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<document id="40C1C5B9BFE06062C8DB2FB026A1A448" ID-DOI="10.1649/0010-065x-68.2.241" ID-ISSN="1938-4394" ID-Zenodo-Dep="10085566" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1622480877462" checkinUser="carolina" docAuthor="Ratcliffe, Brett C." docDate="2014" docId="03DC9113632B516952B9FE00F137FB57" docLanguage="en" docName="TheColeopBull.68.2.241-262.pdf.imf" docOrigin="The Coleopterists Bulletin 68 (2)" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Badelina aterrima" docType="treatment" docVersion="2" lastPageNumber="245" masterDocId="FFE5E96B6328516D5220FFC6F510FFA9" masterDocTitle="A Review Of The Neotropical Genera Badelina Thomson, 1880, Balsameda Thomson, 1880, Guatemalica Neervoort Van De Poll, 1886, And Heterocotinis Martínez, 1948 (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini)" masterLastPageNumber="262" masterPageNumber="241" pageNumber="244" updateTime="1699477508221" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B2F3E5429AC49AB150BE2FCBF3723C28">
<mods:title id="C081168FFB2B5D7D5278A0135C2A8DFF">A Review Of The Neotropical Genera Badelina Thomson, 1880, Balsameda Thomson, 1880, Guatemalica Neervoort Van De Poll, 1886, And Heterocotinis Martínez, 1948 (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini)</mods:title>
</mods:titleInfo>
<mods:name id="DBBCB7107E8393ED2968821775596A91" type="personal">
<mods:role id="BF5FF2175D4CA41603E499F0C6927048">
<mods:roleTerm id="ABA51A26F57903CD1AA7D8617E391F96">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="44724424D41F31E84ACDCE3A65639825">Ratcliffe, Brett C.</mods:namePart>
<mods:affiliation id="FCB0C7E264153E37592C3F5C45FD7D19">Systematics Research Collections W 436 Nebraska Hall, University of Nebraska Lincoln, NE 68588 - 0514, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="31D059DAB13C28ACF037ACD81E29256B" type="email">bratcliffe1@unl.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="B8C7533A18A3435BABCC62DB281214A3">text</mods:typeOfResource>
<mods:relatedItem id="C3F88693EDC0CE35D08CB467B1923786" type="host">
<mods:titleInfo id="B7CFB0CD155536B29EA29419FB6F8770">
<mods:title id="BA1B2F3856DCC0CE8A15A66ADC17867B">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="865924C9AD88E1FD05143F26AE157DA9">
<mods:date id="5781A1F432214211F2CE792026A285A2">2014</mods:date>
<mods:detail id="8604B3A34F5F292B4BC3BC510D1A29D8" type="pubDate">
<mods:number id="CF177736D28F1D2BF6A993DC7035E50E">2014-06-01</mods:number>
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<mods:detail id="187372A2128F94425B53AAB9E239CD92" type="volume">
<mods:number id="EA1F7903D775ED9ABF2A22FB79A186B5">68</mods:number>
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<mods:number id="FB024DA90E00CB5A456D906DFC363C3D">2</mods:number>
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<mods:start id="6D4A62B52EA3A188AF946DD21478E288">241</mods:start>
<mods:end id="E60D0C790FF5C47C72237A26A011AE8B">262</mods:end>
</mods:extent>
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<mods:classification id="83C0D54DEF35B47F1A8D5D2409E0AFEF">journal article</mods:classification>
<mods:identifier id="DD3A2F0C6743AC1974730E091ED4B6EB" type="DOI">10.1649/0010-065x-68.2.241</mods:identifier>
<mods:identifier id="15A454D222D71C59B147C503BC96CA34" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="B76E65CF8BE28D612DF4D726EC566D21" type="Zenodo-Dep">10085566</mods:identifier>
</mods:mods>
<treatment id="03DC9113632B516952B9FE00F137FB57" LSID="urn:lsid:plazi:treatment:03DC9113632B516952B9FE00F137FB57" httpUri="http://treatment.plazi.org/id/03DC9113632B516952B9FE00F137FB57" lastPageId="4" lastPageNumber="245" pageId="3" pageNumber="244">
<subSubSection id="C36F738E632B516E52B9FE00F4D1FE5F" pageId="3" pageNumber="244" type="nomenclature">
<paragraph id="8BCA2005632B516E52B9FE00F4D1FE5F" blockId="3.[153,636,454,502]" pageId="3" pageNumber="244">
<taxonomicName id="4C755B86632B516E52B9FE00F76CFE72" authority="(Gory and Percheron, 1833)" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[153,636,454,475]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632B516E52B9FE00F76CFE72" bold="true" box="[153,636,454,475]" pageId="3" pageNumber="244">
<emphasis id="B901FC17632B516E52B9FE00F45EFE72" bold="true" box="[153,334,454,475]" italics="true" pageId="3" pageNumber="244">Badelina aterrima</emphasis>
(Gory and Percheron, 1833)
</emphasis>
</taxonomicName>
(
<figureCitation id="134E3C80632B516E537BFE27F4AAFE5F" box="[347,442,480,502]" captionStart="Figs" captionStartId="3.[175,215,1710,1729]" captionTargetBox="[154,1169,1031,1688]" captionTargetId="figure-379@3.[151,1172,1025,1704]" captionTargetPageId="3" captionText="Figs. 14. Badelina aterrima. 1) Habitus; 2) Mesometasternal process, lateral view; 34) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085568" httpUri="https://zenodo.org/record/10085568/files/figure.png" pageId="3" pageNumber="244">Figs. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36F738E632B516E52B7FDCAF743FDFF" pageId="3" pageNumber="244" type="reference_group">
<paragraph id="8BCA2005632B516E52B7FDCAF743FDFF" blockId="3.[151,638,523,598]" pageId="3" pageNumber="244">
<taxonomicName id="4C755B86632B516E52B7FDCAF5C6FD92" authority="Gory and Percheron 1833: 343" authorityName="Gory and Percheron" authorityPageNumber="343" authorityYear="1833" class="Insecta" family="Cetoniidae" genus="Gymnetis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632B516E52B7FDCAF44BFD88" box="[151,347,524,545]" italics="true" pageId="3" pageNumber="244">Gymnetis aterrima</emphasis>
<materialsCitation id="3B1D2A58632B516E5345FDCDF5C6FD92" collectionCode="MHNG" country="Brazil" location="Brazil" pageId="3" pageNumber="244" specimenCount="1" typeStatus="holotype">Gory and Percheron 1833: 343</materialsCitation>
</taxonomicName>
(original combination).
<typeStatus id="54CE9EA7632B516E53EFFDE0F73AFD92" box="[463,554,550,571]" pageId="3" pageNumber="244" type="holotype">Holotype</typeStatus>
male at MHNG, examined. Type locality: “
<collectingCountry id="F3626095632B516E5029FD87F752FDFF" box="[521,578,577,598]" name="Brazil" pageId="3" pageNumber="244">Brazil</collectingCountry>
”.
</paragraph>
</subSubSection>
<subSubSection id="C36F738E632B516E528FFDADF104FCB6" pageId="3" pageNumber="244" type="description">
<paragraph id="8BCA2005632B516E528FFDADF104FCB6" blockId="3.[151,639,619,959]" lastBlockId="3.[685,1173,194,959]" pageId="3" pageNumber="244">
<emphasis id="B901FC17632B516E528FFDADF43BFD29" bold="true" box="[175,299,619,640]" pageId="3" pageNumber="244">Description.</emphasis>
Length 24.0
<quantity id="4C8D8DE0632B516E53EDFDADF721FD29" box="[461,561,619,640]" metricMagnitude="-2" metricUnit="m" metricValue="2.78" pageId="3" pageNumber="244" unit="mm" value="27.8">27.8 mm</quantity>
; width across humeri 13.917.0 mm. Color black on dorsal surface; head, pronotum, and elytra velutinous. Venter an enamel grey to yellowish grey with black punctures.
<emphasis id="B901FC17632B516E5322FD13F428FD43" bold="true" box="[258,312,725,746]" pageId="3" pageNumber="244">Head</emphasis>
: Surface with punctures small to moderate in size and density. Clypeus with apex subtruncate to truncate, apex weakly reflexed. Eyes small, interocular width 6.06.5 transverse eye diameters. Antenna with 10 antennomeres, club distinctly longer than antennomeres
<quantity id="4C8D8DE0632B516E5027FC9CF759FCC6" box="[519,585,858,880]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.778" metricValueMin="0.508" pageId="3" pageNumber="244" unit="in" value="4.5" valueMax="7.0" valueMin="2.0">27 in</quantity>
both sexes.
<emphasis id="B901FC17632B516E52C0FCB3F45CFC23" bold="true" box="[224,332,885,906]" pageId="3" pageNumber="244">Pronotum</emphasis>
: Surface sculpturing usually obscured by velutinous covering; when exposed, punctures small, sparse. Sides with slender mar- ginal bead, bead usually not reaching anterior or posterior angles.
<emphasis id="B901FC17632B516E517EFF1BF6B0FF5B" bold="true" box="[862,928,221,242]" pageId="3" pageNumber="244">Elytra</emphasis>
: Surface with sculpturing usually obscured by velutinous covering; when exposed, punctures small, sparse, becoming moderate in size and crescent-shaped behind weak apical umbones. Apices at suture subacutely produced.
<emphasis id="B901FC17632B516E5136FEA4F667FEDE" bold="true" box="[790,887,354,375]" pageId="3" pageNumber="244">Pygidium</emphasis>
: Surface in both sexes with transverse, vermiform strigae, setigerous; setae short, dense, tawny, occasionally all black. In lateral view, profile in males nearly flat in basal two-thirds, convex in apical third; surface in females weakly convex.
<emphasis id="B901FC17632B516E51B9FE20F6C9FE52" bold="true" box="[921,985,486,507]" pageId="3" pageNumber="244">Venter</emphasis>
: Setae long, brown and black mixed, occasionally completely black. Mesometasternal process, in lateral view, protuberant, subparallel to ventral axis of body (
<figureCitation id="134E3C80632B516E566DFDF0F198FDE2" box="[1101,1160,566,587]" captionStart="Figs" captionStartId="3.[175,215,1710,1729]" captionTargetBox="[154,1169,1031,1688]" captionTargetId="figure-379@3.[151,1172,1025,1704]" captionTargetPageId="3" captionText="Figs. 14. Badelina aterrima. 1) Habitus; 2) Mesometasternal process, lateral view; 34) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085568" httpUri="https://zenodo.org/record/10085568/files/figure.png" pageId="3" pageNumber="244">Fig. 2</figureCitation>
), apex rounded in ventral view. Abdominal ventrites
<quantity id="4C8D8DE0632B516E508DFDADF7FDFD29" box="[685,749,619,641]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="3" pageNumber="244" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">15 in</quantity>
both sexes nearly smooth in central third, lateral thirds with sparse, moderately large, crescentshaped punctures.
<emphasis id="B901FC17632B516E5146FD66F686FD1C" bold="true" box="[870,918,672,693]" pageId="3" pageNumber="244">Legs</emphasis>
: Protibia slender, weakly tridentate in both sexes.
<emphasis id="B901FC17632B516E5187FD7DF106FD79" bold="true" box="[935,1046,699,720]" pageId="3" pageNumber="244">Parameres</emphasis>
(
<figureCitation id="134E3C80632B516E5606FD7DF198FD79" box="[1062,1160,698,720]" captionStart="Figs" captionStartId="3.[175,215,1710,1729]" captionTargetBox="[154,1169,1031,1688]" captionTargetId="figure-379@3.[151,1172,1025,1704]" captionTargetPageId="3" captionText="Figs. 14. Badelina aterrima. 1) Habitus; 2) Mesometasternal process, lateral view; 34) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085568" httpUri="https://zenodo.org/record/10085568/files/figure.png" pageId="3" pageNumber="244">Figs. 34</figureCitation>
): Form subrectangular in caudal view; apices broadly lanceolate, curving inwards, and with subapical, stout, spine-like tooth on lateral edge.
</paragraph>
</subSubSection>
<subSubSection id="C36F738E632B516950E5FCE3F456FBF7" lastPageId="4" lastPageNumber="245" pageId="3" pageNumber="244" type="distribution">
<paragraph id="8BCA2005632B516E50E5FCE3F6C1FCFC" blockId="3.[685,1173,194,959]" pageId="3" pageNumber="244">
<emphasis id="B901FC17632B516E50E5FCE3F658FC93" bold="true" box="[709,840,805,826]" pageId="3" pageNumber="244">Distribution.</emphasis>
<taxonomicName id="4C755B86632B516E5175FCE3F114FC93" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[853,1028,805,826]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632B516E5175FCE3F114FC93" box="[853,1028,805,826]" italics="true" pageId="3" pageNumber="244">Badelina aterrima</emphasis>
</taxonomicName>
is known from southern
<collectingCountry id="F3626095632B516E5127FC86F650FCFC" box="[775,832,832,853]" name="Brazil" pageId="3" pageNumber="244">Brazil</collectingCountry>
and
<collectingCountry id="F3626095632B516E5153FC86F6DDFCFC" box="[883,973,832,853]" name="Paraguay" pageId="3" pageNumber="244">Paraguay</collectingCountry>
.
</paragraph>
<paragraph id="8BCA2005632B516950E5FC9CF455FC5D" blockId="3.[685,1173,194,959]" lastBlockId="4.[173,660,911,1729]" lastPageId="4" lastPageNumber="245" pageId="3" pageNumber="244">
<emphasis id="B901FC17632B516E50E5FC9CF66AFCC6" bold="true" box="[709,890,858,879]" pageId="3" pageNumber="244">Locality Records.</emphasis>
<materialsCitation id="3B1D2A58632B516951A8FC9CF450FC5D" collectionCode="BCRC, BMNH, CASC, CMNH, FMNH, MHNG, MLUH, MNHN, RMNH, UFRJ, USNM, ZMHU" country="Brazil" county="No" lastPageId="4" lastPageNumber="245" location="No" municipality="No" pageId="3" pageNumber="244" specimenCount="58" stateProvince="Asuncion">
<specimenCount id="9D73EB8C632B516E51A8FC9CF11DFCC6" box="[904,1037,858,879]" pageId="3" pageNumber="244" type="generic">58 specimens</specimenCount>
from
<collectionCode id="ED64B8C0632B516E566DFC9CF180FCC6" box="[1101,1168,858,879]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="3" pageNumber="244">BCRC</collectionCode>
,
<collectionCode id="ED64B8C0632B516E508DFCB3F7E7FC23" box="[685,759,885,906]" pageId="3" pageNumber="244">BMNH</collectionCode>
,
<collectionCode id="ED64B8C0632B516E5122FCB3F653FC23" box="[770,835,885,906]" pageId="3" pageNumber="244">CASC</collectionCode>
,
<collectionCode id="ED64B8C0632B516E516DFCB3F687FC23" box="[845,919,885,906]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="3" pageNumber="244" type="Museum">CMNH</collectionCode>
,
<collectionCode id="ED64B8C0632B516E5181FCB3F6F8FC23" box="[929,1000,885,906]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="244" type="Museum">FMNH</collectionCode>
,
<collectionCode id="ED64B8C0632B516E51D2FCB3F12DFC23" box="[1010,1085,885,906]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="244" type="Museum">MHNG</collectionCode>
,
<collectionCode id="ED64B8C0632B516E5668FCB3F180FC23" box="[1096,1168,885,906]" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="3" pageNumber="244">MLUH</collectionCode>
,
<collectionCode id="ED64B8C0632B516E508DFC49F7EDFC0D" box="[685,765,911,932]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="244">MNHN</collectionCode>
,
<collectionCode id="ED64B8C0632B516E512CFC49F64BFC0D" box="[780,859,911,932]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="244" type="Museum">RMNH</collectionCode>
,
<collectionCode id="ED64B8C0632B516E514AFC49F6B9FC0D" box="[874,937,911,932]" pageId="3" pageNumber="244">UFRJ</collectionCode>
,
<collectionCode id="ED64B8C0632B516E5198FC49F114FC0D" box="[952,1028,911,932]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="244" type="Museum">USNM</collectionCode>
, and
<collectionCode id="ED64B8C0632B516E5663FC49F181FC0D" box="[1091,1169,911,932]" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="3" pageNumber="244">ZMHU</collectionCode>
.
<emphasis id="B901FC17632B516E508DFC6CF658FC16" bold="true" box="[685,840,938,959]" pageId="3" pageNumber="244">
<collectingCountry id="F3626095632B516E508DFC6CF61BFC16" box="[685,779,938,959]" name="Brazil" pageId="3" pageNumber="244">BRAZIL</collectingCountry>
(57).
</emphasis>
RIO DE JANEIRO (18): Rio de Janeiro,
<collectingCounty id="62AB5889632C51695329FC49F437FC0D" box="[265,295,911,932]" pageId="4" pageNumber="245">No</collectingCounty>
data. SÃO PAULO (1):
<collectingMunicipality id="6BAEBA7F632C51695018FC49F746FC0D" box="[568,598,911,932]" pageId="4" pageNumber="245">No</collectingMunicipality>
data. SANTA CATARINA (4):
<location id="8EAA76DE632C5169539FFC6CF4CDFC16" LSID="urn:lsid:plazi:treatment:03DC9113632B516952B9FE00F137FB57:8EAA76DE632C5169539FFC6CF4CDFC16" box="[447,477,938,959]" country="Brazil" county="No" municipality="No" name="No" pageId="4" pageNumber="245" stateProvince="Asuncion">No</location>
data. NO DATA (34).
<emphasis id="B901FC17632C516952C6FC03F4B0FC73" bold="true" box="[230,416,965,986]" pageId="4" pageNumber="245">
<collectingCountry id="F3626095632C516952C6FC03F47DFC73" box="[230,365,965,986]" name="Paraguay" pageId="4" pageNumber="245">PARAGUAY</collectingCountry>
(1).
</emphasis>
<collectingRegion id="49B1EEE7632C5169538BFC03F719FC73" box="[427,521,965,986]" country="Paraguay" name="Asuncion" pageId="4" pageNumber="245">Asunción</collectingRegion>
(autonomous capital district)
</materialsCitation>
.
</paragraph>
<caption id="DF0A708D632B516E528FF968F120F968" ID-DOI="http://doi.org/10.5281/zenodo.10085568" ID-Zenodo-Dep="10085568" box="[175,1072,1710,1729]" httpUri="https://zenodo.org/record/10085568/files/figure.png" pageId="3" pageNumber="244" startId="3.[175,215,1710,1729]" targetBox="[154,1169,1031,1688]" targetPageId="3">
<paragraph id="8BCA2005632B516E528FF968F120F968" blockId="3.[175,1072,1710,1729]" box="[175,1072,1710,1729]" pageId="3" pageNumber="244">
<emphasis id="B901FC17632B516E528FF968F41BF968" bold="true" box="[175,267,1710,1729]" pageId="3" pageNumber="244">Figs. 14.</emphasis>
<taxonomicName id="4C755B86632B516E5300F968F4AEF969" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[288,446,1710,1728]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="244" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632B516E5300F968F4AEF969" box="[288,446,1710,1728]" italics="true" pageId="3" pageNumber="244">Badelina aterrima</emphasis>
</taxonomicName>
.
<emphasis id="B901FC17632B516E53EAF968F4CDF968" bold="true" box="[458,477,1710,1729]" pageId="3" pageNumber="244">1)</emphasis>
Habitus;
<emphasis id="B901FC17632B516E5013F968F756F968" bold="true" box="[563,582,1710,1729]" pageId="3" pageNumber="244">2)</emphasis>
Mesometasternal process, lateral view;
<emphasis id="B901FC17632B516E5180F968F6DAF968" bold="true" box="[928,970,1710,1729]" pageId="3" pageNumber="244">34)</emphasis>
Parameres.
</paragraph>
</caption>
<caption id="DF0A708D632C516952E5FC8BF15EFCC9" ID-DOI="http://doi.org/10.5281/zenodo.10085570" ID-Zenodo-Dep="10085570" box="[197,1102,845,864]" httpUri="https://zenodo.org/record/10085570/files/figure.png" pageId="4" pageNumber="245" startId="4.[197,237,845,864]" targetBox="[175,1189,189,823]" targetPageId="4">
<paragraph id="8BCA2005632C516952E5FC8BF15EFCC9" blockId="4.[197,1102,845,864]" box="[197,1102,845,864]" pageId="4" pageNumber="245">
<emphasis id="B901FC17632C516952E5FC8BF430FCC9" bold="true" box="[197,288,845,864]" pageId="4" pageNumber="245">Figs. 58.</emphasis>
<taxonomicName id="4C755B86632C51695316FC8BF4CCFCF6" baseAuthorityName="Thomson" baseAuthorityYear="1878" box="[310,476,845,863]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B901FC17632C51695316FC8BF4CCFCF6" box="[310,476,845,863]" italics="true" pageId="4" pageNumber="245">Badelina pygidialis</emphasis>
</taxonomicName>
.
<emphasis id="B901FC17632C516953C8FC8BF4ECFCC9" bold="true" box="[488,508,845,864]" pageId="4" pageNumber="245">5)</emphasis>
Habitus;
<emphasis id="B901FC17632C51695071FC8BF774FCC9" bold="true" box="[593,612,845,864]" pageId="4" pageNumber="245">6)</emphasis>
Mesometasternal process, lateral view;
<emphasis id="B901FC17632C5169519FFC8BF6F9FCC9" bold="true" box="[959,1001,845,864]" pageId="4" pageNumber="245">78)</emphasis>
Parameres.
</paragraph>
</caption>
<paragraph id="8BCA2005632C516952E5FC3CF456FBF7" blockId="4.[173,660,911,1729]" pageId="4" pageNumber="245">
<emphasis id="B901FC17632C516952E5FC3CF4A6FBA6" bold="true" box="[197,438,1018,1039]" pageId="4" pageNumber="245">Temporal Distribution.</emphasis>
January (2), February (1), March (1), October (1), November (3), December (1). Most specimens have little or no collecting data.
</paragraph>
</subSubSection>
<subSubSection id="C36F738E632C516952E5FBA2F137FB57" pageId="4" pageNumber="245" type="diagnosis">
<paragraph id="8BCA2005632C516952E5FBA2F671FBD0" blockId="4.[173,660,911,1729]" lastBlockId="4.[707,1193,911,1278]" pageId="4" pageNumber="245">
<emphasis id="B901FC17632C516952E5FBA2F43DFBD0" bold="true" box="[197,301,1124,1145]" pageId="4" pageNumber="245">Diagnosis.</emphasis>
<taxonomicName id="4C755B86632C5169531BFBA2F4FFFBD0" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[315,495,1124,1145]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632C5169531BFBA2F4FFFBD0" box="[315,495,1124,1145]" italics="true" pageId="4" pageNumber="245">Badelina aterrima</emphasis>
</taxonomicName>
is distinguished by a velutinous, totally black color on the pronotum and elytra (
<figureCitation id="134E3C80632C51695309FB5FF479FB07" box="[297,361,1177,1198]" captionStart="Figs" captionStartId="3.[175,215,1710,1729]" captionTargetBox="[154,1169,1031,1688]" captionTargetId="figure-379@3.[151,1172,1025,1704]" captionTargetPageId="3" captionText="Figs. 14. Badelina aterrima. 1) Habitus; 2) Mesometasternal process, lateral view; 34) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085568" httpUri="https://zenodo.org/record/10085568/files/figure.png" pageId="4" pageNumber="245">Fig. 1</figureCitation>
).
<taxonomicName id="4C755B86632C516953A3FB5FF742FB07" baseAuthorityName="Thomson" baseAuthorityYear="1878" box="[387,594,1177,1198]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B901FC17632C516953A3FB5FF742FB07" box="[387,594,1177,1198]" italics="true" pageId="4" pageNumber="245">Badelina pygidialis</emphasis>
</taxonomicName>
has a brown pronotum and elytra, each with small, black spots (
<figureCitation id="134E3C80632C516952CEFB08F43CFB4A" box="[238,300,1230,1251]" captionStart="Figs" captionStartId="4.[197,237,845,864]" captionTargetBox="[175,1189,189,823]" captionTargetId="figure-445@4.[172,1193,183,837]" captionTargetPageId="4" captionText="Figs. 58. Badelina pygidialis. 5) Habitus; 6) Mesometasternal process, lateral view; 78) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085570" httpUri="https://zenodo.org/record/10085570/files/figure.png" pageId="4" pageNumber="245">Fig. 5</figureCitation>
). The parameres of
<taxonomicName id="4C755B86632C516953D6FB08F777FB4A" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[502,615,1230,1251]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632C516953D6FB08F777FB4A" box="[502,615,1230,1251]" italics="true" pageId="4" pageNumber="245">B. aterrima</emphasis>
</taxonomicName>
and
<taxonomicName id="4C755B86632C5169528DFB2FF435FB57" baseAuthorityName="Thomson" baseAuthorityYear="1878" box="[173,293,1257,1278]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B901FC17632C5169528DFB2FF435FB57" box="[173,293,1257,1278]" italics="true" pageId="4" pageNumber="245">B. pygidialis</emphasis>
</taxonomicName>
are very similar (
<figureCitation id="134E3C80632C516953F4FB2FF73FFB57" box="[468,559,1256,1278]" captionStart="Figs" captionStartId="3.[175,215,1710,1729]" captionTargetBox="[154,1169,1031,1688]" captionTargetId="figure-379@3.[151,1172,1025,1704]" captionTargetPageId="3" captionText="Figs. 14. Badelina aterrima. 1) Habitus; 2) Mesometasternal process, lateral view; 34) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085568" httpUri="https://zenodo.org/record/10085568/files/figure.png" pageId="4" pageNumber="245">Figs. 34</figureCitation>
and
<figureCitation id="134E3C80632C51695042FB2FF797FB57" box="[610,647,1256,1278]" captionStart="Figs" captionStartId="4.[197,237,845,864]" captionTargetBox="[175,1189,189,823]" captionTargetId="figure-445@4.[172,1193,183,837]" captionTargetPageId="4" captionText="Figs. 58. Badelina pygidialis. 5) Habitus; 6) Mesometasternal process, lateral view; 78) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085570" httpUri="https://zenodo.org/record/10085570/files/figure.png" pageId="4" pageNumber="245">78</figureCitation>
), and so I debated whether
<taxonomicName id="4C755B86632C5169538EFAC5F70DFAB1" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[430,541,1283,1304]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632C5169538EFAC5F70DFAB1" box="[430,541,1283,1304]" italics="true" pageId="4" pageNumber="245">B. aterrima</emphasis>
</taxonomicName>
was simply a melanistic phase of
<taxonomicName id="4C755B86632C516953A7FAD8F712FA9A" baseAuthorityName="Thomson" baseAuthorityYear="1878" box="[391,514,1310,1331]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B901FC17632C516953A7FAD8F712FA9A" box="[391,514,1310,1331]" italics="true" pageId="4" pageNumber="245">B. pygidialis</emphasis>
</taxonomicName>
. I believe that the two species are different in spite of the overall similarity of the parameres which, in many
<taxonomicName id="4C755B86632C5169528DFAA8F40EFA2A" box="[173,286,1390,1411]" class="Insecta" family="Cetoniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="tribe" tribe="Gymnetini">Gymnetini</taxonomicName>
, are often not diagnostic anyway. Nevertheless, there are differences in the genitalia. In lateral view, the length of the basal piece is different between the two species. In
<taxonomicName id="4C755B86632C5169503EFA7BF79DFA7B" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[542,653,1469,1490]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632C5169503EFA7BF79DFA7B" box="[542,653,1469,1490]" italics="true" pageId="4" pageNumber="245">B. aterrima</emphasis>
</taxonomicName>
, the basal piece is proportionately shorter, as measured against the length of the parameres, than in
<taxonomicName id="4C755B86632C5169528DF9CBF435F98B" baseAuthorityName="Thomson" baseAuthorityYear="1878" box="[173,293,1549,1570]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B901FC17632C5169528DF9CBF435F98B" box="[173,293,1549,1570]" italics="true" pageId="4" pageNumber="245">B. pygidialis</emphasis>
</taxonomicName>
(
<figureCitation id="134E3C80632C51695315F9CBF469F98B" box="[309,377,1549,1570]" captionStart="Figs" captionStartId="3.[175,215,1710,1729]" captionTargetBox="[154,1169,1031,1688]" captionTargetId="figure-379@3.[151,1172,1025,1704]" captionTargetPageId="3" captionText="Figs. 14. Badelina aterrima. 1) Habitus; 2) Mesometasternal process, lateral view; 34) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085568" httpUri="https://zenodo.org/record/10085568/files/figure.png" pageId="4" pageNumber="245">Figs. 4</figureCitation>
and
<figureCitation id="134E3C80632C5169538DF9CBF4AAF98B" box="[429,442,1549,1570]" captionStart="Figs" captionStartId="4.[197,237,845,864]" captionTargetBox="[175,1189,189,823]" captionTargetId="figure-445@4.[172,1193,183,837]" captionTargetPageId="4" captionText="Figs. 58. Badelina pygidialis. 5) Habitus; 6) Mesometasternal process, lateral view; 78) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085570" httpUri="https://zenodo.org/record/10085570/files/figure.png" pageId="4" pageNumber="245">8</figureCitation>
). And in caudal view, the subapical spine on each paramere is much broader in
<taxonomicName id="4C755B86632C51695337F984F495F9FE" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[279,389,1602,1623]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632C51695337F984F495F9FE" box="[279,389,1602,1623]" italics="true" pageId="4" pageNumber="245">B. aterrima</emphasis>
</taxonomicName>
than the slender spine seen in
<taxonomicName id="4C755B86632C516952E7F99BF42EF9DB" baseAuthorityName="Thomson" baseAuthorityYear="1878" box="[199,318,1629,1650]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B901FC17632C516952E7F99BF42EF9DB" box="[199,318,1629,1650]" italics="true" pageId="4" pageNumber="245">B. pygidialis</emphasis>
</taxonomicName>
(
<figureCitation id="134E3C80632C5169536DF99BF480F9DB" box="[333,400,1629,1650]" captionStart="Figs" captionStartId="3.[175,215,1710,1729]" captionTargetBox="[154,1169,1031,1688]" captionTargetId="figure-379@3.[151,1172,1025,1704]" captionTargetPageId="3" captionText="Figs. 14. Badelina aterrima. 1) Habitus; 2) Mesometasternal process, lateral view; 34) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085568" httpUri="https://zenodo.org/record/10085568/files/figure.png" pageId="4" pageNumber="245">Figs. 3</figureCitation>
and
<figureCitation id="134E3C80632C516953E2F99BF4C0F9DB" box="[450,464,1629,1650]" captionStart="Figs" captionStartId="4.[197,237,845,864]" captionTargetBox="[175,1189,189,823]" captionTargetId="figure-445@4.[172,1193,183,837]" captionTargetPageId="4" captionText="Figs. 58. Badelina pygidialis. 5) Habitus; 6) Mesometasternal process, lateral view; 78) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10085570" httpUri="https://zenodo.org/record/10085570/files/figure.png" pageId="4" pageNumber="245">7</figureCitation>
). Furthermore, there are never any intergrades of body color or pattern as is usually seen in other gymnetine species that have melanistic morphs. I completely immersed several specimens of
<taxonomicName id="4C755B86632C516951B6FC56F115FC0C" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[918,1029,912,933]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632C516951B6FC56F115FC0C" box="[918,1029,912,933]" italics="true" pageId="4" pageNumber="245">B. aterrima</emphasis>
</taxonomicName>
in ethanol for a lengthy period to see if there might be any underlying color or spot pattern typical of
<taxonomicName id="4C755B86632C51695610FC03F1B9FC73" baseAuthorityName="Thomson" baseAuthorityYear="1878" box="[1072,1193,965,986]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B901FC17632C51695610FC03F1B9FC73" box="[1072,1193,965,986]" italics="true" pageId="4" pageNumber="245">B. pygidialis</emphasis>
</taxonomicName>
that would become apparent upon being wetted, but there was nothing other than black. Because of the differences in the genitalia and the totally disparate color forms that have no intergrades, I maintain
<taxonomicName id="4C755B86632C51695113FB8CF6B6FBF6" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[819,934,1098,1119]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632C51695113FB8CF6B6FBF6" box="[819,934,1098,1119]" italics="true" pageId="4" pageNumber="245">B. aterrima</emphasis>
</taxonomicName>
and
<taxonomicName id="4C755B86632C516951FEFB8CF14BFBF6" baseAuthorityName="Thomson" baseAuthorityYear="1878" box="[990,1115,1098,1119]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B901FC17632C516951FEFB8CF14BFBF6" box="[990,1115,1098,1119]" italics="true" pageId="4" pageNumber="245">B. pygidialis</emphasis>
</taxonomicName>
as two separate species.
</paragraph>
<paragraph id="8BCA2005632C516950FBFBB9F137FB57" blockId="4.[707,1193,911,1278]" pageId="4" pageNumber="245">
<emphasis id="B901FC17632C516950FBFBB9F693FB3D" bold="true" box="[731,899,1151,1172]" pageId="4" pageNumber="245">Natural History.</emphasis>
Nothing is known of the life history of these beetles.
<bibRefCitation id="EFE45DF4632C516951E2FB5FF15CFB07" author="Ohaus" box="[962,1100,1177,1198]" pageId="4" pageNumber="245" pagination="193 - 274" refId="ref16255" refString="Ohaus, F. 1900. Bericht uber eine entomologische Reise nach Centralbrasilien. Entomologische Zeitung 61: 193 - 274." type="journal article" year="1900">Ohaus (1900)</bibRefCitation>
reported young larvae of
<taxonomicName id="4C755B86632C51695149FB72F6CEFB60" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[873,990,1204,1225]" class="Insecta" family="Cetoniidae" genus="Badelina" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="245" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis id="B901FC17632C51695149FB72F6CEFB60" box="[873,990,1204,1225]" italics="true" pageId="4" pageNumber="245">G. aterrima</emphasis>
</taxonomicName>
living in detritus in the earth near rotten tree stumps, and larger larvae were found inside the rotten stumps.
</paragraph>
</subSubSection>
</treatment>
</document>