCountry id="F33A47A2FFE52C39FE94F8C2FE28F8EE" box="[374,472,1798,1824]" name="Vietnam" pageId="1" pageNumber="40">Vietnam</collectingCountry>
: provinces 
<collectingRegion id="49E9C9D0FFE52C39FDB9F8C3FD4FF8EF" box="[603,703,1799,1825]" country="Vietnam" name="Bac Kan" pageId="1" pageNumber="40">Bac Kan</collectingRegion>
(Cho Don District), 
<collectingRegion id="49E9C9D0FFE52C39FC50F8C3FBD3F8EF" box="[946,1059,1799,1825]" country="Vietnam" name="Cao Bang" pageId="1" pageNumber="40">Cao Bang</collectingRegion>
(Tra Linh District) and 
<collectingRegion id="49E9C9D0FFE52C39FADFF8C3FA5BF8EE" box="[1341,1451,1798,1825]" country="Vietnam" name="Ha Giang" pageId="1" pageNumber="40">Ha Giang</collectingRegion>
(districts Vi Xuyen and Quan Ba). 
<collectingCountry id="F33A47A2FFE52C39FDECF8EDFDA5F88D" box="[526,597,1833,1859]" name="China" pageId="1" pageNumber="40">China</collectingCountry>
: 
<collectingRegion id="49E9C9D0FFE52C39FD82F8EDFCEEF88D" box="[608,798,1833,1859]" country="China" name="Guangxi" pageId="1" pageNumber="40">Guangxi Zhuang</collectingRegion>
Autonomous Region (Jingxi Xian and Napo Xian) (
<bibRefCitation id="EFBC7AC3FFE52C39FA85F8EDFF16F8A8" author="Liu, Y. &amp; Kono, Y. &amp; Lin, C. - R. &amp; Xu, W. - B. &amp; Peng, C." pageId="1" pageNumber="40" pagination="367 - 373" refId="ref7475" refString="Liu, Y., Kono, Y., Lin, C. - R., Xu, W. - B. &amp; Peng, C. - I (2011) Aspidistra erecta (Asparagaceae), a new species from limestone areas in Guangxi, China. Botanical Studies 52: 367 - 373." type="journal article" year="2011">
Liu 
<emphasis id="B959DB20FFE52C39FA75F8EEFF55F8A8" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
2011
</bibRefCitation>
, 
<bibRefCitation id="EFBC7AC3FFE52C39FF12F888FE03F8A8" author="Vislobokov, N. A. &amp; Fu, L. F. &amp; Wei, Y. G. &amp; Nuraliev, M. S." box="[240,499,1868,1894]" pageId="1" pageNumber="40" pagination="65 - 86" refId="ref8519" refString="Vislobokov, N. A., Fu, L. F., Wei, Y. G. &amp; Nuraliev, M. S. (2021) Leaf epidermal micromorphology in Aspidistra (Asparagaceae): diversity and taxonomic significance. PhytoKeys 185: 65 - 86. https: // doi. org / 10.3897 / phytokeys. 185.72259" type="journal article" year="2021">
Vislobokov 
<emphasis id="B959DB20FFE52C39FE9AF889FE41F8A8" box="[376,433,1868,1894]" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
<paragraph id="8B920732FFE52C39FF5FF8ABFA52F7DB" blockId="1.[136,1451,1659,2070]" pageId="1" pageNumber="40">
<emphasis id="B959DB20FFE52C39FF5FF8ABFE46F847" bold="true" box="[189,438,1903,1929]" pageId="1" pageNumber="40">Conservation status:</emphasis>
— 
<taxonomicName id="4C2D7CB1FFE52C39FE30F8ABFD64F847" box="[466,660,1903,1929]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="40" phylum="Tracheophyta" rank="species" species="erecta">
<emphasis id="B959DB20FFE52C39FE30F8ABFD64F847" box="[466,660,1903,1929]" italics="true" pageId="1" pageNumber="40">Aspidistra erecta</emphasis>
</taxonomicName>
is presently known from several locations in 
<collectingCountry id="F33A47A2FFE52C39FB46F8ABFB17F847" box="[1188,1255,1903,1929]" name="China" pageId="1" pageNumber="40">China</collectingCountry>
(
<bibRefCitation id="EFBC7AC3FFE52C39FB1AF8ABFA57F847" author="Liu, Y. &amp; Kono, Y. &amp; Lin, C. - R. &amp; Xu, W. - B. &amp; Peng, C." box="[1272,1447,1903,1929]" pageId="1" pageNumber="40" pagination="367 - 373" refId="ref7475" refString="Liu, Y., Kono, Y., Lin, C. - R., Xu, W. - B. &amp; Peng, C. - I (2011) Aspidistra erecta (Asparagaceae), a new species from limestone areas in Guangxi, China. Botanical Studies 52: 367 - 373." type="journal article" year="2011">
Liu 
<emphasis id="B959DB20FFE52C39FAC8F8B4FA94F847" box="[1322,1380,1903,1929]" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
2011
</bibRefCitation>
, 
<bibRefCitation id="EFBC7AC3FFE52C39FF6AF856FE7BF862" author="Vislobokov, N. A. &amp; Fu, L. F. &amp; Wei, Y. G. &amp; Nuraliev, M. S." box="[136,395,1938,1964]" pageId="1" pageNumber="40" pagination="65 - 86" refId="ref8519" refString="Vislobokov, N. A., Fu, L. F., Wei, Y. G. &amp; Nuraliev, M. S. (2021) Leaf epidermal micromorphology in Aspidistra (Asparagaceae): diversity and taxonomic significance. PhytoKeys 185: 65 - 86. https: // doi. org / 10.3897 / phytokeys. 185.72259" type="journal article" year="2021">
Vislobokov 
<emphasis id="B959DB20FFE52C39FEF3F857FEBBF862" box="[273,331,1938,1964]" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
2021
</bibRefCitation>
) and four locations in 
<collectingCountry id="F33A47A2FFE52C39FD6CF856FD1DF862" box="[654,749,1938,1964]" name="Vietnam" pageId="1" pageNumber="40">Vietnam</collectingCountry>
with the total estimated extent of occurrence (EOO) less than 
<quantity id="4CD5AAD7FFE52C39FF6AF871FF08F801" box="[136,248,1973,1999]" metricMagnitude="7" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="40" unit="km" value="20000.0">20000 km</quantity>
<superScript id="7C58AA7AFFE52C39FF15F871FF0FF80D" attach="left" box="[247,255,1973,1987]" fontSize="6" pageId="1" pageNumber="40">2</superScript>
and the area of occupancy (AAO) surely less than 
<quantity id="4CD5AAD7FFE52C39FCC9F871FC7DF801" box="[811,909,1973,1999]" metricMagnitude="6" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="40" unit="km" value="2000.0">2000 km</quantity>
<superScript id="7C58AA7AFFE52C39FC6FF871FC65F80D" attach="left" box="[909,917,1973,1987]" fontSize="6" pageId="1" pageNumber="40">2</superScript>
. Taking into consideration severe fragmentation of all the known populations of this species and the continuing decline of habitat quality (iii), we preliminarily assign its conservation status as Vulnerable (VU) following the criteria B1ab(iii) + B2ab(iii) proposed by the 
<bibRefCitation id="EFBC7AC3FFE52C39FAE6F83FFA6DF7DB" author="IUCN Standards &amp; Petitions Committee" box="[1284,1437,2043,2069]" pageId="1" pageNumber="40" refId="ref7014" refString="IUCN Standards and Petitions Committee (2023) Guidelines for using the IUCN Red List categories and criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from: https: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed: 16 June 2023)." type="url" year="2023">IUCN (2023)</bibRefCitation>
.
</paragraph>
<caption id="DF5257BAFFE62C3AFF6AF9D2FF2FF99E" ID-DOI="http://doi.org/10.5281/zenodo.13213474" ID-Zenodo-Dep="13213474" httpUri="https://zenodo.org/record/13213474/files/figure.png" pageId="2" pageNumber="41" startId="2.[136,229,1558,1580]" targetBox="[358,1230,187,1531]" targetPageId="2" targetType="figure">
<paragraph id="8B920732FFE62C3AFF6AF9D2FF2FF99E" blockId="2.[136,1452,1558,1616]" pageId="2" pageNumber="41">
<emphasis id="B959DB20FFE62C3AFF6AF9D2FEF1F9E5" bold="true" box="[136,257,1558,1580]" pageId="2" pageNumber="41">FIGURE 1.</emphasis>
Distribution map of 
<taxonomicName id="4C2D7CB1FFE62C3AFE36F9D2FD8BF9E2" box="[468,635,1558,1580]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="erecta">
<emphasis id="B959DB20FFE62C3AFE36F9D2FD8BF9E2" box="[468,635,1558,1580]" italics="true" pageId="2" pageNumber="41">Aspidistra erecta</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2D7CB1FFE62C3AFD6BF9D3FD03F9E5" box="[649,755,1558,1580]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="globosa">
<emphasis id="B959DB20FFE62C3AFD6BF9D3FD03F9E5" box="[649,755,1558,1580]" italics="true" pageId="2" pageNumber="41">A. globosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2D7CB1FFE62C3AFCC5F9D3FC71F9E2" box="[807,897,1559,1580]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="quanii">
<emphasis id="B959DB20FFE62C3AFCC5F9D3FC71F9E2" box="[807,897,1559,1580]" italics="true" pageId="2" pageNumber="41">A. quanii</emphasis>
</taxonomicName>
(the map is not aimed to show countries in their entire borders).
</paragraph>
</caption>
<paragraph id="8B920732FFE62C3AFF5FF9BBFF0AF7EB" blockId="2.[136,1452,1663,2085]" pageId="2" pageNumber="41">
<emphasis id="B959DB20FFE62C3AFF5FF9BBFEF9F957" bold="true" box="[189,265,1663,1689]" pageId="2" pageNumber="41">Notes:</emphasis>
— 
<taxonomicName id="4C2D7CB1FFE62C3AFEC4F9BBFE14F957" box="[294,484,1663,1689]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="erecta">
<emphasis id="B959DB20FFE62C3AFEC4F9BBFE14F957" box="[294,484,1663,1689]" italics="true" pageId="2" pageNumber="41">Aspidistra erecta</emphasis>
</taxonomicName>
is a distinct species readily recognizable by its slender, ascending to erect stem up to 
<quantity id="4CD5AAD7FFE62C3AFA7FF9BBFF6EF973" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="41" unit="m" value="1.0">1 m</quantity>
high and rather small flowers with bent pistil. The Vietnamese specimens reported here closely meet morphologically the protologue of this species based on the Chinese populations (
<bibRefCitation id="EFBC7AC3FFE62C3AFC8BF903FBE8F92F" author="Liu, Y. &amp; Kono, Y. &amp; Lin, C. - R. &amp; Xu, W. - B. &amp; Peng, C." box="[873,1048,1735,1761]" pageId="2" pageNumber="41" pagination="367 - 373" refId="ref7475" refString="Liu, Y., Kono, Y., Lin, C. - R., Xu, W. - B. &amp; Peng, C. - I (2011) Aspidistra erecta (Asparagaceae), a new species from limestone areas in Guangxi, China. Botanical Studies 52: 367 - 373." type="journal article" year="2011">
Liu 
<emphasis id="B959DB20FFE62C3AFC78F90CFC24F92F" box="[922,980,1735,1761]" italics="true" pageId="2" pageNumber="41">et al.</emphasis>
2011
</bibRefCitation>
). One of them, 
<emphasis id="B959DB20FFE62C3AFB2BF903FF42F8CB" italics="true" pageId="2" pageNumber="41">Averyanov et al. VR 201</emphasis>
, comprises fruiting material, which makes its identification somewhat preliminary. New records of this species reported here expand its known range by more than 
<quantity id="4CD5AAD7FFE62C3AFD3EF8CBFCC3F8E7" box="[732,819,1807,1833]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="41" unit="km" value="120.0">120 km</quantity>
to the south. The presence of 
<taxonomicName id="4C2D7CB1FFE62C3AFB6FF8CBFB03F8E7" box="[1165,1267,1807,1833]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="erecta">
<emphasis id="B959DB20FFE62C3AFB6FF8CBFB03F8E7" box="[1165,1267,1807,1833]" italics="true" pageId="2" pageNumber="41">A. erecta</emphasis>
</taxonomicName>
in 
<collectingCountry id="F33A47A2FFE62C3AFAFBF8CBFA88F8E7" box="[1305,1400,1807,1833]" name="Vietnam" pageId="2" pageNumber="41">Vietnam</collectingCountry>
was quite predictable, given the proximity of its locus classicus to the China-Vietnam border and the similarity of the limestone habitats of northern 
<collectingCountry id="F33A47A2FFE62C3AFE00F893FDB1F8BF" box="[482,577,1879,1905]" name="Vietnam" pageId="2" pageNumber="41">Vietnam</collectingCountry>
to those in Guangxi. Indeed, all four Vietnamese populations are confined to karst formations, just like the type and the 
<typeStatus id="5496B990FFE62C3AFDBDF8BFFD4FF85B" box="[607,703,1915,1941]" pageId="2" pageNumber="41" type="paratype">paratype</typeStatus>
gatherings. Flowers of 
<taxonomicName id="4C2D7CB1FFE62C3AFC27F8BFFBD9F85B" box="[965,1065,1915,1941]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="erecta">
<emphasis id="B959DB20FFE62C3AFC27F8BFFBD9F85B" box="[965,1065,1915,1941]" italics="true" pageId="2" pageNumber="41">A. erecta</emphasis>
</taxonomicName>
are surprisingly similar to those of 
<taxonomicName id="4C2D7CB1FFE62C3AFF6AF85BFDECF877" authority="Tillich (2005: 321)" authorityName="Tillich" authorityPageNumber="321" authorityYear="2005" box="[136,540,1951,1978]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="marasmioides">
<emphasis id="B959DB20FFE62C3AFF6AF85BFEB3F877" box="[136,323,1951,1977]" italics="true" pageId="2" pageNumber="41">A. marasmioides</emphasis>
<bibRefCitation id="EFBC7AC3FFE62C3AFEAAF85BFDECF877" author="Tillich, H. - J." box="[328,540,1951,1978]" pageId="2" pageNumber="41" pagination="313 - 338" refId="ref7954" refString="Tillich, H. - J. (2005) A key for Aspidistra (Ruscaceae), including fifteen new species from Vietnam. Feddes Repertorium 116 (5 - 6): 313 - 338. https: // doi. org / 10.1002 / fedr. 200511076" type="journal article" year="2005">Tillich (2005: 321)</bibRefCitation>
</taxonomicName>
. 
<bibRefCitation id="EFBC7AC3FFE62C3AFDC6F85BFCC6F877" author="Averyanov, L. V. &amp; Nguyen, K. S. &amp; Tran, T. H. &amp; Averyanova, A. L. &amp; Maisak, T. V. &amp; Nguyen, H. T." box="[548,822,1951,1977]" pageId="2" pageNumber="41" refId="ref6380" refString="Averyanov, L. V., Nguyen, K. S., Tran, T. H., Averyanova, A. L., Maisak, T. V. &amp; Nguyen, H. T. (2020 c) Plant diversity, flora and vegetation of Bat Dai Son Mountain area, northern Vietnam. Strata, Saint-Petersburg, 561 pp." type="book" year="2020">
Averyanov 
<emphasis id="B959DB20FFE62C3AFD41F864FD2AF877" box="[675,730,1951,1977]" italics="true" pageId="2" pageNumber="41">et al.</emphasis>
(2020c)
</bibRefCitation>
tentatively synonymized 
<taxonomicName id="4C2D7CB1FFE62C3AFBB0F85BFB44F877" box="[1106,1204,1951,1977]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="erecta">
<emphasis id="B959DB20FFE62C3AFBB0F85BFB44F877" box="[1106,1204,1951,1977]" italics="true" pageId="2" pageNumber="41">A. erecta</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C2D7CB1FFE62C3AFB13F85BFA5BF877" authorityName="Tillich" authorityYear="2005" box="[1265,1451,1951,1977]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="marasmioides">
<emphasis id="B959DB20FFE62C3AFB13F85BFA5BF877" box="[1265,1451,1951,1977]" italics="true" pageId="2" pageNumber="41">A. marasmioides</emphasis>
</taxonomicName>
while reporting the fruiting specimen 
<emphasis id="B959DB20FFE62C3AFDCEF807FCC8F813" box="[556,824,1987,2013]" italics="true" pageId="2" pageNumber="41">Averyanov et al. VR 201</emphasis>
. However, these two species possess strikingly different growth form: 
<taxonomicName id="4C2D7CB1FFE62C3AFEC0F823FE2EF7CF" authorityName="Tillich" authorityYear="2005" box="[290,478,2023,2049]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="41" phylum="Tracheophyta" rank="species" species="marasmioides">
<emphasis id="B959DB20FFE62C3AFEC0F823FE2EF7CF" box="[290,478,2023,2049]" italics="true" pageId="2" pageNumber="41">A. marasmioides</emphasis>
</taxonomicName>
is characterized by exclusively plagiotropic shoots, which is typical for most species of the genus.
</paragraph>
<caption id="DF5257BAFFE72C3BFF6AF88EFC4FF7DA" ID-DOI="http://doi.org/10.5281/zenodo.13213476" ID-Zenodo-Dep="13213476" httpUri="https://zenodo.org/record/13213476/files/figure.png" pageId="3" pageNumber="42" startId="3.[136,229,1866,1888]" targetBox="[243,1344,190,1841]" targetPageId="3" targetType="figure">
<paragraph id="8B920732FFE72C3BFF6AF88EFC4FF7DA" blockId="3.[136,1452,1866,2068]" pageId="3" pageNumber="42">
<emphasis id="B959DB20FFE72C3BFF6AF88EFF0EF891" bold="true" box="[136,254,1866,1888]" pageId="3" pageNumber="42">FIGURE 2.</emphasis>
<taxonomicName id="4C2D7CB1FFE72C3BFEE6F88EFE58F8AE" box="[260,424,1866,1888]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="42" phylum="Tracheophyta" rank="species" species="erecta">
<emphasis id="B959DB20FFE72C3BFEE6F88EFE58F8AE" box="[260,424,1866,1888]" italics="true" pageId="3" pageNumber="42">Aspidistra erecta</emphasis>
</taxonomicName>
. 
<emphasis id="B959DB20FFE72C3BFE57F88EFE3CF891" bold="true" box="[437,460,1866,1887]" pageId="3" pageNumber="42">A.</emphasis>
Flowering plant. 
<emphasis id="B959DB20FFE72C3BFD9AF88EFD7EF891" bold="true" box="[632,654,1866,1887]" pageId="3" pageNumber="42">B.</emphasis>
Apical portion of stem. 
<emphasis id="B959DB20FFE72C3BFC9BF88EFC60F8AE" bold="true" box="[889,912,1866,1888]" pageId="3" pageNumber="42">C.</emphasis>
Leaves, adaxial and abaxial side. 
<emphasis id="B959DB20FFE72C3BFB38F88EFAFEF891" bold="true" box="[1242,1294,1866,1887]" pageId="3" pageNumber="42">D, E.</emphasis>
Flower bud and open flower, side view. 
<emphasis id="B959DB20FFE72C3BFE85F8AAFE86F84D" bold="true" box="[359,374,1902,1923]" pageId="3" pageNumber="42">F</emphasis>
. Flower on shoot, front view. 
<emphasis id="B959DB20FFE72C3BFD73F8AAFD5AF84A" bold="true" box="[657,682,1902,1924]" pageId="3" pageNumber="42">G.</emphasis>
Flower, side and half-side views. 
<emphasis id="B959DB20FFE72C3BFC09F8AAFBF4F84D" bold="true" box="[1003,1028,1902,1923]" pageId="3" pageNumber="42">H.</emphasis>
Inflorescence bracts, abaxial side. 
<emphasis id="B959DB20FFE72C3BFAACF8AAFAADF84D" bold="true" box="[1358,1373,1902,1923]" pageId="3" pageNumber="42">I.</emphasis>
Flower, front view (perianth lobes removed), showing stamens and stigma. 
<emphasis id="B959DB20FFE72C3BFCF6F856FCD6F869" bold="true" box="[788,806,1938,1959]" pageId="3" pageNumber="42">J.</emphasis>
Sagittal section of flower, front and side views. 
<emphasis id="B959DB20FFE72C3BFB1FF856FAE6F869" bold="true" box="[1277,1302,1938,1959]" pageId="3" pageNumber="42">K.</emphasis>
Perianth lobes, adaxial and abaxial surface. 
<emphasis id="B959DB20FFE72C3BFE7BF872FE5FF805" bold="true" box="[409,431,1974,1995]" pageId="3" pageNumber="42">L.</emphasis>
Peduncle and sagittally dissected flower (with perianth lobes removed), showing intact pistil. 
<emphasis id="B959DB20FFE72C3BFAD9F872FAA7F805" bold="true" box="[1339,1367,1974,1995]" pageId="3" pageNumber="42">M.</emphasis>
Stamen, front and side view. 
<emphasis id="B959DB20FFE72C3BFEAFF81EFE95F821" bold="true" box="[333,357,2010,2031]" pageId="3" pageNumber="42">N.</emphasis>
Pistil, side view. 
<emphasis id="B959DB20FFE72C3BFDEDF81EFDD8F83E" bold="true" box="[527,552,2010,2032]" pageId="3" pageNumber="42">O.</emphasis>
Pistil, view from above. 
<emphasis id="B959DB20FFE72C3BFCFEF81EFCDFF821" bold="true" box="[796,815,2010,2031]" pageId="3" pageNumber="42">P.</emphasis>
Sagittal section of pistil. 
<emphasis id="B959DB20FFE72C3BFBC4F81FFF5AF7DA" italics="true" pageId="3" pageNumber="42">Bui Van Huong, Truong Ba Vuong, BVH s.n.</emphasis>
Photos by Bui Van Huong, correction and design by L. Averyanov and T. Maisak.
</paragraph>
</caption>
<paragraph id="8B920732FFE02C3CFF5FFF5AFBCBFD1A" blockId="4.[136,1452,158,725]" pageId="4" pageNumber="43">
<emphasis id="B959DB20FFE02C3CFF5FFF5AFE2AFF77" bold="true" box="[189,474,158,185]" pageId="4" pageNumber="43">Specimens examined:—</emphasis>
<materialsCitation id="3B450D6FFFE02C3CFE38FF5AFC6CFECE" collectingDate="2023-11-29" collectionCode="VNM, VNMN, LE" collectorName="Bui Van Huong &amp; Truong Ba Vuong" country="Vietnam" location="Ban Thi Commune" municipality="Cho Don District" pageId="4" pageNumber="43" specimenCode="LE01124750" specimenCount="1" stateProvince="Bac Kan Province">
<emphasis id="B959DB20FFE02C3CFE38FF5AFD25FF76" bold="true" box="[474,725,158,184]" pageId="4" pageNumber="43">
<collectingCountry id="F33A47A2FFE02C3CFE38FF5AFD93FF76" box="[474,611,158,184]" name="Vietnam" pageId="4" pageNumber="43">VIETNAM</collectingCountry>
. 
<collectingRegion id="49E9C9D0FFE02C3CFD8DFF5AFCB3FF76" box="[623,835,158,184]" country="Vietnam" name="Bac Kan" pageId="4" pageNumber="43">Bac Kan Province</collectingRegion>
</emphasis>
: 
<collectingMunicipality id="6BF69D48FFE02C3CFCAFFF5AFBFFFF76" box="[845,1039,158,185]" pageId="4" pageNumber="43">Cho Don District</collectingMunicipality>
, 
<location id="8EF251E9FFE02C3CFBFBFF5BFB01FF76" LSID="urn:lsid:plazi:treatment:0384B624FFE52C3CFF6AF9D5FBCBFD1A:8EF251E9FFE02C3CFBFBFF5BFB01FF76" box="[1049,1265,158,185]" country="Vietnam" municipality="Cho Don District" name="Ban Thi Commune" pageId="4" pageNumber="43" stateProvince="Bac Kan Province">Ban Thi Commune</location>
, terrestrial plant, in secondary forest on karst rocks, 
<date id="FF9321F2FFE02C3CFDF0FF06FD1FFF12" box="[530,751,194,220]" pageId="4" pageNumber="43" value="2023-11-29">
<collectingDate id="EFD7D81AFFE02C3CFDF0FF06FD1FFF12" box="[530,751,194,220]" pageId="4" pageNumber="43" value="2023-11-29">29 November 2023</collectingDate>
</date>
, 
<emphasis id="B959DB20FFE02C3CFD18FF07FB1EFF12" box="[762,1262,195,220]" italics="true" pageId="4" pageNumber="43">
<collectorName id="26D862E4FFE02C3CFD18FF07FC59FF12" box="[762,937,195,220]" pageId="4" pageNumber="43">Bui Van Huong</collectorName>
, 
<collectorName id="26D862E4FFE02C3CFC57FF07FB8EFF12" box="[949,1150,195,220]" pageId="4" pageNumber="43">Truong Ba Vuong</collectorName>
, BVH s.n.
</emphasis>
(
<collectionCode id="ED3C9FF7FFE02C3CFB1CFF06FAB4FF12" box="[1278,1348,194,220]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="4" pageNumber="43" type="Herbarium">VNM</collectionCode>
, 
<collectionCode id="ED3C9FF7FFE02C3CFAADFF07FA57FF13" box="[1359,1447,195,221]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="4" pageNumber="43">VNMN</collectionCode>
, photos 
<collectionCode id="ED3C9FF7FFE02C3CFF38FF23FF0CFECF" box="[218,252,231,257]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="43">LE</collectionCode>
<specimenCode id="DB8BAF49FFE02C3CFEE0FF22FE63FECF" box="[258,403,230,257]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="43">LE01124750</specimenCode>
https://en.herbariumle.ru/?t=occ&amp;id=208858)
</materialsCitation>
; 
<materialsCitation id="3B450D6FFFE02C3CFC4BFF22FC11FE5E" collectingDate="2018-11-29" collectionCode="LE" collectorName="Bleddyn Wynn-Jones" country="Vietnam" elevation="781" latitude="22.85" location="Quang Vinh Municipality" longLatPrecision="1258" longitude="106.23333" municipality="Tra Linh District" pageId="4" pageNumber="43" specimenCode="LE01073384" specimenCount="1" stateProvince="Cao Bang Province">
<collectingRegion id="49E9C9D0FFE02C3CFC4BFF22FB7DFECE" box="[937,1165,230,256]" country="Vietnam" name="Cao Bang" pageId="4" pageNumber="43">
<emphasis id="B959DB20FFE02C3CFC4BFF22FBEEFECE" bold="true" box="[937,1054,230,256]" pageId="4" pageNumber="43">Cao Bang</emphasis>
Province
</collectingRegion>
: 
<collectingMunicipality id="6BF69D48FFE02C3CFB75FF23FAA6FECE" box="[1175,1366,230,257]" pageId="4" pageNumber="43">Tra Linh District</collectingMunicipality>
, 
<location id="8EF251E9FFE02C3CFA83FF22FEA7FEEA" LSID="urn:lsid:plazi:treatment:0384B624FFE52C3CFF6AF9D5FBCBFD1A:8EF251E9FFE02C3CFA83FF22FEA7FEEA" country="Vietnam" latitude="22.85" longLatPrecision="1258" longitude="106.23333" municipality="Tra Linh District" name="Quang Vinh Municipality" pageId="4" pageNumber="43" stateProvince="Cao Bang Province">Quang Vinh Municipality</location>
, about 
<locationDeviation id="9A201E10FFE02C3CFE45FECEFD6EFEEB" box="[423,670,266,293]" location="Cao Bang" pageId="4" pageNumber="43">
<quantity id="4CD5AAD7FFE02C3CFE45FECEFE1EFEEA" box="[423,494,266,292]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="43" unit="km" value="21.0">21 km</quantity>
N of Cao Bang
</locationDeviation>
town, 
<geoCoordinate id="EE1961F5FFE02C3CFD09FECEFCB7FEEB" box="[747,839,266,293]" degrees="22" direction="north" minutes="51" orientation="latitude" pageId="4" pageNumber="43" precision="925" value="22.85">22°51'N</geoCoordinate>
<geoCoordinate id="EE1961F5FFE02C3CFCACFECEFC48FEEA" box="[846,952,266,292]" degrees="106" direction="east" minutes="14" orientation="longitude" pageId="4" pageNumber="43" precision="925" value="106.23333">106°14'E</geoCoordinate>
, 
<elevation id="0000E001FFE02C3CFC26FECEFBB7FEEA" box="[964,1095,266,293]" metricMagnitude="2" metricUnit="m" metricValue="7.81" pageId="4" pageNumber="43" unit="m" value="781.0">
<quantity id="4CD5AAD7FFE02C3CFC26FECEFBFBFEEB" box="[964,1035,266,293]" metricMagnitude="2" metricUnit="m" metricValue="7.81" pageId="4" pageNumber="43" unit="m" value="781.0">781 m</quantity>
a.s.l.
</elevation>
, growing in limestone rubble at the edge of steep evergreen primary forest, a species with erect slender bamboo-like rhizomes, leaves of this collection with small scattered white spots, 
<date id="FF9321F2FFE02C3CFE16FE96FD3CFEA2" box="[500,716,338,364]" pageId="4" pageNumber="43" value="2018-11-29">
<collectingDate id="EFD7D81AFFE02C3CFE16FE96FD3CFEA2" box="[500,716,338,364]" pageId="4" pageNumber="43" value="2018-11-29">29 November 2018</collectingDate>
</date>
, 
<emphasis id="B959DB20FFE02C3CFD37FE96FBB9FEA2" box="[725,1097,338,364]" italics="true" pageId="4" pageNumber="43">
<collectorName id="26D862E4FFE02C3CFD37FE96FC30FEA2" box="[725,960,338,364]" pageId="4" pageNumber="43">Bleddyn Wynn-Jones</collectorName>
, BWJ 16147
</emphasis>
(private herbarium of B. Wynn-Jones, photos 
<collectionCode id="ED3C9FF7FFE02C3CFECAFEB3FEBAFE5F" box="[296,330,375,401]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="43">LE</collectionCode>
<specimenCode id="DB8BAF49FFE02C3CFEB1FEB2FE15FE5E" box="[339,485,374,400]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="43">LE01073384</specimenCode>
https://en.herbariumle.ru/?t=occ&amp;id=16551)
</materialsCitation>
; living plant from the same gathering in Crûg Farm Nursery, UK; 
<materialsCitation id="3B450D6FFFE02C3CFE53FE5EFA87FDA6" collectingDate="2018-04-15" collectionCode="LE" collectorName="L. Averyanov &amp; Nguyen Sinh Khang &amp; Chuong Quang Ngan &amp; T. Maisak" country="Vietnam" elevation="1252" latitude="23.100723" location="Lung Vay Village" longLatPrecision="1" longitude="105.051254" municipality="Quan Ba District" pageId="4" pageNumber="43" specimenCode="LE01054358" specimenCount="1" stateProvince="Ha Giang Province">
<collectingRegion id="49E9C9D0FFE02C3CFE53FE5EFD6AFE7A" box="[433,666,410,436]" country="Vietnam" name="Ha Giang" pageId="4" pageNumber="43">
<emphasis id="B959DB20FFE02C3CFE53FE5EFDD8FE7A" bold="true" box="[433,552,410,436]" pageId="4" pageNumber="43">Ha Giang</emphasis>
Province
</collectingRegion>
: 
<collectingMunicipality id="6BF69D48FFE02C3CFD44FE5EFC9DFE7A" box="[678,877,410,437]" pageId="4" pageNumber="43">Quan Ba District</collectingMunicipality>
, Bat Dai Son Nature Reserve, Can Ty Commune, 
<location id="8EF251E9FFE02C3CFF6AFE7BFEA4FE16" LSID="urn:lsid:plazi:treatment:0384B624FFE52C3CFF6AF9D5FBCBFD1A:8EF251E9FFE02C3CFF6AFE7BFEA4FE16" box="[136,340,446,473]" country="Vietnam" latitude="23.100723" longLatPrecision="1" longitude="105.051254" municipality="Quan Ba District" name="Lung Vay Village" pageId="4" pageNumber="43" stateProvince="Ha Giang Province">Lung Vay Village</location>
, around point 
<geoCoordinate id="EE1961F5FFE02C3CFE18FE7AFD62FE16" box="[506,658,446,472]" degrees="23" direction="north" minutes="06" orientation="latitude" pageId="4" pageNumber="43" precision="1" seconds="02.6" value="23.100723">23°06'02.6''N</geoCoordinate>
<geoCoordinate id="EE1961F5FFE02C3CFD78FE7AFCCFFE17" box="[666,831,446,473]" degrees="105" direction="east" minutes="03" orientation="longitude" pageId="4" pageNumber="43" precision="1" seconds="04.5" value="105.051254">105°03'04.5''E</geoCoordinate>
, 
<elevation id="0000E001FFE02C3CFCAFFE7AFBD8FE16" box="[845,1064,446,473]" metricMagnitude="3" metricUnit="m" metricValue="1.2525" metricValueMax="1.305" metricValueMin="1.2" pageId="4" pageNumber="43" unit="m" value="1252.5" valueMax="1305.0" valueMin="1200.0">
<quantity id="4CD5AAD7FFE02C3CFCAFFE7AFC1AFE17" box="[845,1002,446,473]" metricMagnitude="3" metricUnit="m" metricValue="1.2525" metricValueMax="1.305" metricValueMin="1.2" pageId="4" pageNumber="43" unit="m" value="1252.5" valueMax="1305.0" valueMin="1200.0">1200–1305 m</quantity>
a.s.l.
</elevation>
, karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops, erect terrestrial herb with semi-woody stem to 
<quantity id="4CD5AAD7FFE02C3CFCADFDC2FC65FDEF" box="[847,917,518,545]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="43" unit="cm" value="40.0">40 cm</quantity>
tall among shady rocks, leaves uniform green, fruits brown, globular prickly-papillose, 
<quantity id="4CD5AAD7FFE02C3CFDACFDEEFD5BFD8B" box="[590,683,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="43" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
in diameter, not common, 
<date id="FF9321F2FFE02C3CFC34FDEEFB84FD8A" box="[982,1140,554,581]" pageId="4" pageNumber="43" value="2018-04-15">
<collectingDate id="EFD7D81AFFE02C3CFC34FDEEFB84FD8A" box="[982,1140,554,581]" pageId="4" pageNumber="43" value="2018-04-15">15 April 2018</collectingDate>
</date>
, 
<emphasis id="B959DB20FFE02C3CFB9DFDEFFD4AFDA6" italics="true" pageId="4" pageNumber="43">
<collectorName id="26D862E4FFE02C3CFB9DFDEFFAE3FD8B" box="[1151,1299,555,581]" pageId="4" pageNumber="43">L. Averyanov</collectorName>
, 
<collectorName id="26D862E4FFE02C3CFAFFFDEFFF25FDA6" pageId="4" pageNumber="43">Nguyen Sinh Khang</collectorName>
, 
<collectorName id="26D862E4FFE02C3CFF01FD8AFE28FDA6" box="[227,472,590,617]" pageId="4" pageNumber="43">Chuong Quang Ngan</collectorName>
, 
<collectorName id="26D862E4FFE02C3CFE04FD8BFDA8FDA6" box="[486,600,590,616]" pageId="4" pageNumber="43">T. Maisak</collectorName>
, VR 201
</emphasis>
(
<collectionCode id="ED3C9FF7FFE02C3CFD29FD8AFD1FFDA6" box="[715,751,590,616]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="43">LE</collectionCode>
<specimenCode id="DB8BAF49FFE02C3CFD1AFD8AFC7AFDA7" box="[760,906,590,617]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="43">LE01054358</specimenCode>
https://en.herbariumle.ru/?t=occ&amp;id=4690)
</materialsCitation>
; 
<materialsCitation id="3B450D6FFFE02C3CFA6AFD8AFBC7FD1A" collectingDate="2016-11-30" collectionCode="HNU, SGN, LE" collectorName="Luu Hong Truong &amp; Nguyen Anh Duc &amp; Le Khac Quyet" country="Vietnam" county="Vi Xuyen District" elevation="770" latitude="22.847765" location="Dong Van Karst Plateau - Du Gia National Park" longLatPrecision="1" longitude="105.120255" municipality="Du Gia National Park" pageId="4" pageNumber="43" specimenCode="LE01124770" specimenCount="1" stateProvince="Ha Giang Province">
<collectingRegion id="49E9C9D0FFE02C3CFA6AFD8AFEB0FD42" country="Vietnam" name="Ha Giang" pageId="4" pageNumber="43">
<emphasis id="B959DB20FFE02C3CFA6AFD8AFF21FD42" bold="true" pageId="4" pageNumber="43">Ha Giang</emphasis>
Province
</collectingRegion>
: 
<collectingCounty id="62F37FBEFFE02C3CFEA8FDB6FDE5FD42" box="[330,533,626,653]" pageId="4" pageNumber="43">Vi Xuyen District</collectingCounty>
, 
<location id="8EF251E9FFE02C3CFDC2FDB7FBB1FD42" LSID="urn:lsid:plazi:treatment:0384B624FFE52C3CFF6AF9D5FBCBFD1A:8EF251E9FFE02C3CFDC2FDB7FBB1FD42" box="[544,1089,626,653]" country="Vietnam" county="Vi Xuyen District" latitude="22.847765" longLatPrecision="1" longitude="105.120255" municipality="Du Gia National Park" name="Dong Van Karst Plateau - Du Gia National Park" pageId="4" pageNumber="43" stateProvince="Ha Giang Province">Dong Van Karst Plateau—Du Gia National Park</location>
, 
<location id="8EF251E9FFE02C3CFBAEFDB6FB09FD42" LSID="urn:lsid:plazi:treatment:0384B624FFE52C3CFF6AF9D5FBCBFD1A:8EF251E9FFE02C3CFBAEFDB6FB09FD42" box="[1100,1273,626,652]" country="Vietnam" county="Vi Xuyen District" latitude="22.847765" longLatPrecision="1" longitude="105.120255" municipality="Du Gia National Park" name="Khau Ca forest" pageId="4" pageNumber="43" stateProvince="Ha Giang Province">Khau Ca forest</location>
, 
<geoCoordinate id="EE1961F5FFE02C3CFAE6FDB6FA5CFD43" box="[1284,1452,626,653]" degrees="22" direction="north" minutes="50" orientation="latitude" pageId="4" pageNumber="43" precision="1" seconds="51.95" value="22.847765">22°50'51.95&quot;N</geoCoordinate>
<geoCoordinate id="EE1961F5FFE02C3CFF6AFD52FEC3FD7E" box="[136,307,662,689]" degrees="105" direction="east" minutes="7" orientation="longitude" pageId="4" pageNumber="43" precision="1" seconds="12.91" value="105.120255">105° 7'12.91&quot;E</geoCoordinate>
, 
<elevation id="0000E001FFE02C3CFEDCFD52FDDBFD7E" box="[318,555,662,689]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="4" pageNumber="43" unit="m" value="770.0">
elevation 
<quantity id="4CD5AAD7FFE02C3CFE49FD52FE00FD7F" box="[427,496,662,689]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="4" pageNumber="43" unit="m" value="770.0">770 m</quantity>
a.s.l.
</elevation>
, 
<date id="FF9321F2FFE02C3CFDD5FD52FCFFFD7E" box="[567,783,662,688]" pageId="4" pageNumber="43" value="2016-11-30">
<collectingDate id="EFD7D81AFFE02C3CFDD5FD52FCFFFD7E" box="[567,783,662,688]" pageId="4" pageNumber="43" value="2016-11-30">30 November 2016</collectingDate>
</date>
, 
<emphasis id="B959DB20FFE02C3CFCFBFD53FA5BFD7E" box="[793,1451,662,689]" italics="true" pageId="4" pageNumber="43">
<collectorName id="26D862E4FFE02C3CFCFBFD53FC12FD7E" box="[793,994,663,688]" pageId="4" pageNumber="43">Luu Hong Truong</collectorName>
, 
<collectorName id="26D862E4FFE02C3CFC0EFD53FB5AFD7E" box="[1004,1194,662,688]" pageId="4" pageNumber="43">Nguyen Anh Duc</collectorName>
, 
<collectorName id="26D862E4FFE02C3CFB57FD53FAADFD7F" box="[1205,1373,662,689]" pageId="4" pageNumber="43">Le Khac Quyet</collectorName>
, HG31
</emphasis>
(
<collectionCode id="ED3C9FF7FFE02C3CFF72FD7EFF21FD1A" box="[144,209,698,724]" collectionName="HNU" country="China" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="4" pageNumber="43">HNU</collectionCode>
, 
<collectionCode id="ED3C9FF7FFE02C3CFF3EFD7EFEE7FD1A" box="[220,279,698,724]" country="0" httpUri="http://grbio.org/cool/fyzf-afsy" name="Southern Institute of Ecology" pageId="4" pageNumber="43" type="Herbarium">SGN</collectionCode>
, photos 
<collectionCode id="ED3C9FF7FFE02C3CFE96FD7FFE66FD1B" box="[372,406,699,725]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="43">LE</collectionCode>
<specimenCode id="DB8BAF49FFE02C3CFE7FFD7EFDDEFD1A" box="[413,558,698,724]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="43">LE01124770</specimenCode>
https://en.herbariumle.ru/?t=occ&amp;id=211192)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>