404 lines
48 KiB
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404 lines
48 KiB
XML
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<mods:title id="C9F8F88B098AC8F165CA5CB89C818D8D">Seven new taxa from the butterfly subtribe Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) with revisional notes on Harjesia Forster, 1964 and Pseudeuptychia Forster, 1964</mods:title>
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<mods:namePart id="FB3738BC12ACD228A6583A530F3C82FC">Nakahara, Shinichi</mods:namePart>
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<mods:namePart id="CCECE7D761DFB14B97BD9CE92E930227">Willmott, Keith R.</mods:namePart>
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<mods:namePart id="B9CA7F9552E4E6387D1656CCA4AC11E4">Mielke, Olaf H. H.</mods:namePart>
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<mods:namePart id="C6D94FE2D7A7653A86FF8BB8E579D52F">Schwartz, Johanna</mods:namePart>
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<mods:namePart id="04619E857C41E4733918E89B384BFDC2">Zacca, Thamara</mods:namePart>
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<mods:namePart id="CBD33FBDEFEBCA535C92E44E9672EF9F">Espeland, Marianne</mods:namePart>
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<mods:namePart id="3AB527E392FC3D07FFBC895FDED215AC">Lamas, Gerardo</mods:namePart>
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<mods:typeOfResource id="FDFE9D428150EF281CDC965FD2C931FE">text</mods:typeOfResource>
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<mods:title id="6A89761089E2D2EC028748D0DE81E73A">Insecta Mundi</mods:title>
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<mods:part id="E02DA655492F0F2724F46C26A1A53EB7">
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<mods:date id="D665D14ABF811EC9D00C8D7635C72D60">2018</mods:date>
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<mods:number id="E5BC77BCD51A6403B01355D7AE5C1EA1">2018-07-27</mods:number>
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<treatment id="537987D27841FFF06C99FF79FBCAECFC" ID-DOI="http://doi.org/10.5281/zenodo.3716571" ID-GBIF-Taxon="162485155" ID-Zenodo-Dep="3716571" LSID="urn:lsid:plazi:treatment:537987D27841FFF06C99FF79FBCAECFC" httpUri="http://treatment.plazi.org/id/537987D27841FFF06C99FF79FBCAECFC" lastPageNumber="7" pageId="8" pageNumber="7">
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<subSubSection id="93CA654F7841FFF06C99FF79FE15E993" box="[192,497,217,241]" pageId="8" pageNumber="7" type="nomenclature">
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<paragraph id="DB6F36C47841FFF06C99FF79FE15E993" blockId="8.[192,497,217,270]" box="[192,497,217,241]" pageId="8" pageNumber="7">
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<heading id="802781A87841FFF06C99FF79FE15E993" bold="true" box="[192,497,217,241]" fontSize="10" level="2" pageId="8" pageNumber="7" reason="3">
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<taxonomicName id="1CD04D477841FFF06C99FF79FE15E993" ID-CoL="92B6C" authority="Forster, 1964" authorityName=": Forster" authorityYear="1964" box="[192,497,217,241]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="E9A4EAD67841FFF06C99FF79FE15E993" bold="true" box="[192,497,217,241]" pageId="8" pageNumber="7">
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<emphasis id="E9A4EAD67841FFF06C99FF79FED3E993" bold="true" box="[192,311,217,241]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
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<bibRefCitation id="BF414B357841FFF06D67FF79FE15E993" author="Forster, W." box="[318,497,217,241]" pageId="8" pageNumber="7" pagination="51 - 188" refId="ref18607" refString="Forster, W. 1964. Beitrage zur Kenntnis der Insektenfauna Boliviens XIX Lepidoptera III. Satyridae. Veroffentlichungen der Zoologischen Staatssammlung Munchen 8: 51 - 188." type="journal article" year="1964">Forster, 1964</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="93CA654F7841FFF06C99FF58FE0DE86F" box="[192,489,248,270]" pageId="8" pageNumber="7" type="reference_group">
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<paragraph id="DB6F36C47841FFF06C99FF58FE0DE86F" blockId="8.[192,497,217,270]" box="[192,489,248,270]" pageId="8" pageNumber="7">
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<treatmentCitationGroup id="FBC011EA7841FFF06C99FF58FE0DE86F" box="[192,489,248,270]" pageId="8" pageNumber="7">
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<treatmentCitation id="5A7110D57841FFF06C99FF58FE02E86F" author="Forster, W." box="[192,486,248,270]" page="78" pageId="8" pageNumber="7" year="1964">
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<taxonomicName id="1CD04D477841FFF06C99FF58FE02E86F" ID-CoL="92B6C" authority=": Forster 1964: 78" authorityName=": Forster" authorityPageNumber="78" authorityYear="1964" box="[192,486,248,270]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="E9A4EAD67841FFF06C99FF58FEFBE86F" box="[192,287,248,269]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
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:
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<bibRefCitation id="BF414B357841FFF06D75FF58FE02E86F" author="Forster, W." box="[300,486,248,270]" pageId="8" pageNumber="7" pagination="51 - 188" refId="ref18607" refString="Forster, W. 1964. Beitrage zur Kenntnis der Insektenfauna Boliviens XIX Lepidoptera III. Satyridae. Veroffentlichungen der Zoologischen Staatssammlung Munchen 8: 51 - 188." type="journal article" year="1964">Forster 1964: 78</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="93CA654F7841FFF06C99FE86FAFBE85C" box="[192,1311,294,318]" pageId="8" pageNumber="7" type="type_taxon">
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<paragraph id="DB6F36C47841FFF06C99FE86FAFBE85C" blockId="8.[192,1311,294,318]" box="[192,1311,294,318]" pageId="8" pageNumber="7">
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<emphasis id="E9A4EAD67841FFF06C99FE86FE93E85C" bold="true" box="[192,375,294,318]" pageId="8" pageNumber="7">
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<typeStatus id="046B88667841FFF06C99FE86FEE0E85C" box="[192,260,294,318]" pageId="8" pageNumber="7">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="1CD04D477841FFF06D27FE87FCDEE85C" authority="Moschler 1877: 325" authorityName="Moschler" authorityPageNumber="325" authorityYear="1877" box="[382,826,294,318]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="E9A4EAD67841FFF06D27FE87FDA6E85C" box="[382,578,294,318]" italics="true" pageId="8" pageNumber="7">Taygetis blanda</emphasis>
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<bibRefCitation id="BF414B357841FFF06E13FE86FCDEE85C" author="Moschler, H. B." box="[586,826,294,318]" pageId="8" pageNumber="7" pagination="293 - 352" refId="ref19667" refString="Moschler, H. B. 1877. Beitrage zur Schmetterlings-Fauna von Surinam. Verhandlungen der kaiserlichkoniglichen zoologisch-botanischen Gesellschaft in Wien 26 (1): 293 - 352." type="journal article" year="1877">Möschler 1877: 325</bibRefCitation>
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</taxonomicName>
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, pl. 3, fig. 12. ― by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="93CA654F7841FFF06CB1FEF7FBC4EAAD" pageId="8" pageNumber="7" type="discussion">
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<paragraph id="DB6F36C47841FFF06CB1FEF7FA44EBCC" blockId="8.[192,1441,342,975]" pageId="8" pageNumber="7">
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<bibRefCitation id="BF414B357841FFF06CB1FEF7FE7CE80C" author="Forster, W." box="[232,408,342,366]" pageId="8" pageNumber="7" pagination="51 - 188" refId="ref18607" refString="Forster, W. 1964. Beitrage zur Kenntnis der Insektenfauna Boliviens XIX Lepidoptera III. Satyridae. Veroffentlichungen der Zoologischen Staatssammlung Munchen 8: 51 - 188." type="journal article" year="1964">Forster (1964)</bibRefCitation>
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erected
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<taxonomicName id="1CD04D477841FFF06E5BFEF7FD8FE80C" authorityName=": Forster" authorityYear="1964" box="[514,619,343,366]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="E9A4EAD67841FFF06E5BFEF7FD8FE80C" box="[514,619,343,366]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
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</taxonomicName>
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to accommodate two species,
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<taxonomicName id="1CD04D477841FFF06F86FEF7FA8DE80C" authority="Moschler, 1877" authorityName="Moschler" authorityYear="1877" box="[991,1385,342,366]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="E9A4EAD67841FFF06F86FEF7FB40E80C" box="[991,1188,342,366]" italics="true" pageId="8" pageNumber="7">Taygetis blanda</emphasis>
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<bibRefCitation id="BF414B357841FFF068F7FEF6FA8DE80C" author="Moschler, H. B." box="[1198,1385,342,366]" pageId="8" pageNumber="7" pagination="293 - 352" refId="ref19667" refString="Moschler, H. B. 1877. Beitrage zur Schmetterlings-Fauna von Surinam. Verhandlungen der kaiserlichkoniglichen zoologisch-botanischen Gesellschaft in Wien 26 (1): 293 - 352." type="journal article" year="1877">Möschler, 1877</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="1CD04D477841FFF06C99FED6FDA6E8EC" authority="Butler, 1867" authorityName="Butler" authorityYear="1867" box="[192,578,374,398]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="obscura">
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<emphasis id="E9A4EAD67841FFF06C99FED6FE46E8EC" box="[192,418,374,398]" italics="true" pageId="8" pageNumber="7">Euptychia obscura</emphasis>
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<bibRefCitation id="BF414B357841FFF06DF1FED6FDA6E8EC" author="Butler, A. G." box="[424,578,374,398]" pageId="8" pageNumber="7" pagination="458 - 504" refId="ref18109" refString="Butler, A. G. 1867. A monograph of the genus Euptychia, a numerous race of butterflies belonging to the family Satyridae; with descriptions of sixty species new to science, and notes to their affinities, etc. Proceedings of the Zoological Society of London 1866 (3): 458 - 504." type="journal article" year="1867">Butler, 1867</bibRefCitation>
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</taxonomicName>
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, and designated
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<taxonomicName id="1CD04D477841FFF06F52FED7FC64E8EC" authorityName="Moschler" authorityYear="1877" box="[779,896,374,398]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="E9A4EAD67841FFF06F52FED7FC64E8EC" box="[779,896,374,398]" italics="true" pageId="8" pageNumber="7">T. blanda</emphasis>
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</taxonomicName>
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as the
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<typeStatus id="046B88667841FFF06F8CFED7FBEEE8ED" box="[981,1034,375,399]" pageId="8" pageNumber="7">type</typeStatus>
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species of the genus, even though
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<taxonomicName id="1CD04D477841FFF06C99FE37FEDDE8CC" authorityName="Moschler" authorityYear="1877" box="[192,313,406,430]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="E9A4EAD67841FFF06C99FE37FEDDE8CC" box="[192,313,406,430]" italics="true" pageId="8" pageNumber="7">T. blanda</emphasis>
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</taxonomicName>
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was considered a subspecies of
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<taxonomicName id="1CD04D477841FFF06E9EFE37FBFDE8CC" authority="Bates, 1866" authorityName="Bates" authorityYear="1866" box="[711,1049,406,430]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="sylvia">
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<emphasis id="E9A4EAD67841FFF06E9EFE37FC9BE8CC" box="[711,895,406,430]" italics="true" pageId="8" pageNumber="7">Taygetis sylvia</emphasis>
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Bates, 1866
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</taxonomicName>
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by
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<bibRefCitation id="BF414B357841FFF0681EFE37FAE3E8CD" author="Weymer, G." box="[1095,1287,407,431]" pageId="8" pageNumber="7" pagination="173 - 283" refId="ref20448" refString="Weymer, G. 1910 - 1913. 4. Familie: Satyridae. In: A. Seitz (ed.). Die Gross-Schmetterlinge der Erde. Stuttgart, A. Kernen. 5: 173 - 283." type="journal article" year="1910">Weymer (1910)</bibRefCitation>
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. Concerning
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<taxonomicName id="1CD04D477841FFF06C99FE17FE07E8AC" authority=", Forster (1964)" authorityName=", Forster" authorityYear="1964" box="[192,483,438,462]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="E9A4EAD67841FFF06C99FE17FECDE8AC" box="[192,297,439,462]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
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,
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<bibRefCitation id="BF414B357841FFF06D6EFE17FE07E8AC" author="Forster, W." box="[311,483,438,462]" pageId="8" pageNumber="7" pagination="51 - 188" refId="ref18607" refString="Forster, W. 1964. Beitrage zur Kenntnis der Insektenfauna Boliviens XIX Lepidoptera III. Satyridae. Veroffentlichungen der Zoologischen Staatssammlung Munchen 8: 51 - 188." type="journal article" year="1964">Forster (1964)</bibRefCitation>
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</taxonomicName>
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stated “Characteristic for this genus is mainly the male genitalia with the narrow tip of the valva”, providing no comparison with other genera. Subsequently,
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<bibRefCitation id="BF414B357841FFF068D7FE77FAD0E88D" author="Lamas, G." box="[1166,1332,471,495]" pageId="8" pageNumber="7" pagination="217 - 223" refId="ref19473" refString="Lamas, G. 2004. Nymphalidae. Satyrinae. Tribe Satyrini. Subtribe Euptychiina, pp. 217 - 223. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea-Papilionoidea. In: J. B. Heppner (ed.). Atlas of Neotropical Lepidoptera. Volume 5 A. Association for Tropical Lepidoptera, Scientific Publishers; Gainesville, FL. [xxxvi] + 439 p." type="book chapter" year="2004">Lamas (2004)</bibRefCitation>
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followed this classification and also included
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<taxonomicName id="1CD04D477841FFF06E2AFE56FC32EB6D" authority="Butler, 1870" authorityName="Butler" authorityYear="1870" box="[627,982,502,527]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="oreba">
|
|
<emphasis id="E9A4EAD67841FFF06E2AFE56FCDDEB6C" box="[627,825,502,526]" italics="true" pageId="8" pageNumber="7">Euptychia oreba</emphasis>
|
|
Butler, 1870
|
|
</taxonomicName>
|
|
(regarding
|
|
<taxonomicName id="1CD04D477841FFF0683BFE56FF1FEB4D" authority="Butler, 1870" authorityName="Butler" authorityYear="1870" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="gulnare">
|
|
<emphasis id="E9A4EAD67841FFF0683BFE56FAA7EB6C" box="[1122,1347,502,526]" italics="true" pageId="8" pageNumber="7">Euptychia gulnare</emphasis>
|
|
Butler, 1870
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="1CD04D477841FFF06D6DFDB6FD54EB4D" authority="Weymer, 1911" authorityName="Weymer" authorityYear="1911" box="[308,688,534,559]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="civica">
|
|
<emphasis id="E9A4EAD67841FFF06D6DFDB6FE18EB4C" box="[308,508,534,558]" italics="true" pageId="8" pageNumber="7">Euptychia civica</emphasis>
|
|
Weymer, 1911
|
|
</taxonomicName>
|
|
(
|
|
<emphasis id="E9A4EAD67841FFF06EE7FDB7FC88EB4C" box="[702,876,534,558]" italics="true" pageId="8" pageNumber="7">nomen nudum</emphasis>
|
|
) as junior subjective synonyms of this species),
|
|
<taxonomicName id="1CD04D477841FFF06C99FD96FDBAEB2D" authority="Weymer, 1911" authorityName="Weymer" authorityYear="1911" box="[192,606,566,591]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="griseola">
|
|
<emphasis id="E9A4EAD67841FFF06C99FD96FE41EB2C" box="[192,421,566,590]" italics="true" pageId="8" pageNumber="7">Euptychia griseola</emphasis>
|
|
Weymer, 1911
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="1CD04D477841FFF06EC4FD97FC3BEB2C" authority="Kheil, 1896" authorityName=": Kheil" authorityYear="1896" box="[669,991,566,590]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="vrazi">
|
|
<emphasis id="E9A4EAD67841FFF06EC4FD97FCADEB2C" box="[669,841,567,590]" italics="true" pageId="8" pageNumber="7">Taygetis vrazi</emphasis>
|
|
Kheil, 1896
|
|
</taxonomicName>
|
|
as members of
|
|
<taxonomicName id="1CD04D477841FFF068F8FD97FAEEEB2C" authorityName=", Forster" authorityYear="1964" box="[1185,1290,567,590]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF068F8FD97FAEEEB2C" box="[1185,1290,567,590]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
, in addition to proposing
|
|
<taxonomicName id="1CD04D477841FFF06D04FDF6FD10EB0D" authority="Weymer, 1911" authorityName="Weymer" authorityYear="1911" box="[349,756,598,623]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="eremita">
|
|
<emphasis id="E9A4EAD67841FFF06D04FDF6FDDAEB0C" box="[349,574,598,622]" italics="true" pageId="8" pageNumber="7">Euptychia eremita</emphasis>
|
|
Weymer, 1911
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="1CD04D477841FFF06F68FDF7FB48EB0D" authority="Ribeiro, 1931" authorityName="Ribeiro" authorityYear="1931" box="[817,1196,598,623]" class="Insecta" family="Nymphalidae" genus="Taygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="indecisa">
|
|
<emphasis id="E9A4EAD67841FFF06F68FDF7FBE7EB0C" box="[817,1027,598,622]" italics="true" pageId="8" pageNumber="7">Taygetis indecisa</emphasis>
|
|
Ribeiro, 1931
|
|
</taxonomicName>
|
|
as junior subjective synonyms of
|
|
<taxonomicName id="1CD04D477841FFF06D03FDD7FE3CEBEC" authorityName="Butler" authorityYear="1867" box="[346,472,630,654]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="obscura">
|
|
<emphasis id="E9A4EAD67841FFF06D03FDD7FE3CEBEC" box="[346,472,630,654]" italics="true" pageId="8" pageNumber="7">E. obscura</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<emphasis id="E9A4EAD67841FFF06E48FDD7FD71EBEC" box="[529,661,630,654]" italics="true" pageId="8" pageNumber="7">
|
|
<taxonomicName id="1CD04D477841FFF06E48FDD7FD74EBEC" authorityName="Butler" authorityYear="1870" box="[529,656,630,654]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="gulnare">E. gulnare</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
respectively.
|
|
<bibRefCitation id="BF414B357841FFF06F61FDD7FC38EBED" author="Lamas, G." box="[824,988,631,655]" pageId="8" pageNumber="7" pagination="217 - 223" refId="ref19473" refString="Lamas, G. 2004. Nymphalidae. Satyrinae. Tribe Satyrini. Subtribe Euptychiina, pp. 217 - 223. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea-Papilionoidea. In: J. B. Heppner (ed.). Atlas of Neotropical Lepidoptera. Volume 5 A. Association for Tropical Lepidoptera, Scientific Publishers; Gainesville, FL. [xxxvi] + 439 p." type="book chapter" year="2004">Lamas (2004)</bibRefCitation>
|
|
also listed four undescribed species of
|
|
<taxonomicName id="1CD04D477841FFF06C99FD37FECDEBCC" authorityName=", Forster" authorityYear="1964" box="[192,297,663,686]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06C99FD37FECDEBCC" box="[192,297,663,686]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
, of which two (
|
|
<taxonomicName id="1CD04D477841FFF06D84FD37FD93EBCC" authority="Nakahara & Willmott & Mielke & Schwartz & Zacca & Espeland & Lamas, 2018" authorityName="Nakahara & Willmott & Mielke & Schwartz & Zacca & Espeland & Lamas" authorityYear="2018" box="[477,631,663,686]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="argentata" status="sp. nov.">
|
|
<emphasis id="E9A4EAD67841FFF06D84FD37FD93EBCC" box="[477,631,663,686]" italics="true" pageId="8" pageNumber="7">H. argentata</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="E9A4EAD67841FFF06E24FD36FD27EBCC" bold="true" box="[637,707,662,686]" pageId="8" pageNumber="7">
|
|
<taxonomicNameLabel id="F29757AD7841FFF06E24FD36FD27EBCC" box="[637,707,662,686]" pageId="8" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
|
|
</emphasis>
|
|
and
|
|
<taxonomicName id="1CD04D477841FFF06EA4FD37FC34EBCC" authority="Nakahara & Willmott & Mielke & Schwartz & Zacca & Espeland & Lamas, 2018" authorityName="Nakahara & Willmott & Mielke & Schwartz & Zacca & Espeland & Lamas" authorityYear="2018" box="[765,976,663,686]" class="Insecta" family="Nymphalidae" genus="Orotaygetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="surui" status="n. gen. et n. sp.">
|
|
<emphasis id="E9A4EAD67841FFF06EA4FD37FC34EBCC" box="[765,976,663,686]" italics="true" pageId="8" pageNumber="7">Orotaygetis surui</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="E9A4EAD67841FFF06F8FFD36FB78EBCC" bold="true" box="[982,1180,662,686]" pageId="8" pageNumber="7">
|
|
<taxonomicNameLabel id="F29757AD7841FFF06F8FFD36FB78EBCC" box="[982,1180,662,686]" pageId="8" pageNumber="7" rank="species">n. gen. et n. sp.</taxonomicNameLabel>
|
|
</emphasis>
|
|
) are described below.
|
|
</paragraph>
|
|
<paragraph id="DB6F36C47841FFF06CB1FD16FBC4EAAD" blockId="8.[192,1441,342,975]" pageId="8" pageNumber="7">
|
|
<bibRefCitation id="BF414B357841FFF06CB1FD16FE34EBAC" author="Freitas, A. V. L. & E. Barbosa & K. R. Willmott & N. Wahlberg & G. Lamas" box="[232,464,694,718]" pageId="8" pageNumber="7" pagination="157 - 165" refId="ref18763" refString="Freitas, A. V. L., E. Barbosa, K. R. Willmott, N. Wahlberg, and G. Lamas. 2016. ' Species' from two different butterfly genera combined into one: description of a new genus of Euptychiina (Nymphalidae: Satyrinae) with unusually variable wing pattern. Revista Brasileira de Entomologia 60 (2): 157 - 165." type="journal article" year="2016">Freitas et al. (2016)</bibRefCitation>
|
|
recently treated
|
|
<taxonomicName id="1CD04D477841FFF06EC0FD17FCF2EBAC" authorityName="Weymer" authorityYear="1911" box="[665,790,694,718]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="griseola">
|
|
<emphasis id="E9A4EAD67841FFF06EC0FD17FCF2EBAC" box="[665,790,694,718]" italics="true" pageId="8" pageNumber="7">E. griseola</emphasis>
|
|
</taxonomicName>
|
|
as a synonym of
|
|
<taxonomicName id="1CD04D477841FFF06F84FD17FAFCEBAD" authority="Weymer, 1911" authorityName="Weymer" authorityYear="1911" box="[989,1304,695,719]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="punctata">
|
|
<emphasis id="E9A4EAD67841FFF06F84FD17FB83EBAC" box="[989,1127,695,718]" italics="true" pageId="8" pageNumber="7">E. punctata</emphasis>
|
|
Weymer, 1911
|
|
</taxonomicName>
|
|
and moved this taxon out of
|
|
<taxonomicName id="1CD04D477841FFF06DD7FD77FE13EB8C" authorityName=", Forster" authorityYear="1964" box="[398,503,727,750]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06DD7FD77FE13EB8C" box="[398,503,727,750]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
by placing it in its own monotypic genus,
|
|
<taxonomicName id="1CD04D477841FFF06FAFFD77FA78EB8C" authority="Freitas and Barbosa, 2016" authorityName="Freitas and Barbosa" authorityYear="2016" box="[1014,1436,726,750]" class="Insecta" family="Nymphalidae" genus="Sepona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06FAFFD77FBABEB8C" box="[1014,1103,727,750]" italics="true" pageId="8" pageNumber="7">Sepona</emphasis>
|
|
<bibRefCitation id="BF414B357841FFF0680FFD76FA78EB8C" author="Freitas, A. V. L. & E. Barbosa & K. R. Willmott & N. Wahlberg & G. Lamas" box="[1110,1436,726,750]" pageId="8" pageNumber="7" pagination="157 - 165" refId="ref18763" refString="Freitas, A. V. L., E. Barbosa, K. R. Willmott, N. Wahlberg, and G. Lamas. 2016. ' Species' from two different butterfly genera combined into one: description of a new genus of Euptychiina (Nymphalidae: Satyrinae) with unusually variable wing pattern. Revista Brasileira de Entomologia 60 (2): 157 - 165." type="journal article" year="2016">Freitas and Barbosa, 2016</bibRefCitation>
|
|
</taxonomicName>
|
|
, based on molecular and morphological data. As discussed above,
|
|
<taxonomicName id="1CD04D477841FFF06F90FD56FB46EA6C" authorityName="Nakahara & Willmott & Mielke & Schwartz & Zacca & Espeland & Lamas" authorityYear="2018" baseAuthorityName="Kheil" baseAuthorityYear="1896" box="[969,1186,758,782]" class="Insecta" family="Nymphalidae" genus="Pseudodebis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="vrazi" status="comb. nov.">
|
|
<emphasis id="E9A4EAD67841FFF06F90FD56FB46EA6C" box="[969,1186,758,782]" italics="true" pageId="8" pageNumber="7">Pseudodebis vrazi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="E9A4EAD67841FFF068FEFD56FAFCEA6C" bold="true" box="[1191,1304,758,782]" pageId="8" pageNumber="7">
|
|
<taxonomicNameLabel id="F29757AD7841FFF068FEFD56FAFCEA6C" box="[1191,1304,758,782]" pageId="8" pageNumber="7" rank="species">n. comb.</taxonomicNameLabel>
|
|
</emphasis>
|
|
is removed from
|
|
<taxonomicName id="1CD04D477841FFF06CA6FCB7FE8CEA4C" authorityName=", Forster" authorityYear="1964" box="[255,360,791,814]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06CA6FCB7FE8CEA4C" box="[255,360,791,814]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
and considered conspecific with one of the new species of
|
|
<taxonomicName id="1CD04D477841FFF06841FCB7FB65EA4C" authorityName=", Forster" authorityYear="1964" box="[1048,1153,791,814]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06841FCB7FB65EA4C" box="[1048,1153,791,814]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
listed by
|
|
<bibRefCitation id="BF414B357841FFF068AAFCB7FA78EA4D" author="Lamas, G." box="[1267,1436,791,815]" pageId="8" pageNumber="7" pagination="217 - 223" refId="ref19473" refString="Lamas, G. 2004. Nymphalidae. Satyrinae. Tribe Satyrini. Subtribe Euptychiina, pp. 217 - 223. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea-Papilionoidea. In: J. B. Heppner (ed.). Atlas of Neotropical Lepidoptera. Volume 5 A. Association for Tropical Lepidoptera, Scientific Publishers; Gainesville, FL. [xxxvi] + 439 p." type="book chapter" year="2004">Lamas (2004)</bibRefCitation>
|
|
. Three species, namely
|
|
<taxonomicName id="1CD04D477841FFF06D85FC97FD40EA2C" baseAuthorityName="Moschler" baseAuthorityYear="1877" box="[476,676,822,846]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="blanda">
|
|
<emphasis id="E9A4EAD67841FFF06D85FC97FD40EA2C" box="[476,676,822,846]" italics="true" pageId="8" pageNumber="7">Harjesia blanda</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="1CD04D477841FFF06EECFC97FCDFEA2C" authorityName="Butler" authorityYear="1867" box="[693,827,822,846]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="obscura">
|
|
<emphasis id="E9A4EAD67841FFF06EECFC97FCDFEA2C" box="[693,827,822,846]" italics="true" pageId="8" pageNumber="7">H. obscura</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="1CD04D477841FFF06F22FC97FBFCEA2C" authority="Nakahara & Willmott & Mielke & Schwartz & Zacca & Espeland & Lamas, 2018" authorityName="Nakahara & Willmott & Mielke & Schwartz & Zacca & Espeland & Lamas" authorityYear="2018" box="[891,1048,823,846]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="argentata" status="sp. nov.">
|
|
<emphasis id="E9A4EAD67841FFF06F22FC97FBFCEA2C" box="[891,1048,823,846]" italics="true" pageId="8" pageNumber="7">H. argentata</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="E9A4EAD67841FFF0687BFC96FB8FEA2C" bold="true" box="[1058,1131,822,846]" pageId="8" pageNumber="7">
|
|
<taxonomicNameLabel id="F29757AD7841FFF0687BFC96FB8FEA2C" box="[1058,1131,822,846]" pageId="8" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
|
|
</emphasis>
|
|
, do form a monophyletic group with a high support (bs = 99, see
|
|
<figureCitation id="43EB2A417841FFF06EF1FCF6FD0FEA0D" box="[680,747,854,879]" captionStart="Figure 1" captionStartId="24.[195,287,1840,1864]" captionTargetId="graphics@24.[181,1058,697,1588]" captionTargetPageId="24" captionText="Figure 1. Phylogeny of selected taxa of the “Taygetis clade” sensu Peña et al. (2010) based on maximum likelihood with support shown as non-parametric bootstrap. Taxa discussed or described in this paper are highlighted in bold." figureDoi="http://doi.org/10.5281/zenodo.3708154" httpUri="https://zenodo.org/record/3708154/files/figure.png" pageId="8" pageNumber="7">Fig. 1</figureCitation>
|
|
), and we regard
|
|
<taxonomicName id="1CD04D477841FFF06FE2FCF7FBC0EA0C" authorityName=", Forster" authorityYear="1964" box="[955,1060,855,878]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06FE2FCF7FBC0EA0C" box="[955,1060,855,878]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
as a small monophyletic genus to accommodate these three species. Finally, the remaining species in
|
|
<taxonomicName id="1CD04D477841FFF0686CFCD7FA44EAED" authority="(sensu Lamas 2004)" baseAuthorityName="sensu Lamas" baseAuthorityYear="2004" box="[1077,1440,887,911]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF0686CFCD7FB7AEAEC" box="[1077,1182,887,910]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
(
|
|
<emphasis id="E9A4EAD67841FFF068E8FCD7FB11EAEC" box="[1201,1269,887,910]" italics="true" pageId="8" pageNumber="7">sensu</emphasis>
|
|
<bibRefCitation id="BF414B357841FFF068A6FCD7FA7CEAED" author="Lamas, G." box="[1279,1432,887,911]" pageId="8" pageNumber="7" pagination="217 - 223" refId="ref19473" refString="Lamas, G. 2004. Nymphalidae. Satyrinae. Tribe Satyrini. Subtribe Euptychiina, pp. 217 - 223. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea-Papilionoidea. In: J. B. Heppner (ed.). Atlas of Neotropical Lepidoptera. Volume 5 A. Association for Tropical Lepidoptera, Scientific Publishers; Gainesville, FL. [xxxvi] + 439 p." type="book chapter" year="2004">Lamas 2004</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
appear not to belong in the clade containing the
|
|
<typeStatus id="046B88667841FFF06F50FC37FCDAEACD" box="[777,830,919,943]" pageId="8" pageNumber="7">type</typeStatus>
|
|
for the genus (see statement above), and thus will need to be transferred to another genus (Espeland et al. unpubl. data).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="93CA654F7841FFF06C99FC46FBCAECFC" pageId="8" pageNumber="7" type="diagnosis">
|
|
<paragraph id="DB6F36C47841FFF06C99FC46FBCAECFC" blockId="8.[192,1440,998,1439]" pageId="8" pageNumber="7">
|
|
<emphasis id="E9A4EAD67841FFF06C99FC46FD5DEA9C" bold="true" box="[192,697,998,1023]" pageId="8" pageNumber="7">Systematic placement and diagnosis.</emphasis>
|
|
<taxonomicName id="1CD04D477841FFF06E99FC47FCCDEA9C" baseAuthorityName="sensu Lamas" baseAuthorityYear="2004" box="[704,809,999,1022]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06E99FC47FCCDEA9C" box="[704,809,999,1022]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
appears to be a member of a clade consisting of
|
|
<taxonomicName id="1CD04D477841FFF06935FC47FE23ED7D" authority="Gray, 1973" authorityName="Gray" authorityYear="1973" class="Insecta" family="Nymphalidae" genus="Forsterinaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06935FC47FEDDED7C" italics="true" pageId="8" pageNumber="7">Forsterinaria</emphasis>
|
|
Gray, 1973
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="1CD04D477841FFF06D88FBA7FCE9ED7D" authority="Forster, 1964" authorityName="Forster" authorityYear="1964" box="[465,781,1031,1055]" class="Insecta" family="Nymphalidae" genus="Posttaygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06D88FBA7FD87ED7C" box="[465,611,1031,1054]" italics="true" pageId="8" pageNumber="7">Posttaygetis</emphasis>
|
|
<bibRefCitation id="BF414B357841FFF06E31FBA7FCE9ED7D" author="Forster, W." box="[616,781,1031,1055]" pageId="8" pageNumber="7" pagination="51 - 188" refId="ref18607" refString="Forster, W. 1964. Beitrage zur Kenntnis der Insektenfauna Boliviens XIX Lepidoptera III. Satyridae. Veroffentlichungen der Zoologischen Staatssammlung Munchen 8: 51 - 188." type="journal article" year="1964">Forster, 1964</bibRefCitation>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="1CD04D477841FFF06F12FBA7FB68ED7D" authority="Forster, 1964" authorityName="Forster" authorityYear="1964" box="[843,1164,1031,1055]" class="Insecta" family="Nymphalidae" genus="Parataygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06F12FBA7FC01ED7C" box="[843,997,1031,1054]" italics="true" pageId="8" pageNumber="7">Parataygetis</emphasis>
|
|
<bibRefCitation id="BF414B357841FFF06FB3FBA7FB68ED7D" author="Forster, W." box="[1002,1164,1031,1055]" pageId="8" pageNumber="7" pagination="51 - 188" refId="ref18607" refString="Forster, W. 1964. Beitrage zur Kenntnis der Insektenfauna Boliviens XIX Lepidoptera III. Satyridae. Veroffentlichungen der Zoologischen Staatssammlung Munchen 8: 51 - 188." type="journal article" year="1964">Forster, 1964</bibRefCitation>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="43EB2A417841FFF068C2FBA7FB39ED7D" box="[1179,1245,1031,1055]" captionStart="Figure 1" captionStartId="24.[195,287,1840,1864]" captionTargetId="graphics@24.[181,1058,697,1588]" captionTargetPageId="24" captionText="Figure 1. Phylogeny of selected taxa of the “Taygetis clade” sensu Peña et al. (2010) based on maximum likelihood with support shown as non-parametric bootstrap. Taxa discussed or described in this paper are highlighted in bold." figureDoi="http://doi.org/10.5281/zenodo.3708154" httpUri="https://zenodo.org/record/3708154/files/figure.png" pageId="8" pageNumber="7">Fig. 1</figureCitation>
|
|
), but support in this part of the tree is low.
|
|
<taxonomicName id="1CD04D477841FFF06E5CFB87FD8AED5C" baseAuthorityName="sensu Lamas" baseAuthorityYear="2004" box="[517,622,1063,1086]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06E5CFB87FD8AED5C" box="[517,622,1063,1086]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="1CD04D477841FFF06F25FB87FBFBED5C" authorityName="Gray" authorityYear="1973" box="[892,1055,1063,1086]" class="Insecta" family="Nymphalidae" genus="Forsterinaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06F25FB87FBFBED5C" box="[892,1055,1063,1086]" italics="true" pageId="8" pageNumber="7">Forsterinaria</emphasis>
|
|
</taxonomicName>
|
|
by having a ringed ocellus with white pupil in VFW cell M
|
|
<subScript id="475434817841FFF06E58FBF6FDEEED06" attach="left" box="[513,522,1110,1124]" fontSize="6" pageId="8" pageNumber="7">1</subScript>
|
|
and often in adjacent cells, whereas these ocelli appear just as white pupils or a white patch in
|
|
<taxonomicName id="1CD04D477841FFF06DE9FBC7FDB7ED1C" authorityName="Gray" authorityYear="1973" box="[432,595,1127,1150]" class="Insecta" family="Nymphalidae" genus="Forsterinaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06DE9FBC7FDB7ED1C" box="[432,595,1127,1150]" italics="true" pageId="8" pageNumber="7">Forsterinaria</emphasis>
|
|
</taxonomicName>
|
|
; ocelli in VHW cells M
|
|
<subScript id="475434817841FFF06F32FBD6FC90EDE6" attach="left" box="[875,884,1142,1156]" fontSize="6" pageId="8" pageNumber="7">2</subScript>
|
|
and M
|
|
<subScript id="475434817841FFF06F93FBD6FC37EDE6" attach="left" box="[970,979,1142,1156]" fontSize="6" pageId="8" pageNumber="7">3</subScript>
|
|
are ringed, and in
|
|
<taxonomicName id="1CD04D477841FFF068E7FBC7FAC3ED1C" baseAuthorityName="sensu Lamas" baseAuthorityYear="2004" box="[1214,1319,1127,1150]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF068E7FBC7FAC3ED1C" box="[1214,1319,1127,1150]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
these are large, whereas in
|
|
<taxonomicName id="1CD04D477841FFF06DC7FB27FDA5EDFC" authorityName="Gray" authorityYear="1973" box="[414,577,1159,1182]" class="Insecta" family="Nymphalidae" genus="Forsterinaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06DC7FB27FDA5EDFC" box="[414,577,1159,1182]" italics="true" pageId="8" pageNumber="7">Forsterinaria</emphasis>
|
|
</taxonomicName>
|
|
they are small or appear just as simple white pupils; the brachia are about half the length of the uncus in
|
|
<taxonomicName id="1CD04D477841FFF06E24FB07FD02EDDC" baseAuthorityName="sensu Lamas" baseAuthorityYear="2004" box="[637,742,1191,1214]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06E24FB07FD02EDDC" box="[637,742,1191,1214]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
(except for
|
|
<taxonomicName id="1CD04D477841FFF06F2BFB07FC11EDDC" authorityName="Butler" authorityYear="1867" box="[882,1013,1190,1214]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="obscura">
|
|
<emphasis id="E9A4EAD67841FFF06F2BFB07FC11EDDC" box="[882,1013,1190,1214]" italics="true" pageId="8" pageNumber="7">H. obscura</emphasis>
|
|
</taxonomicName>
|
|
, in which the brachia are reduced), whereas the brachia are absent or reduced in
|
|
<taxonomicName id="1CD04D477841FFF06EBEFB67FC6EEDBC" authorityName="Gray" authorityYear="1973" box="[743,906,1223,1246]" class="Insecta" family="Nymphalidae" genus="Forsterinaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06EBEFB67FC6EEDBC" box="[743,906,1223,1246]" italics="true" pageId="8" pageNumber="7">Forsterinaria</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<taxonomicName id="1CD04D477841FFF06FC1FB67FBE5EDBC" baseAuthorityName="sensu Lamas" baseAuthorityYear="2004" box="[920,1025,1223,1246]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06FC1FB67FBE5EDBC" box="[920,1025,1223,1246]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="1CD04D477841FFF06957FB67FA44EDBC" authorityName="Forster" authorityYear="1964" box="[1294,1440,1223,1246]" class="Insecta" family="Nymphalidae" genus="Posttaygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06957FB67FA44EDBC" box="[1294,1440,1223,1246]" italics="true" pageId="8" pageNumber="7">Posttaygetis</emphasis>
|
|
</taxonomicName>
|
|
by the absence of a horizontal creamy band in the VHW discal cell that also extends along M
|
|
<subScript id="475434817841FFF06972FB56FAD0EC66" attach="left" box="[1323,1332,1270,1284]" fontSize="6" pageId="8" pageNumber="7">3</subScript>
|
|
, in addition to the absence of a reduced and basally placed ocellus in VHW cell M
|
|
<subScript id="475434817841FFF06816FAB6FBBCEC46" attach="left" box="[1103,1112,1302,1316]" fontSize="6" pageId="8" pageNumber="7">2</subScript>
|
|
, whereas these characters are present in
|
|
<taxonomicName id="1CD04D477841FFF06D2DFA87FDE2EC5C" authorityName="Forster" authorityYear="1964" box="[372,518,1319,1342]" class="Insecta" family="Nymphalidae" genus="Posttaygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06D2DFA87FDE2EC5C" box="[372,518,1319,1342]" italics="true" pageId="8" pageNumber="7">Posttaygetis</emphasis>
|
|
</taxonomicName>
|
|
. As mentioned above, the brachia are about half the length of the uncus in
|
|
<taxonomicName id="1CD04D477841FFF06C99FAE7FECDEC3C" baseAuthorityName="sensu Lamas" baseAuthorityYear="2004" box="[192,297,1351,1374]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06C99FAE7FECDEC3C" box="[192,297,1351,1374]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
(except for
|
|
<taxonomicName id="1CD04D477841FFF06DE8FAE7FDD6EC3C" authorityName="Butler" authorityYear="1867" box="[433,562,1350,1374]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="obscura">
|
|
<emphasis id="E9A4EAD67841FFF06DE8FAE7FDD6EC3C" box="[433,562,1350,1374]" italics="true" pageId="8" pageNumber="7">H. obscura</emphasis>
|
|
</taxonomicName>
|
|
), whereas the brachia are absent or reduced in
|
|
<taxonomicName id="1CD04D477841FFF0683FFAE7FB13EC3C" authorityName="Forster" authorityYear="1964" box="[1126,1271,1351,1374]" class="Insecta" family="Nymphalidae" genus="Posttaygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF0683FFAE7FB13EC3C" box="[1126,1271,1351,1374]" italics="true" pageId="8" pageNumber="7">Posttaygetis</emphasis>
|
|
</taxonomicName>
|
|
. Among other characters,
|
|
<taxonomicName id="1CD04D477841FFF06D09FAC7FE5DEC1C" baseAuthorityName="sensu Lamas" baseAuthorityYear="2004" box="[336,441,1383,1406]" class="Insecta" family="Nymphalidae" genus="Harjesia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06D09FAC7FE5DEC1C" box="[336,441,1383,1406]" italics="true" pageId="8" pageNumber="7">Harjesia</emphasis>
|
|
</taxonomicName>
|
|
is distinguished from
|
|
<taxonomicName id="1CD04D477841FFF06E95FAC7FC82EC1C" authorityName="Forster" authorityYear="1964" box="[716,870,1383,1406]" class="Insecta" family="Nymphalidae" genus="Parataygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06E95FAC7FC82EC1C" box="[716,870,1383,1406]" italics="true" pageId="8" pageNumber="7">Parataygetis</emphasis>
|
|
</taxonomicName>
|
|
by having a narrow, rounded apical process of the valva, whereas the distal side is broad and serrated in
|
|
<taxonomicName id="1CD04D477841FFF06FD7FA27FBCCECFC" authorityName="Forster" authorityYear="1964" box="[910,1064,1415,1438]" class="Insecta" family="Nymphalidae" genus="Parataygetis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E9A4EAD67841FFF06FD7FA27FBCCECFC" box="[910,1064,1415,1438]" italics="true" pageId="8" pageNumber="7">Parataygetis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |