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<document id="C4CD4D5171989EB9E77C57F6006D3700" ID-CLB-Dataset="6085" ID-DOI="10.11646/phytotaxa.31.1.1" ID-GBIF-Dataset="2185a8cf-c15c-448b-b8c0-6055976408ed" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894648" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1622649568931" checkinUser="felipe" docAuthor="Gonzales, Jasivia &amp; Kessler, Michael" docDate="2011" docId="0380F37CFFEB7F0299A6E63AD7E1F825" docLanguage="en" docName="Phytotaxa.31.1-54.pdf.imf" docOrigin="Phytotaxa 31" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Sticherus strictissimus Copeland 1947" docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="FFB98B04FFC57F2C9931E37FD712FFAA" masterDocTitle="A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="47" updateTime="1698998516251" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9F35766F9B5575E5E91EA456EB71540D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="440EA3D11551ABF17A27FF0969DD23B8">
<mods:title id="4D93E6F022DE31655DE6C2D86FB2ECC0">A synopsis of the Neotropical species of Sticherus (Gleicheniaceae), with descriptions of nine new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="A194158F312EF688D22FCE2A3B48C4D6">Gonzales, Jasivia</mods:namePart>
<mods:affiliation id="D35A1BF215CD0C7EBDD35C9559AE52F6">Herbario Nacional de Bolivia, Casilla 10077, La Paz, Bolivia. Systematic Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland. E-mail: michael. kessler @ systbot. uzh. ch</mods:affiliation>
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<mods:namePart id="5032FD5001AFD01B9C13770F60385DAD">Kessler, Michael</mods:namePart>
</mods:name>
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<mods:title id="DD8DDB6BF55CBD1E199D5E0B056BED45">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="425A02621ED64ECAB5C6FB066DFC6C90">
<mods:date id="D698585945C2ED38F4BD9A1FC88ADFA8">2011</mods:date>
<mods:detail id="193692E8AA7BEF8FCB7D684EE6960C2A" type="pubDate">
<mods:number id="748376C02926C75F48085D9C93056C6B">2011-12-31</mods:number>
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<mods:detail id="CF707F877B2693ECBCFFFD2E7412A396" type="volume">
<mods:number id="72B02C292B070A24378CFB099AFA40DC">31</mods:number>
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<mods:start id="74B68CF4CCB3EB67592EDBD5751E573A">1</mods:start>
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<mods:classification id="C2BF493C8BEA25B0B4C06F4CF2A076FC">journal article</mods:classification>
<mods:identifier id="C9160319F34E51E4BDE5F99FE28ABD4A" type="CLB-Dataset">6085</mods:identifier>
<mods:identifier id="1E9D665C13534CC7340CFC6A721BACB0" type="DOI">10.11646/phytotaxa.31.1.1</mods:identifier>
<mods:identifier id="5FBEBE08BE24EBCDD4E96B7FD44F8359" type="GBIF-Dataset">2185a8cf-c15c-448b-b8c0-6055976408ed</mods:identifier>
<mods:identifier id="4693A61BB1C3C29D67D2D7989099A251" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="C702179AFEF5FD689B35FA779E216520" type="Zenodo-Dep">4894648</mods:identifier>
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<treatment id="0380F37CFFEB7F0299A6E63AD7E1F825" ID-DOI="http://doi.org/10.5281/zenodo.4894800" ID-GBIF-Taxon="182742320" ID-Zenodo-Dep="4894800" LSID="urn:lsid:plazi:treatment:0380F37CFFEB7F0299A6E63AD7E1F825" httpUri="http://treatment.plazi.org/id/0380F37CFFEB7F0299A6E63AD7E1F825" lastPageNumber="47" pageId="46" pageNumber="47">
<subSubSection id="C33311E1FFEB7F0299A6E63AD5E7FAF5" box="[151,757,1349,1375]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8B96426AFFEB7F0299A6E63AD5E7FAF5" blockId="46.[151,757,1349,1375]" box="[151,757,1349,1375]" pageId="46" pageNumber="47">
<heading id="D0DEF506FFEB7F0299A6E63AD5E7FAF5" box="[151,757,1349,1375]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="3">
<treatmentCitationGroup id="AB396544FFEB7F0299A6E63AD5E7FAF5" box="[151,757,1349,1375]" pageId="46" pageNumber="47">
<treatmentCitation id="0A88647BFFEB7F0299A6E63AD5E7FAF5" author="Copeland, E. B." box="[151,757,1349,1375]" page=")" pageId="46" pageNumber="47" year="1947">
<taxonomicName id="4C2939E9FFEB7F0299A6E63AD5E7FAF5" ID-CoL="52HXV" authority="(Christ) Copeland (1947: 28)" authorityName="Copeland" authorityPageNumber="28" authorityYear="1947" baseAuthorityName="Christ" box="[151,757,1349,1375]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="strictissimus">
<emphasis id="B95D9E78FFEB7F0299A6E63AD6B3FAF5" bold="true" box="[151,417,1349,1375]" italics="true" pageId="46" pageNumber="47">Sticherus strictissimus</emphasis>
(Christ)
<bibRefCitation id="EFB83F9BFFEB7F029B39E63AD5E7FAF5" author="Copeland, E. B." box="[520,757,1349,1375]" pageId="46" pageNumber="47" pagination="1 - 247" refId="ref30844" refString="Copeland, E. B. (1947) Genera filicum. The genera of ferns. Annales Cryptogamici et Phytopathologici 5: 1 - 247." type="journal article" year="1947">Copeland (1947: 28)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33311E1FFEB7F0299A6E6EDD3F7FA03" box="[151,1253,1426,1449]" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="8B96426AFFEB7F0299A6E6EDD3F7FA03" blockId="46.[151,1436,1426,1935]" box="[151,1253,1426,1449]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB396544FFEB7F0299A6E6EDD3F7FA03" box="[151,1253,1426,1449]" pageId="46" pageNumber="47">
<treatmentCitation id="0A88647BFFEB7F0299A6E6EDD54BFA03" author="Christ, H." box="[151,601,1426,1449]" page=")" pageId="46" pageNumber="47" year="1905">
<taxonomicName id="4C2939E9FFEB7F0299A6E6EDD54BFA03" ID-CoL="6KH3C" authority="Christ (1905 a: 13)" authorityName="Christ" authorityPageNumber="13" authorityYear="1905" box="[151,601,1426,1449]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="strictissima">
<emphasis id="B95D9E78FFEB7F0299A6E6EDD69FFA03" box="[151,397,1426,1449]" italics="true" pageId="46" pageNumber="47">Gleichenia strictissima</emphasis>
<bibRefCitation id="EFB83F9BFFEB7F0298A5E6EDD54BFA03" author="Christ, H." box="[404,601,1426,1449]" pageId="46" pageNumber="47" pagination="1 - 16" refId="ref30509" refString="Christ, H. (1905 a) Primitiae florae costaricensis. Bulletin de l'Herbier Boissier ser. 2, 5: 1 - 16." type="journal article" year="1905">Christ (1905a: 13)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="0A88647BFFEB7F029B54E6EDD3F2FA03" author="Underwood, L. M." box="[613,1248,1426,1449]" page=")" pageId="46" pageNumber="47" year="1907">
<taxonomicName id="4C2939E9FFEB7F029B54E6EDD3F2FA03" ID-CoL="6CTMT" authority="(Christ) Underwood (1907: 261)" authorityName="Underwood" authorityPageNumber="261" authorityYear="1907" baseAuthorityName="Christ" box="[613,1248,1426,1449]" class="Polypodiopsida" family="Gleicheniaceae" genus="Dicranopteris" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="strictissima">
<emphasis id="B95D9E78FFEB7F029B54E6EDD46EFA03" box="[613,892,1426,1449]" italics="true" pageId="46" pageNumber="47">Dicranopteris strictissima</emphasis>
(Christ)
<bibRefCitation id="EFB83F9BFFEB7F029AEDE6EDD3F2FA03" author="Underwood, L. M." box="[988,1248,1426,1449]" pageId="46" pageNumber="47" pagination="243 - 262" refId="ref34344" refString="Underwood, L. M. (1907) American Ferns VIII. A preliminary review of the North American Gleicheniaceae. Bulletin of the Torrey Botanical Club 34: 243 - 262." type="journal article" year="1907">Underwood (1907: 261)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33311E1FFEB7F0299F6E6CDD470FA63" box="[199,866,1458,1482]" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8B96426AFFEB7F0299F6E6CDD470FA63" blockId="46.[151,1436,1426,1935]" box="[199,866,1458,1482]" pageId="46" pageNumber="47">
<emphasis id="B95D9E78FFEB7F0299F6E6CDD615FA60" bold="true" box="[199,263,1458,1482]" pageId="46" pageNumber="47">Type:</emphasis>
—
<materialsCitation id="3B414837FFEB7F029813E6CDD43BFA63" ID-GBIF-Occurrence="3131398306" box="[290,809,1458,1481]" collectionCode="P" country="Costa Rica" location="Werckle" pageId="46" pageNumber="47" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F33E02FAFFEB7F029813E6CDD6D6FA63" box="[290,452,1458,1481]" name="Costa Rica" pageId="46" pageNumber="47">COSTA RICA</collectingCountry>
.
<emphasis id="B95D9E78FFEB7F0298FCE6CDD542FA63" box="[461,592,1458,1481]" italics="true" pageId="46" pageNumber="47">
<location id="8EF614B1FFEB7F0298FCE6CDD533FA63" LSID="urn:lsid:plazi:treatment:0380F37CFFEB7F0299A6E63AD7E1F825:8EF614B1FFEB7F0298FCE6CDD533FA63" box="[461,545,1458,1481]" country="Costa Rica" name="Werckle" pageId="46" pageNumber="47">Wercklé</location>
215
</emphasis>
(
<typeStatus id="5492FCC8FFEB7F029B6FE6CDD5AFFA63" box="[606,701,1458,1481]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
<collectionCode id="ED38DAAFFFEB7F029BF2E6CDD5C0FA63" box="[707,722,1458,1481]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="46" pageNumber="47" type="Herbarium">P</collectionCode>
, isotype
</materialsCitation>
<collectingCountry id="F33E02FAFFEB7F029A01E6CDD444FA63" box="[816,854,1458,1481]" name="United States of America" pageId="46" pageNumber="47">US</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C33311E1FFEB7F0299A6E6ADD3AEFA43" box="[151,1212,1490,1514]" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="8B96426AFFEB7F0299A6E6ADD3AEFA43" blockId="46.[151,1436,1426,1935]" box="[151,1212,1490,1514]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB396544FFEB7F0299A6E6ADD3AEFA43" box="[151,1212,1490,1514]" pageId="46" pageNumber="47">
<treatmentCitation id="0A88647BFFEB7F0299A6E6ADD531FA43" author="Christ, H." box="[151,547,1490,1513]" page=")" pageId="46" pageNumber="47" year="1906">
<taxonomicName id="4C2939E9FFEB7F0299A6E6ADD531FA43" ID-CoL="3G76V" authority="Christ (1906: 6)" authorityName="Christ" authorityPageNumber="6" authorityYear="1906" box="[151,547,1490,1513]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="glaucina">
<emphasis id="B95D9E78FFEB7F0299A6E6ADD662FA43" box="[151,368,1490,1513]" italics="true" pageId="46" pageNumber="47">Gleichenia glaucina</emphasis>
<materialsCitation id="3B414837FFEB7F029847E6ADD531FA43" ID-GBIF-Occurrence="3131398304" box="[374,547,1490,1513]" collectorName="La Palma" country="Costa Rica" location="COSTA RICA" pageId="46" pageNumber="47" specimenCount="1">
<bibRefCitation id="EFB83F9BFFEB7F029847E6ADD531FA43" author="Christ, H." box="[374,547,1490,1513]" pageId="46" pageNumber="47" pagination="279 - 288" refId="ref30561" refString="Christ, H. (1906) Primitiae florae costaricensis. Bulletin de l'Herbier Boissier ser. 2, 6: 279 - 288." type="journal article" year="1906">Christ (1906: 6)</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="B95D9E78FFEB7F029B1FE6ADD57DFA40" bold="true" box="[558,623,1490,1514]" pageId="46" pageNumber="47">
<typeStatus id="5492FCC8FFEB7F029B1FE6ADD574FA40" box="[558,614,1490,1514]" pageId="46" pageNumber="47">Type</typeStatus>
:
</emphasis>
—
<collectingCountry id="F33E02FAFFEB7F029BB8E6ADD438FA43" box="[649,810,1490,1513]" name="Costa Rica" pageId="46" pageNumber="47">COSTA RICA</collectingCountry>
.
<collectorName id="26DC27BCFFEB7F029A05E6ADD48EFA43" box="[820,924,1490,1513]" pageId="46" pageNumber="47">La Palma</collectorName>
,
<emphasis id="B95D9E78FFEB7F029A96E6ADD335FA43" box="[935,1063,1490,1513]" italics="true" pageId="46" pageNumber="47">Wercklé s.n.</emphasis>
(not located).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33311E1FFEB7F0299F7E68DD7E1F825" pageId="46" pageNumber="47" type="distribution">
<paragraph id="8B96426AFFEB7F0299F7E68DD562F9F0" blockId="46.[151,1436,1426,1935]" pageId="46" pageNumber="47">
<emphasis id="B95D9E78FFEB7F0299F7E68DD532F9A6" bold="true" box="[198,544,1522,1548]" pageId="46" pageNumber="47">Distribution and ecology:—</emphasis>
Mountains of
<collectingCountry id="F33E02FAFFEB7F029BF4E68DD455F9A6" box="[709,839,1522,1548]" name="Guatemala" pageId="46" pageNumber="47">Guatemala</collectingCountry>
,
<collectingCountry id="F33E02FAFFEB7F029A63E68DD4C7F9A6" box="[850,981,1522,1548]" name="Costa Rica" pageId="46" pageNumber="47">Costa Rica</collectingCountry>
,
<collectingCountry id="F33E02FAFFEB7F029AD0E68DD34AF9A6" box="[993,1112,1522,1548]" name="Colombia" pageId="46" pageNumber="47">Colombia</collectingCountry>
,
<collectingCountry id="F33E02FAFFEB7F029D52E68DD3CDF9A6" box="[1123,1247,1522,1548]" name="Venezuela" pageId="46" pageNumber="47">Venezuela</collectingCountry>
, and
<collectingCountry id="F33E02FAFFEB7F029C2AE68DD26DF9A6" box="[1307,1407,1522,1548]" name="Ecuador" pageId="46" pageNumber="47">Ecuador</collectingCountry>
. It is locally common in
<collectingCountry id="F33E02FAFFEB7F0298AAE566D50EF999" box="[411,540,1561,1587]" name="Costa Rica" pageId="46" pageNumber="47">Costa Rica</collectingCountry>
and
<collectingCountry id="F33E02FAFFEB7F029B69E566D5AFF999" box="[600,701,1561,1587]" name="Ecuador" pageId="46" pageNumber="47">Ecuador</collectingCountry>
, but apparently rare elsewhere; grows in disturbed habitats in humid montane forests at
<quantity id="4CD1EF8FFFEB7F0298F6E53FD57EF9F0" box="[455,620,1600,1626]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.4" metricValueMin="1.4" pageId="46" pageNumber="47" unit="m" value="1900.0" valueMax="2400.0" valueMin="1400.0">1400–2400 m</quantity>
.
</paragraph>
<paragraph id="8B96426AFFEB7F0299F7E519D7E1F825" blockId="46.[151,1436,1426,1935]" pageId="46" pageNumber="47">
<emphasis id="B95D9E78FFEB7F0299F7E519D621F92A" bold="true" box="[198,307,1638,1664]" pageId="46" pageNumber="47">Notes:—</emphasis>
<taxonomicName id="4C2939E9FFEB7F029802E518D53CF92A" box="[307,558,1639,1664]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="stictissimus">
<emphasis id="B95D9E78FFEB7F029802E518D53CF92A" box="[307,558,1639,1664]" italics="true" pageId="46" pageNumber="47">Sticherus stictissimus</emphasis>
</taxonomicName>
has usually been considered to be conspecific with
<taxonomicName id="4C2939E9FFEB7F029DBFE518D227F92A" authorityName="Nakai" authorityYear="1950" baseAuthorityName="Mett." box="[1166,1333,1639,1664]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="rubiginosus">
<emphasis id="B95D9E78FFEB7F029DBFE518D227F92A" box="[1166,1333,1639,1664]" italics="true" pageId="46" pageNumber="47">S. rubiginosus</emphasis>
</taxonomicName>
and was thought to be just a dwarf form of that species. In fact,
<bibRefCitation id="EFB83F9BFFEB7F029A64E5F2D3C2F90D" author="Ostergaard A. &amp; Ollgaard, E." box="[853,1232,1677,1703]" pageId="46" pageNumber="47" pagination="107 - 169" refId="ref33047" refString="Ostergaard A., Ollgaard, E. &amp; B. (2001) Gleicheniaceae. In Harling, G. &amp; Andersson, L. (eds.), Flora of Ecuador 66: 107 - 169." type="journal article" year="2001">Østergaard &amp; Øllgaard (2001)</bibRefCitation>
believed that
<taxonomicName id="4C2939E9FFEB7F029CB4E5F2D63BF967" authorityName="Copeland" authorityYear="1947" baseAuthorityName="Christ" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="strictissimus">
<emphasis id="B95D9E78FFEB7F029CB4E5F2D63BF967" italics="true" pageId="46" pageNumber="47">S. strictissimus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2939E9FFEB7F029852E5CBD548F967" authorityName="Bosco" authorityYear="1938" box="[355,602,1716,1741]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenia" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="salesiana">
<emphasis id="B95D9E78FFEB7F029852E5CBD548F967" box="[355,602,1716,1741]" italics="true" pageId="46" pageNumber="47">Gleichenia salesiana</emphasis>
</taxonomicName>
(see under doubtful names) represented precociously fertile specimens of
<taxonomicName id="4C2939E9FFEB7F029989E5A4D677F95E" authorityName="Nakai" authorityYear="1950" baseAuthorityName="Mett." box="[184,357,1755,1780]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="rubiginosus">
<emphasis id="B95D9E78FFEB7F029989E5A4D677F95E" box="[184,357,1755,1780]" italics="true" pageId="46" pageNumber="47">S. rubiginosus</emphasis>
</taxonomicName>
. However, in addition to size,
<taxonomicName id="4C2939E9FFEB7F029BECE5A4D481F95E" authorityName="Copeland" authorityYear="1947" baseAuthorityName="Christ" box="[733,915,1755,1780]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="strictissimus">
<emphasis id="B95D9E78FFEB7F029BECE5A4D481F95E" box="[733,915,1755,1780]" italics="true" pageId="46" pageNumber="47">S. strictissimus</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="4C2939E9FFEB7F029D5FE5A4D209F95E" authorityName="Nakai" authorityYear="1950" baseAuthorityName="Mett." box="[1134,1307,1755,1780]" class="Polypodiopsida" family="Gleicheniaceae" genus="Sticherus" kingdom="Plantae" order="Gleicheniales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="rubiginosus">
<emphasis id="B95D9E78FFEB7F029D5FE5A4D209F95E" box="[1134,1307,1755,1780]" italics="true" pageId="46" pageNumber="47">S. rubiginosus</emphasis>
</taxonomicName>
by having shorter segments with pointed apices, less densely scaly rhizomes, simple to forked, thick, whitish filamentous scales on the midveins, obscure veinlets with thick, whitish filamentous scales, and abundant papillae on segment surfaces. Based on all these differences, it seems best to consider both entities as distinct species.
</paragraph>
</subSubSection>
</treatment>
</document>
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