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<document id="83DAA3F35167C4FD63687DA7F6383D4E" ID-CLB-Dataset="7212" ID-DOI="10.11646/zootaxa.4958.1.33" ID-GBIF-Dataset="00f4ae66-2696-4a36-be77-6de24695dcab" ID-ISSN="1175-5326" ID-Zenodo-Dep="4692469" ID-ZooBank="D586C16A-7D57-4BE6-A1DA-2CF7703E60BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618472714310" checkinUser="plazi" docAuthor="Roell, Talita, Brugnera, Ricardo &amp; Lemaître, Valérie A." docDate="2021" docId="894E3E5CD565FF9EF9AE9014B475F985" docLanguage="en" docName="zootaxa.4958.1.33.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Podisus graziae Brugnera, Roell &amp; Lemaitre 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="578" masterDocId="75774624D563FF96F939972DB623FF8C" masterDocTitle="Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae)" masterLastPageNumber="584" masterPageNumber="570" pageNumber="576" updateTime="1698936925256" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2F9D6FD28904F95385A503234B8446E7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8CE5EAE916689049D07CA60D7129296A">
<mods:title id="429ABB875B60904BDA5798363E8038FA">Lost and found-Discovery of the presumed lost type of Arma pallipes Dallas new synonymy and description of two new species of Podisus Herrich-Schäffer (Hemiptera: Pentatomidae: Asopinae)</mods:title>
</mods:titleInfo>
<mods:name id="35199C6009F4E8144F03D14CF1D6FD34" type="personal">
<mods:role id="A8FA0178028461D12A54A20266F15CDE">
<mods:roleTerm id="4715A0831D85984DB53CDFAEFD51BFCC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B13FD42845F578BC9560EF36D22936E">Roell, Talita</mods:namePart>
</mods:name>
<mods:name id="C50EE89F8DECEA4FB1F8666CEF21FAB1" type="personal">
<mods:role id="5E1C37814C77B679FD4CFC62A8E5162D">
<mods:roleTerm id="FEB7D4A61C633233B300F503444D3DCF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD5AB6FD8BBCAC50B19810B6F60AB971">Brugnera, Ricardo</mods:namePart>
<mods:affiliation id="3AE4344AE4679A6A66678C63FC7C0210">Universidade Federal do Rio Grande do Sul, Instituto de Biociências, Departamento de Zoologia, Av. Bento Gonçalves, 9500, Agronomia, 91501 - 970 Porto Alegre, RS, Brasil.</mods:affiliation>
</mods:name>
<mods:name id="79759E037BB41F91AFBCDFE9CD655999" type="personal">
<mods:role id="49E114C9AB74469B7BD38906BE229374">
<mods:roleTerm id="530404D901FAFB6CA280BB71EF693C47">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9572CC56C78826C18ABD80EEAAC9EBA7">Lemaître, Valérie A.</mods:namePart>
<mods:affiliation id="1558A25C7114705C68858E8F2D5A6E97">Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8399F425F981139042A0D680106BA621">text</mods:typeOfResource>
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<mods:titleInfo id="4100421ED8520D1F4A1F014C64DF76C8">
<mods:title id="E10BD472EE967F276FDE9201CB4C847C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="481E7332AA01B4D3721ECDB0385BCC12">
<mods:date id="1E3C3725A890B0F5D56A59F0A4AA0207">2021</mods:date>
<mods:detail id="3A13B759C7865B9321D114630C0985B5" type="pubDate">
<mods:number id="12C7F0F64171EE61F803E7FF451BBAB2">2021-04-14</mods:number>
</mods:detail>
<mods:detail id="DE31502CE8743D3620BAAAE839B3E8CC" type="volume">
<mods:number id="F60207C6A91D98C54B4E488CD7FAB410">4958</mods:number>
</mods:detail>
<mods:detail id="5D3B5328F37BC24223A38DDD597E3A91" type="issue">
<mods:number id="26CA34A60132E6AC82DD07C12AB829E8">1</mods:number>
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<mods:start id="C3B00B73D72D11D49CF0009BB811D391">570</mods:start>
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<mods:classification id="EB5E788F63D641EB64AC405939E36E46">journal article</mods:classification>
<mods:identifier id="E14A13B54C82B0BE920A5BF7B54668C1" type="CLB-Dataset">7212</mods:identifier>
<mods:identifier id="1A2965C7CD0DED1D30CADF4D8E8AD58F" type="DOI">10.11646/zootaxa.4958.1.33</mods:identifier>
<mods:identifier id="0DDE92FAC514E5364D172452AF9632B9" type="GBIF-Dataset">00f4ae66-2696-4a36-be77-6de24695dcab</mods:identifier>
<mods:identifier id="59E0974FA6E0A757E3B1F012FEE0329D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D9F8BE1E0579669F455F0E94C4972371" type="Zenodo-Dep">4692469</mods:identifier>
<mods:identifier id="38C911F13100995B720FE62231C20BB4" type="ZooBank">D586C16A-7D57-4BE6-A1DA-2CF7703E60BC</mods:identifier>
</mods:mods>
<treatment id="894E3E5CD565FF9EF9AE9014B475F985" ID-DOI="http://doi.org/10.5281/zenodo.4711395" ID-GBIF-Taxon="181531982" ID-Zenodo-Dep="4711395" LSID="urn:lsid:plazi:treatment:894E3E5CD565FF9EF9AE9014B475F985" httpUri="http://treatment.plazi.org/id/894E3E5CD565FF9EF9AE9014B475F985" lastPageId="8" lastPageNumber="578" pageId="6" pageNumber="576">
<subSubSection id="49FDDCC1D565FF90F9AE9014B52EF8DF" box="[151,781,1849,1876]" pageId="6" pageNumber="576" type="nomenclature">
<paragraph id="01588F4AD565FF90F9AE9014B52EF8DF" blockId="6.[151,799,1849,1912]" box="[151,781,1849,1876]" pageId="6" pageNumber="576">
<heading id="5A103826D565FF90F9AE9014B52EF8DF" bold="true" box="[151,781,1849,1876]" fontSize="11" level="1" pageId="6" pageNumber="576" reason="1">
<emphasis id="33935358D565FF90F9AE9014B52EF8DF" bold="true" box="[151,781,1849,1876]" pageId="6" pageNumber="576">
<taxonomicName id="C6E7F4C9D565FF90F9AE9014B48BF8D8" authority="Brugnera, Roell &amp; Lemaitre" authorityName="Brugnera, Roell &amp; Lemaitre" authorityYear="2021" box="[151,680,1849,1876]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="576" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="33935358D565FF90F9AE9014B76FF8DF" bold="true" box="[151,332,1849,1876]" italics="true" pageId="6" pageNumber="576">Podisus graziae</emphasis>
Brugnera, Roell &amp; Lemaître
</taxonomicName>
,
<taxonomicNameLabel id="28A0EE23D565FF90FB8D9014B52EF8DF" box="[692,781,1849,1875]" pageId="6" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D565FF90F9AE9070B53CF8F4" box="[151,799,1885,1912]" pageId="6" pageNumber="576" type="description">
<paragraph id="01588F4AD565FF90F9AE9070B53CF8F4" blockId="6.[151,799,1849,1912]" box="[151,799,1885,1912]" pageId="6" pageNumber="576">
(
<figureCitation id="99DC93CFD565FF90F9A69070B703F8FB" box="[159,288,1885,1911]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="6" pageNumber="576">Figs 10–12</figureCitation>
,
<figureCitation id="99DC93CFD565FF90F8159070B757F8FB" box="[300,372,1885,1911]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="6" pageNumber="576">19–21</figureCitation>
,
<figureCitation id="99DC93CFD565FF90F8B99070B7EAF8FB" box="[384,457,1885,1911]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="6" pageNumber="576">28–30</figureCitation>
,
<figureCitation id="99DC93CFD565FF90F8ED9070B43FF8FB" box="[468,540,1885,1911]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="6" pageNumber="576">36–37</figureCitation>
,
<figureCitation id="99DC93CFD565FF90FB119070B452F8F4" box="[552,625,1885,1912]" captionStart="FIGURES 40–48" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[260,1315,181,1919]" captionTargetId="figure-20@10.[260,1328,181,1919]" captionTargetPageId="10" captionText="FIGURES 40–48. Phallus of Podisus ventralis (40–42), Podisus graziae sp. n. (43–45), and Podisus thomasi sp. n. (46–48) in posterior (40, 43, 46), lateral (41, 44, 47) and anterior (42, 45, 48) views. Abbreviations: bt, basal theca; cl, conjunctival lobes; ts, thecal shield; v, vesica. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692479" httpUri="https://zenodo.org/record/4692479/files/figure.png" pageId="6" pageNumber="576">43–45</figureCitation>
,
<figureCitation id="99DC93CFD565FF90FB459070B4E6F8F4" box="[636,709,1885,1912]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="6" pageNumber="576">51–52</figureCitation>
,
<figureCitation id="99DC93CFD565FF90FBE99070B534F8F4" box="[720,791,1885,1912]" captionStart="FIGURES 55–60" captionStartId="12.[151,264,1290,1314]" captionTargetBox="[151,1436,453,1267]" captionTargetId="figure-150@12.[151,1436,453,1267]" captionTargetPageId="12" captionText="FIGURES 55–60. Live specimens of Podisus graziae sp. n. (55–57), and Podisus thomasi sp. n. (58–60). 55–57, photos by Ricardo Brugnera, Porto Alegre, Brazil. 58–60, photos by Erika Schulz, Mindo, Pichincha, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.4692483" httpUri="https://zenodo.org/record/4692483/files/figure.png" pageId="6" pageNumber="576">55–57</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D565FF91F9AE9088B78EFECD" lastPageId="7" lastPageNumber="577" pageId="6" pageNumber="576" type="materials_examined">
<paragraph id="01588F4AD565FF90F9AE9088B246F868" blockId="6.[151,1437,1957,2020]" pageId="6" pageNumber="576">
<materialsCitation id="B18F8517D565FF90F9AE9088B246F868" ID-GBIF-Occurrence="3083796305" collectingDate="2017-07-14" collectingDateMax="2017-07-16" collectingDateMin="2017-07-14" collectionCode="MZSP" collectorName="Pano &amp; J. Avendano &amp; R. Brugnera &amp; V. Castro" country="Brazil" latitude="-28.558887" location="Sao Domingos do Sul" longLatPrecision="20" longitude="-51.836388" municipality="Santa Gema" pageId="6" pageNumber="576" specimenCount="1" typeStatus="holotype">
<emphasis id="33935358D565FF90F9AE9088B728F833" bold="true" box="[151,267,1957,1983]" pageId="6" pageNumber="576">
<typeStatus id="DE5C31E8D565FF90F9AE9088B725F833" box="[151,262,1957,1983]" pageId="6" pageNumber="576" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Labels:
<collectingCountry id="79F0CFDAD565FF90F88F9088B43FF833" box="[438,540,1957,1983]" name="Brazil" pageId="6" pageNumber="576">BRASIL</collectingCountry>
, RS,
<location id="0438D991D565FF90FB609088B570F833" LSID="urn:lsid:plazi:treatment:894E3E5CD565FF9EF9AE9014B475F985:0438D991D565FF90FB609088B570F833" box="[601,851,1957,1983]" country="Brazil" latitude="-28.558887" longLatPrecision="20" longitude="-51.836388" municipality="Santa Gema" name="Sao Domingos do Sul" pageId="6" pageNumber="576">São Domingos do Sul</location>
,
<collectingMunicipality id="E13C1530D565FF90FA599088B5CFF833" box="[864,1004,1957,1983]" pageId="6" pageNumber="576">Santa Gema</collectingMunicipality>
,
<geoCoordinate id="64D3E98DD565FF90FACE9088B2A1F833" box="[1015,1154,1957,1983]" degrees="28" direction="south" minutes="33" orientation="latitude" pageId="6" pageNumber="576" precision="15" seconds="32" value="-28.558887">28°33’32’’S</geoCoordinate>
,
<geoCoordinate id="64D3E98DD565FF90FDB69088B309F84C" box="[1167,1322,1957,1984]" degrees="51" direction="west" minutes="50" orientation="longitude" pageId="6" pageNumber="576" precision="15" seconds="11" value="-51.836388">51° 50’11’’W</geoCoordinate>
.
<date id="7559A98AD565FF90FC0F9088B6F0F86F" pageId="6" pageNumber="576" value="2017-07-14" valueMax="2017-07-16" valueMin="2017-07-14">
<collectingDate id="651D5062D565FF90FC0F9088B6F0F86F" pageId="6" pageNumber="576" value="2017-07-14" valueMax="2017-07-16" valueMin="2017-07-14">14-16 vii 2017</collectingDate>
</date>
.
<collectorName id="AC12EA9CD565FF90F9E490E7B736F868" box="[221,277,1994,2020]" pageId="6" pageNumber="576">Pano</collectorName>
de batida.
<collectorName id="AC12EA9CD565FF90F8A990E7B438F86F" box="[400,539,1993,2020]" pageId="6" pageNumber="576">J. Avendaño</collectorName>
,
<collectorName id="AC12EA9CD565FF90FB1F90E7B48CF868" box="[550,687,1994,2020]" pageId="6" pageNumber="576">R. Brugnera</collectorName>
&amp;
<collectorName id="AC12EA9CD565FF90FBEB90E7B519F86F" box="[722,826,1993,2020]" pageId="6" pageNumber="576">V. Castro</collectorName>
col.. (
<collectionCode id="67F6178FD565FF90FAB990E4B5EFF86F" box="[896,972,1993,2019]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="576" type="Museum">MZSP</collectionCode>
) (
<figureCitation id="99DC93CFD565FF90FADA90E4B27EF868" box="[995,1117,1993,2020]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="6" pageNumber="576">Figs 10–l2</figureCitation>
)
</materialsCitation>
</paragraph>
<paragraph id="01588F4AD564FF91F9FE97BAB78EFECD" blockId="7.[151,1437,150,933]" pageId="7" pageNumber="577">
<materialsCitation id="B18F8517D564FF91F9FE97BAB2A9FF3D" ID-GBIF-Occurrence="3083796328" box="[199,1162,150,177]" collectionCode="JEE" collectorName="D. H. Habeck" country="Brazil" location="Curitiba" pageId="7" pageNumber="577" specimenCount="1" specimenCount-female="1" stateProvince="Parana" typeStatus="paratype">
<emphasis id="33935358D564FF91F9FE97BAB401FF3D" bold="true" box="[199,546,150,177]" pageId="7" pageNumber="577">
<typeStatus id="DE5C31E8D564FF91F9FE97BAB762FF3D" box="[199,321,151,177]" pageId="7" pageNumber="577" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="79F0CFDAD564FF91F87797BBB79DFF3C" box="[334,446,150,176]" name="Brazil" pageId="7" pageNumber="577">BRAZIL</collectingCountry>
:
<collectingRegion id="C32341A8D564FF91F8F297BBB401FF3D" box="[459,546,150,177]" country="Brazil" name="Parana" pageId="7" pageNumber="577">Paraná</collectingRegion>
</emphasis>
,
<specimenCount id="17E144C3D564FF91FB0897BAB477FF3D" box="[561,596,151,177]" pageId="7" pageNumber="577" type="female">1♀</specimenCount>
,
<location id="0438D991D564FF91FB5A97BAB4E1FF3D" LSID="urn:lsid:plazi:treatment:894E3E5CD565FF9EF9AE9014B475F985:0438D991D564FF91FB5A97BAB4E1FF3D" box="[611,706,151,177]" country="Brazil" name="Curitiba" pageId="7" pageNumber="577" stateProvince="Parana">Curitiba</location>
, 12-III-year?,
<collectorName id="AC12EA9CD564FF91FA5D97BAB220FF3D" box="[868,1027,151,177]" pageId="7" pageNumber="577">D. H. Habeck</collectorName>
, col., (
<collectionCode id="67F6178FD564FF91FD6997BAB2A0FF3D" box="[1104,1155,151,177]" pageId="7" pageNumber="577">JEE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B18F8517D564FF91FDAE97BBB219FF59" ID-GBIF-Occurrence="3083796324" collectionCode="AMNH" collectorName="F. Plaumann" country="Brazil" latitude="-27.183332" location="Nova Teutonia" pageId="7" pageNumber="577" specimenCount="1" specimenCount-female="1" stateProvince="Santa Catarina" typeStatus="paratype">
<emphasis id="33935358D564FF91FDAE97BBB36CFF3C" bold="true" box="[1175,1359,150,176]" pageId="7" pageNumber="577">
<collectingRegion id="C32341A8D564FF91FDAE97BBB36CFF3C" box="[1175,1359,150,176]" country="Brazil" name="Santa Catarina" pageId="7" pageNumber="577">Santa Catarina</collectingRegion>
</emphasis>
,
<collectingRegion id="C32341A8D564FF91FC6797BAB3BFFF3D" box="[1374,1436,151,177]" country="Brazil" name="Ceara" pageId="7" pageNumber="577">Seara</collectingRegion>
[
<location id="0438D991D564FF91F9A79796B769FF59" LSID="urn:lsid:plazi:treatment:894E3E5CD565FF9EF9AE9014B475F985:0438D991D564FF91F9A79796B769FF59" box="[158,330,186,213]" country="Brazil" latitude="-27.183332" name="Nova Teutonia" pageId="7" pageNumber="577" stateProvince="Santa Catarina">Nova Teutônia</location>
],
<specimenCount id="17E144C3D564FF91F8639796B75EFF59" box="[346,381,187,213]" pageId="7" pageNumber="577" type="female">1♀</specimenCount>
,
<geoCoordinate id="64D3E98DD564FF91F8B09796B7C5FF59" box="[393,486,187,213]" degrees="27" direction="south" minutes="11" orientation="latitude" pageId="7" pageNumber="577" precision="925" value="-27.183332">27°11’S</geoCoordinate>
,
<geoCoordinate id="64D3E98DD564FF91F8C89797B479FF59" box="[497,602,186,213]" pageId="7" pageNumber="577">52° 23’ L</geoCoordinate>
, 06-XIII-1948,
<collectorName id="AC12EA9CD564FF91FA319796B5B1FF59" box="[776,914,187,213]" pageId="7" pageNumber="577">F. Plaumann</collectorName>
col., (
<collectionCode id="67F6178FD564FF91FAEF9796B211FF59" box="[982,1074,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="7" pageNumber="577">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B18F8517D564FF91FD7A9796B433FF75" ID-GBIF-Occurrence="3083796302" collectingDate="1980-02-28" collectionCode="DBTC" collectorName="D. B. Thomas" country="Brazil" location="Sao Lourenco" pageId="7" pageNumber="577" specimenCount="1" stateProvince="Santa Catarina" typeStatus="paratype">
<location id="0438D991D564FF91FD7A9796B2C2FF59" LSID="urn:lsid:plazi:treatment:894E3E5CD565FF9EF9AE9014B475F985:0438D991D564FF91FD7A9796B2C2FF59" box="[1091,1249,187,213]" country="Brazil" name="Sao Lourenco" pageId="7" pageNumber="577" stateProvince="Santa Catarina">São Lourenço</location>
,
<quantity id="C61F22AFD564FF91FDD29796B311FF59" box="[1259,1330,187,213]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="7" pageNumber="577" unit="mi" value="20.0">20 mi.</quantity>
W,
<date id="7559A98AD564FF91FC629796B6F1FF75" pageId="7" pageNumber="577" value="1980-02-28">
<collectingDate id="651D5062D564FF91FC629796B6F1FF75" pageId="7" pageNumber="577" value="1980-02-28">28-II- 1980</collectingDate>
</date>
,
<collectorName id="AC12EA9CD564FF91F9E497F2B751FF75" box="[221,370,223,249]" pageId="7" pageNumber="577">D.B. Thomas</collectorName>
col., (
<collectionCode id="67F6178FD564FF91F88E97F2B42BFF75" box="[439,520,223,249]" pageId="7" pageNumber="577">DBTC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B18F8517D564FF91FB2297F3B2D4FF75" ID-GBIF-Occurrence="3083796306" box="[539,1271,222,249]" collectingDate="2017-07-14" collectingDateMax="2017-07-16" collectingDateMin="2017-07-14" collectionCode="UFRGS" collectorName="Pano &amp; J. Avendano &amp; R. Brugnera &amp; V. Castro" country="Brazil" latitude="-28.558887" location="Sao Domingos do Sul" longLatPrecision="20" longitude="-51.836388" municipality="Santa Gema" pageId="7" pageNumber="577" specimenCount="20" specimenCount-female="10" specimenCount-male="10" stateProvince="Rio Grande do Sul" typeStatus="paratype">
<emphasis id="33935358D564FF91FB2297F3B4DAFF74" bold="true" box="[539,761,222,249]" pageId="7" pageNumber="577">
<collectingRegion id="C32341A8D564FF91FB2297F3B4DAFF74" box="[539,761,222,249]" country="Brazil" name="Rio Grande do Sul" pageId="7" pageNumber="577">Rio Grande do Sul</collectingRegion>
</emphasis>
<specimenCount id="17E144C3D564FF91FA3997F2B519FF75" box="[768,826,223,249]" pageId="7" pageNumber="577" type="male">10 ♂</specimenCount>
,
<specimenCount id="17E144C3D564FF91FA7C97F2B55EFF75" box="[837,893,223,249]" pageId="7" pageNumber="577" type="female">10 ♀</specimenCount>
, same data as holotype (
<collectionCode id="67F6178FD564FF91FDB797F2B2CCFF75" box="[1166,1263,223,249]" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="7" pageNumber="577">UFRGS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B18F8517D564FF91FC3B97F2B793FE91" ID-GBIF-Occurrence="3083796303" collectingDate="2017-07-14" collectingDateMax="2017-07-16" collectingDateMin="2017-07-14" collectionCode="MZSP" collectorName="Pano &amp; J. Avendano &amp; R. Brugnera &amp; V. Castro" country="Brazil" latitude="-28.558887" location="Sao Domingos do Sul" longLatPrecision="20" longitude="-51.836388" municipality="Santa Gema" pageId="7" pageNumber="577" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Rio Grande do Sul" typeStatus="paratype">
<specimenCount id="17E144C3D564FF91FC3B97F2B30EFF75" box="[1282,1325,223,249]" pageId="7" pageNumber="577" type="male">3 ♂</specimenCount>
,
<specimenCount id="17E144C3D564FF91FC0097F2B37FFF75" box="[1337,1372,223,249]" pageId="7" pageNumber="577" type="female">3♀</specimenCount>
same data as holotype (
<collectionCode id="67F6178FD564FF91F861962EB784FE91" box="[344,423,259,285]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="577" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B18F8517D564FF91F883962EB550FE91" ID-GBIF-Occurrence="3083796313" box="[442,883,259,285]" collectingDate="2006-01" collectionCode="MCNZ" country="Brazil" location="Fepagro" municipality="Maquine" pageId="7" pageNumber="577" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul" typeStatus="paratype">
<specimenCount id="17E144C3D564FF91F883962EB7F2FE91" box="[442,465,259,285]" pageId="7" pageNumber="577" type="male">♂</specimenCount>
,
<collectingMunicipality id="E13C1530D564FF91F8E2962EB462FE91" box="[475,577,259,285]" pageId="7" pageNumber="577">Maquiné</collectingMunicipality>
,
<location id="0438D991D564FF91FB72962EB489FE91" LSID="urn:lsid:plazi:treatment:894E3E5CD565FF9EF9AE9014B475F985:0438D991D564FF91FB72962EB489FE91" box="[587,682,259,285]" country="Brazil" municipality="Maquine" name="Fepagro" pageId="7" pageNumber="577" stateProvince="Rio Grande do Sul">Fepagro</location>
,
<date id="7559A98AD564FF91FB8D962EB522FE91" box="[692,769,259,285]" pageId="7" pageNumber="577" value="2006-01">
<collectingDate id="651D5062D564FF91FB8D962EB522FE91" box="[692,769,259,285]" pageId="7" pageNumber="577" value="2006-01">I-2006</collectingDate>
</date>
, (
<collectionCode id="67F6178FD564FF91FA2A962EB549FE91" box="[787,874,259,285]" country="Brazil" httpUri="http://grbio.org/cool/wyk1-wacz" name="Porto Alegre, Museu de Ciencias Naturais da Fundacao Zoo-Botanica do Rio Grande do Sul" pageId="7" pageNumber="577">MCNZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B18F8517D564FF91FA45962FB78AFECD" ID-GBIF-Occurrence="3083796319" collectingDate="1989-10-21" collectingDateMax="1989-10-23" collectingDateMin="1989-10-21" collectionCode="JEE" collectorName="G. Arriagada" country="Paraguay" location="Alto Parana" pageId="7" pageNumber="577" specimenCount="3" specimenCount-female="3" stateProvince="Alto Parana" typeStatus="paratype">
<emphasis id="33935358D564FF91FA45962FB290FE91" bold="true" box="[892,1203,258,285]" pageId="7" pageNumber="577">
<collectingCountry id="79F0CFDAD564FF91FA45962FB239FE90" box="[892,1050,258,284]" name="Paraguay" pageId="7" pageNumber="577">PARAGUAY</collectingCountry>
:
<collectingRegion id="C32341A8D564FF91FD1A962FB290FE91" box="[1059,1203,258,285]" country="Paraguay" name="Alto Parana" pageId="7" pageNumber="577">Alto Paraná</collectingRegion>
</emphasis>
,
<specimenCount id="17E144C3D564FF91FDF9962EB2CBFE91" box="[1216,1256,259,285]" pageId="7" pageNumber="577" type="female">3 ♀</specimenCount>
,
<date id="7559A98AD564FF91FDCC962EB3BBFE91" box="[1269,1432,259,285]" pageId="7" pageNumber="577" value="1989-10-21" valueMax="1989-10-23" valueMin="1989-10-21">
<collectingDate id="651D5062D564FF91FDCC962EB3BBFE91" box="[1269,1432,259,285]" pageId="7" pageNumber="577" value="1989-10-21" valueMax="1989-10-23" valueMin="1989-10-21">21-23-X-1989</collectingDate>
</date>
,
<collectorName id="AC12EA9CD564FF91F9AE960AB70BFECD" box="[151,296,295,321]" pageId="7" pageNumber="577">G. Arriagada</collectorName>
col., (
<collectionCode id="67F6178FD564FF91F857960AB782FECD" box="[366,417,295,321]" pageId="7" pageNumber="577">JEE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D564FF91F9FE9666B2C5FE21" pageId="7" pageNumber="577" type="etymology">
<paragraph id="01588F4AD564FF91F9FE9666B2C5FE21" blockId="7.[151,1437,150,933]" pageId="7" pageNumber="577">
<emphasis id="33935358D564FF91F9FE9666B76DFEE9" bold="true" box="[199,334,331,357]" pageId="7" pageNumber="577">Etymology.</emphasis>
We are pleased to dedicate this new species to Dr Jocelia Grazia (UFRGS) in recognition of her outstanding contributions to the knowledge of the
<taxonomicName id="C6E7F4C9D564FF91FBF19642B56CFE05" box="[712,847,367,393]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="7" pageNumber="577" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
, especially to that of Neotropical Pentatomoidea, the passion for which she has, over the years, graciously communicated to her many grateful students.
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D564FF91F9FE969AB793FE5D" box="[199,432,438,465]" pageId="7" pageNumber="577" type="materials_examined">
<paragraph id="01588F4AD564FF91F9FE969AB793FE5D" blockId="7.[151,1437,150,933]" box="[199,432,438,465]" pageId="7" pageNumber="577">
<emphasis id="33935358D564FF91F9FE969AB745FE5C" bold="true" box="[199,358,438,465]" pageId="7" pageNumber="577">Type locality.</emphasis>
<collectingCountry id="79F0CFDAD564FF91F855969AB793FE5D" box="[364,432,439,465]" name="Brazil" pageId="7" pageNumber="577">Brazil</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D564FF91F9FE96F7B723FDED" pageId="7" pageNumber="577" type="diagnosis">
<paragraph id="01588F4AD564FF91F9FE96F7B723FDED" blockId="7.[151,1437,150,933]" pageId="7" pageNumber="577">
<emphasis id="33935358D564FF91F9FE96F7B761FE78" bold="true" box="[199,322,474,500]" pageId="7" pageNumber="577">Diagnosis.</emphasis>
Anterolateral margins of pronotum calloused, crenulated, paler than disc; humeral angles angular, without a posterior tooth, directed laterally. Apex of scutellum concolourous with disc (
<figureCitation id="99DC93CFD564FF91FD5696D2B2E5FD95" box="[1135,1222,511,537]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="7" pageNumber="577">Figs 10</figureCitation>
,
<figureCitation id="99DC93CFD564FF91FDEA96D2B33FFD95" box="[1235,1308,511,537]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="7" pageNumber="577">19, 21</figureCitation>
). Apices of parameres obtuse (
<figureCitation id="99DC93CFD564FF91F856950EB7D0FDB1" box="[367,499,547,573]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="7" pageNumber="577">Figs 28–30</figureCitation>
,
<figureCitation id="99DC93CFD564FF91FB38950EB402FDB2" box="[513,545,547,574]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="7" pageNumber="577">36</figureCitation>
). Posterior margins of valvifers VIII slightly sinuous, posterior angles obtuse (
<figureCitation id="99DC93CFD564FF91F999956AB6D0FDED" box="[160,243,582,609]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="7" pageNumber="577">Fig. 51</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D564FF9EF9FE9547B6C8FB2E" lastPageId="8" lastPageNumber="578" pageId="7" pageNumber="577" type="description">
<paragraph id="01588F4AD564FF91F9FE9547B5C8FD09" blockId="7.[151,1437,150,933]" box="[199,1003,618,645]" pageId="7" pageNumber="577">
<emphasis id="33935358D564FF91F9FE9547B77AFD08" bold="true" box="[199,345,618,644]" pageId="7" pageNumber="577">Description.</emphasis>
Body length 10.57–11.07 mm (
<specimenCount id="17E144C3D564FF91FBF99546B4F1FD09" box="[704,722,619,645]" pageId="7" pageNumber="577" type="female">♀</specimenCount>
) and 8.00–9.70 mm (
<specimenCount id="17E144C3D564FF91FAFF9546B5FEFD09" box="[966,989,619,645]" pageId="7" pageNumber="577" type="male">♂</specimenCount>
).
</paragraph>
<paragraph id="01588F4AD564FF91F9FE95A2B5FAFD7D" blockId="7.[151,1437,150,933]" pageId="7" pageNumber="577">
Body castaneous, anterior half of lateral margins of pronotum forming a marginal wide carina, lighter than rest of pronotum. A thin central line mostly without punctures or lighter than rest of dorsum extends from anterior margin of pronotum to posterior margin of scutellum (
<figureCitation id="99DC93CFD564FF91FBF295FAB501FD7D" box="[715,802,727,753]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="7" pageNumber="577">Figs 10</figureCitation>
,
<figureCitation id="99DC93CFD564FF91FA1795FAB555FD7D" box="[814,886,727,753]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="7" pageNumber="577">19, 21</figureCitation>
,
<figureCitation id="99DC93CFD564FF91FABB95FBB5EFFD7D" box="[898,972,726,753]" captionStart="FIGURES 55–60" captionStartId="12.[151,264,1290,1314]" captionTargetBox="[151,1436,453,1267]" captionTargetId="figure-150@12.[151,1436,453,1267]" captionTargetPageId="12" captionText="FIGURES 55–60. Live specimens of Podisus graziae sp. n. (55–57), and Podisus thomasi sp. n. (58–60). 55–57, photos by Ricardo Brugnera, Porto Alegre, Brazil. 58–60, photos by Erika Schulz, Mindo, Pichincha, Ecuador." figureDoi="http://doi.org/10.5281/zenodo.4692483" httpUri="https://zenodo.org/record/4692483/files/figure.png" pageId="7" pageNumber="577">55–57</figureCitation>
).
</paragraph>
<paragraph id="01588F4AD564FF91F9FE95D6B79AFC29" blockId="7.[151,1437,150,933]" pageId="7" pageNumber="577">
Head subrectangular, mandibular plates uniformly punctured, clypeus as long as or slightly surpassing apex of mandibular plates; ocelli lying behind an imaginary line through posterior margins of compound eyes; antennal tubercles partially visible from above, proportion of antennomeres: I&lt;II&gt;III&lt;IV&gt;V (
<figureCitation id="99DC93CFD564FF91FD7B946EB2BBFCD1" box="[1090,1176,835,861]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="7" pageNumber="577">Figs 10</figureCitation>
,
<figureCitation id="99DC93CFD564FF91FD9D946EB2CEFCD1" box="[1188,1261,835,861]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="7" pageNumber="577">19, 21</figureCitation>
); bucculae evanescent posteriorly, apex rounded; labium extending to posterior margin of metasternum, proportion of labiomeres: I&lt;II&gt;III&gt;IV (
<figureCitation id="99DC93CFD564FF91F81394A6B7A3FC29" box="[298,384,907,933]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="7" pageNumber="577">Figs 11</figureCitation>
,
<figureCitation id="99DC93CFD564FF91F8B594A6B78FFC29" box="[396,428,907,933]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="7" pageNumber="577">20</figureCitation>
).
</paragraph>
<caption id="5598DFC2D564FF91F9AE91B9B6CFF978" ID-DOI="http://doi.org/10.5281/zenodo.4692477" ID-Zenodo-Dep="4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="7" pageNumber="577" startId="7.[151,264,1684,1708]" targetBox="[151,1436,957,1660]" targetPageId="7">
<paragraph id="01588F4AD564FF91F9AE91B9B6CFF978" blockId="7.[151,1436,1684,1781]" pageId="7" pageNumber="577">
<emphasis id="33935358D564FF91F9AE91B9B770F920" bold="true" box="[151,339,1684,1708]" pageId="7" pageNumber="577">FIGURES 34–39.</emphasis>
Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of
<taxonomicName id="C6E7F4C9D564FF91FD0491B9B2C8F920" box="[1085,1259,1684,1708]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="577" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D564FF91FD0491B9B2C8F920" box="[1085,1259,1684,1708]" italics="true" pageId="7" pageNumber="577">Podisus ventralis</emphasis>
</taxonomicName>
(34–35),
<taxonomicName id="C6E7F4C9D564FF91FC7591B9B6C0F95C" authority="Roell &amp; Brugnera &amp; Lemaître, 2021" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="577" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="33935358D564FF91FC7591B9B6C0F95C" italics="true" pageId="7" pageNumber="577">Podisus graziae</emphasis>
</taxonomicName>
<emphasis id="33935358D564FF91F9D29194B706F95D" bold="true" box="[235,293,1721,1745]" pageId="7" pageNumber="577">
<taxonomicNameLabel id="28A0EE23D564FF91F9D29194B706F95D" box="[235,293,1721,1745]" pageId="7" pageNumber="577" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(36–37), and
<taxonomicName id="C6E7F4C9D564FF91F88E9195B47CF95C" authority="Roell &amp; Brugnera &amp; Lemaître, 2021" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[439,607,1720,1744]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="577" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="33935358D564FF91F88E9195B47CF95C" box="[439,607,1720,1744]" italics="true" pageId="7" pageNumber="577">Podisus thomasi</emphasis>
</taxonomicName>
<emphasis id="33935358D564FF91FB5E9194B482F95D" bold="true" box="[615,673,1721,1745]" pageId="7" pageNumber="577">
<taxonomicNameLabel id="28A0EE23D564FF91FB5E9194B482F95D" box="[615,673,1721,1745]" pageId="7" pageNumber="577" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="01588F4AD564FF9EF9FE900EB357FE91" blockId="7.[151,1437,1827,2033]" lastBlockId="8.[151,1437,151,1545]" lastPageId="8" lastPageNumber="578" pageId="7" pageNumber="577">
Pronotum hexagonal, uniformly punctured; cicatrices flat; anterior and lateral margins concave in dorsal view, lateral margins slightly crenulated and lighter than disc on anterior half, anterior half of lateral margins preceded by a marginal sulcus; humeral angles angular, slightly produced. Scutellum longer than wide, reaching an imaginary line connecting fifth connexives medially; frenal margins longer than post frenal margins. Coria longer than scutellum, each attaining abdominal segment IV, uniformly punctured; hemelytral membranes well surpassing apex of abdomen, with apex spotted in brown (
<figureCitation id="99DC93CFD564FF91FB6F90FAB48DF87D" box="[598,686,2007,2033]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="7" pageNumber="577">Figs 10</figureCitation>
,
<figureCitation id="99DC93CFD564FF91FB8590FAB524F87D" box="[700,775,2007,2033]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="7" pageNumber="577">19, 21</figureCitation>
). Pro-, meso- and metasterna covered by small thin setae; prosternum with a weak median carina; mesosternum carinated longitudinally; limit between mesosternum and metasternum tumid; metasternum flat. Pro-, meso-, and metapleura sparsely punctured, evaporatoria on posterior margin of each mesopleura; metapleural evaporatoria surrounding each ostiolar peritreme; each ostiolar peritreme discal-type, elongated, with a darker spot on apical angle; each ostiolar opening directed laterally (
<figureCitation id="99DC93CFD56BFF9EFDDF962EB31FFE91" box="[1254,1340,259,285]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="8" pageNumber="578">Figs 11</figureCitation>
,
<figureCitation id="99DC93CFD56BFF9EFC7E962EB344FE91" box="[1351,1383,259,285]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="8" pageNumber="578">20</figureCitation>
).
</paragraph>
<paragraph id="01588F4AD56BFF9EF9FE960AB320FEE9" blockId="8.[151,1437,151,1545]" pageId="8" pageNumber="578">
Abdomen sparsely punctured, posterolateral angles of each connexivum projected as small spines, longer on seventh segment. Mesial tubercle projecting from sternite III not surpassing metacoxae (
<figureCitation id="99DC93CFD56BFF9EFD4D9666B2E9FEE9" box="[1140,1226,331,357]" captionStart="FIGURES 1–15" captionStartId="3.[151,264,1887,1911]" captionTargetBox="[162,1427,749,1855]" captionTargetId="figure-281@3.[151,1435,741,1864]" captionTargetPageId="3" captionText="FIGURES 1–15. Type specimens examined, in ventral and dorsal views, with their respective labels. 1–3, Arma ventralis (NH- MUK); 4–6, Arma pallipes (NHMUK); 7–9, Podisus neniator (SDEI); 10–12; Podisus graziae sp. n. (MZSP); 13–15, Podisus thomasi sp. n. (NHMUK). 7–9, photographed by Angelika Weirauch (Senckenberg Deutsches Entomologisches Institut); original photos cropped, light levels and contrast adjusted. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.4692471" httpUri="https://zenodo.org/record/4692471/files/figure.png" pageId="8" pageNumber="578">Figs 11</figureCitation>
,
<figureCitation id="99DC93CFD56BFF9EFDEF9666B2D5FEE9" box="[1238,1270,331,357]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="8" pageNumber="578">20</figureCitation>
).
</paragraph>
<paragraph id="01588F4AD56BFF9EF9FE9642B5B8FE05" blockId="8.[151,1437,151,1545]" box="[199,923,367,393]" pageId="8" pageNumber="578">
<emphasis id="33935358D56BFF9EF9FE9642B728FE05" bold="true" box="[199,267,367,393]" pageId="8" pageNumber="578">Male.</emphasis>
Without glandular patches on ventral surface of abdomen.
</paragraph>
<paragraph id="01588F4AD56BFF9EF9FE96BEB406FD42" blockId="8.[151,1437,151,1545]" pageId="8" pageNumber="578">
Genitalia. Pygophore bowl-shaped, opened posteriorly, setose, setae denser on ventral rim (
<figureCitation id="99DC93CFD56BFF9EFDFB96BEB366FE21" box="[1218,1349,403,429]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="8" pageNumber="578">Figs 28–30</figureCitation>
); dorsal rim concave with 1+1 dorsal projections near segment X (
<figureCitation id="99DC93CFD56BFF9EFA1A969AB553FE5D" box="[803,880,439,465]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="8" pageNumber="578">Fig 28</figureCitation>
, dr, dp) ventral rim concave, with 1+1 small ventral projections near posterolateral angles (
<figureCitation id="99DC93CFD56BFF9EFB4096F6B4D8FE79" box="[633,763,475,501]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="8" pageNumber="578">Figs 29, 30</figureCitation>
, vr, vp, pa); surface between inferior and superior layers of ventral rim excavated (
<figureCitation id="99DC93CFD56BFF9EF8A196D2B430FD95" box="[408,531,511,537]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="8" pageNumber="578">Fig. 29, 30</figureCitation>
, il, sl). Posterolateral angles acute (
<figureCitation id="99DC93CFD56BFF9EFAA496D2B23DFD95" box="[925,1054,511,537]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="8" pageNumber="578">Figs 28–30</figureCitation>
, pa). Segment X tubular, ventrally directed. Head of parameres divided into two arms, each apex obtuse, well visible posteriorly (
<figureCitation id="99DC93CFD56BFF9EFD8A950EB317FDB1" box="[1203,1332,547,573]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="8" pageNumber="578">Figs 28–30</figureCitation>
,
<figureCitation id="99DC93CFD56BFF9EFC06950EB37EFDB1" box="[1343,1373,547,573]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="8" pageNumber="578">36</figureCitation>
, par). Pseudoclasper C-shaped, outer margins sinuous, placed dorsolaterally to parameres, with globose projections and elevations (
<figureCitation id="99DC93CFD56BFF9EF8239546B7BCFD09" box="[282,415,619,645]" captionStart="FIGURES 25–33" captionStartId="6.[151,264,1635,1659]" captionTargetBox="[151,1436,381,1612]" captionTargetId="figure-110@6.[151,1436,381,1612]" captionTargetPageId="6" captionText="FIGURES 25–33. Male pygophore of Podisus ventralis (25–27), Podisus graziae sp. n. (28–30), and Podisus thomasi sp. n. (31–33) in dorsal (25, 28, 31), posterior (26, 29, 32) and ventral (27, 30, 33) views. Abbreviations: dp, dorsal projection; dr, dorsal rim; pa, posterolateral angle; il, inferior layer of ventral rim; par, paramere; pc, pseudoclasper; sl, superior layer of ventral rim; vp, ventral projection; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4692475" httpUri="https://zenodo.org/record/4692475/files/figure.png" pageId="8" pageNumber="578">Figs 28–30</figureCitation>
,
<figureCitation id="99DC93CFD56BFF9EF8949546B7E8FD09" box="[429,459,619,645]" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="8" pageNumber="578">37</figureCitation>
, pc). Phallus. Basal theca and thecal shield subequal in length, thecal shield slightly wider than basal theca (
<figureCitation id="99DC93CFD56BFF9EF8A295A2B43EFD25" box="[411,541,654,681]" captionStart="FIGURES 40–48" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[260,1315,181,1919]" captionTargetId="figure-20@10.[260,1328,181,1919]" captionTargetPageId="10" captionText="FIGURES 40–48. Phallus of Podisus ventralis (40–42), Podisus graziae sp. n. (43–45), and Podisus thomasi sp. n. (46–48) in posterior (40, 43, 46), lateral (41, 44, 47) and anterior (42, 45, 48) views. Abbreviations: bt, basal theca; cl, conjunctival lobes; ts, thecal shield; v, vesica. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692479" httpUri="https://zenodo.org/record/4692479/files/figure.png" pageId="8" pageNumber="578">Figs 43–45</figureCitation>
), vesica golf club-shaped in lateral view (
<figureCitation id="99DC93CFD56BFF9EFADE95A2B219FD25" box="[999,1082,655,681]" captionStart="FIGURES 40–48" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[260,1315,181,1919]" captionTargetId="figure-20@10.[260,1328,181,1919]" captionTargetPageId="10" captionText="FIGURES 40–48. Phallus of Podisus ventralis (40–42), Podisus graziae sp. n. (43–45), and Podisus thomasi sp. n. (46–48) in posterior (40, 43, 46), lateral (41, 44, 47) and anterior (42, 45, 48) views. Abbreviations: bt, basal theca; cl, conjunctival lobes; ts, thecal shield; v, vesica. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692479" httpUri="https://zenodo.org/record/4692479/files/figure.png" pageId="8" pageNumber="578">Fig. 44</figureCitation>
, v), apices of conjunctival lobes without sculptures (
<figureCitation id="99DC93CFD56BFF9EF84D959EB7D5FD41" box="[372,502,690,717]" captionStart="FIGURES 40–48" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[260,1315,181,1919]" captionTargetId="figure-20@10.[260,1328,181,1919]" captionTargetPageId="10" captionText="FIGURES 40–48. Phallus of Podisus ventralis (40–42), Podisus graziae sp. n. (43–45), and Podisus thomasi sp. n. (46–48) in posterior (40, 43, 46), lateral (41, 44, 47) and anterior (42, 45, 48) views. Abbreviations: bt, basal theca; cl, conjunctival lobes; ts, thecal shield; v, vesica. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692479" httpUri="https://zenodo.org/record/4692479/files/figure.png" pageId="8" pageNumber="578">Figs 43–45</figureCitation>
, cl).
</paragraph>
<paragraph id="01588F4AD56BFF9EF9FE95FAB6C8FB2E" blockId="8.[151,1437,151,1545]" pageId="8" pageNumber="578">
<emphasis id="33935358D56BFF9EF9FE95FAB706FD7D" bold="true" box="[199,293,727,753]" pageId="8" pageNumber="578">Female.</emphasis>
Genitalia. Genital plates positioned ventroposteriorly (
<figureCitation id="99DC93CFD56BFF9EFAB695FAB5C7FD7D" box="[911,996,727,753]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="8" pageNumber="578">Fig. 20</figureCitation>
); valvifers VIII subtriangular, posterior margins slightly sinuous, posterior angles obtuse, sutural margins straight, juxtaposed (
<figureCitation id="99DC93CFD56BFF9EFD5295D6B29EFC99" box="[1131,1213,762,789]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 51</figureCitation>
, vf
<subScript id="9D638D0FD56BFF9EFDD89420B2CAFC97" attach="left" box="[1249,1257,781,795]" fontSize="6" pageId="8" pageNumber="578">8</subScript>
). Laterotergites VIII triangular with spiracles on proximal angle (
<figureCitation id="99DC93CFD56BFF9EFBF19432B53FFCB5" box="[712,796,798,825]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 51</figureCitation>
, la
<subScript id="9D638D0FD56BFF9EFA07941CB565FCB3" attach="right" box="[830,838,817,831]" fontSize="6" pageId="8" pageNumber="578">8</subScript>
). Exposed portion of valvifers IX rectangular, wider than long (
<figureCitation id="99DC93CFD56BFF9EF828946EB746FCD1" box="[273,357,834,861]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 51</figureCitation>
, vf
<subScript id="9D638D0FD56BFF9EF8B09478B7B2FCEF" attach="left" box="[393,401,853,867]" fontSize="6" pageId="8" pageNumber="578">9</subScript>
). Exposed portion of laterotergites IX tongue-like (
<figureCitation id="99DC93CFD56BFF9EFAE1946EB20FFCD1" box="[984,1068,834,861]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 51</figureCitation>
, la
<subScript id="9D638D0FD56BFF9EFD749478B276FCEF" attach="left" box="[1101,1109,853,867]" fontSize="6" pageId="8" pageNumber="578">9</subScript>
), surpassing segment X, and almost attaining mediotergite VIII. Segment X subquadrate (
<figureCitation id="99DC93CFD56BFF9EFA7B944AB5B6FC0D" box="[834,917,870,897]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 51</figureCitation>
, X). Inner portion of valvifers IX projected in 1+1 straight elongated arms with acute apices (
<figureCitation id="99DC93CFD56BFF9EFB9194A6B4D8FC29" box="[680,763,906,933]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, vf
<subScript id="9D638D0FD56BFF9EFA2694B0B504FC27" attach="right" box="[799,807,925,939]" fontSize="6" pageId="8" pageNumber="578">9</subScript>
). Valvulae IX with 1+1 small lunate secondary thickenings (
<figureCitation id="99DC93CFD56BFF9EF9EC9482B70BFC45" box="[213,296,942,969]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, tva
<subScript id="9D638D0FD56BFF9EF86F94ECB77DFC43" attach="left" box="[342,350,961,975]" fontSize="6" pageId="8" pageNumber="578">9</subScript>
). Ring sclerites inconspicuous, small, circular, placed posteriorly to thickening of vaginal intima (
<figureCitation id="99DC93CFD56BFF9EF99994FEB6D0FC61" box="[160,243,978,1005]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, rs, tvi); thickening of vaginal intima short, obtuse (
<figureCitation id="99DC93CFD56BFF9EFA7C94FEB5BBFC61" box="[837,920,978,1005]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, tvi); arcus placed anteriorly of thickening of vaginal intima (
<figureCitation id="99DC93CFD56BFF9EF87294DAB7BCFB9D" box="[331,415,1014,1041]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, ar). Vesicular area almost 2.5 times longer than length of pars intermedialis plus capsula seminalis (
<figureCitation id="99DC93CFD56BFF9EF8299336B742FBB9" box="[272,353,1050,1077]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, va, pi, cs). Inner and median ducts of vesicular areas of uniform diameter (
<figureCitation id="99DC93CFD56BFF9EFD919336B2D9FBB9" box="[1192,1274,1050,1077]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, id, md), outer duct constricted basally (
<figureCitation id="99DC93CFD56BFF9EF8899312B421FBD5" box="[432,514,1086,1113]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, od). Pars intermedialis tubular, shorter and narrower than capsula seminalis; capsula seminalis oval, elongated, smooth (
<figureCitation id="99DC93CFD56BFF9EFB6C934EB484FBF1" box="[597,679,1122,1149]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, pi, cs). Proximal and distal annular crests of similar width (
<figureCitation id="99DC93CFD56BFF9EFC7C934EB3B4FBF1" box="[1349,1431,1122,1149]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
, pa, da).
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D56BFF9EF9FE9387B4A7FB49" box="[199,644,1194,1222]" pageId="8" pageNumber="578" type="distribution">
<paragraph id="01588F4AD56BFF9EF9FE9387B4A7FB49" blockId="8.[151,1437,151,1545]" box="[199,644,1194,1222]" pageId="8" pageNumber="578">
<emphasis id="33935358D56BFF9EF9FE9387B77AFB48" bold="true" box="[199,345,1194,1220]" pageId="8" pageNumber="578">Distribution</emphasis>
:
<collectingCountry id="79F0CFDAD56BFF9EF8519386B78FFB49" box="[360,428,1195,1221]" name="Brazil" pageId="8" pageNumber="578">Brazil</collectingCountry>
(South),
<collectingCountry id="79F0CFDAD56BFF9EFB2F9386B4A3FB49" box="[534,640,1195,1221]" name="Paraguay" pageId="8" pageNumber="578">Paraguay</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="49FDDCC1D56BFF9EF9FE93E2B475F985" pageId="8" pageNumber="578" type="discussion">
<paragraph id="01588F4AD56BFF9EF9FE93E2B475F985" blockId="8.[151,1437,151,1545]" pageId="8" pageNumber="578">
<emphasis id="33935358D56BFF9EF9FE93E2B718FB65" bold="true" box="[199,315,1231,1257]" pageId="8" pageNumber="578">Remarks.</emphasis>
The external morphology of
<taxonomicName id="C6E7F4C9D56BFF9EFBB293E2B4DFFB64" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[651,764,1231,1256]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="33935358D56BFF9EFBB293E2B4DFFB64" box="[651,764,1231,1256]" italics="true" pageId="8" pageNumber="578">P. graziae</emphasis>
</taxonomicName>
resembles that of
<taxonomicName id="C6E7F4C9D56BFF9EFAF793E2B26EFB65" box="[974,1101,1231,1257]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D56BFF9EFAF793E2B26EFB65" box="[974,1101,1231,1257]" italics="true" pageId="8" pageNumber="578">P. ventralis</emphasis>
</taxonomicName>
. Neither species have a paler spot on the post-frenal lobe of the scutellum (
<figureCitation id="99DC93CFD56BFF9EFBAF93DEB534FA81" box="[662,791,1267,1293]" captionStart="FIGURES 16–24" captionStartId="4.[151,264,1975,1999]" captionTargetBox="[173,1413,182,1952]" captionTargetId="figure-20@4.[173,1414,181,1952]" captionTargetPageId="4" captionText="FIGURES 16–24. Podisus ventralis (16–18), Podisus graziae sp. n. (19–21), and Podisus thomasi sp. n. (22–24) in dorsal and ventral views, and detail of head and pronotum. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692473" httpUri="https://zenodo.org/record/4692473/files/figure.png" pageId="8" pageNumber="578">Figs 16, 19</figureCitation>
) and both present parameres with obtuse apices (
<figureCitation id="99DC93CFD56BFF9EFC7993DEB694FABE" captionStart="FIGURES 34–39" captionStartId="7.[151,264,1684,1708]" captionTargetBox="[151,1436,957,1660]" captionTargetId="figure-331@7.[151,1436,957,1661]" captionTargetPageId="7" captionText="FIGURES 34–39. Male right parameres (34, 36, 38) and right pseudoclaspers (35, 37, 39) of Podisus ventralis (34–35), Podisus graziae sp. n. (36–37), and Podisus thomasi sp. n. (38–39). Abbreviations: im, inner margin; om, outer margin. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692477" httpUri="https://zenodo.org/record/4692477/files/figure.png" pageId="8" pageNumber="578">Figs 34, 36</figureCitation>
). However, the female external and internal genitalia are very different; in
<taxonomicName id="C6E7F4C9D56BFF9EFD3B923AB2A3FABD" box="[1026,1152,1303,1329]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D56BFF9EFD3B923AB2A3FABD" box="[1026,1152,1303,1329]" italics="true" pageId="8" pageNumber="578">P. ventralis</emphasis>
</taxonomicName>
, the posterior margins of valvifers VIII are strongly sinuous, forming a V-projection on the posterior angles over valvifers IX (
<figureCitation id="99DC93CFD56BFF9EFDC19216B369FAD9" box="[1272,1354,1339,1365]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 49</figureCitation>
), while in
<taxonomicName id="C6E7F4C9D56BFF9EF98D9272B707FAF4" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[180,292,1375,1400]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="33935358D56BFF9EF98D9272B707FAF4" box="[180,292,1375,1400]" italics="true" pageId="8" pageNumber="578">P. graziae</emphasis>
</taxonomicName>
these posterior margins are slightly sinuous, with obtuse posterior angles (
<figureCitation id="99DC93CFD56BFF9EFD579272B2E1FAF5" box="[1134,1218,1374,1401]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 51</figureCitation>
); the vesicular area is longer in
<taxonomicName id="C6E7F4C9D56BFF9EF82792AEB7BFFA11" box="[286,412,1411,1437]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D56BFF9EF82792AEB7BFFA11" box="[286,412,1411,1437]" italics="true" pageId="8" pageNumber="578">P. ventralis</emphasis>
</taxonomicName>
than in
<taxonomicName id="C6E7F4C9D56BFF9EF8C092AEB44AFA10" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[505,617,1411,1436]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="33935358D56BFF9EF8C092AEB44AFA10" box="[505,617,1411,1436]" italics="true" pageId="8" pageNumber="578">P. graziae</emphasis>
</taxonomicName>
(
<figureCitation id="99DC93CFD56BFF9EFB4192AEB4D8FA11" box="[632,763,1410,1437]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Figs 50, 52</figureCitation>
), and the outer duct of the vesicular area is constricted only in
<taxonomicName id="C6E7F4C9D56BFF9EF98C928AB705FA4C" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[181,294,1447,1472]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="33935358D56BFF9EF98C928AB705FA4C" box="[181,294,1447,1472]" italics="true" pageId="8" pageNumber="578">P. graziae</emphasis>
</taxonomicName>
(
<figureCitation id="99DC93CFD56BFF9EF80F928AB7A8FA4D" box="[310,395,1446,1473]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,1249,1273]" captionTargetBox="[151,1435,181,1224]" captionTargetId="figure-32@11.[151,1435,181,1225]" captionTargetPageId="11" captionText="FIGURES 49–54. External (49, 51, 53) and internal (50, 52, 54) female genitalia of Podisus ventralis (49–50), Podisus graziae sp. n. (51–52), and Podisus thomasi sp. n. (53–54). Abbreviations: ar, arcus; cs, capsula seminalis; da, distal annular crest; id, inner duct; la9, laterotergites IX; md, median duct; od, outer duct; pa, proximal annular crest; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; tva 9, thickening of valvulae IX; va, vesicular area; vf8, valvifers VIII; vf9, valvifers IX; X, segment X. Scale bars: 49, 51, 53 = 0.5 mm; 50, 52, 54 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4692481" httpUri="https://zenodo.org/record/4692481/files/figure.png" pageId="8" pageNumber="578">Fig. 52</figureCitation>
). Besides that, these two species are allopatric in the Neotropical region;
<taxonomicName id="C6E7F4C9D56BFF9EFDF5928AB368FA4D" box="[1228,1355,1447,1473]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="33935358D56BFF9EFDF5928AB368FA4D" box="[1228,1355,1447,1473]" italics="true" pageId="8" pageNumber="578">P. ventralis</emphasis>
</taxonomicName>
can be found in the northern regions of South America (
<collectingCountry id="79F0CFDAD56BFF9EFB8292E6B510FA69" box="[699,819,1483,1509]" name="Venezuela" pageId="8" pageNumber="578">Venezuela</collectingCountry>
,
<collectingCountry id="79F0CFDAD56BFF9EFA0692E6B593FA69" box="[831,944,1483,1509]" name="Colombia" pageId="8" pageNumber="578">Colombia</collectingCountry>
,
<collectingCountry id="79F0CFDAD56BFF9EFA8592E6B23DFA6A" box="[956,1054,1483,1510]" name="Ecuador" pageId="8" pageNumber="578">Ecuador</collectingCountry>
), while
<taxonomicName id="C6E7F4C9D56BFF9EFD4F92E6B2C5FA68" authorityName="Roell &amp; Brugnera &amp; Lemaître" authorityYear="2021" box="[1142,1254,1483,1508]" class="Insecta" family="Pentatomidae" genus="Podisus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="578" phylum="Arthropoda" rank="species" species="graziae">
<emphasis id="33935358D56BFF9EFD4F92E6B2C5FA68" box="[1142,1254,1483,1508]" italics="true" pageId="8" pageNumber="578">P. graziae</emphasis>
</taxonomicName>
is distributed in the southern regions (
<collectingCountry id="79F0CFDAD56BFF9EF8B092C2B7F1F985" box="[393,466,1519,1545]" name="Brazil" pageId="8" pageNumber="578">Brazil</collectingCountry>
,
<collectingCountry id="79F0CFDAD56BFF9EF8E492C2B469F985" box="[477,586,1519,1545]" name="Paraguay" pageId="8" pageNumber="578">Paraguay</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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