163 lines
18 KiB
XML
163 lines
18 KiB
XML
<document id="A1D5A792E41CA3DAF2870DFD00B82F57" ID-CLB-Dataset="6085" ID-DOI="10.11646/phytotaxa.31.1.1" ID-GBIF-Dataset="2185a8cf-c15c-448b-b8c0-6055976408ed" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894648" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1622649568931" checkinUser="felipe" docAuthor="Gonzales, Jasivia & Kessler, Michael" docDate="2011" docId="0380F37CFFEE7F0799A6E718D25DF915" docLanguage="en" docName="Phytotaxa.31.1-54.pdf.imf" docOrigin="Phytotaxa 31" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Sticherus rubiginosus Nakai 1950" docType="treatment" docVersion="7" lastPageNumber="44" masterDocId="FFB98B04FFC57F2C9931E37FD712FFAA" masterDocTitle="A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="44" updateTime="1698998516251" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2FC7DDBAEFA6F1A62A7F1233332FE783">A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species</mods:title>
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</mods:titleInfo>
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<mods:name id="408914343C79D5ACF4823A287573DADA" type="personal">
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<mods:roleTerm id="B05957FAD70D1D488F4774B64A2946A4">Author</mods:roleTerm>
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<mods:namePart id="9CCDD9481E25FBA1D4BF667966EC3A62">Gonzales, Jasivia</mods:namePart>
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<mods:affiliation id="1E61DFFB5F498B4E6288DFA10FF1B4FE">Herbario Nacional de Bolivia, Casilla 10077, La Paz, Bolivia. Systematic Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland. E-mail: michael. kessler @ systbot. uzh. ch</mods:affiliation>
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<mods:namePart id="018A33136887893F2AAFDFB41EDADC7E">Kessler, Michael</mods:namePart>
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<mods:title id="D5B9927B89416DCFFF67175ABE52F7EF">Phytotaxa</mods:title>
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<mods:part id="05641C70C0FEBF92B3BD1F88A3EDAC2A">
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<mods:date id="5BA262C59E0C3FC7E14B9703060EC3DC">2011</mods:date>
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<mods:number id="B5B0EBCA552C17B12FAF251D25343368">2011-12-31</mods:number>
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<mods:number id="E569A29C4F4DBBEF03C2AAE8E6A90EDB">31</mods:number>
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<mods:classification id="6EB62E3189BFABA3FF02128C168C43D1">journal article</mods:classification>
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<treatment id="0380F37CFFEE7F0799A6E718D25DF915" ID-DOI="http://doi.org/10.5281/zenodo.4894792" ID-GBIF-Taxon="182742328" ID-Zenodo-Dep="4894792" LSID="urn:lsid:plazi:treatment:0380F37CFFEE7F0799A6E718D25DF915" httpUri="http://treatment.plazi.org/id/0380F37CFFEE7F0799A6E718D25DF915" lastPageNumber="44" pageId="43" pageNumber="44">
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<subSubSection id="C33311E1FFEE7F0799A6E718D5DAFB28" box="[151,712,1127,1154]" pageId="43" pageNumber="44" type="nomenclature">
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<treatmentCitation id="0A88647BFFEE7F0799A6E718D5DAFB28" author="Nakai, T." box="[151,712,1127,1154]" page=")" pageId="43" pageNumber="44" year="1950">
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<taxonomicName id="4C2939E9FFEE7F0799A6E718D5DAFB28" ID-CoL="52HXN" authority="(Mett.) Nakai (1950: 28)" authorityName="Nakai" authorityPageNumber="28" authorityYear="1950" baseAuthorityName="Mett." box="[151,712,1127,1154]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="rubiginosus">
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<emphasis id="B95D9E78FFEE7F0799A6E718D68CFB2B" bold="true" box="[151,414,1127,1153]" italics="true" pageId="43" pageNumber="44">Sticherus rubiginosus</emphasis>
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(Mett.)
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<bibRefCitation id="EFB83F9BFFEE7F0798CEE717D5DAFB28" author="Nakai, T." box="[511,712,1128,1154]" pageId="43" pageNumber="44" pagination="1 - 71" refId="ref32966" refString="Nakai, T. (1950) A new classification of Gleicheniales. Bulletin of the National Science Museum 29: 1 - 71." type="journal article" year="1950">Nakai (1950: 28)</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C33311E1FFEE7F0799A6E7CAD3D7FB66" box="[151,1221,1205,1228]" pageId="43" pageNumber="44" type="reference_group">
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<paragraph id="8B96426AFFEE7F0799A6E7CAD3D7FB66" blockId="43.[151,1436,1205,1727]" box="[151,1221,1205,1228]" pageId="43" pageNumber="44">
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<treatmentCitationGroup id="AB396544FFEE7F0799A6E7CAD3D7FB66" box="[151,1221,1205,1228]" pageId="43" pageNumber="44">
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<treatmentCitation id="0A88647BFFEE7F0799A6E7CAD56CFB66" author="Mettenius, G." box="[151,638,1205,1228]" page=")" pageId="43" pageNumber="44" year="1864">
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<taxonomicName id="4C2939E9FFEE7F0799A6E7CAD56CFB66" ID-CoL="6KH3H" authority="Mettenius (1864: 267)" authorityName="Mettenius" authorityPageNumber="267" authorityYear="1864" box="[151,638,1205,1228]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="rubiginosa">
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<emphasis id="B95D9E78FFEE7F0799A6E7CAD694FB66" box="[151,390,1205,1228]" italics="true" pageId="43" pageNumber="44">Gleichenia rubiginosa</emphasis>
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<bibRefCitation id="EFB83F9BFFEE7F0798BDE7CAD56CFB66" author="Mettenius, G." box="[396,638,1205,1228]" pageId="43" pageNumber="44" pagination="193 - 271" refId="ref32777" refString="Mettenius, G. (1864) Prodromus florae novo-granatensis ou Enumeration des plantes de la Nouvelle-Grenade, Filices. Annales des Sciences Naturelles; Botanique ser. 5, 2: 193 - 271." type="journal article" year="1864">Mettenius (1864: 267)</bibRefCitation>
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</taxonomicName>
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.
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<treatmentCitation id="0A88647BFFEE7F079BB9E7CAD3D3FB66" author="Maxon, W. R." box="[648,1217,1205,1228]" page=")" pageId="43" pageNumber="44" year="1922">
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<taxonomicName id="4C2939E9FFEE7F079BB9E7CAD3D3FB66" ID-CoL="35SX3" authority="(Mett.) Maxon (1922 a: 50)" authorityName="Maxon" authorityPageNumber="50" authorityYear="1922" baseAuthorityName="Mett." box="[648,1217,1205,1228]" class="Polypodiopsida" family="Gleicheniaceae" genus="Dicranopteris" kingdom="Plantae" order="Gleicheniales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="rubiginosa">
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<emphasis id="B95D9E78FFEE7F079BB9E7CAD485FB66" box="[648,919,1205,1228]" italics="true" pageId="43" pageNumber="44">Dicranopteris rubiginosa</emphasis>
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(Mett.)
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<bibRefCitation id="EFB83F9BFFEE7F079AC1E7CAD3D3FB66" author="Maxon, W. R." box="[1008,1217,1205,1228]" pageId="43" pageNumber="44" pagination="33 - 63" refId="ref32535" refString="Maxon, W. R. (1922 a) Studies of tropical American fern - No. 7. Contributions from the United States National Herbarium 24: 33 - 63, i - viii, pl. 11 - 20." type="journal article" year="1922">Maxon (1922a: 50)</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<subSubSection id="C33311E1FFEE7F0799F7E7AAD3A3FB45" box="[198,1201,1237,1263]" pageId="43" pageNumber="44" type="materials_examined">
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<paragraph id="8B96426AFFEE7F0799F7E7AAD3A3FB45" blockId="43.[151,1436,1205,1727]" box="[198,1201,1237,1263]" pageId="43" pageNumber="44">
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<emphasis id="B95D9E78FFEE7F0799F7E7AAD639FB45" bold="true" box="[198,299,1237,1263]" pageId="43" pageNumber="44">Type:—</emphasis>
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<materialsCitation id="3B414837FFEE7F07981AE7AAD361FB45" ID-GBIF-Occurrence="3131398335" box="[299,1139,1237,1263]" collectionCode="B" country="Colombia" elevation="1650" location="Puente Nacional" municipality="Lindig" pageId="43" pageNumber="44" specimenCount="1" typeStatus="holotype">
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<collectingCountry id="F33E02FAFFEE7F07981AE7AAD6D9FB45" box="[299,459,1237,1263]" name="Colombia" pageId="43" pageNumber="44">COLOMBIA</collectingCountry>
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.
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<location id="8EF614B1FFEE7F0798E4E7AAD58BFB45" LSID="urn:lsid:plazi:treatment:0380F37CFFEE7F0799A6E718D25DF915:8EF614B1FFEE7F0798E4E7AAD58BFB45" box="[469,665,1237,1263]" country="Colombia" municipality="Lindig" name="Puente Nacional" pageId="43" pageNumber="44">Puente Nacional</location>
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,
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</quantity>
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,
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<emphasis id="B95D9E78FFEE7F079A3DE7AAD46FFB44" box="[780,893,1237,1262]" italics="true" pageId="43" pageNumber="44">
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<collectingMunicipality id="6BF2D810FFEE7F079A3DE7AAD44BFB44" box="[780,857,1237,1262]" pageId="43" pageNumber="44">Lindig</collectingMunicipality>
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71
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</emphasis>
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(
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<collectionCode id="ED38DAAFFFEE7F079ACDE7AAD300FB45" box="[1020,1042,1237,1263]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="43" pageNumber="44" type="Herbarium">B</collectionCode>
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, isotype
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</materialsCitation>
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<collectingCountry id="F33E02FAFFEE7F079D4BE7AAD3B6FB45" box="[1146,1188,1237,1263]" name="United States of America" pageId="43" pageNumber="44">US</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33311E1FFEE7F0799F7E784D25DF915" pageId="43" pageNumber="44" type="distribution">
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<paragraph id="8B96426AFFEE7F0799F7E784D5F7FAC9" blockId="43.[151,1436,1205,1727]" pageId="43" pageNumber="44">
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<emphasis id="B95D9E78FFEE7F0799F7E784D53CFABF" bold="true" box="[198,558,1275,1301]" pageId="43" pageNumber="44">Distribution and ecology:—</emphasis>
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Locally common in mountains of
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<collectingCountry id="F33E02FAFFEE7F079AEFE783D37AFABC" box="[990,1128,1276,1302]" name="Costa Rica" pageId="43" pageNumber="44">Costa Rica</collectingCountry>
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,
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<collectingCountry id="F33E02FAFFEE7F079D4BE783D21CFABC" box="[1146,1294,1276,1302]" name="Puerto Rico" pageId="43" pageNumber="44">Puerto Rico</collectingCountry>
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,
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<collectingCountry id="F33E02FAFFEE7F079C10E783D28AFABC" box="[1313,1432,1276,1302]" name="Colombia" pageId="43" pageNumber="44">Colombia</collectingCountry>
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,
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<collectingCountry id="F33E02FAFFEE7F0799A6E65DD604FA96" box="[151,278,1314,1340]" name="Venezuela" pageId="43" pageNumber="44">Venezuela</collectingCountry>
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,
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<collectingCountry id="F33E02FAFFEE7F079812E65DD69BFA96" box="[291,393,1314,1340]" name="Ecuador" pageId="43" pageNumber="44">Ecuador</collectingCountry>
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,
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<collectingCountry id="F33E02FAFFEE7F0798A7E65DD6C2FA96" box="[406,464,1314,1340]" name="Peru" pageId="43" pageNumber="44">Peru</collectingCountry>
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, and
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<collectingCountry id="F33E02FAFFEE7F079B23E65DD57DFA96" box="[530,623,1314,1340]" name="Bolivia" pageId="43" pageNumber="44">Bolivia</collectingCountry>
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. Found in open habitats in humid montane forests, along roads, on landslides, and near streams, (800–)
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<quantity id="4CD1EF8FFFEE7F079B0BE636D5F3FAC9" box="[570,737,1353,1379]" metricMagnitude="3" metricUnit="m" metricValue="2.325" metricValueMax="3.25" metricValueMin="1.4" pageId="43" pageNumber="44" unit="m" value="2325.0" valueMax="3250.0" valueMin="1400.0">1400–3250 m</quantity>
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.
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</paragraph>
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<paragraph id="8B96426AFFEE7F0799F7E610D25DF915" blockId="43.[151,1436,1205,1727]" pageId="43" pageNumber="44">
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<emphasis id="B95D9E78FFEE7F0799F7E610D621FA23" bold="true" box="[198,307,1391,1417]" pageId="43" pageNumber="44">Notes:—</emphasis>
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<taxonomicName id="4C2939E9FFEE7F079802E60FD520FA23" authorityName="Nakai" authorityYear="1950" baseAuthorityName="Mett." box="[307,562,1392,1417]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="rubiginosus">
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<emphasis id="B95D9E78FFEE7F079802E60FD520FA23" box="[307,562,1392,1417]" italics="true" pageId="43" pageNumber="44">Sticherus rubiginosus</emphasis>
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</taxonomicName>
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belongs to a group of five closely related species, which also includes
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<taxonomicName id="4C2939E9FFEE7F079CB7E60FD7ECFA1A" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="brittonii">
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<emphasis id="B95D9E78FFEE7F079CB7E60FD7ECFA1A" italics="true" pageId="43" pageNumber="44">S. brittonii</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C2939E9FFEE7F079821E6E8D6DCFA1A" authorityName="Copeland" authorityYear="1947" baseAuthorityName="Christ" box="[272,462,1431,1456]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="strictissimus">
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||
<emphasis id="B95D9E78FFEE7F079821E6E8D6DCFA1A" box="[272,462,1431,1456]" italics="true" pageId="43" pageNumber="44">S. strictissimus</emphasis>
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||
</taxonomicName>
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,
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<taxonomicName id="4C2939E9FFEE7F0798D1E6E8D567FA1A" authorityName="Ching" authorityYear="1940" baseAuthorityName="Kaulf." box="[480,629,1431,1456]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="farinosus">
|
||
<emphasis id="B95D9E78FFEE7F0798D1E6E8D567FA1A" box="[480,629,1431,1456]" italics="true" pageId="43" pageNumber="44">S. farinosus</emphasis>
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||
</taxonomicName>
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||
, and
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<taxonomicName id="4C2939E9FFEE7F079B8CE6E8D476FA1A" authorityName="Lellinger" authorityYear="1984" baseAuthorityName="Maxon & C. V. Morton" box="[701,868,1431,1456]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="boliviensis">
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||
<emphasis id="B95D9E78FFEE7F079B8CE6E8D476FA1A" box="[701,868,1431,1456]" italics="true" pageId="43" pageNumber="44">S. boliviensis</emphasis>
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||
</taxonomicName>
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. All five species have been considered to be conspecific (e.g., by
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<bibRefCitation id="EFB83F9BFFEE7F0798BAE6C2D596FA7D" author="Tryon, R. M. & Stolze, R. G." box="[395,644,1469,1495]" pageId="43" pageNumber="44" pagination="1 - 122" refId="ref34267" refString="Tryon, R. M. & Stolze, R. G. (1989) Pteridophyta of Peru. Part II. 13. Pteridaceae - 15. Dennstaedtiaceae. Fieldiana Botany, n. s. 22: 1 - 122." type="book chapter" year="1989">Tryon & Stolze 1989</bibRefCitation>
|
||
,
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<bibRefCitation id="EFB83F9BFFEE7F079BA1E6C2D4F0FA7D" author="Ostergaard A. & Ollgaard, E." box="[656,994,1469,1495]" pageId="43" pageNumber="44" pagination="107 - 169" refId="ref33047" refString="Ostergaard A., Ollgaard, E. & B. (2001) Gleicheniaceae. In Harling, G. & Andersson, L. (eds.), Flora of Ecuador 66: 107 - 169." type="journal article" year="2001">Østergaard & Øllgaard 2001</bibRefCitation>
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||
). They all have lanceolate branch and bud scales with a marginal to central pattern of scale darkening, but they differ in the degree of separation of the segments, the shape of the midvein scales, the prominence of the veinlets, and to some degree, in the pattern of scale darkening. Most collections from the complex surrounding
|
||
<taxonomicName id="4C2939E9FFEE7F079D09E54ED3F4F9E0" authorityName="Nakai" authorityYear="1950" baseAuthorityName="Mett." box="[1080,1254,1585,1610]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="rubiginosus">
|
||
<emphasis id="B95D9E78FFEE7F079D09E54ED3F4F9E0" box="[1080,1254,1585,1610]" italics="true" pageId="43" pageNumber="44">S. rubiginosus</emphasis>
|
||
</taxonomicName>
|
||
can readily be placed into the five species we propose. However, there are some specimens, especially from
|
||
<collectingCountry id="F33E02FAFFEE7F079DD1E527D22BF9D8" box="[1248,1337,1624,1650]" name="Bolivia" pageId="43" pageNumber="44">Bolivia</collectingCountry>
|
||
, that are either intermediate between species or show a unique combination of characters. Since these specimens do not form a uniform group or show an indication of hybridity, they cannot be delimited as a distinct species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |