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<document id="60C7E24B0C87ACC365D0C23D828F2C2B" ID-CLB-Dataset="6085" ID-DOI="10.11646/phytotaxa.31.1.1" ID-GBIF-Dataset="2185a8cf-c15c-448b-b8c0-6055976408ed" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894648" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1622649568931" checkinUser="felipe" docAuthor="Gonzales, Jasivia &amp; Kessler, Michael" docDate="2011" docId="0380F37CFFEC7F0699A6E5EDD256FE36" docLanguage="en" docName="Phytotaxa.31.1-54.pdf.imf" docOrigin="Phytotaxa 31" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Sticherus Gonzales &amp; Kessler, 2011, comb. nov." docType="treatment" docVersion="7" lastPageNumber="43" masterDocId="FFB98B04FFC57F2C9931E37FD712FFAA" masterDocTitle="A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="42" updateTime="1698998516251" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="291D1987B3B78E26C3868859AEB78DA0">A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species</mods:title>
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<mods:namePart id="DBE26EC83CA445389EB1EAA4BC1E9651">Gonzales, Jasivia</mods:namePart>
<mods:affiliation id="AA616DB8F87F673F9C7C05C528E61C77">Herbario Nacional de Bolivia, Casilla 10077, La Paz, Bolivia. Systematic Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland. E-mail: michael. kessler @ systbot. uzh. ch</mods:affiliation>
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<mods:namePart id="9E048D6537D9D7541D5E2414AA051CD8">Kessler, Michael</mods:namePart>
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<mods:title id="909427F82C65EF85875A71C1E7D12D67">Phytotaxa</mods:title>
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<mods:date id="8C3EF305ECF77685156E4F99EDB73593">2011</mods:date>
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<treatment id="0380F37CFFEC7F0699A6E5EDD256FE36" ID-DOI="http://doi.org/10.5281/zenodo.4894611" ID-GBIF-Taxon="182742275" ID-Zenodo-Dep="4894611" LSID="urn:lsid:plazi:treatment:0380F37CFFEC7F0699A6E5EDD256FE36" httpUri="http://treatment.plazi.org/id/0380F37CFFEC7F0699A6E5EDD256FE36" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<subSubSection id="C33311E1FFEC7F0599A6E5EDD31BF906" box="[151,1033,1682,1708]" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="8B96426AFFEC7F0599A6E5EDD31BF906" blockId="41.[151,1033,1682,1708]" box="[151,1033,1682,1708]" pageId="41" pageNumber="42">
<heading id="D0DEF506FFEC7F0599A6E5EDD31BF906" bold="true" box="[151,1033,1682,1708]" fontSize="10" level="2" pageId="41" pageNumber="42" reason="2">
<emphasis id="B95D9E78FFEC7F0599A6E5EDD31BF906" bold="true" box="[151,1033,1682,1708]" pageId="41" pageNumber="42">
<emphasis id="B95D9E78FFEC7F0599A6E5EDD6D8F906" bold="true" box="[151,458,1682,1708]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="4C2939E9FFEC7F0599A6E5EDD615F906" ID-CoL="645MH" authorityName="Presl" authorityYear="1836" box="[151,263,1682,1708]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="genus">Sticherus</taxonomicName>
× pseudobifidus
</emphasis>
(Jermy &amp; T.G.Walker) J.Gonzales,
<emphasis id="B95D9E78FFEC7F059AB9E5EDD31BF906" bold="true" box="[904,1033,1682,1708]" italics="true" pageId="41" pageNumber="42">
<taxonomicNameLabel id="A26E2303FFEC7F059AB9E5EDD31BF906" box="[904,1033,1682,1708]" pageId="41" pageNumber="42" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33311E1FFEC7F0599A6E59FD5C6F95D" box="[151,724,1760,1783]" pageId="41" pageNumber="42" type="reference_group">
<paragraph id="8B96426AFFEC7F0599A6E59FD5C6F95D" blockId="41.[151,1436,1760,2011]" box="[151,724,1760,1783]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="AB396544FFEC7F0599A6E59FD5C6F95D" box="[151,724,1760,1783]" pageId="41" pageNumber="42">
<emphasis id="B95D9E78FFEC7F0599A6E59FD6BCF95D" box="[151,430,1760,1783]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="4C2939E9FFEC7F0599A6E59FD61EF95D" ID-CoL="62SHP" authorityName="Smith" authorityYear="1793" box="[151,268,1760,1783]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="genus">Gleichenia</taxonomicName>
×pseudobifida
</emphasis>
<treatmentCitation id="0A88647BFFEC7F059884E59FD5C2F95D" author="Jermy, A. C. &amp; Walker, T. G." box="[437,720,1760,1783]" page="2" pageId="41" pageNumber="42" year="1985">
<bibRefCitation id="EFB83F9BFFEC7F059884E59FD5C2F95D" author="Jermy, A. C. &amp; Walker, T. G." box="[437,720,1760,1783]" pageId="41" pageNumber="42" pagination="258 - 264" refId="ref31812" refString="Jermy, A. C. &amp; Walker, T. G. (1985) Cytotaxonomic studies of the ferns of Trinidad. Bulletin of the British Museum Natural History Botany 13: 185 - 191, 258 - 264." type="journal article" year="1985">Jermy &amp; Walker (1985: 2)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33311E1FFEC7F0599F7E580D65FF8EA" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="8B96426AFFEC7F0599F7E580D65FF8EA" blockId="41.[151,1436,1760,2011]" pageId="41" pageNumber="42">
<emphasis id="B95D9E78FFEC7F0599F7E580D639F8B3" bold="true" box="[198,299,1791,1817]" pageId="41" pageNumber="42">Type:—</emphasis>
<materialsCitation id="3B414837FFEC7F05981AE47FD242F8B3" ID-GBIF-Occurrence="3131398397" box="[299,1360,1792,1818]" collectionCode="T" country="Trinidad and Tobago" elevation="450" latitude="10.966666" location="Arima-Blanchisseouse Road" longLatPrecision="1296" longitude="-61.166668" pageId="41" pageNumber="42" specimenCount="1">
<collectingCountry id="F33E02FAFFEC7F05981AE47FD6AEF8B0" box="[299,444,1792,1818]" name="Trinidad and Tobago" pageId="41" pageNumber="42">TRINIDAD</collectingCountry>
.
<location id="8EF614B1FFEC7F0598FDE47FD431F8B0" LSID="urn:lsid:plazi:treatment:0380F37CFFEC7F0699A6E5EDD256FE36:8EF614B1FFEC7F0598FDE47FD431F8B0" box="[460,803,1792,1818]" country="Trinidad and Tobago" latitude="10.966666" longLatPrecision="1296" longitude="-61.166668" name="Arima-Blanchisseouse Road" pageId="41" pageNumber="42">Arima-Blanchisseouse Road</location>
, ca.
<geoCoordinate id="EE1D24ADFFEC7F059A55E47FD4CBF8B0" box="[868,985,1792,1818]" degrees="10" direction="north" minutes="58" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="10.966666">10°58 N</geoCoordinate>
,
<geoCoordinate id="EE1D24ADFFEC7F059ADAE47FD377F8B0" box="[1003,1125,1792,1818]" degrees="61" direction="west" minutes="10" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-61.166668">61°10 W</geoCoordinate>
,
<quantity id="4CD1EF8FFFEC7F059D47E47FD3DBF8B0" box="[1142,1225,1792,1818]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="41" pageNumber="42" unit="m" value="450.0">
<elevation id="0004A559FFEC7F059D47E47FD3DBF8B0" box="[1142,1225,1792,1818]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="41" pageNumber="42" unit="m" value="450.0">450 m</elevation>
</quantity>
,
<emphasis id="B95D9E78FFEC7F059DEAE47FD242F8B3" box="[1243,1360,1792,1817]" italics="true" pageId="41" pageNumber="42">
Walker,
<collectionCode id="ED38DAAFFFEC7F059C0FE47FD242F8B3" box="[1342,1360,1792,1817]" name="Tavera, Department of Geology and Geophysics" pageId="41" pageNumber="42">T</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="B95D9E78FFEC7F059C61E47FD246F8B3" box="[1360,1364,1792,1817]" italics="true" pageId="41" pageNumber="42">.</emphasis>
<materialsCitation id="3B414837FFEC7F059C50E47FD658F8EA" ID-GBIF-Occurrence="3131398376" collectionCode="BM" country="Trinidad and Tobago" pageId="41" pageNumber="42" specimenCount="1" typeStatus="holotype">
<emphasis id="B95D9E78FFEC7F059C50E47FD28EF8B3" box="[1377,1436,1792,1817]" italics="true" pageId="41" pageNumber="42">6414</emphasis>
(
<typeStatus id="5492FCC8FFEC7F0599AEE459D615F8EA" box="[159,263,1830,1856]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
<collectionCode id="ED38DAAFFFEC7F05983FE459D652F8EA" box="[270,320,1830,1856]" country="United Kingdom" name="Bristol Museum" pageId="41" pageNumber="42">BM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33311E1FFEC7F0699F7E432D256FE36" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="distribution">
<paragraph id="8B96426AFFEC7F0599F7E432D205F871" blockId="41.[151,1436,1760,2011]" pageId="41" pageNumber="42">
<emphasis id="B95D9E78FFEC7F0599F7E432D532F8CD" bold="true" box="[198,544,1869,1895]" pageId="41" pageNumber="42">Distribution and ecology:—</emphasis>
Best known in
<collectingCountry id="F33E02FAFFEC7F059BE1E432D425F8CD" box="[720,823,1869,1895]" name="Trinidad and Tobago" pageId="41" pageNumber="42">Trinidad</collectingCountry>
, where it is fairly common along roadsides in humid forests on the northern mountain range at
<quantity id="4CD1EF8FFFEC7F059BBDE40BD407F824" box="[652,789,1908,1934]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="4.5" metricValueMin="2.0" pageId="41" pageNumber="42" unit="m" value="325.0" valueMax="450.0" valueMin="200.0">200450 m</quantity>
, but also known from scattered specimens from
<collectingCountry id="F33E02FAFFEC7F059C69E40BD7C0F81E" name="Costa Rica" pageId="41" pageNumber="42">Costa Rica</collectingCountry>
,
<collectingCountry id="F33E02FAFFEC7F0599EEE4E5D651F81E" box="[223,323,1946,1972]" name="Panama" pageId="41" pageNumber="42">Panama</collectingCountry>
,
<collectingCountry id="F33E02FAFFEC7F059860E4E5D686F81E" box="[337,404,1946,1972]" name="Cuba" pageId="41" pageNumber="42">Cuba</collectingCountry>
,
<collectingCountry id="F33E02FAFFEC7F059890E4E5D525F81E" box="[417,567,1946,1972]" name="Puerto Rico" pageId="41" pageNumber="42">Puerto Rico</collectingCountry>
,
<collectingCountry id="F33E02FAFFEC7F059B74E4E5D5D2F81E" box="[581,704,1946,1972]" name="Dominica" pageId="41" pageNumber="42">Dominica</collectingCountry>
,
<collectingCountry id="F33E02FAFFEC7F059BFCE4E5D42AF81E" box="[717,824,1946,1972]" name="Grenada" pageId="41" pageNumber="42">Grenada</collectingCountry>
,
<collectingCountry id="F33E02FAFFEC7F059A77E4E5D4D3F81E" box="[838,961,1946,1972]" name="Colombia" pageId="41" pageNumber="42">Colombia</collectingCountry>
, northern and southeastern
<collectingCountry id="F33E02FAFFEC7F059C26E4E5D28AF81E" box="[1303,1432,1946,1972]" name="Venezuela" pageId="41" pageNumber="42">Venezuela</collectingCountry>
,
<collectingCountry id="F33E02FAFFEC7F0599A6E4BED7E7F871" box="[151,245,1985,2011]" name="Guyana" pageId="41" pageNumber="42">Guyana</collectingCountry>
,
<collectingCountry id="F33E02FAFFEC7F059830E4BED676F871" box="[257,356,1985,2011]" name="Ecuador" pageId="41" pageNumber="42">Ecuador</collectingCountry>
, and
<collectingCountry id="F33E02FAFFEC7F059890E4BED6C9F871" box="[417,475,1985,2011]" name="Peru" pageId="41" pageNumber="42">Peru</collectingCountry>
. It is likely to be more common and widespread than currently known.
</paragraph>
<paragraph id="8B96426AFFEF7F0699F7E3E8D256FE36" blockId="42.[151,1436,151,412]" pageId="42" pageNumber="43">
<emphasis id="B95D9E78FFEF7F0699F7E3E8D621FF1B" bold="true" box="[198,307,151,177]" pageId="42" pageNumber="43">Notes:—</emphasis>
This triploid hybrid between
<taxonomicName id="4C2939E9FFEF7F069BBBE3E7D431FF1B" authorityName="J. Gonzales" authorityYear="2011" baseAuthorityName="Jermy &amp; T. G. Walker" baseAuthorityYear="2011" box="[650,803,152,177]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="interjectus">
<emphasis id="B95D9E78FFEF7F069BBBE3E7D431FF1B" box="[650,803,152,177]" italics="true" pageId="42" pageNumber="43">S. interjectus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2939E9FFEF7F069A6DE3E7D313FF1B" authorityName="J. Gonzales" authorityYear="2011" baseAuthorityName="Desv." box="[860,1025,152,177]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ferrugineus">
<emphasis id="B95D9E78FFEF7F069A6DE3E7D313FF1B" box="[860,1025,152,177]" italics="true" pageId="42" pageNumber="43">S. ferrugineus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFB83F9BFFEF7F069DEBE3E7D7F7FF72" author="Jermy, A. C. &amp; Walker, T. G." pageId="42" pageNumber="43" pagination="258 - 264" refId="ref31812" refString="Jermy, A. C. &amp; Walker, T. G. (1985) Cytotaxonomic studies of the ferns of Trinidad. Bulletin of the British Museum Natural History Botany 13: 185 - 191, 258 - 264." type="journal article" year="1985">Jermy &amp; Walker (1985)</bibRefCitation>
on the basis of cytotaxonomic studies of the ferns of
<collectingCountry id="F33E02FAFFEF7F069A51E3C1D4DAFF72" box="[864,968,190,216]" name="Trinidad and Tobago" pageId="42" pageNumber="43">Trinidad</collectingCountry>
. It is similar in size to
<taxonomicName id="4C2939E9FFEF7F069DE4E3C0D27DFF72" authorityName="J. Gonzales" authorityYear="2011" baseAuthorityName="Jermy &amp; T. G. Walker" baseAuthorityYear="2011" box="[1237,1391,191,216]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="interjectus">
<emphasis id="B95D9E78FFEF7F069DE4E3C0D27DFF72" box="[1237,1391,191,216]" italics="true" pageId="42" pageNumber="43">S. interjectus</emphasis>
</taxonomicName>
but has a much denser indument abaxially on the segments, thus approaching
<taxonomicName id="4C2939E9FFEF7F069AC5E39AD38AFF54" authorityName="J. Gonzales" authorityYear="2011" baseAuthorityName="Desv." box="[1012,1176,229,254]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ferrugineus">
<emphasis id="B95D9E78FFEF7F069AC5E39AD38AFF54" box="[1012,1176,229,254]" italics="true" pageId="42" pageNumber="43">S. ferrugineus</emphasis>
</taxonomicName>
. On the other hand,
<emphasis id="B95D9E78FFEF7F069CB7E39AD647FE8F" italics="true" pageId="42" pageNumber="43">S. × pseudobifidus</emphasis>
differs from
<taxonomicName id="4C2939E9FFEF7F0698DEE273D581FE8F" authorityName="J. Gonzales" authorityYear="2011" baseAuthorityName="Desv." box="[495,659,268,293]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ferrugineus">
<emphasis id="B95D9E78FFEF7F0698DEE273D581FE8F" box="[495,659,268,293]" italics="true" pageId="42" pageNumber="43">S. ferrugineus</emphasis>
</taxonomicName>
in having irregularly darkened parts on the bud and branch scales, whereas
<taxonomicName id="4C2939E9FFEF7F069837E24DD6A4FEE1" authorityName="J. Gonzales" authorityYear="2011" baseAuthorityName="Desv." box="[262,438,306,331]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ferrugineus">
<emphasis id="B95D9E78FFEF7F069837E24DD6A4FEE1" box="[262,438,306,331]" italics="true" pageId="42" pageNumber="43">S. ferrugineus</emphasis>
</taxonomicName>
has translucent or only basally darkened scales.
<taxonomicName id="4C2939E9FFEF7F069D15E24DD385FEE1" authorityName="Presl" authorityYear="1836" box="[1060,1175,306,331]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B95D9E78FFEF7F069D15E24DD385FEE1" box="[1060,1175,306,331]" italics="true" pageId="42" pageNumber="43">Sticherus</emphasis>
</taxonomicName>
×
<emphasis id="B95D9E78FFEF7F069D8DE24DD27BFEE1" box="[1212,1385,306,331]" italics="true" pageId="42" pageNumber="43">pseudobifidus</emphasis>
has conspicuously irregular segment lengths, a character shared by
<emphasis id="B95D9E78FFEF7F069AB7E226D487FED8" box="[902,917,345,370]" italics="true" pageId="42" pageNumber="43">S</emphasis>
. ×
<emphasis id="B95D9E78FFEF7F069A8CE226D351FED8" box="[957,1091,345,370]" italics="true" pageId="42" pageNumber="43">subremotus</emphasis>
. Also, both hybrid taxa have strikingly unequally developed 2
<superScript id="7C5CEF22FFEF7F069B29E2FFD53AFE24" attach="left" box="[536,552,384,398]" fontSize="6" pageId="42" pageNumber="43">nd</superScript>
branches, i.e., the opposite pairs of branches have different lengths.
</paragraph>
</subSubSection>
</treatment>
</document>