treatments-xml/data/03/F1/5D/03F15D3E8C03FFFE3ED3FA02AC6AFCAE.xml
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<document id="D9BCF26BF0EAA3704314128F08C50E70" ID-DOI="10.5281/zenodo.187344" ID-GBIF-Dataset="229e851e-7107-4628-b823-7bf467cc98ad" ID-ISSN="1175-5326" ID-Zenodo-Dep="187344" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461067443105" checkinUser="plazi" docAuthor="Martins-Neto, Rafael Gioia &amp; Tassi, Lara Vaz" docDate="2009" docId="03F15D3E8C03FFFE3ED3FA02AC6AFCAE" docLanguage="en" docName="zt02080p037.pdf" docOrigin="Zootaxa 2080" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cearagrylloides perforatorius Martins-Neto, n. comb." docType="treatment" docVersion="10" lastPageNumber="28" masterDocId="FFC825468C05FFF93E44FFFEAD14FFB2" masterDocTitle="The Orthoptera (Ensifera) from the Santana formation (Early Cretaceous, Northeast Brazil): A statistical and paleoecological approach, with description of new taxa" masterLastPageNumber="37" masterPageNumber="21" pageNumber="27" updateTime="1698583641591" updateUser="plazi">
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<mods:title id="9051799D10F3E91058BFE75DB2A1EB95">The Orthoptera (Ensifera) from the Santana formation (Early Cretaceous, Northeast Brazil): A statistical and paleoecological approach, with description of new taxa</mods:title>
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<mods:namePart id="35A0146403148E23F769906C93E7D384">Martins-Neto, Rafael Gioia</mods:namePart>
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<mods:namePart id="2509BA14CA465333DD3778508FCE4CA8">Tassi, Lara Vaz</mods:namePart>
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<mods:date id="70533028C1963206FBE1C1DB1B0841F3">2009</mods:date>
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<treatment id="03F15D3E8C03FFFE3ED3FA02AC6AFCAE" ID-DOI="http://doi.org/10.5281/zenodo.5628401" ID-GBIF-Taxon="119615097" ID-Zenodo-Dep="5628401" LSID="urn:lsid:plazi:treatment:03F15D3E8C03FFFE3ED3FA02AC6AFCAE" httpUri="http://treatment.plazi.org/id/03F15D3E8C03FFFE3ED3FA02AC6AFCAE" lastPageId="7" lastPageNumber="28" pageId="6" pageNumber="27">
<subSubSection id="C342BFA38C03FFFF3ED3FA02ACAFF98B" pageId="6" pageNumber="27" type="nomenclature">
<paragraph id="8BE7EC288C03FFFF3ED3FA02AE32F9A4" blockId="6.[151,806,1532,1593]" box="[151,806,1532,1558]" pageId="6" pageNumber="27">
<heading id="D0AF5B448C03FFFF3ED3FA02AE32F9A4" bold="true" box="[151,806,1532,1558]" fontSize="11" level="1" pageId="6" pageNumber="27" reason="1">
<emphasis id="B92C303A8C03FFFF3ED3FA02AE32F9A4" bold="true" box="[151,806,1532,1558]" pageId="6" pageNumber="27">
<taxonomicName id="4C5897AB8C03FFFF3ED3FA02AFBBF9A4" authority="Martins-Neto" authorityName="Martins-Neto" box="[151,687,1532,1558]" class="Insecta" family="Cooloolidae" genus="Cearagrylloides" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="perforatorius" status="comb. nov.">
<emphasis id="B92C303A8C03FFFF3ED3FA02ACE8F9A4" bold="true" box="[151,508,1532,1558]" italics="true" pageId="6" pageNumber="27">Cearagrylloides perforatorius</emphasis>
Martins-Neto
</taxonomicName>
,
<taxonomicNameLabel id="A21F8D418C03FFFF3CFFFA02AE32F9A4" box="[699,806,1532,1558]" pageId="6" pageNumber="27">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7EC288C03FFFF3ED3F9E1ACAFF98B" blockId="6.[151,806,1532,1593]" box="[151,443,1567,1593]" pageId="6" pageNumber="27">
(Pl. IIB;
<figureCitation id="1363F0AD8C03FFFF3EBAF9E1AC5AF98B" box="[254,334,1567,1593]" captionStart="FIGURE 2" captionStartId="8.[151,256,1117,1141]" captionTargetBox="[185,1418,192,1067]" captionTargetId="figure@8.[167,1419,160,1093]" captionTargetPageId="8" captionText="FIGURE 2. Fore wing (A-C), ovipositor (D-G) and hind legs (H-K) details of the Santana Formation Cearagryllinae (females). A, D, H) Cearagrylloides perforatorius n. gen. et n. comb., drawn from the specimen RGMN- 506 (A), CV- 6015 (D, H); B, E) Cearagrylloides microcephalus n. gen. et n. comb., drawn from the specimen RGMN- 508 (B), and GP / 1 T- 1680 (E); C, G, K) Paracearagryllus poliacanthus n. gen. et n. comb., drawn from the specimen RGMN- 338 (C, G), and from holotype (K); F) Cearagrylloides previstus n. gen. et n. comb., drawn from holotype. Scale bar 1 mm." httpUri="https://zenodo.org/record/187346/files/figure.png" pageId="6" pageNumber="27">Figs. 2</figureCitation>
A, D, H.)
</paragraph>
</subSubSection>
<subSubSection id="C342BFA38C03FFFF3ED3F992A954F941" pageId="6" pageNumber="27" type="key">
<paragraph id="8BE7EC288C03FFFF3ED3F992AFA7F931" blockId="6.[151,691,1644,1667]" box="[151,691,1644,1667]" pageId="6" pageNumber="27">
<keyLead id="30A94F1D8C03FFFF3ED3F992AFA7F931" box="[151,691,1644,1667]" pageId="6" pageNumber="27">
<key id="FFC9BB198C03FFFF3ED3F992AFA7F931" box="[151,691,1644,1667]" pageId="6" pageNumber="27">
<keyStep id="30ACF48D8C03FFFF3ED3F992AFA7F931" box="[151,691,1644,1667]" pageId="6" pageNumber="27">
1.
<taxonomicName id="4C5897AB8C03FFFF3EF5F992AFBBF931" authority="Martins-Neto, 1991" authorityName="Martins-Neto" authorityYear="1991" box="[177,687,1644,1667]" class="Insecta" family="Cooloolidae" genus="Cearagryllus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="perforatorius">
<emphasis id="B92C303A8C03FFFF3EF5F992ACC6F931" box="[177,466,1644,1667]" italics="true" pageId="6" pageNumber="27">Cearagryllus perforatorius</emphasis>
<bibRefCitation id="EFC991D98C03FFFF3F9EF992AFBBF931" author="Martins-Neto" box="[474,687,1644,1667]" pageId="6" pageNumber="36" refString="Martins-Neto, R. G. (1991) Sistematica dos Ensifera (Insecta, Orthopteroida) da Formacao Santana, Cretaceo Inferior do Nordeste do Brasil Acta Geologica Leopoldensia, 32 (14), 3 - 162." type="journal article" year="1991">Martins-Neto, 1991</bibRefCitation>
</taxonomicName>
.
</keyStep>
</key>
</keyLead>
</paragraph>
<paragraph id="8BE7EC288C03FFFF3ED3F94CA954F941" blockId="6.[151,1436,1714,2011]" pageId="6" pageNumber="27">
<emphasis id="B92C303A8C03FFFF3ED3F94CAC80F97E" bold="true" box="[151,404,1714,1740]" pageId="6" pageNumber="27">Specimens included.</emphasis>
CV-6015 (
<typeStatus id="54E3528A8C03FFFF3C50F94CAF95F97E" box="[532,641,1714,1740]" pageId="6" pageNumber="27" type="holotype">holotype</typeStatus>
), DGM-6281 (
<typeStatus id="54E3528A8C03FFFF3D68F94CAE83F97E" box="[812,919,1714,1740]" pageId="6" pageNumber="27" type="paratype">paratype</typeStatus>
), CD-I-41, CV-1020, and the supplementary material originally included, RGMN-662, CV-6015, CV-2429, and RGMN-901.
</paragraph>
</subSubSection>
<subSubSection id="C342BFA38C03FFFE3ED3F8FEADDBFF6A" lastPageId="7" lastPageNumber="28" pageId="6" pageNumber="27" type="description">
<paragraph id="8BE7EC288C03FFFF3ED3F8FEAEF9F8A8" blockId="6.[151,1436,1714,2011]" box="[151,1005,1792,1818]" pageId="6" pageNumber="27">Additional specimens included. CV-1980, RGMN-506, and RGMN-507.</paragraph>
<paragraph id="8BE7EC288C03FFFF3E82F8D8AEEEF8F3" blockId="6.[151,1436,1714,2011]" box="[198,1018,1830,1857]" pageId="6" pageNumber="27">
<emphasis id="B92C303A8C03FFFF3E82F8D8ACD3F8F2" bold="true" box="[198,455,1830,1856]" pageId="6" pageNumber="27">Specimens excluded.</emphasis>
CV-1699, RGMN-203, CD-I-123, and CV-257.
</paragraph>
<paragraph id="8BE7EC288C03FFFE3E82F8B3ADDBFF6A" blockId="6.[151,1436,1714,2011]" lastBlockId="7.[151,1437,151,796]" lastPageId="7" lastPageNumber="28" pageId="6" pageNumber="27">
<emphasis id="B92C303A8C03FFFF3E82F8B3ACD0F8D5" bold="true" box="[198,452,1869,1895]" pageId="6" pageNumber="27">Emended diagnosis.</emphasis>
Females with robust body, varying from
<quantity id="4CA041CD8C03FFFF3D83F8B3A97DF8D5" box="[967,1129,1869,1895]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="6" pageNumber="27" unit="mm" value="30.5" valueMax="32.0" valueMin="29.0">29 to 32 mm</quantity>
length; fore wing length varying from
<quantity id="4CA041CD8C03FFFF3F7CF88AACC6F83C" box="[312,466,1908,1934]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="6" pageNumber="27" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">28 to 32 mm</quantity>
; ovipositor length varying from
<quantity id="4CA041CD8C03FFFF3D0BF88AAEFDF83C" box="[847,1001,1908,1934]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.6" metricValueMin="3.2" pageId="6" pageNumber="27" unit="mm" value="34.0" valueMax="36.0" valueMin="32.0">32 to 36 mm</quantity>
. Ovipositor 1.1 times longer than the body and/or the fore wing length. Relatively large and dorsally rounded head. Fore wing about three times longer than wide;
<emphasis id="B92C303A8C03FFFF3F29F83FAC88F869" bold="true" box="[365,412,1985,2011]" pageId="6" pageNumber="27">ScP</emphasis>
sigmoid, distally converging towards the costal margin and notably thickened basally;
<emphasis id="B92C303A8C02FFFE3ED3FF69ADB8FF03" bold="true" box="[151,172,151,177]" pageId="7" pageNumber="28">R</emphasis>
parallel to
<emphasis id="B92C303A8C02FFFE3F73FF69AC70FF03" bold="true" box="[311,356,151,177]" pageId="7" pageNumber="28">ScP</emphasis>
.
<emphasis id="B92C303A8C02FFFE3F30FF69ACB7FF03" bold="true" box="[372,419,151,177]" pageId="7" pageNumber="28">MP</emphasis>
two-branched; around nine
<emphasis id="B92C303A8C02FFFE3CBDFF69AE22FF03" bold="true" box="[761,822,151,177]" pageId="7" pageNumber="28">MP2</emphasis>
secondary branches.
<emphasis id="B92C303A8C02FFFE3A78FF69A97EFF03" bold="true" box="[1084,1130,151,177]" pageId="7" pageNumber="28">r-m</emphasis>
present, connecting
<emphasis id="B92C303A8C02FFFE3B23FF69A868FF03" bold="true" box="[1383,1404,151,177]" pageId="7" pageNumber="28">R</emphasis>
to
<emphasis id="B92C303A8C02FFFE3ED3FF40ADDCFF6A" bold="true" box="[151,200,190,216]" pageId="7" pageNumber="28">MA</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C342BFA38C02FFFE3E82FF1BAE75FEFE" pageId="7" pageNumber="28" type="discussion">
<paragraph id="8BE7EC288C02FFFE3E82FF1BAE75FEFE" blockId="7.[151,1437,151,796]" pageId="7" pageNumber="28">
<emphasis id="B92C303A8C02FFFE3E82FF1BAC46FF4D" bold="true" box="[198,338,229,255]" pageId="7" pageNumber="28">Discussion.</emphasis>
Following the examination of new supplementary material (especially well-preserved wings), several apomorphies of this species can be recognized, such as the sigmoid
<emphasis id="B92C303A8C02FFFE3A55FEF5A929FE97" bold="true" box="[1041,1085,267,293]" pageId="7" pageNumber="28">ScP</emphasis>
, thickened at its base, as well as the presence of
<emphasis id="B92C303A8C02FFFE3F2BFECCAC88FEFE" bold="true" box="[367,412,306,332]" pageId="7" pageNumber="28">r-m</emphasis>
, confirmed in all observed specimens.
</paragraph>
</subSubSection>
<subSubSection id="C342BFA38C02FFFE3E82FEA6A81FFDE9" pageId="7" pageNumber="28" type="description">
<paragraph id="8BE7EC288C02FFFE3E82FEA6A81FFDE9" blockId="7.[151,1437,151,796]" pageId="7" pageNumber="28">
<emphasis id="B92C303A8C02FFFE3E82FEA6AC43FEC0" bold="true" box="[198,343,344,370]" pageId="7" pageNumber="28">Description</emphasis>
(supplementary material RGMN-506). Body and forewing length
<quantity id="4CA041CD8C02FFFE3A2FFEA7A9D6FEC1" box="[1131,1218,345,371]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="28" unit="mm" value="32.0">32 mm</quantity>
. Ovipositor length as preserved
<quantity id="4CA041CD8C02FFFE3F75FE7EAC90FE28" box="[305,388,384,410]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="28" unit="mm" value="32.0">32 mm</quantity>
(total length ca.
<quantity id="4CA041CD8C02FFFE3C09FE7EAFB1FE28" box="[589,677,384,410]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="28" unit="mm" value="36.0">36 mm</quantity>
). Fore tibia smooth with a relatively long apical spur. Fore wing (
<figureCitation id="1363F0AD8C02FFFE3EE4FE58ADF3FE72" box="[160,231,422,448]" captionStart="FIGURE 3" captionStartId="13.[151,258,883,907]" captionTargetBox="[152,1413,184,882]" captionTargetId="figure@13.[151,1436,153,887]" captionTargetPageId="13" captionText="FIGURE 3. Dispersion curves for Araripe cearagryllids. A and B fore wing length and ovipositor, respectively for Euclydes ramosfernandesi Martins-Neto, 2007 (Stenopelmatoidea), a possible mimetic Orthoptera of cearagryllids." httpUri="https://zenodo.org/record/187347/files/figure.png" pageId="7" pageNumber="28">Fig. 3</figureCitation>
A) three times longer than wide, with wide costal area, narrowing towards the apex, filled by rather long cross-veins and accessory veinlets forming a mosaic of heterogeneous cells.
<emphasis id="B92C303A8C02FFFE3A21FE33A981FE55" bold="true" box="[1125,1173,461,487]" pageId="7" pageNumber="28">ScP</emphasis>
sigmoid, thickened at its base, converging towards the costal margin around a quarter wing-length from the apex.
<emphasis id="B92C303A8C02FFFE3A8AFE0DA9F7FDBF" bold="true" box="[1230,1251,499,525]" pageId="7" pageNumber="28">R</emphasis>
parallel to
<emphasis id="B92C303A8C02FFFE3B2DFE0DA881FDBF" bold="true" box="[1385,1429,499,525]" pageId="7" pageNumber="28">ScP</emphasis>
.
<emphasis id="B92C303A8C02FFFE3ED3FDE4ADA6FD86" bold="true" box="[151,178,538,564]" pageId="7" pageNumber="28">M</emphasis>
three-branched. Presence of
<emphasis id="B92C303A8C02FFFE3C4CFDE4AF21FD86" bold="true" box="[520,565,538,564]" pageId="7" pageNumber="28">r-m</emphasis>
connecting
<emphasis id="B92C303A8C02FFFE3C82FDE4AFCFFD86" bold="true" box="[710,731,538,564]" pageId="7" pageNumber="28">R</emphasis>
to
<emphasis id="B92C303A8C02FFFE3D45FDE4AE26FD86" bold="true" box="[769,818,538,564]" pageId="7" pageNumber="28">MA</emphasis>
, situated at the mid length of the wing. At least nine
<emphasis id="B92C303A8C02FFFE3ED3FDBEADC7FDE8" bold="true" box="[151,211,576,602]" pageId="7" pageNumber="28">MP2</emphasis>
secondary branches, all connected by cross-veins, forming a mosaic of heterogeneous cells.
</paragraph>
</subSubSection>
<subSubSection id="C342BFA38C02FFFE3E82FD99AC6AFCAE" pageId="7" pageNumber="28" type="discussion">
<paragraph id="8BE7EC288C02FFFE3E82FD99AC6AFCAE" blockId="7.[151,1437,151,796]" pageId="7" pageNumber="28">
<emphasis id="B92C303A8C02FFFE3E82FD99AC54FD33" bold="true" box="[198,320,615,641]" pageId="7" pageNumber="28">Remarks.</emphasis>
The specimen CV-1980, an isolated fore wing, was figured by
<bibRefCitation id="EFC991D98C02FFFE3A3EFD96A861FD30" author="Martins-Neto" box="[1146,1397,616,642]" pageId="7" pageNumber="36" refString="Martins-Neto, R. G. (1991) Sistematica dos Ensifera (Insecta, Orthopteroida) da Formacao Santana, Cretaceo Inferior do Nordeste do Brasil Acta Geologica Leopoldensia, 32 (14), 3 - 162." type="journal article" year="1991">Martins-Neto (1991)</bibRefCitation>
as belonging to
<taxonomicName id="4C5897AB8C02FFFE3F01FD71AFB5FD1A" box="[325,673,655,680]" class="Insecta" family="Cooloolidae" genus="Cearagryllus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="poliacanthus">
<emphasis id="B92C303A8C02FFFE3F01FD71AFB5FD1A" box="[325,673,655,680]" italics="true" pageId="7" pageNumber="28">Cearagryllus poliacanthus</emphasis>
</taxonomicName>
. Its wing venation is, in fact, virtually identical to that of
<taxonomicName id="4C5897AB8C02FFFE3ED3FD4BAFE1FD7D" authority="Martins-Neto (1991)" authorityName="Martins-Neto" authorityYear="1991" box="[151,757,693,719]" class="Insecta" family="Cooloolidae" genus="Cearagrylloides" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="perforatorius">
<emphasis id="B92C303A8C02FFFE3ED3FD4BACECFD7C" box="[151,504,693,718]" italics="true" pageId="7" pageNumber="28">Cearagrylloides perforatorius</emphasis>
<bibRefCitation id="EFC991D98C02FFFE3C44FD4BAFE1FD7D" author="Martins-Neto" box="[512,757,693,719]" pageId="7" pageNumber="36" refString="Martins-Neto, R. G. (1991) Sistematica dos Ensifera (Insecta, Orthopteroida) da Formacao Santana, Cretaceo Inferior do Nordeste do Brasil Acta Geologica Leopoldensia, 32 (14), 3 - 162." type="journal article" year="1991">Martins-Neto (1991)</bibRefCitation>
</taxonomicName>
, to which species it is now transferred. The specimen CV- 257 does not exhibit sufficient diagnostic characters to be included in the species, being now considered
<taxonomicName id="4C5897AB8C02FFFE3ED3FCFCAC42FCA9" box="[151,342,770,795]" class="Insecta" family="Cooloolidae" genus="Cearagrylloides" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B92C303A8C02FFFE3ED3FCFCAC42FCA9" box="[151,342,770,795]" italics="true" pageId="7" pageNumber="28">Cearagrylloides</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
</treatment>
</document>