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<mods:title id="72607D06EEEE2245592AA7AB06B358D1">Five new species of the genus Hermonassa Walker, 1865 from Xizang Autonomous Region, China (Lepidoptera, Noctuidae, Noctuinae)</mods:title>
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<mods:namePart id="A69F48E9A25CA775613BEC7005261B5E">Gao, Biao</mods:namePart>
<mods:affiliation id="61F1677FDACF8F7413D7F265715B42CE">Northeast Forestry University, School of Forestry, Harbin 150040, China</mods:affiliation>
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<mods:namePart id="2373EF89B8FD4F93BF35E020D119C089">Han, Hui-Lin</mods:namePart>
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<mods:affiliation id="09B66A3ADC9AFA4EAFF9D1B50F7968BF">Northeast Forestry University, School of Forestry, Harbin 150040, China</mods:affiliation>
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<mods:namePart id="C2D6CF8C074D0E75C51F95FA413A2249">Kononenko, Vladimir S.</mods:namePart>
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<mods:affiliation id="BA93FF33A43696A51B0BE8AEEAEFCCDC">Northeast Asia Biodiversity Research Center, Northeast Forestry University, Harbin 15004, China</mods:affiliation>
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<mods:namePart id="996D22D0DB83E2A18948B320919DCBD8">Pan, Zhao-Hui</mods:namePart>
<mods:affiliation id="E234FCACE4239BED918EE41894A89321">Northeast Forestry University, Ministry of Education, Key Laboratory of Sustainable Forest Ecosystem Management, Harbin 150040, China</mods:affiliation>
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<subSubSection id="FDF1D0DCFCC3EFD614BB2108D6105970" pageId="0" pageNumber="35" type="nomenclature">
<paragraph id="11C0B8908F3EF6744CFAB69CE9AB0E8B" pageId="0" pageNumber="35">
<taxonomicName id="3905C34F76868E13DCBCDB7A47FCB078" LSID="979210C1-BC8F-564C-AF70-1370A203BB94" authority="Sugi, 1995" authorityName="Sugi" authorityYear="1995" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="shizukoae">Hermonassa shizukoae Sugi, 1995</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="589F5CAD30AD101C2D4BEFD039E7662F" pageId="0" pageNumber="35" type="description">
<paragraph id="EC4CDEEB622CE8863DB50A23A0BAFF6F" pageId="0" pageNumber="35">
<figureCitation id="3124A4E81F0F55576200F5659CFCB36F" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">Figs 10</figureCitation>
<figureCitation id="40F0C8EDF71A68454987D5490EDFA23C" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">, 21</figureCitation>
<figureCitation id="FC456689EF00D707D2B5AC3EC30EDF2E" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">, 22</figureCitation>
<figureCitation id="BB533B3E35263ABB4C78B6C887B4756D" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">, 27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="78E650183C58539DCC300D326A087479" pageId="0" pageNumber="35" type="reference_group">
<paragraph id="B97D1ECCF6CE150D52144E071C19A795" pageId="0" pageNumber="35">
<taxonomicName id="5B710DAF031BB6789BE69BD5E7E09FCC" authorityName="Sugi" authorityYear="1995" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="shizukoae">Hermonassa shizukoae</taxonomicName>
Sugi, 1995, Tinea 14, (suppl. 2): 92, pl. 117, fig. 17;
<bibRefCitation id="C254063D30AF28851B982B39598049BB" author="Hreblay, M" editor="Haruta, T" journalOrPublisher="V. Tinea 15 (suppl. 1)" pageId="0" pageNumber="35" pagination="117 - 310" refId="B13" refString="Hreblay, M, Ronkay, L, 1998. Noctuidae from Nepal. In: Haruta, T, Ed., Moths of Nepal Pt. V. Tinea 15 (suppl. 1): 117 - 310" title="Noctuidae from Nepal." volumeTitle="Moths of Nepal Pt." year="1998">Hreblay and Ronkay 1998</bibRefCitation>
: 133 (junior synonym of
<taxonomicName id="DD5BDCBC1711B2E6EEA5EB80FAFB50C9" family="Noctuidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">H. renifera</taxonomicName>
Chen, 1991);
<bibRefCitation id="3DBE3FB9E777A516381A4DE3F90D058C" author="Kovacs, ST" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="35" pagination="297 - 319" refId="B16" refString="Kovacs, ST, Ronkay, G, Ronkay, L, 2018. Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species. Revue Suisse de Zoologie 126 (2): 297 - 319" title="Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species." volume="126" year="2018">
<normalizedToken id="C8D58122CDF4D7945990D4771D78E05C" originalValue="Kovács">Kovacs</normalizedToken>
et al. 2018
</bibRefCitation>
: 306, pl. 3, figs 5-8, genit. figs 15, 16 (distinct species).
</paragraph>
</subSubSection>
<subSubSection id="903E9852CDEE6D54E962D1D2A97C6D16" pageId="0" pageNumber="35" type="type material">
<paragraph id="37A2C7AAA00402A595C9F5BACE545838" pageId="0" pageNumber="35">Type material.</paragraph>
<paragraph id="ACD7D7C6347B40F4FB545D05226C32A3" pageId="0" pageNumber="35">
<emphasis id="9FD16997A35DA9C6139BD7ACD142A17B" bold="true" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
: ♂, India, West Bengal, Sandadakphu, ca 50 km NW of Darjiling, 3600 m, 14.viii.1985(W.Thomas), genitalia slide No SS-5155 &quot;Shigero Sugi Collection&quot;,
<normalizedToken id="D902DDD953A348C7F7A011F6F657C096" originalValue="“HOLOTYPE”">&quot;HOLOTYPE&quot;</normalizedToken>
(red label),
<taxonomicName id="5568BF5921FB1C400957EACA6F773DC8" authorityName="Sugi" authorityYear="1995" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="shizukoae">
<emphasis id="1BBAEBC74E2836A5FDF7BA5F75EB8313" italics="true" pageId="0" pageNumber="35">Hermonassa shizukoae</emphasis>
</taxonomicName>
Sugi ♂, det. S. Sugi, 1995&quot; (coll. NIAES), examined.
</paragraph>
</subSubSection>
<subSubSection id="0D17132CB06FA33F4E034C253D9D02F4" pageId="0" pageNumber="35" type="distribution">
<paragraph id="3373F2A609FC638F88AF78CDF7B1E548" pageId="0" pageNumber="35">Distribution and biology.</paragraph>
<paragraph id="BBDC399088C5F6DC81D5A183422BDC8E" pageId="0" pageNumber="35">
The species is distributed in north India and Nepal where it is rather common at altitude ca 3600 m (
<bibRefCitation id="1B1D15AFFF759BDC1E57C3F7F724571B" author="Kovacs, ST" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="35" pagination="297 - 319" refId="B16" refString="Kovacs, ST, Ronkay, G, Ronkay, L, 2018. Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species. Revue Suisse de Zoologie 126 (2): 297 - 319" title="Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species." volume="126" year="2018">
<normalizedToken id="519E304488D893FDA7B78F6637480403" originalValue="Kovács">Kovacs</normalizedToken>
et al. 2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="36C93E8D2FF9F57B5C76135898271F65" pageId="0" pageNumber="35" type="remarks">
<paragraph id="426BF4853EBB1B33B3E76E2F53A851C7" pageId="0" pageNumber="35">Remarks.</paragraph>
<paragraph id="1866A7C7B69A943E7C789E4424B84DA7" pageId="0" pageNumber="35">
The species is the southern Himalayan representative of
<taxonomicName id="8F00FCB9F8A39295C8D64B4F32926B71" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
<emphasis id="F1948C3CFDB8CEE3CCB444F8F376C8FB" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
</taxonomicName>
-
<taxonomicName id="94044FFE92B15BCBF9BDE2DB43B21AE2" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
<emphasis id="C826D909B4FE9A04DC68204621E6A652" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
</taxonomicName>
species pair (
<bibRefCitation id="C833997C61612FA1C0E7D660404A0473" author="Kovacs, ST" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="35" pagination="297 - 319" refId="B16" refString="Kovacs, ST, Ronkay, G, Ronkay, L, 2018. Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species. Revue Suisse de Zoologie 126 (2): 297 - 319" title="Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species." volume="126" year="2018">
<normalizedToken id="7990FF6DE7F6FD411CFE9B98DFBCB232" originalValue="Kovács">Kovacs</normalizedToken>
et al. 2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8E7629EBFD772A505D3B6B3A623BA507" pageId="0" pageNumber="35" type="the cuprina species group">
<paragraph id="FB3468BFC35B9433E8F20B1A1D5453AA" pageId="0" pageNumber="35">
The
<taxonomicName id="22F4FA5057C45E1D83CFA8E9DABDFBFE" authorityName="Moore" authorityYear="1882" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa cuprina" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="cuprina">
<emphasis id="E4C1440F908B817D736868A1EA9AD21A" italics="true" pageId="0" pageNumber="35">Hermonassa cuprina</emphasis>
</taxonomicName>
species group
</paragraph>
<paragraph id="EF0C4A36DD66A15EDF4D4B353146D749" pageId="0" pageNumber="35">
The
<taxonomicName id="5C45DE568FF4416BA3B5C12195C89B44" authorityName="Moore" authorityYear="1882" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa cuprina" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="cuprina">
<emphasis id="C451BA65E541EC088B65B89777A46524" italics="true" pageId="0" pageNumber="35">Hermonassa cuprina</emphasis>
</taxonomicName>
species group includes two species,
<taxonomicName id="86C5CB80F9366C33AD64D2FEA50C585D" family="Noctuidae" kingdom="Animalia" lsidName="H. cuprina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="cuprina">
<emphasis id="7ABEFE982B8FAD39260D042290BBDCE4" italics="true" pageId="0" pageNumber="35">H. cuprina</emphasis>
</taxonomicName>
and
<taxonomicName id="FC2B4DC8529868739D8835BD43D0D22D" family="Noctuidae" kingdom="Animalia" lsidName="H. brunneocuprina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="brunneocuprina">
<emphasis id="1411768E7A6DD0F75AC5C83ED91D9352" italics="true" pageId="0" pageNumber="35">H. brunneocuprina</emphasis>
</taxonomicName>
sp. nov. This species group can be characterised by the uniform external appearance with a relatively large size, the narrow forewings, cupper-brown or brown colouration and by the forewing pattern with more or less clearly expressed thin basal, antemedial, and postmedial transverse lines and the presence of black contrasting claviform, and the orbicular and reniform stigmata. The main diagnostic features of the male genitalia are the relatively short, flat and apically dilated, spatulate uncus; juxta small, three leaved, dart-shaped, large U-shaped vinculum, the relatively narrow simple valva, thin, upcurved harpe, and the slightly curved aedeagus with a sclerotised comb in the carina. The female genitalia are characterised by the deeply split antevaginal plate, deep cup-like antrum, a thin, medium length ductus seminalis, and the presence of two longitudinal signa on the corpus bursae.
</paragraph>
</subSubSection>
</treatment>
</document>
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