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<document id="3554A587ED98B801D84E04DE0F972585" ID-DOI="10.5852/ejt.2024.925.2465" ID-ISSN="2118-9773" ID-Zenodo-Dep="10824968" ID-ZooBank="5848AF9A-F01D-45F1-8626-E489AC15EE7F" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1710590167793" checkinUser="plazi" docAuthor="Génier, François &amp; Darling, James D. G." docDate="2024" docId="03DC87B7EF07FF99FE30FE0ECC4CE761" docLanguage="en" docName="EJT.2024.925.220-267.pdf" docOrigin="European Journal of Taxonomy 925" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2465/10955" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Homocopris williami Darling &amp; Genier 2024, sp. nov." docType="treatment" docUuid="94FAF15A-7945-460D-BF15-69FB3D05CBD0" docUuidSource="ZooBank" docVersion="8" lastPageNumber="259" masterDocId="FFE5FFCFEF25FFBEFFCFFF9FC93BE277" masterDocTitle="Revision of the South American genera Andinocopris new genus and Homocopris Burmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae: Homocoprini new tribe)" masterLastPageNumber="267" masterPageNumber="220" pageNumber="254" updateTime="1710953094570" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3A449A6D17A2F5E89F9F3B74154040F3">Revision of the South American genera Andinocopris new genus and Homocopris Burmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae: Homocoprini new tribe)</mods:title>
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<mods:namePart id="291F7720B95051631C88C261FEFC031F">Génier, François</mods:namePart>
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<mods:affiliation id="C4C7B7A8BE2225FC595A1442D3D51270">Beaty Center for Species Discovery, Canadian Museum of Nature, 1740 Pink Road, Gatineau, Québec, J 9 J 3 N 7, Canada.</mods:affiliation>
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<mods:namePart id="D3DDF8BC2B79A12F91221828D5E6AD14">Darling, James D. G.</mods:namePart>
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<mods:affiliation id="577F2004FA725E690D71299D3C56960F">Beaty Center for Species Discovery, Canadian Museum of Nature, 1740 Pink Road, Gatineau, Québec, J 9 J 3 N 7, Canada.</mods:affiliation>
<mods:nameIdentifier id="342BE71E0BFDF7613990E86B60FAB64B" type="email">james21darling@gmail.com</mods:nameIdentifier>
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<mods:date id="6799B035381F706D5199653FDE9F649A">2024</mods:date>
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<paragraph id="8BCA36A1EF07FF9CFE30FE0ECD0EE3DC" blockId="34.[364,1223,400,498]" box="[511,1077,400,427]" pageId="34" pageNumber="254">
<heading id="D08281CDEF07FF9CFE30FE0ECD0EE3DC" box="[511,1077,400,427]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="254" reason="2">
<taxonomicName id="4C754D22EF07FF9CFE30FE0ECAE8E3DC" authority="Darling &amp; Genier" authorityName="Darling &amp; Genier" authorityYear="2024" box="[511,979,400,427]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="254" phylum="Arthropoda" rank="species" species="williami" status="sp. nov.">
<emphasis id="B901EAB3EF07FF9CFE30FE0ECBC6E3DC" bold="true" box="[511,765,400,427]" italics="true" pageId="34" pageNumber="254">Homocopris williami</emphasis>
Darling &amp; Génier
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<taxonomicNameLabel id="A23257C8EF07FF9CFC15FE0ECD0EE3DC" box="[986,1077,401,427]" pageId="34" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BCA36A1EF07FF9CFD9AFE48CAE5E386" blockId="34.[364,1223,400,498]" box="[597,990,471,498]" pageId="34" pageNumber="254">
<figureCitation id="134E2A24EF07FF9CFD9AFE48CBE2E386" box="[597,729,471,497]" captionStart="Figs 9–12" captionStartId="11.[189,242,1928,1954]" captionTargetBox="[207,1370,321,1842]" captionTargetId="figure-18@11.[207,1378,265,1880]" captionTargetPageId="11" captionText="Figs 9–12. Dorsal habitus. 9. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 10. Homocopris torulosus, specimen ♀ (repository unknown). 11. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 12. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824974" httpUri="https://zenodo.org/record/10824974/files/figure.png" pageId="34" pageNumber="254">Figs 11–12</figureCitation>
, 
<figureCitation id="134E2A24EF07FF9CFD2AFE48CA09E386" box="[741,818,471,497]" captionStart="Figs 21–24" captionStartId="16.[189,242,1928,1954]" captionTargetBox="[210,1378,289,1875]" captionTargetId="figure-12@16.[210,1378,265,1875]" captionTargetPageId="16" captionText="Figs 21–24. Head and pronotum, lateral view.21. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 22. Homocopris torulosus, specimen ♀ (repository unknown). 23. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 24. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824982" httpUri="https://zenodo.org/record/10824982/files/figure.png" pageId="34" pageNumber="254">23–24</figureCitation>
, 
<figureCitation id="134E2A24EF07FF9CFCF2FE48CA66E386" box="[829,861,471,497]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="34" pageNumber="254">30</figureCitation>
, 
<figureCitation id="134E2A24EF07FF9CFCA6FE48CAB2E386" box="[873,905,471,497]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="34" pageNumber="254">36</figureCitation>
, 
<figureCitation id="134E2A24EF07FF9CFC5AFE48CA8EE385" box="[917,949,471,498]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling &amp; Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="34" pageNumber="254">55</figureCitation>
, 
<figureCitation id="134E2A24EF07FF9CFC0EFE48CAE5E386" box="[961,990,471,497]" captionStart="Fig" captionStartId="37.[189,232,1952,1978]" captionTargetBox="[189,1398,998,1904]" captionTargetId="figure-178@37.[189,1398,997,1904]" captionTargetPageId="37" captionText="Fig. 66. Homocopris grossiorum Darling &amp; Génier sp. nov. and Homocopris williami Darling &amp; Génier sp. nov., distribution map. Legend and scale bar included in the top and bottom right corners, respectively." figureDoi="http://doi.org/10.5281/zenodo.10825000" httpUri="https://zenodo.org/record/10825000/files/figure.png" pageId="34" pageNumber="254">66</figureCitation>
</paragraph>
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<treatmentCitationGroup id="AB65118FEF07FF9CFF72FD86CA22E044" box="[189,793,537,563]" pageId="34" pageNumber="254">
<taxonomicName id="4C754D22EF07FF9CFF72FD86C8B0E044" ID-CoL="7HRQS" box="[189,395,537,563]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="254" phylum="Arthropoda" rank="species" species="torulosus">
<emphasis id="B901EAB3EF07FF9CFF72FD86C8B0E044" box="[189,395,537,563]" italics="true" pageId="34" pageNumber="254">Pinotus torulosus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AD410B0EF07FF9CFE67FD86CB54E044" author="Lange R. B." box="[424,623,537,563]" page="313" pageId="34" pageNumber="254" year="1947">
<bibRefCitation id="EFE44B50EF07FF9CFE67FD86CB54E044" author="Lange R. B." box="[424,623,537,563]" pageId="34" pageNumber="254" pagination="305 - 315" refId="ref26605" refString="Lange R. B. 1947. Ensaio da zoogeografia dos Scarabaeidae do Parana com algumas notas eto-ecologicas. Arquivos do Museu Paranaense 6: 305 - 315." type="journal article" year="1947">Lange 1947: 313</bibRefCitation>
</treatmentCitation>
(distribution).
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</paragraph>
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<subSubSection id="C36F652AEF07FF9CFF72FDC5CAA8E158" pageId="34" pageNumber="254" type="diagnosis">
<paragraph id="8BCA36A1EF07FF9CFF72FDC5C8F2E003" blockId="34.[189,457,602,628]" box="[189,457,602,628]" pageId="34" pageNumber="254">
<heading id="D08281CDEF07FF9CFF72FDC5C8F2E003" bold="true" box="[189,457,602,628]" fontSize="11" level="3" pageId="34" pageNumber="254" reason="3">
<emphasis id="B901EAB3EF07FF9CFF72FDC5C8F2E003" bold="true" box="[189,457,602,628]" pageId="34" pageNumber="254">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36A1EF07FF9CFF72FD16CAA8E158" blockId="34.[188,1401,649,816]" pageId="34" pageNumber="254">
Dorsal ocular width subequal to one-sixth interocular distance; anteromedial pronotal edge curvature tends to be continuous; posterior pronotal surface uniformly punctate; elytral interstriae punctation tends to be visible at 10 × magnification; metasternal depression divided anteriorly. 
<specimenCount id="9D73FD28EF07FF9CFB88FD4FCD66E09D" box="[1095,1117,720,746]" count="1" pageId="34" pageNumber="254" type="male">♂</specimenCount>
pronotal ridge with two sets of distinct bilaterally paired tubercles in large individuals; paramere apex evenly rounded; FLP endophallite elongate, with two indentations along right edge.
</paragraph>
</subSubSection>
<subSubSection id="C36F652AEF07FF9CFF72FCC8CB75E1B5" pageId="34" pageNumber="254" type="etymology">
<paragraph id="8BCA36A1EF07FF9CFF72FCC8C87FE106" blockId="34.[189,324,855,881]" box="[189,324,855,881]" pageId="34" pageNumber="254">
<heading id="D08281CDEF07FF9CFF72FCC8C87FE106" bold="true" box="[189,324,855,881]" fontSize="11" level="3" pageId="34" pageNumber="254" reason="3">
<emphasis id="B901EAB3EF07FF9CFF72FCC8C87FE106" bold="true" box="[189,324,855,881]" pageId="34" pageNumber="254">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36A1EF07FF9CFF72FC1ACB75E1B5" blockId="34.[189,1398,901,962]" pageId="34" pageNumber="254">
<emphasis id="B901EAB3EF07FF9CFF72FC1AC81BE1E8" box="[189,288,901,927]" italics="true" pageId="34" pageNumber="254">Williami</emphasis>
is the Latin singular genitive form of William. This species is named in honour of the second author’s brother, William Darling.
</paragraph>
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<subSubSection id="C36F652AEF07FF9CFF72FC75CB41E758" pageId="34" pageNumber="254" type="materials_examined">
<paragraph id="8BCA36A1EF07FF9CFF72FC75C8D7E673" blockId="34.[189,492,1002,1028]" box="[189,492,1002,1028]" pageId="34" pageNumber="254">
<heading id="D08281CDEF07FF9CFF72FC75C8D7E673" bold="true" box="[189,492,1002,1028]" fontSize="11" level="3" pageId="34" pageNumber="254" reason="3">
<emphasis id="B901EAB3EF07FF9CFF72FC75C8D7E673" bold="true" box="[189,492,1002,1028]" pageId="34" pageNumber="254">Name-bearing type data</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36A1EF07FF9CFF72FB87CD27E6EB" blockId="34.[189,1398,1048,1181]" pageId="34" pageNumber="254">
<typeStatus id="54CE8803EF07FF9CFF72FB87C811E645" box="[189,298,1048,1074]" pageId="34" pageNumber="254" type="holotype">Holotype</typeStatus>
<specimenCount id="9D73FD28EF07FF9CFEFCFB86C872E644" box="[307,329,1049,1075]" count="1" pageId="34" pageNumber="254" type="male">♂</specimenCount>
(
<collectionCode id="ED64AE64EF07FF9CFE94FB86C890E644" box="[347,427,1049,1075]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="254" type="Museum">MZSP</collectionCode>
) (
<figureCitation id="134E2A24EF07FF9CFE0AFB87CB24E644" box="[453,543,1048,1075]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling &amp; Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="34" pageNumber="254">Fig. 55</figureCitation>
): “ 
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: 
<collectingRegion id="49B1F843EF07FF9CFD73FB86CA64E644" box="[700,863,1049,1077]" country="Brazil" name="Sao Paulo" pageId="34" pageNumber="254">SÃO PAULO</collectingRegion>
| 
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SE Mogi
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das Cruzes | 
<collectingMunicipality id="6BAEACDBEF07FF9CFB1DFB86CC49E644" box="[1234,1394,1048,1075]" pageId="34" pageNumber="254">Serra do Mar</collectingMunicipality>
, Est. Biol. | 
<location id="8EAA607AEF07FF9CFE8AFBA3C88BE621" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF07FF9CFE8AFBA3C88BE621" box="[325,432,1084,1110]" country="Brazil" municipality="Serra do Mar" name="Boraceia" pageId="34" pageNumber="254" stateProvince="Sao Paulo">Boracéia</location>
, 
<quantity id="4C8D9B44EF07FF9CFE71FBA3CB02E621" box="[446,569,1084,1110]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="34" pageNumber="254" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
<elevation id="0058D192EF07FF9CFE71FBA3CB02E621" box="[446,569,1084,1110]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="34" pageNumber="254" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900m</elevation>
</quantity>
, 28-30. | 
<date id="FFCB1061EF07FF9CFD64FBA3CA30E621" box="[683,779,1084,1110]" pageId="34" pageNumber="254" value="1997-04">
<collectingDate id="EF8FE989EF07FF9CFD64FBA3CA30E621" box="[683,779,1084,1110]" pageId="34" pageNumber="254" value="1997-04">IV.1997</collectingDate>
</date>
, 
<collectorName id="26805377EF07FF9CFCD6FBA3CABCE621" box="[793,903,1084,1110]" pageId="34" pageNumber="254">F. Génier</collectorName>
&amp; 
<collectorName id="26805377EF07FF9CFC7FFBA3CACDE621" box="[944,1014,1084,1110]" pageId="34" pageNumber="254">S. Ide</collectorName>
| 
<emphasis id="B901EAB3EF07FF9CFBDEFBA2CD12E621" box="[1041,1065,1085,1110]" italics="true" pageId="34" pageNumber="254">ex</emphasis>
. 
<collectingMethod id="52344EB6EF07FF9CFBF5FBA3CDFCE621" box="[1082,1223,1084,1110]" pageId="34" pageNumber="254">carrion trap</collectingMethod>
, cloud forest”; “ 
<collectingRegion id="49B1F843EF07FF9CFF06FBC0C80CE60E" box="[201,311,1119,1145]" country="Brazil" name="WORLD" pageId="34" pageNumber="254">WORLD</collectingRegion>
| SCARAB. | 
<collectingRegion id="49B1F843EF07FF9CFE14FBC0CB49E60E" box="[475,626,1119,1145]" country="Brazil" name="DATABASE" pageId="34" pageNumber="254">DATABASE</collectingRegion>
| WSD00035694” [barcode label]; “ 
<typeStatus id="54CE8803EF07FF9CFBD6FBC0CD8CE60E" box="[1049,1207,1119,1145]" pageId="34" pageNumber="254" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D73FD28EF07FF9CFB73FBC0CDE9E60E" box="[1212,1234,1119,1145]" count="1" pageId="34" pageNumber="254" type="male">♂</specimenCount>
| 
<taxonomicName id="4C754D22EF07FF9CFB2AFBFFCC4DE60E" authorityName="Burmeister" authorityYear="1846" box="[1253,1398,1120,1145]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAB3EF07FF9CFB2AFBFFCC4DE60E" box="[1253,1398,1120,1145]" italics="true" pageId="34" pageNumber="254">Homocopris</emphasis>
</taxonomicName>
| 
<taxonomicName id="4C754D22EF07FF9CFF05FB1DC811E6EB" authorityName="Darling &amp; Genier" authorityYear="2024" box="[202,298,1154,1180]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="254" phylum="Arthropoda" rank="species" species="williami">
<emphasis id="B901EAB3EF07FF9CFF05FB1DC811E6EB" box="[202,298,1154,1180]" italics="true" pageId="34" pageNumber="254">williami</emphasis>
</taxonomicName>
| Des. Darling &amp; Génier, 2018” [on red card with black border].
</paragraph>
<paragraph id="8BCA36A1EF07FF9CFF72FB5BC866E6AA" blockId="34.[189,349,1219,1246]" box="[189,349,1219,1246]" pageId="34" pageNumber="254">
<heading id="D08281CDEF07FF9CFF72FB5BC866E6AA" bold="true" box="[189,349,1219,1246]" fontSize="11" level="3" pageId="34" pageNumber="254" reason="3">
<emphasis id="B901EAB3EF07FF9CFF72FB5BC866E6AA" bold="true" box="[189,349,1219,1246]" pageId="34" pageNumber="254">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36A1EF07FF9CFF72FB6CCB41E758" blockId="34.[189,1399,1265,1328]" pageId="34" pageNumber="254">
Estação Biológica de Boracéia, 
<geoCoordinate id="EE415066EF07FF9CFDA3FB6ECA31E77B" box="[620,778,1265,1293]" degrees="23" direction="south" minutes="38" orientation="latitude" pageId="34" pageNumber="254" precision="15" seconds="55" value="-23.648611">23°38′55″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF07FF9CFCEAFB6ECAF7E77A" box="[805,972,1265,1293]" degrees="45" direction="west" minutes="52" orientation="longitude" pageId="34" pageNumber="254" precision="15" seconds="20" value="-45.87222">45°52′20″ W</geoCoordinate>
, 
<quantity id="4C8D9B44EF07FF9CFC29FB6DCD42E77A" box="[998,1145,1266,1293]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="34" pageNumber="254" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800–900 m</quantity>
, Salesópolis, Serra do Mar Mountains, 
<collectingRegion id="49B1F843EF07FF9CFE69FA8ACB24E758" box="[422,543,1301,1327]" country="Brazil" name="Sao Paulo" pageId="34" pageNumber="254">São Paulo</collectingRegion>
, 
<collectingCountry id="F3627631EF07FF9CFDE4FA8ACB4DE758" box="[555,630,1301,1327]" name="Brazil" pageId="34" pageNumber="254">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F652AEF07FF9DFF72FAC8CDB8E607" lastPageId="35" lastPageNumber="255" pageId="34" pageNumber="254" type="materials_examined">
<paragraph id="8BCA36A1EF07FF9CFF72FAC8C857E707" blockId="34.[189,364,1366,1393]" box="[189,364,1366,1393]" pageId="34" pageNumber="254">
<heading id="D08281CDEF07FF9CFF72FAC8C857E707" bold="true" box="[189,364,1366,1393]" fontSize="11" level="3" pageId="34" pageNumber="254" reason="3">
<emphasis id="B901EAB3EF07FF9CFF72FAC8C857E707" bold="true" box="[189,364,1366,1393]" pageId="34" pageNumber="254">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B1D3CFCEF07FF9CFF25FA1ACB40E47E" collectedFrom="cloud forest" collectingDate="1997-04-28" collectingDateMax="1997-04-30" collectingDateMin="1997-04-28" collectingMethod="dung trap" collectionCode="MZSP" collectorName="F. Genier &amp; S. Ide" country="Brazil" county="Salesopolis" elevation="850" latitude="-23.648611" location="Serra do Mar" longLatPrecision="21" longitude="-45.87222" municipality="Estacao Biologica de Boraceia" pageId="34" pageNumber="254" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="holotype">
<paragraph id="8BCA36A1EF07FF9CFF25FA1AC867E7E8" blockId="34.[189,1399,1413,1545]" box="[234,348,1413,1439]" pageId="34" pageNumber="254">
<emphasis id="B901EAB3EF07FF9CFF25FA1AC867E7E8" bold="true" box="[234,348,1413,1439]" pageId="34" pageNumber="254">
<typeStatus id="54CE8803EF07FF9CFF25FA1AC867E7E8" box="[234,348,1413,1439]" pageId="34" pageNumber="254" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCA36A1EF07FF9CFF72FA36CB40E47E" blockId="34.[189,1399,1413,1545]" pageId="34" pageNumber="254">
<collectingCountry id="F3627631EF07FF9CFF72FA36C81FE7B4" box="[189,292,1449,1475]" name="Brazil" pageId="34" pageNumber="254">BRASIL</collectingCountry>
• 
<specimenCount id="9D73FD28EF07FF9CFE8AFA36C860E7B4" box="[325,347,1449,1475]" count="1" pageId="34" pageNumber="254" type="male">♂</specimenCount>
(
<figureCitation id="134E2A24EF07FF9CFEA0FA37C8F1E7B4" box="[367,458,1448,1475]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling &amp; Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="34" pageNumber="254">Fig. 55</figureCitation>
); 
<collectingRegion id="49B1F843EF07FF9CFE2BFA36CB5AE7B5" box="[484,609,1448,1475]" country="Brazil" name="Sao Paulo" pageId="34" pageNumber="254">São Paulo</collectingRegion>
, 
<collectingCounty id="62AB4E2DEF07FF9CFDBFFA37CBC2E7B5" box="[624,761,1448,1474]" pageId="34" pageNumber="254">Salesópolis</collectingCounty>
, 
<collectingMunicipality id="6BAEACDBEF07FF9CFCC7FA36CD46E7B5" box="[776,1149,1448,1475]" pageId="34" pageNumber="254">Estação Biológica de Boracéia</collectingMunicipality>
, 
<locationDeviation id="9A782F83EF07FF9CFB42FA37C879E791" pageId="34" pageNumber="254">
<quantity id="4C8D9B44EF07FF9CFB42FA37CDE1E7B5" box="[1165,1242,1448,1475]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="254" unit="km" value="50.0">50 km</quantity>
SE of Mogi das Cruzes
</locationDeviation>
, 
<location id="8EAA607AEF07FF9CFE9FFA53C8C8E791" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF07FF9CFE9FFA53C8C8E791" box="[336,499,1484,1510]" country="Brazil" county="Salesopolis" latitude="-23.648611" longLatPrecision="21" longitude="-45.87222" municipality="Estacao Biologica de Boraceia" name="Serra do Mar" pageId="34" pageNumber="254" stateProvince="Sao Paulo">Serra do Mar</location>
; [
<geoCoordinate id="EE415066EF07FF9CFDC7FA53CBA7E791" box="[520,668,1482,1510]" degrees="23" direction="south" minutes="38" orientation="latitude" pageId="34" pageNumber="254" precision="15" seconds="55" value="-23.648611">23°38′55″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF07FF9CFD64FA54CA70E791" box="[683,843,1482,1510]" degrees="45" direction="west" minutes="52" orientation="longitude" pageId="34" pageNumber="254" precision="15" seconds="20" value="-45.87222">45°52′20″ W</geoCoordinate>
]; 
<quantity id="4C8D9B44EF07FF9CFCAEFA53CAD0E791" box="[865,1003,1484,1510]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="34" pageNumber="254" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
<elevation id="0058D192EF07FF9CFCAEFA53CAD0E791" box="[865,1003,1484,1510]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="34" pageNumber="254" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800–900 m</elevation>
</quantity>
; 
<date id="FFCB1061EF07FF9CFC35FA53CDFDE791" box="[1018,1222,1484,1510]" pageId="34" pageNumber="254" value="1997-04-28" valueMax="1997-04-30" valueMin="1997-04-28">
<collectingDate id="EF8FE989EF07FF9CFC35FA53CDFDE791" box="[1018,1222,1484,1510]" pageId="34" pageNumber="254" value="1997-04-28" valueMax="1997-04-30" valueMin="1997-04-28">28–30 Apr. 1997</collectingDate>
</date>
; 
<collectorName id="26805377EF07FF9CFB1BFA53CC78E791" box="[1236,1347,1484,1510]" pageId="34" pageNumber="254">F. Génier</collectorName>
and 
<collectorName id="26805377EF07FF9CFF72FA70C83EE47E" box="[189,261,1519,1545]" pageId="34" pageNumber="254">S. Ide</collectorName>
; 
<collectedFrom id="3869755CEF07FF9CFEDEFA70C8A4E47E" box="[273,415,1519,1545]" pageId="34" pageNumber="254">cloud forest</collectedFrom>
, 
<collectingMethod id="52344EB6EF07FF9CFE64FA70CB25E47E" box="[427,542,1519,1545]" pageId="34" pageNumber="254">dung trap</collectingMethod>
; 
<collectionCode id="ED64AE64EF07FF9CFDE5FA70CB4CE47E" box="[554,631,1519,1545]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="254" type="Museum">MZSP</collectionCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="3B1D3CFCEF07FF9CFF25F9AFCA51E419" collectingDate="1997-04-28" collectingDateMax="1997-04-30" collectingDateMin="1997-04-28" collectionCode="MZSP" collectorName=" F. Genier &amp; S. Ide" country="Brazil" county="Salesopolis" elevation="850" latitude="-23.648611" location="BRASIL" longLatPrecision="21" longitude="-45.87222" municipality="Estacao Biologica de Boraceia" pageId="34" pageNumber="254" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="allotype">
<paragraph id="8BCA36A1EF07FF9CFF25F9AFC86FE43D" blockId="34.[189,874,1584,1646]" box="[234,340,1584,1610]" pageId="34" pageNumber="254">
<emphasis id="B901EAB3EF07FF9CFF25F9AFC86FE43D" bold="true" box="[234,340,1584,1610]" pageId="34" pageNumber="254">
<typeStatus id="54CE8803EF07FF9CFF25F9AFC86FE43D" box="[234,340,1584,1610]" pageId="34" pageNumber="254" type="allotype">Allotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCA36A1EF07FF9CFF72F9CBCA51E419" blockId="34.[189,874,1584,1646]" box="[189,874,1619,1646]" pageId="34" pageNumber="254">
<collectingCountry id="F3627631EF07FF9CFF72F9CBC81FE419" box="[189,292,1620,1646]" name="Brazil" pageId="34" pageNumber="254">BRASIL</collectingCountry>
• 
<specimenCount id="9D73FD28EF07FF9CFEF2F9CBC86FE419" box="[317,340,1620,1646]" count="1" pageId="34" pageNumber="254" type="female">♀</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED64AE64EF07FF9CFCD6F9CBCA5DE419" box="[793,870,1620,1646]" collectionName="Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="254" type="Museum">MZSP</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BCA36A1EF07FF9CFF72F90ACBFFE4D8" blockId="34.[189,708,1685,1712]" box="[189,708,1685,1712]" pageId="34" pageNumber="254">
<emphasis id="B901EAB3EF07FF9CFF72F90AC8C2E4D8" bold="true" box="[189,505,1685,1711]" pageId="34" pageNumber="254">Other material examined</emphasis>
(
<specimenCount id="9D73FD28EF07FF9CFDC7F90ACB67E4C7" box="[520,604,1685,1712]" count="55" pageId="34" pageNumber="254" type="male">55 ♂♂</specimenCount>
, 
<specimenCount id="9D73FD28EF07FF9CFDA6F90ACB83E4D8" box="[617,696,1685,1711]" count="32" pageId="34" pageNumber="254" type="female">32 ♀♀</specimenCount>
)
</paragraph>
<materialsCitation id="3B1D3CFCEF07FF9CFF25F95BCB96E553" collectingDate="1884-04" collectionCode="MNHN" collectorName="P. Germain" country="Brazil" county="Nova Friburgo" latitude="-22.383333" location="Haut Rio Macae" longLatPrecision="1260" longitude="-42.466667" municipality="Upper Macae River" pageId="34" pageNumber="254" specimenCount="5" specimenCount-male="5" stateProvince="Rio de Janeiro" typeStatus="paratype">
<paragraph id="8BCA36A1EF07FF9CFF25F95BC85CE4A9" blockId="34.[189,1399,1732,2004]" box="[234,359,1732,1758]" pageId="34" pageNumber="254">
<emphasis id="B901EAB3EF07FF9CFF25F95BC85CE4A9" bold="true" box="[234,359,1732,1758]" pageId="34" pageNumber="254">
<typeStatus id="54CE8803EF07FF9CFF25F95BC85CE4A9" box="[234,359,1732,1758]" pageId="34" pageNumber="254" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCA36A1EF07FF9DFF72F978CDB8E607" blockId="34.[189,1399,1732,2004]" lastBlockId="35.[187,1399,263,1136]" lastPageId="35" lastPageNumber="255" pageId="34" pageNumber="254">
<collectingCountry id="F3627631EF07FF9CFF72F978C81DE576" box="[189,294,1767,1793]" name="Brazil" pageId="34" pageNumber="254">BRAZIL</collectingCountry>
– 
<emphasis id="B901EAB3EF07FF9CFE8EF979C8CDE577" bold="true" box="[321,502,1766,1793]" pageId="34" pageNumber="254">
<collectingRegion id="49B1F843EF07FF9CFE8EF979C8CDE577" box="[321,502,1766,1793]" country="Brazil" name="Rio de Janeiro" pageId="34" pageNumber="254">Rio de Janeiro</collectingRegion>
</emphasis>
• 
<specimenCount id="9D73FD28EF07FF9CFDC1F979CB68E576" box="[526,595,1766,1793]" count="5" pageId="34" pageNumber="254" type="male">5 ♂♂</specimenCount>
; 
<collectingCounty id="62AB4E2DEF07FF9CFD90F978CA35E576" box="[607,782,1767,1793]" pageId="34" pageNumber="254">Nova Friburgo</collectingCounty>
, 
<collectingMunicipality id="6BAEACDBEF07FF9CFCD6F978CAC4E576" box="[793,1023,1767,1793]" country="Brazil" county="Nova Friburgo" latitude="-22.383333" longLatPrecision="1260" longitude="-42.466667" municipality="Upper Macae River" name="Upper Macae River" pageId="34" pageNumber="254" stateProvince="Rio de Janeiro">Upper Macaé River</collectingMunicipality>
(“ 
<location id="8EAA607AEF07FF9CFBD2F978CDE7E576" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF07FF9CFBD2F978CDE7E576" box="[1053,1244,1767,1793]" country="Brazil" county="Nova Friburgo" latitude="-22.383333" longLatPrecision="1260" longitude="-42.466667" municipality="Upper Macae River" name="Haut Rio Macae" pageId="34" pageNumber="254" stateProvince="Rio de Janeiro">Haut Rio Macaé</location>
”); [
<geoCoordinate id="EE415066EF07FF9CFAC6F978CC4AE576" box="[1289,1393,1765,1793]" degrees="22" direction="south" minutes="23" orientation="latitude" pageId="34" pageNumber="254" precision="925" value="-22.383333">22°23′ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF07FF9CFF72F895C80BE553" box="[189,304,1800,1828]" degrees="42" direction="west" minutes="28" orientation="longitude" pageId="34" pageNumber="254" precision="925" value="-42.466667">42°28′ W</geoCoordinate>
]; 
<date id="FFCB1061EF07FF9CFE8EF895C882E553" box="[321,441,1802,1828]" pageId="34" pageNumber="254" value="1884-04">
<collectingDate id="EF8FE989EF07FF9CFE8EF895C882E553" box="[321,441,1802,1828]" pageId="34" pageNumber="254" value="1884-04">Apr. 1884</collectingDate>
</date>
; 
<collectorName id="26805377EF07FF9CFE0CF895CB73E553" box="[451,584,1802,1828]" pageId="34" pageNumber="254">P. Germain</collectorName>
; 
<collectionCode id="ED64AE64EF07FF9CFD9CF895CB96E553" box="[595,685,1802,1828]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="254">MNHN</collectionCode>
• 
<materialsCitation id="3B1D3CFCEF07FF9CFD0DF895CC48E553" box="[706,1395,1802,1828]" collectionCode="MNHN" country="Brazil" pageId="34" pageNumber="254" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<specimenCount id="9D73FD28EF07FF9CFD0DF895CBCBE553" box="[706,752,1802,1828]" count="1" pageId="34" pageNumber="254" type="male">1 ♂</specimenCount>
; unspecified locality]: [no date]; [anonymous]; 
<collectionCode id="ED64AE64EF07FF9CFADAF895CC48E553" box="[1301,1395,1802,1828]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="254">MNHN</collectionCode>
</materialsCitation>
. – 
<materialsCitation id="3B1D3CFCEF07FF9CFF1CF8B2CC10E530" box="[211,1323,1836,1864]" collectingDate="2001-09-30" collectionCode="CEMT" collectorName="Arioli" country="Brazil" latitude="-28.783333" location="Nova Prata" longLatPrecision="1230" longitude="-51.6" pageId="34" pageNumber="254" specimenCount="2" specimenCount-female="2" stateProvince="Rio Grande do Sul" typeStatus="paratype">
<emphasis id="B901EAB3EF07FF9CFF1CF8B2C887E530" bold="true" box="[211,444,1837,1863]" pageId="34" pageNumber="254">
<collectingRegion id="49B1F843EF07FF9CFF1CF8B2C887E530" box="[211,444,1837,1863]" country="Brazil" name="Rio Grande do Sul" pageId="34" pageNumber="254">Rio Grande do Sul</collectingRegion>
</emphasis>
• 
<specimenCount id="9D73FD28EF07FF9CFE1BF8B2CB2CE53F" box="[468,535,1837,1864]" count="2" pageId="34" pageNumber="254" type="female">2 ♀♀</specimenCount>
; 
<location id="8EAA607AEF07FF9CFDEAF8B1CB97E53F" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF07FF9CFDEAF8B1CB97E53F" box="[549,684,1838,1864]" country="Brazil" latitude="-28.783333" longLatPrecision="1230" longitude="-51.6" name="Nova Prata" pageId="34" pageNumber="254" stateProvince="Rio Grande do Sul">Nova Prata</location>
; [
<geoCoordinate id="EE415066EF07FF9CFD70F8B2CA1CE530" box="[703,807,1836,1864]" degrees="28" direction="south" minutes="47" orientation="latitude" pageId="34" pageNumber="254" precision="925" value="-28.783333">28°47′ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF07FF9CFCFBF8B2CA93E53F" box="[820,936,1836,1864]" degrees="51" direction="west" minutes="36" orientation="longitude" pageId="34" pageNumber="254" precision="925" value="-51.6">51°36′ W</geoCoordinate>
]; 
<date id="FFCB1061EF07FF9CFC73F8B2CD63E530" box="[956,1112,1837,1863]" pageId="34" pageNumber="254" value="2001-09-30">
<collectingDate id="EF8FE989EF07FF9CFC73F8B2CD63E530" box="[956,1112,1837,1863]" pageId="34" pageNumber="254" value="2001-09-30">30 Sep. 2001</collectingDate>
</date>
; C. 
<collectorName id="26805377EF07FF9CFB4AF8B2CDF4E530" box="[1157,1231,1837,1863]" pageId="34" pageNumber="254">Arioli</collectorName>
; 
<collectionCode id="ED64AE64EF07FF9CFB15F8B2CC10E530" box="[1242,1323,1837,1863]" pageId="34" pageNumber="254">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF07FF9CFA8CF8B2C82BE5F9" collectingDate="2008-01-14" collectionCode="CEMT" collectorName="Audino" country="Brazil" latitude="-29.446943" location="Sao Francisco de Paula" longLatPrecision="20" longitude="-50.579166" pageId="34" pageNumber="254" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Rio Grande do Sul" typeStatus="paratype">
<specimenCount id="9D73FD28EF07FF9CFA8CF8B2CC4AE53F" box="[1347,1393,1837,1864]" count="1" pageId="34" pageNumber="254" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D73FD28EF07FF9CFF72F8CFC838E51C" box="[189,259,1872,1899]" count="4" pageId="34" pageNumber="254" type="female">4 ♀♀</specimenCount>
; 
<location id="8EAA607AEF07FF9CFEDBF8CFCB0BE51D" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF07FF9CFEDBF8CFCB0BE51D" box="[276,560,1872,1898]" country="Brazil" latitude="-29.446943" longLatPrecision="20" longitude="-50.579166" name="Sao Francisco de Paula" pageId="34" pageNumber="254" stateProvince="Rio Grande do Sul">São Francisco de Paula</location>
; [
<geoCoordinate id="EE415066EF07FF9CFD89F8CFCBE0E51D" box="[582,731,1871,1899]" degrees="29" direction="south" minutes="26" orientation="latitude" pageId="34" pageNumber="254" precision="15" seconds="49" value="-29.446943">29°26′49″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF07FF9CFD25F8CFCAB0E51C" box="[746,907,1871,1899]" degrees="50" direction="west" minutes="34" orientation="longitude" pageId="34" pageNumber="254" precision="15" seconds="45" value="-50.579166">50°34′45″ W</geoCoordinate>
]; 
<date id="FFCB1061EF07FF9CFC6DF8CFCD04E51D" box="[930,1087,1872,1899]" pageId="34" pageNumber="254" value="2008-01-14">
<collectingDate id="EF8FE989EF07FF9CFC6DF8CFCD04E51D" box="[930,1087,1872,1899]" pageId="34" pageNumber="254" value="2008-01-14">14 Jan. 2008</collectingDate>
</date>
; L. 
<collectorName id="26805377EF07FF9CFBBFF8CFCDF7E51D" box="[1136,1228,1872,1898]" pageId="34" pageNumber="254">Audino</collectorName>
; human dung; 
<collectionCode id="ED64AE64EF07FF9CFF72F8EBC82BE5F9" box="[189,272,1908,1934]" pageId="34" pageNumber="254">CEMT</collectionCode>
</materialsCitation>
. – 
<materialsCitation id="3B1D3CFCEF07FF9CFEFEF8ECC8BEE5C6" collectingDate="2001-12" collectionCode="CEMT" collectorName="Arioli" country="Brazil" latitude="-28.341667" location="Bom Jardim da Serra" longLatPrecision="20" longitude="-49.625" pageId="34" pageNumber="254" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<emphasis id="B901EAB3EF07FF9CFEFEF8ECC8CBE5FA" bold="true" box="[305,496,1907,1933]" pageId="34" pageNumber="254">
<collectingRegion id="49B1F843EF07FF9CFEFEF8ECC8CBE5FA" box="[305,496,1907,1933]" country="Brazil" name="Santa Catarina" pageId="34" pageNumber="254">Santa Catarina</collectingRegion>
</emphasis>
• 
<specimenCount id="9D73FD28EF07FF9CFDC6F8EBCB0CE5F9" box="[521,567,1908,1934]" count="1" pageId="34" pageNumber="254" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D73FD28EF07FF9CFD8BF8EBCBBCE5F9" box="[580,647,1908,1934]" count="2" pageId="34" pageNumber="254" type="female">2 ♀♀</specimenCount>
; 
<location id="8EAA607AEF07FF9CFD5AF8EBCAA9E5F9" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF07FF9CFD5AF8EBCAA9E5F9" box="[661,914,1907,1934]" country="Brazil" latitude="-28.341667" longLatPrecision="20" longitude="-49.625" name="Bom Jardim da Serra" pageId="34" pageNumber="254" stateProvince="Santa Catarina">Bom Jardim da Serra</location>
; [
<geoCoordinate id="EE415066EF07FF9CFC6AF8EBCD0CE5F9" box="[933,1079,1906,1934]" degrees="28" direction="south" minutes="20" orientation="latitude" pageId="34" pageNumber="254" precision="15" seconds="30" value="-28.341667">28°20′30″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF07FF9CFB8BF8EBCDD9E5F9" box="[1092,1250,1906,1934]" degrees="49" direction="west" minutes="37" orientation="longitude" pageId="34" pageNumber="254" precision="15" seconds="30" value="-49.625">49°37′30″ W</geoCoordinate>
]; 
<date id="FFCB1061EF07FF9CFB39F8EBCC49E5F9" box="[1270,1394,1908,1934]" pageId="34" pageNumber="254" value="2001-12">
<collectingDate id="EF8FE989EF07FF9CFB39F8EBCC49E5F9" box="[1270,1394,1908,1934]" pageId="34" pageNumber="254" value="2001-12">Dec. 2001</collectingDate>
</date>
; C. 
<collectorName id="26805377EF07FF9CFF11F808C813E5C6" box="[222,296,1943,1969]" pageId="34" pageNumber="254">Arioli</collectorName>
; 
<collectionCode id="ED64AE64EF07FF9CFEFBF808C8BEE5C6" box="[308,389,1943,1969]" pageId="34" pageNumber="254">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF07FF9CFE51F808CD92E5C6" box="[414,1193,1943,1969]" collectingDate="2002-01" collectionCode="CEMT" collectorName="Arioli" country="Brazil" latitude="-28.341667" location="Bom Jardim da Serra" longLatPrecision="20" longitude="-49.625" pageId="34" pageNumber="254" specimenCount="3" specimenCount-male="3" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF07FF9CFE51F808C8D8E5C6" box="[414,483,1943,1969]" count="3" pageId="34" pageNumber="254" type="male">3 ♂♂</specimenCount>
; same locality as for preceding; 
<date id="FFCB1061EF07FF9CFCAEF808CAEEE5C6" box="[865,981,1943,1969]" pageId="34" pageNumber="254" value="2002-01">
<collectingDate id="EF8FE989EF07FF9CFCAEF808CAEEE5C6" box="[865,981,1943,1969]" pageId="34" pageNumber="254" value="2002-01">Jan. 2002</collectingDate>
</date>
; C. 
<collectorName id="26805377EF07FF9CFBCCF808CD76E5C6" box="[1027,1101,1943,1969]" pageId="34" pageNumber="254">Arioli</collectorName>
; 
<collectionCode id="ED64AE64EF07FF9CFB97F808CD92E5C6" box="[1112,1193,1943,1969]" pageId="34" pageNumber="254">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF07FF9CFB0CF808CAB9E5A3" collectingDate="2001-12-28" collectionCode="CEMT" collectorName="Arioli" country="Brazil" latitude="-28.341667" location="Bom Jardim da Serra" longLatPrecision="20" longitude="-49.625" pageId="34" pageNumber="254" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF07FF9CFB0CF808CDCAE5C6" box="[1219,1265,1943,1969]" count="1" pageId="34" pageNumber="254" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D73FD28EF07FF9CFB31F808CC10E5C6" box="[1278,1323,1943,1969]" count="1" pageId="34" pageNumber="254" type="female">1 ♀</specimenCount>
; same locality as for preceding; 
<date id="FFCB1061EF07FF9CFE35F825CB99E5A3" box="[506,674,1978,2004]" pageId="34" pageNumber="254" value="2001-12-28">
<collectingDate id="EF8FE989EF07FF9CFE35F825CB99E5A3" box="[506,674,1978,2004]" pageId="34" pageNumber="254" value="2001-12-28">28 Dec. 2001</collectingDate>
</date>
; C. 
<collectorName id="26805377EF07FF9CFD17F825CA19E5A3" box="[728,802,1978,2004]" pageId="34" pageNumber="254">Arioli</collectorName>
; 
<collectionCode id="ED64AE64EF07FF9CFCFEF825CAB9E5A3" box="[817,898,1978,2004]" pageId="34" pageNumber="254">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF07FF9DFC6BF825CB18E354" collectingDate="2002-01-03" collectionCode="CEMT" collectorName="Arioli" country="Brazil" lastPageId="35" lastPageNumber="255" latitude="-28.341667" location="Bom Jardim da Serra" longLatPrecision="20" longitude="-49.625" pageId="34" pageNumber="254" specimenCount="4" specimenCount-female="4" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF07FF9CFC6BF825CAD0E5A3" box="[932,1003,1978,2004]" count="4" pageId="34" pageNumber="254" type="female">4 ♀♀</specimenCount>
; same locality as for preceding; 
<date id="FFCB1061EF06FF9DFF72FE96C871E354" box="[189,330,265,291]" pageId="35" pageNumber="255" value="2002-01-03">
<collectingDate id="EF8FE989EF06FF9DFF72FE96C871E354" box="[189,330,265,291]" pageId="35" pageNumber="255" value="2002-01-03">3 Jan. 2002</collectingDate>
</date>
; C. 
<collectorName id="26805377EF06FF9DFEB4FE96C8FEE354" box="[379,453,265,291]" pageId="35" pageNumber="255">Arioli</collectorName>
; 
<collectionCode id="ED64AE64EF06FF9DFE1DFE96CB18E354" box="[466,547,265,291]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFD8FFE96C8A1E331" collectingDate="2011-02" collectionCode="CEMT" collectorName="Campos" country="Brazil" latitude="-27.383333" location="Campos Novo" longLatPrecision="1237" longitude="-51.2" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFD8FFE96CB4AE354" box="[576,625,265,291]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; 
<location id="8EAA607AEF06FF9DFDB0FE96CA16E354" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFDB0FE96CA16E354" box="[639,813,265,291]" country="Brazil" latitude="-27.383333" longLatPrecision="1237" longitude="-51.2" name="Campos Novo" pageId="35" pageNumber="255" stateProvince="Santa Catarina">Campos Novo</location>
; [
<geoCoordinate id="EE415066EF06FF9DFC8DFE96CA96E354" box="[834,941,263,291]" degrees="27" direction="south" minutes="23" orientation="latitude" pageId="35" pageNumber="255" precision="925" value="-27.383333">27°23′ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFC74FE96CD09E354" box="[955,1074,263,292]" degrees="51" direction="west" minutes="12" orientation="longitude" pageId="35" pageNumber="255" precision="925" value="-51.2">51°12′ W</geoCoordinate>
]; 
<date id="FFCB1061EF06FF9DFB87FE96CDFAE354" box="[1096,1217,265,291]" pageId="35" pageNumber="255" value="2011-02">
<collectingDate id="EF8FE989EF06FF9DFB87FE96CDFAE354" box="[1096,1217,265,291]" pageId="35" pageNumber="255" value="2011-02">Feb. 2011</collectingDate>
</date>
; R.C. 
<collectorName id="26805377EF06FF9DFAC1FE96CC49E354" box="[1294,1394,265,291]" pageId="35" pageNumber="255">Campos</collectorName>
; 
<collectingMethod id="52344EB6EF06FF9DFF72FEB3C807E330" box="[189,316,300,327]" pageId="35" pageNumber="255">pitfall trap</collectingMethod>
; 
<collectionCode id="ED64AE64EF06FF9DFE86FEB3C8A1E331" box="[329,410,300,326]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFE7BFEB3CC4DE331" box="[436,1398,300,327]" collectingDate="2013-02" collectionCode="CEMT" collectorName="Campos" country="Brazil" latitude="-27.383333" location="Campos Novo" longLatPrecision="1237" longitude="-51.2" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFE7BFEB3C8DFE330" box="[436,484,300,327]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; same locality as for preceding; 
<date id="FFCB1061EF06FF9DFCAAFEB3CAE5E331" box="[869,990,300,326]" pageId="35" pageNumber="255" value="2013-02">
<collectingDate id="EF8FE989EF06FF9DFCAAFEB3CAE5E331" box="[869,990,300,326]" pageId="35" pageNumber="255" value="2013-02">Feb. 2013</collectingDate>
</date>
; R.C. 
<collectorName id="26805377EF06FF9DFBE6FEB3CDB6E331" box="[1065,1165,300,326]" pageId="35" pageNumber="255">Campos</collectorName>
; 
<collectingMethod id="52344EB6EF06FF9DFB56FEB3CC22E330" box="[1177,1305,300,327]" pageId="35" pageNumber="255">pitfall trap</collectingMethod>
; 
<collectionCode id="ED64AE64EF06FF9DFAEAFEB3CC4DE331" box="[1317,1398,300,326]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFF00FED0C875E3FA" collectingDate="2012-08-06" collectionCode="CEMT" collectorName="Brandl" country="Brazil" elevation="881" latitude="-26.716667" location="Monte Castelo" longLatPrecision="1240" longitude="-50.316666" pageId="35" pageNumber="255" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFF00FED0C9C6E31D" box="[207,253,335,362]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D73FD28EF06FF9DFEC6FED0C80DE31D" box="[265,310,335,362]" count="1" pageId="35" pageNumber="255" type="female">1 ♀</specimenCount>
; 
<location id="8EAA607AEF06FF9DFE8AFECFC8CEE31E" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFE8AFECFC8CEE31E" box="[325,501,335,362]" country="Brazil" latitude="-26.716667" longLatPrecision="1240" longitude="-50.316666" name="Monte Castelo" pageId="35" pageNumber="255" stateProvince="Santa Catarina">Monte Castelo</location>
; [
<geoCoordinate id="EE415066EF06FF9DFDC8FED0CB4BE31E" box="[519,624,334,362]" degrees="26" direction="south" minutes="43" orientation="latitude" pageId="35" pageNumber="255" precision="925" value="-26.716667">26°43′ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFDB2FED0CBC9E31D" box="[637,754,334,362]" degrees="50" direction="west" minutes="19" orientation="longitude" pageId="35" pageNumber="255" precision="925" value="-50.316666">50°19′ W</geoCoordinate>
]; 
<quantity id="4C8D9B44EF06FF9DFCC9FED0CA68E31D" box="[774,851,335,362]" metricMagnitude="2" metricUnit="m" metricValue="8.809999999999999" pageId="35" pageNumber="255" unit="m" value="881.0">
<elevation id="0058D192EF06FF9DFCC9FED0CA68E31D" box="[774,851,335,362]" metricMagnitude="2" metricUnit="m" metricValue="8.809999999999999" pageId="35" pageNumber="255" unit="m" value="881.0">881 m</elevation>
</quantity>
; 
<date id="FFCB1061EF06FF9DFC90FED0CAC8E31E" box="[863,1011,335,362]" pageId="35" pageNumber="255" value="2012-08-06">
<collectingDate id="EF8FE989EF06FF9DFC90FED0CAC8E31E" box="[863,1011,335,362]" pageId="35" pageNumber="255" value="2012-08-06">6 Aug. 2012</collectingDate>
</date>
; A.L. 
<collectorName id="26805377EF06FF9DFBF5FED0CDB5E31E" box="[1082,1166,335,361]" pageId="35" pageNumber="255">Brandl</collectorName>
; mata nativa, 
<collectingMethod id="52344EB6EF06FF9DFAFFFED0C9D5E3FA" pageId="35" pageNumber="255">pitfall trap</collectingMethod>
; 
<collectionCode id="ED64AE64EF06FF9DFF32FEECC875E3FA" box="[253,334,371,397]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFEABFEECCC19E3FA" box="[356,1314,369,397]" collectionCode="CAS" collectorName="F. Plaumann" country="Brazil" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Ceara" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFEABFEECC8A8E3FA" box="[356,403,371,397]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; 
<collectingRegion id="49B1F843EF06FF9DFE6DFEECC8DDE3FA" box="[418,486,371,397]" country="Brazil" name="Ceara" pageId="35" pageNumber="255">Seara</collectingRegion>
, 
<location id="8EAA607AEF06FF9DFE3AFEECCB93E3FB" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFE3AFEECCB93E3FB" box="[501,680,370,397]" country="Brazil" latitude="-27.183332" longLatPrecision="1238" longitude="-52.383335" name="Nova Teutonia" pageId="35" pageNumber="255" stateProvince="Ceara">Nova Teutônia</location>
; [
<geoCoordinate id="EE415066EF06FF9DFD72FEECCA1CE3FA" box="[701,807,369,397]" degrees="27" direction="south" minutes="11" orientation="latitude" pageId="35" pageNumber="255" precision="925" value="-27.183332">27°11′ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFCF8FEEDCA95E3FA" box="[823,942,369,397]" degrees="52" direction="west" minutes="23" orientation="longitude" pageId="35" pageNumber="255" precision="925" value="-52.383335">52°23′ W</geoCoordinate>
]; [no date]; 
<collectorName id="26805377EF06FF9DFB8DFEECCDE1E3FB" box="[1090,1242,370,397]" pageId="35" pageNumber="255">F. Plaumann</collectorName>
; 
<collectionCode id="ED64AE64EF06FF9DFB26FEECCC19E3FA" box="[1257,1314,371,397]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="35" pageNumber="255" type="Museum">CAS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFA8FFEECCA68E3C7" collectionCode="CAS" collectorName="Jun. &amp; F. Plaumann" country="Brazil" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFA8FFEECCC4AE3FA" box="[1344,1393,371,397]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; same locality as for preceding; 
<collectorName id="26805377EF06FF9DFDFBFE09CB53E3C7" box="[564,616,406,432]" pageId="35" pageNumber="255">Jun.</collectorName>
; 
<collectorName id="26805377EF06FF9DFDBAFE09CA36E3C7" box="[629,781,406,432]" pageId="35" pageNumber="255">F. Plaumann</collectorName>
; 
<collectionCode id="ED64AE64EF06FF9DFCD5FE09CA68E3C7" box="[794,851,406,432]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="35" pageNumber="255" type="Museum">CAS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFCA0FE09C8AEE3A4" collectionCode="CAS" collectorName="Sep. &amp; F. Plaumann" country="Brazil" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" pageId="35" pageNumber="255" specimenCount="2" specimenCount-male="2" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFCA0FE09CA8DE3C7" box="[879,950,406,432]" count="2" pageId="35" pageNumber="255" type="male">2 ♂♂</specimenCount>
; same locality as for preceding; 
<collectorName id="26805377EF06FF9DFAF4FE09CC49E3C7" box="[1339,1394,406,432]" pageId="35" pageNumber="255">Sep.</collectorName>
; 
<collectorName id="26805377EF06FF9DFF72FE26C869E3A4" box="[189,338,441,467]" pageId="35" pageNumber="255">F. Plaumann</collectorName>
; 
<collectionCode id="ED64AE64EF06FF9DFE93FE26C8AEE3A4" box="[348,405,441,467]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="35" pageNumber="255" type="Museum">CAS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFE62FE26CD4BE3A4" box="[429,1136,441,467]" collectionCode="CAS" collectorName="Nov. &amp; F. Plaumann" country="Brazil" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFE62FE26C8E0E3A4" box="[429,475,441,467]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; same locality as for preceding; 
<collectorName id="26805377EF06FF9DFC9EFE26CAB7E3A4" box="[849,908,441,467]" pageId="35" pageNumber="255">Nov.</collectorName>
; 
<collectorName id="26805377EF06FF9DFC58FE26CD17E3A4" box="[919,1068,441,467]" pageId="35" pageNumber="255">F. Plaumann</collectorName>
; 
<collectionCode id="ED64AE64EF06FF9DFBF8FE26CD4BE3A4" box="[1079,1136,441,467]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="35" pageNumber="255" type="Museum">CAS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFB48FE26C8CDE381" collectionCode="GHCM" collectorName="Nov. &amp; F. Plaumann" country="Brazil" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFB48FE26CD8EE3A4" box="[1159,1205,441,467]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; same collection data for preceding; 
<collectionCode id="ED64AE64EF06FF9DFE51FE43C8CDE381" box="[414,502,476,502]" pageId="35" pageNumber="255">GHCM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFDC2FE43C835E06E" collectingDate="2012-11-25" collectionCode="CEMT" collectorName="J. A. Bogoni" country="Brazil" latitude="-27.983334" location="Urubici" longLatPrecision="1234" longitude="-49.583332" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFDC2FE43CB00E380" box="[525,571,476,503]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; 
<location id="8EAA607AEF06FF9DFD89FE43CB98E381" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFD89FE43CB98E381" box="[582,675,476,502]" country="Brazil" latitude="-27.983334" longLatPrecision="1234" longitude="-49.583332" name="Urubici" pageId="35" pageNumber="255" stateProvince="Santa Catarina">Urubici</location>
; [
<geoCoordinate id="EE415066EF06FF9DFD79FE43CA26E381" box="[694,797,475,503]" degrees="27" direction="south" minutes="59" orientation="latitude" pageId="35" pageNumber="255" precision="925" value="-27.983334">27°59′ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFCE6FE43CAA6E380" box="[809,925,475,503]" degrees="49" direction="west" minutes="35" orientation="longitude" pageId="35" pageNumber="255" precision="925" value="-49.583332">49°35′ W</geoCoordinate>
]; 
<date id="FFCB1061EF06FF9DFC60FE43CD74E381" box="[943,1103,476,503]" pageId="35" pageNumber="255" value="2012-11-25">
<collectingDate id="EF8FE989EF06FF9DFC60FE43CD74E381" box="[943,1103,476,503]" pageId="35" pageNumber="255" value="2012-11-25">25 Nov. 2012</collectingDate>
</date>
; 
<collectorName id="26805377EF06FF9DFB95FE43CDD1E381" box="[1114,1258,476,502]" pageId="35" pageNumber="255">J.A. Bogoni</collectorName>
; 
<collectingMethod id="52344EB6EF06FF9DFB3BFE43CC49E380" box="[1268,1394,476,503]" pageId="35" pageNumber="255">pitfall trap</collectingMethod>
; 
<collectionCode id="ED64AE64EF06FF9DFF72FE60C835E06E" box="[189,270,511,537]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFEEBFE60C889E04A" collectingDate="2015-12-17" collectionCode="CEMT" collectorName="Silva" country="Brazil" elevation="1300" latitude="-28.139166" location="Urubici" longLatPrecision="20" longitude="-49.630833" pageId="35" pageNumber="255" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFEEBFE60C86AE06D" box="[292,337,511,538]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D73FD28EF06FF9DFE92FE60C8B3E06D" box="[349,392,511,538]" count="1" pageId="35" pageNumber="255" type="female">1 ♀</specimenCount>
; 
<location id="8EAA607AEF06FF9DFE59FE60C8C8E06E" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFE59FE60C8C8E06E" box="[406,499,511,537]" country="Brazil" latitude="-28.139166" longLatPrecision="20" longitude="-49.630833" name="Urubici" pageId="35" pageNumber="255" stateProvince="Santa Catarina">Urubici</location>
; [
<geoCoordinate id="EE415066EF06FF9DFDCAFE60CBBCE06E" box="[517,647,510,538]" degrees="28" direction="south" minutes="8" orientation="latitude" pageId="35" pageNumber="255" precision="15" seconds="21" value="-28.139166">28°8′21″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFD5CFE60CA0BE06D" box="[659,816,510,538]" degrees="49" direction="west" minutes="37" orientation="longitude" pageId="35" pageNumber="255" precision="15" seconds="51" value="-49.630833">49°37′51″ W</geoCoordinate>
]; 
<quantity id="4C8D9B44EF06FF9DFC8DFE60CAA6E06D" box="[834,925,511,538]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="35" pageNumber="255" unit="m" value="1300.0">
<elevation id="0058D192EF06FF9DFC8DFE60CAA6E06D" box="[834,925,511,538]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="35" pageNumber="255" unit="m" value="1300.0">1300 m</elevation>
</quantity>
; 
<date id="FFCB1061EF06FF9DFC67FE60CD7DE06D" box="[936,1094,511,538]" pageId="35" pageNumber="255" value="2015-12-17">
<collectingDate id="EF8FE989EF06FF9DFC67FE60CD7DE06D" box="[936,1094,511,538]" pageId="35" pageNumber="255" value="2015-12-17">17 Dec. 2015</collectingDate>
</date>
; P.G. da 
<collectorName id="26805377EF06FF9DFB65FE60CDD1E06E" box="[1194,1258,511,537]" pageId="35" pageNumber="255">Silva</collectorName>
; 
<collectingMethod id="52344EB6EF06FF9DFB3AFE60CC48E06D" box="[1269,1395,511,538]" pageId="35" pageNumber="255">pitfall trap</collectingMethod>
: human feces; 
<collectionCode id="ED64AE64EF06FF9DFEAEFDBCC889E04A" box="[353,434,547,573]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFE03FDBCCB7EE017" collectingDate="2015-11-20" collectionCode="CEMT" collectorName="Silva" country="Brazil" elevation="1600" latitude="-28.158888" location="Urubici" longLatPrecision="20" longitude="-49.630276" pageId="35" pageNumber="255" specimenCount="1" specimenCount-female="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFE03FDBCC8C2E04A" box="[460,505,547,573]" count="1" pageId="35" pageNumber="255" type="female">1 ♀</specimenCount>
; 
<location id="8EAA607AEF06FF9DFDC7FDBDCB5EE04B" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFDC7FDBDCB5EE04B" box="[520,613,546,572]" country="Brazil" latitude="-28.158888" longLatPrecision="20" longitude="-49.630276" name="Urubici" pageId="35" pageNumber="255" stateProvince="Santa Catarina">Urubici</location>
; [
<geoCoordinate id="EE415066EF06FF9DFDB6FDBCCBC6E04A" box="[633,765,545,573]" degrees="28" direction="south" minutes="9" orientation="latitude" pageId="35" pageNumber="255" precision="15" seconds="32" value="-28.158888">28°9′32″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFCC5FDBCCA92E04A" box="[778,937,545,573]" degrees="49" direction="west" minutes="37" orientation="longitude" pageId="35" pageNumber="255" precision="15" seconds="49" value="-49.630276">49°37′49″ W</geoCoordinate>
]; 
<quantity id="4C8D9B44EF06FF9DFC72FDBDCD21E04A" box="[957,1050,546,573]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="35" pageNumber="255" unit="m" value="1600.0">
<elevation id="0058D192EF06FF9DFC72FDBDCD21E04A" box="[957,1050,546,573]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="35" pageNumber="255" unit="m" value="1600.0">1600 m</elevation>
</quantity>
; 
<date id="FFCB1061EF06FF9DFBE9FDBCCDF2E04A" box="[1062,1225,546,573]" pageId="35" pageNumber="255" value="2015-11-20">
<collectingDate id="EF8FE989EF06FF9DFBE9FDBCCDF2E04A" box="[1062,1225,546,573]" pageId="35" pageNumber="255" value="2015-11-20">20 Nov. 2015</collectingDate>
</date>
; P.G. da 
<collectorName id="26805377EF06FF9DFAFDFDBDCC49E04B" box="[1330,1394,546,572]" pageId="35" pageNumber="255">Silva</collectorName>
; 
<collectingMethod id="52344EB6EF06FF9DFF72FDD9C804E017" box="[189,319,582,608]" pageId="35" pageNumber="255">pitfall trap</collectingMethod>
: human feces; 
<collectionCode id="ED64AE64EF06FF9DFE3BFDD9CB7EE017" box="[500,581,582,608]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFDACFDD9CB7AE0F4" collectingDate="2015-12-17" collectionCode="CEMT" collectorName="Silva" country="Brazil" elevation="1600" latitude="-28.158888" location="Urubici" longLatPrecision="20" longitude="-49.630276" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFDACFDD9CBAFE017" box="[611,660,582,608]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; same locality as for preceding; 
<date id="FFCB1061EF06FF9DFBD1FDD9CDF8E016" box="[1054,1219,582,609]" pageId="35" pageNumber="255" value="2015-12-17">
<collectingDate id="EF8FE989EF06FF9DFBD1FDD9CDF8E016" box="[1054,1219,582,609]" pageId="35" pageNumber="255" value="2015-12-17">17 Dec. 2015</collectingDate>
</date>
; P.G. da 
<collectorName id="26805377EF06FF9DFAFDFDD9CC49E017" box="[1330,1394,582,608]" pageId="35" pageNumber="255">Silva</collectorName>
; 
<collectingMethod id="52344EB6EF06FF9DFF72FDF6C805E0F4" box="[189,318,617,643]" pageId="35" pageNumber="255">pitfall trap</collectingMethod>
: human feces; 
<collectionCode id="ED64AE64EF06FF9DFE3FFDF6CB7AE0F4" box="[496,577,617,643]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFD92FDF6CB00E0D1" collectingDate="2016-04-23" collectionCode="CEMT" collectorName="Silva" country="Brazil" elevation="1600" latitude="-28.158888" location="Urubici" longLatPrecision="20" longitude="-49.630276" pageId="35" pageNumber="255" specimenCount="2" specimenCount-male="2" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFD92FDF6CB98E0F4" box="[605,675,617,643]" count="2" pageId="35" pageNumber="255" type="male">2 ♂♂</specimenCount>
; same locality as for preceding; 
<date id="FFCB1061EF06FF9DFBE7FDF6CDFDE0F4" box="[1064,1222,617,643]" pageId="35" pageNumber="255" value="2016-04-23">
<collectingDate id="EF8FE989EF06FF9DFBE7FDF6CDFDE0F4" box="[1064,1222,617,643]" pageId="35" pageNumber="255" value="2016-04-23">23 Apr. 2016</collectingDate>
</date>
; P.G. da 
<collectorName id="26805377EF06FF9DFAFDFDF6CC49E0F4" box="[1330,1394,617,643]" pageId="35" pageNumber="255">Silva</collectorName>
; 
<collectingMethod id="52344EB6EF06FF9DFF72FD13C807E0D0" box="[189,316,652,679]" pageId="35" pageNumber="255">pitfall trap</collectingMethod>
: human feces; 
<collectionCode id="ED64AE64EF06FF9DFE27FD13CB00E0D1" box="[488,571,652,678]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
. – 
<materialsCitation id="3B1D3CFCEF06FF9DFD95FD13CC5DE0D1" box="[602,1382,652,679]" collectingDate="1997-04-28" collectingDateMax="1997-04-30" collectingDateMin="1997-04-28" collectionCode="CEMT" collectorName=" F. Genier &amp; S. Ide" country="Brazil" county="Salesopolis" elevation="850" latitude="-23.648611" location="Serra do Mar" longLatPrecision="21" longitude="-45.87222" municipality="Estacao Biologica de Boraceia" pageId="35" pageNumber="255" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<emphasis id="B901EAB3EF06FF9DFD95FD13CBEDE0D1" bold="true" box="[602,726,652,678]" pageId="35" pageNumber="255">
<collectingRegion id="49B1F843EF06FF9DFD95FD13CBEDE0D1" box="[602,726,652,678]" country="Brazil" name="Sao Paulo" pageId="35" pageNumber="255">São Paulo</collectingRegion>
</emphasis>
• 
<specimenCount id="9D73FD28EF06FF9DFD21FD13CA27E0D0" box="[750,796,652,679]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D73FD28EF06FF9DFCE7FD13CA6FE0D0" box="[808,852,652,679]" count="1" pageId="35" pageNumber="255" type="female">1 ♀</specimenCount>
; same collection data as for 
<typeStatus id="54CE8803EF06FF9DFB50FD13CC32E0D1" box="[1183,1289,652,678]" pageId="35" pageNumber="255">holotype</typeStatus>
; 
<collectionCode id="ED64AE64EF06FF9DFADAFD13CC5DE0D1" box="[1301,1382,652,678]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFF72FD30C868E09A" collectingDate="1997-04-28" collectingDateMax="1997-04-30" collectingDateMin="1997-04-28" collectionCode="CMNC" collectorName="Genier &amp; Ide" country="Brazil" county="Salesopolis" elevation="850" latitude="-23.648611" location="Serra do Mar" longLatPrecision="21" longitude="-45.87222" municipality="Estacao Biologica de Boraceia" pageId="35" pageNumber="255" specimenCount="10" specimenCount-female="2" specimenCount-male="8" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFF72FD30C83AE0BD" box="[189,257,687,714]" count="8" pageId="35" pageNumber="255" type="male">8 ♂♂</specimenCount>
, 
<specimenCount id="9D73FD28EF06FF9DFEC2FD30C86BE0BD" box="[269,336,687,714]" count="2" pageId="35" pageNumber="255" type="female">2 ♀♀</specimenCount>
; same locality as for 
<typeStatus id="54CE8803EF06FF9DFD86FD30CB88E0BE" box="[585,691,687,713]" pageId="35" pageNumber="255">holotype</typeStatus>
; 
<date id="FFCB1061EF06FF9DFD70FD30CABDE0BE" box="[703,902,687,714]" pageId="35" pageNumber="255" value="1997-04-28" valueMax="1997-04-30" valueMin="1997-04-28">
<collectingDate id="EF8FE989EF06FF9DFD70FD30CABDE0BE" box="[703,902,687,714]" pageId="35" pageNumber="255" value="1997-04-28" valueMax="1997-04-30" valueMin="1997-04-28">28–30 Apr. 1997</collectingDate>
</date>
, F. 
<collectorName id="26805377EF06FF9DFC61FD30CAC5E0BE" box="[942,1022,687,713]" pageId="35" pageNumber="255">Génier</collectorName>
and S. 
<collectorName id="26805377EF06FF9DFB9AFD30CD45E0BE" box="[1109,1150,687,713]" pageId="35" pageNumber="255">Ide</collectorName>
; cloud forest, 
<collectingMethod id="52344EB6EF06FF9DFAECFD30C9D5E09A" pageId="35" pageNumber="255">carrion trap</collectingMethod>
; 
<collectionCode id="ED64AE64EF06FF9DFF32FD4CC868E09A" box="[253,339,723,749]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="35" pageNumber="255">CMNC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFEBDFD4CCB08E167" collectingDate="1997-04-28" collectingDateMax="1997-04-30" collectingDateMin="1997-04-28" collectionCode="CMNC" collectorName="Genier &amp; Ide" country="Brazil" county="Salesopolis" elevation="850" latitude="-23.648611" location="Serra do Mar" longLatPrecision="21" longitude="-45.87222" municipality="Estacao Biologica de Boraceia" pageId="35" pageNumber="255" specimenCount="20" specimenCount-female="8" specimenCount-male="12" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFEBDFD4CC8FCE09A" box="[370,455,723,749]" count="12" pageId="35" pageNumber="255" type="male">12 ♂♂</specimenCount>
, 
<specimenCount id="9D73FD28EF06FF9DFE19FD4CCB27E09A" box="[470,540,723,749]" count="8" pageId="35" pageNumber="255" type="female">8 ♀♀</specimenCount>
; same locality as for 
<typeStatus id="54CE8803EF06FF9DFCEBFD4DCAB5E09B" box="[804,910,722,748]" pageId="35" pageNumber="255">holotype</typeStatus>
; 
<date id="FFCB1061EF06FF9DFC52FD4CCD50E09A" box="[925,1131,723,749]" pageId="35" pageNumber="255" value="1997-04-28" valueMax="1997-04-30" valueMin="1997-04-28">
<collectingDate id="EF8FE989EF06FF9DFC52FD4CCD50E09A" box="[925,1131,723,749]" pageId="35" pageNumber="255" value="1997-04-28" valueMax="1997-04-30" valueMin="1997-04-28">28–30 Apr. 1997</collectingDate>
</date>
; F. 
<collectorName id="26805377EF06FF9DFB56FD4DCDD2E09B" box="[1177,1257,722,748]" pageId="35" pageNumber="255">Génier</collectorName>
and S. 
<collectorName id="26805377EF06FF9DFA86FD4DCC49E09B" box="[1353,1394,722,748]" pageId="35" pageNumber="255">Ide</collectorName>
; cloud forest, 
<collectingMethod id="52344EB6EF06FF9DFE94FD69C8F4E167" box="[347,463,758,784]" pageId="35" pageNumber="255">dung trap</collectingMethod>
; 
<collectionCode id="ED64AE64EF06FF9DFE12FD69CB08E167" box="[477,563,758,784]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="35" pageNumber="255">CMNC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFD9FFD69CBB9E144" collectingDate="1997-04-28" collectingDateMax="1997-04-30" collectingDateMin="1997-04-28" collectionCode="CMNC" collectorName="Genier &amp; Ide" country="Brazil" county="Salesopolis" elevation="850" latitude="-23.648611" location="Serra do Mar" longLatPrecision="21" longitude="-45.87222" municipality="Estacao Biologica de Boraceia" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFD9FFD69CBBAE167" box="[592,641,758,784]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; same locality as for 
<typeStatus id="54CE8803EF06FF9DFC4CFD69CAD6E167" box="[899,1005,758,784]" pageId="35" pageNumber="255">holotype</typeStatus>
; 
<date id="FFCB1061EF06FF9DFC34FD69CDFCE167" box="[1019,1223,758,784]" pageId="35" pageNumber="255" value="1997-04-28" valueMax="1997-04-30" valueMin="1997-04-28">
<collectingDate id="EF8FE989EF06FF9DFC34FD69CDFCE167" box="[1019,1223,758,784]" pageId="35" pageNumber="255" value="1997-04-28" valueMax="1997-04-30" valueMin="1997-04-28">28–30 Apr. 1997</collectingDate>
</date>
; F. 
<collectorName id="26805377EF06FF9DFB3CFD69CC78E167" box="[1267,1347,758,784]" pageId="35" pageNumber="255">Génier</collectorName>
and S. 
<collectorName id="26805377EF06FF9DFF14FC86C83FE144" box="[219,260,793,819]" pageId="35" pageNumber="255">Ide</collectorName>
; cloud forest, 
<collectingMethod id="52344EB6EF06FF9DFE62FC86CB1AE144" box="[429,545,793,819]" pageId="35" pageNumber="255">feces trap</collectingMethod>
; 
<collectionCode id="ED64AE64EF06FF9DFDE3FC86CBB9E144" box="[556,642,793,819]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="35" pageNumber="255">CMNC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFD56FC86CA23E121" collectingDate="1965-09-22" collectionCode="CEMT" country="Brazil" latitude="-23.648611" location="Estacao Biologica de Boraceia" longLatPrecision="21" longitude="-45.87222" municipality="Salesopolis" pageId="35" pageNumber="255" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFD56FC86CBFEE144" box="[665,709,793,819]" count="1" pageId="35" pageNumber="255" type="female">1 ♀</specimenCount>
; 
<collectingMunicipality id="6BAEACDBEF06FF9DFD1DFC86CA60E144" box="[722,859,793,819]" pageId="35" pageNumber="255">Salesópolis</collectingMunicipality>
, 
<location id="8EAA607AEF06FF9DFCA9FC86CDF5E144" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFCA9FC86CDF5E144" box="[870,1230,793,819]" country="Brazil" latitude="-23.648611" longLatPrecision="21" longitude="-45.87222" municipality="Salesopolis" name="Estacao Biologica de Boraceia" pageId="35" pageNumber="255" stateProvince="Sao Paulo">Estação Biológica de Boracéia</location>
; [
<geoCoordinate id="EE415066EF06FF9DFB2FFC86CC4AE144" box="[1248,1393,791,819]" degrees="23" direction="south" minutes="38" orientation="latitude" pageId="35" pageNumber="255" precision="15" seconds="55" value="-23.648611">23°38′55″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFF72FCA3C867E120" box="[189,348,827,855]" degrees="45" direction="west" minutes="52" orientation="longitude" pageId="35" pageNumber="255" precision="15" seconds="20" value="-45.87222">45°52′20″ W</geoCoordinate>
]; 
<date id="FFCB1061EF06FF9DFEBEFCA3CB34E120" box="[369,527,828,855]" pageId="35" pageNumber="255" value="1965-09-22">
<collectingDate id="EF8FE989EF06FF9DFEBEFCA3CB34E120" box="[369,527,828,855]" pageId="35" pageNumber="255" value="1965-09-22">22 Sep. 1965</collectingDate>
</date>
; [anonymous]; 
<collectionCode id="ED64AE64EF06FF9DFD08FCA3CA23E121" box="[711,792,828,854]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFCFCFCA3CB3BE10E" collectingDate="1965-10-06" collectionCode="CEMT" country="Brazil" latitude="-23.648611" location="Estacao Biologica de Boraceia" longLatPrecision="21" longitude="-45.87222" municipality="Salesopolis" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFCFCFCA3CA58E120" box="[819,867,828,855]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; same locality as for preceding; 
<date id="FFCB1061EF06FF9DFB2CFCA3CC49E120" box="[1251,1394,828,855]" pageId="35" pageNumber="255" value="1965-10-06">
<collectingDate id="EF8FE989EF06FF9DFB2CFCA3CC49E120" box="[1251,1394,828,855]" pageId="35" pageNumber="255" value="1965-10-06">6 Oct. 1965</collectingDate>
</date>
; [anonymous]; dung; 
<collectionCode id="ED64AE64EF06FF9DFE60FCC0CB3BE10E" box="[431,512,863,889]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFDD9FCC0C894E1EA" collectingDate="2012-09-21" collectionCode="CEMT" collectorName="Diaz" country="Brazil" latitude="-23.648611" location="Atlantic Forest" longLatPrecision="21" longitude="-45.87222" municipality="Salesopolis" pageId="35" pageNumber="255" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFDD9FCC0CB7AE10D" box="[534,577,863,890]" count="1" pageId="35" pageNumber="255" type="female">1 ♀</specimenCount>
; same locality as for preceding; 
<date id="FFCB1061EF06FF9DFC78FCC0CD69E10E" box="[951,1106,863,889]" pageId="35" pageNumber="255" value="2012-09-21">
<collectingDate id="EF8FE989EF06FF9DFC78FCC0CD69E10E" box="[951,1106,863,889]" pageId="35" pageNumber="255" value="2012-09-21">21 Sep. 2012</collectingDate>
</date>
; A. 
<collectorName id="26805377EF06FF9DFBB2FCC0CD83E10E" box="[1149,1208,863,889]" pageId="35" pageNumber="255">Díaz</collectorName>
; 
<location id="8EAA607AEF06FF9DFB0EFCC0CC54E10D" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFB0EFCC0CC54E10D" box="[1217,1391,863,890]" country="Brazil" name="Atlantic Forest" pageId="35" pageNumber="255" stateProvince="Sao Paulo">Atlantic Forest</location>
, human feces; 
<collectionCode id="ED64AE64EF06FF9DFE91FC1CC894E1EA" box="[350,431,899,925]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFE09FC1CCB3BE1B7" collectingDate="1926-04" collectionCode="MZSP" country="Brazil" latitude="-23.783333" location="Estacao Biologica do Alto da Serra" longLatPrecision="1254" longitude="-46.3" municipality="Santo Andre" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFE09FC1CC8CFE1EA" box="[454,500,899,925]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="6BAEACDBEF06FF9DFE30FC1CCBA8E1EB" box="[511,659,898,925]" pageId="35" pageNumber="255">Santo André</collectingMunicipality>
, 
<location id="8EAA607AEF06FF9DFD54FC1CCD0DE1EA" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFD54FC1CCD0DE1EA" box="[667,1078,898,925]" country="Brazil" latitude="-23.783333" longLatPrecision="1254" longitude="-46.3" municipality="Santo Andre" name="Estacao Biologica do Alto da Serra" pageId="35" pageNumber="255" stateProvince="Sao Paulo">Estação Biológica do Alto da Serra</location>
; [
<geoCoordinate id="EE415066EF06FF9DFB87FC1ECD89E1EA" box="[1096,1202,897,925]" degrees="23" direction="south" minutes="47" orientation="latitude" pageId="35" pageNumber="255" precision="925" value="-23.783333">23°47′ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFB71FC1ECC09E1EA" box="[1214,1330,897,925]" degrees="46" direction="west" minutes="18" orientation="longitude" pageId="35" pageNumber="255" precision="925" value="-46.3">46°18′ W</geoCoordinate>
]; 
<date id="FFCB1061EF06FF9DFA8CFC1CC9C0E1B7" pageId="35" pageNumber="255" value="1926-04">
<collectingDate id="EF8FE989EF06FF9DFA8CFC1CC9C0E1B7" pageId="35" pageNumber="255" value="1926-04">Apr. 1926</collectingDate>
</date>
; [anonymous]; 
<collectionCode id="ED64AE64EF06FF9DFE7BFC39CB3BE1B7" box="[436,512,934,960]" collectionName="Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="35" pageNumber="255" type="Museum">MZSP</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFDD3FC39CD15E194" collectingDate="2012-05-13" collectionCode="CEMT" collectorName="M. Boutefeu" country="Brazil" elevation="580" latitude="-24.066387" location="Parque Estadual Carlos Botelho" longLatPrecision="21" longitude="-47.994167" municipality="Sao Miguel Arcanjo" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFDD3FC39CB77E1B7" box="[540,588,934,960]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="6BAEACDBEF06FF9DFD95FC39CA76E1B7" box="[602,845,934,960]" pageId="35" pageNumber="255">São Miguel Arcanjo</collectingMunicipality>
, 
<location id="8EAA607AEF06FF9DFC96FC39CDE3E1B7" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFC96FC39CDE3E1B7" box="[857,1240,934,960]" country="Brazil" latitude="-24.066387" longLatPrecision="21" longitude="-47.994167" municipality="Sao Miguel Arcanjo" name="Parque Estadual Carlos Botelho" pageId="35" pageNumber="255" stateProvince="Sao Paulo">Parque Estadual Carlos Botelho</location>
; [
<geoCoordinate id="EE415066EF06FF9DFB23FC3BCC4AE1B7" box="[1260,1393,932,960]" degrees="24" direction="south" minutes="3" orientation="latitude" pageId="35" pageNumber="255" precision="15" seconds="59" value="-24.066387">24°3′59″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFF72FC58C866E194" box="[189,349,967,995]" degrees="47" direction="west" minutes="59" orientation="longitude" pageId="35" pageNumber="255" precision="15" seconds="39" value="-47.994167">47°59′39″ W</geoCoordinate>
]; 
<quantity id="4C8D9B44EF06FF9DFEBBFC56C8F8E194" box="[372,451,969,996]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="35" pageNumber="255" unit="m" value="580.0">
<elevation id="0058D192EF06FF9DFEBBFC56C8F8E194" box="[372,451,969,996]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="35" pageNumber="255" unit="m" value="580.0">580 m</elevation>
</quantity>
; 
<date id="FFCB1061EF06FF9DFE1DFC56CB4DE194" box="[466,630,969,995]" pageId="35" pageNumber="255" value="2012-05-13">
<collectingDate id="EF8FE989EF06FF9DFE1DFC56CB4DE194" box="[466,630,969,995]" pageId="35" pageNumber="255" value="2012-05-13">13 May 2012</collectingDate>
</date>
; 
<collectorName id="26805377EF06FF9DFD4BFC56CA25E194" box="[644,798,969,995]" pageId="35" pageNumber="255">M. Boutefeu</collectorName>
; 
<taxonomicName id="4C754D22EF06FF9DFCE3FC55CABEE194" authorityName="Brisson" authorityYear="1762" box="[812,901,970,995]" class="Mammalia" family="Tapiridae" genus="Tapirus" kingdom="Animalia" order="Perissodactyla" pageId="35" pageNumber="255" phylum="Chordata" rank="genus">
<emphasis id="B901EAB3EF06FF9DFCE3FC55CABEE194" box="[812,901,970,995]" italics="true" pageId="35" pageNumber="255">Tapirus</emphasis>
</taxonomicName>
feces; 
<collectionCode id="ED64AE64EF06FF9DFC12FC56CD15E194" box="[989,1070,969,995]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFB84FC56CA40E671" collectingDate="2012-05-15" collectionCode="CEMT" collectorName="Boutefeu" country="Brazil" elevation="580" latitude="-24.066387" location="Parque Estadual Carlos Botelho" longLatPrecision="21" longitude="-47.994167" municipality="Sao Miguel Arcanjo" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFB84FC56CD47E194" box="[1099,1148,969,995]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; same locality as for preceding; 
<date id="FFCB1061EF06FF9DFEF1FC73C8E2E671" box="[318,473,1004,1031]" pageId="35" pageNumber="255" value="2012-05-15">
<collectingDate id="EF8FE989EF06FF9DFEF1FC73C8E2E671" box="[318,473,1004,1031]" pageId="35" pageNumber="255" value="2012-05-15">15 May 2012</collectingDate>
</date>
; M. 
<collectorName id="26805377EF06FF9DFDC5FC73CB42E671" box="[522,633,1004,1030]" pageId="35" pageNumber="255">Boutefeu</collectorName>
; 
<taxonomicName id="4C754D22EF06FF9DFD4FFC72CBE2E671" authorityName="Brisson" authorityYear="1762" box="[640,729,1005,1030]" class="Mammalia" family="Tapiridae" genus="Tapirus" kingdom="Animalia" order="Perissodactyla" pageId="35" pageNumber="255" phylum="Chordata" rank="genus">
<emphasis id="B901EAB3EF06FF9DFD4FFC72CBE2E671" box="[640,729,1005,1030]" italics="true" pageId="35" pageNumber="255">Tapirus</emphasis>
</taxonomicName>
feces; 
<collectionCode id="ED64AE64EF06FF9DFCE5FC73CA40E671" box="[810,891,1004,1030]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFC40FC73CC5FE65E" collectingDate="2012-01-29" collectionCode="CEMT" collectorName="E. Bovy" country="Brazil" elevation="806" latitude="-24.06111" location="Parque Estadual Carlos Botelho" longLatPrecision="21" longitude="-47.97889" municipality="Sao Miguel Arcanjo" pageId="35" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFC40FC73CA87E670" box="[911,956,1004,1031]" count="1" pageId="35" pageNumber="255" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="6BAEACDBEF06FF9DFC09FC73CD89E671" box="[966,1202,1004,1031]" pageId="35" pageNumber="255">São Miguel Arcanjo</collectingMunicipality>
, 
<location id="8EAA607AEF06FF9DFB74FC72C84AE65E" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFB74FC72C84AE65E" country="Brazil" latitude="-24.06111" longLatPrecision="21" longitude="-47.97889" municipality="Sao Miguel Arcanjo" name="Parque Estadual Carlos Botelho" pageId="35" pageNumber="255" stateProvince="Sao Paulo">Parque Estadual Carlos Botelho</location>
; [
<geoCoordinate id="EE415066EF06FF9DFE4BFB91CB30E65E" box="[388,523,1038,1066]" degrees="24" direction="south" minutes="3" orientation="latitude" pageId="35" pageNumber="255" precision="15" seconds="40" value="-24.06111">24°3′40″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFDD6FB91CB82E65D" box="[537,697,1038,1066]" degrees="47" direction="west" minutes="58" orientation="longitude" pageId="35" pageNumber="255" precision="15" seconds="44" value="-47.97889">47°58′44″ W</geoCoordinate>
]; 
<quantity id="4C8D9B44EF06FF9DFD01FB90CA27E65D" box="[718,796,1039,1066]" metricMagnitude="2" metricUnit="m" metricValue="8.059999999999999" pageId="35" pageNumber="255" unit="m" value="806.0">
<elevation id="0058D192EF06FF9DFD01FB90CA27E65D" box="[718,796,1039,1066]" metricMagnitude="2" metricUnit="m" metricValue="8.059999999999999" pageId="35" pageNumber="255" unit="m" value="806.0">806 m</elevation>
</quantity>
; 
<date id="FFCB1061EF06FF9DFCE6FB90CAFFE65E" box="[809,964,1039,1066]" pageId="35" pageNumber="255" value="2012-01-29">
<collectingDate id="EF8FE989EF06FF9DFCE6FB90CAFFE65E" box="[809,964,1039,1066]" pageId="35" pageNumber="255" value="2012-01-29">29 Jan. 2012</collectingDate>
</date>
; 
<collectorName id="26805377EF06FF9DFC1EFB8FCD08E65D" box="[977,1075,1040,1066]" pageId="35" pageNumber="255">E. Bovy</collectorName>
(1); human feces; 
<collectionCode id="ED64AE64EF06FF9DFADCFB90CC5FE65E" box="[1299,1380,1039,1065]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFF72FBACCBAFE607" collectingDate="2012-01-29" collectionCode="CEMT" collectorName="E. Bovy" country="Brazil" elevation="823" latitude="-24.062777" location="Parque Estadual Carlos Botelho" longLatPrecision="21" longitude="-47.97889" municipality="Sao Miguel Arcanjo" pageId="35" pageNumber="255" specimenCount="3" specimenCount-female="3" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFF72FBACC9C4E63A" box="[189,255,1075,1101]" count="3" pageId="35" pageNumber="255" type="female">3 ♀♀</specimenCount>
; 
<collectingMunicipality id="6BAEACDBEF06FF9DFEC3FBACC8C2E63B" box="[268,505,1074,1101]" pageId="35" pageNumber="255">São Miguel Arcanjo</collectingMunicipality>
, 
<location id="8EAA607AEF06FF9DFDCCFBACCA42E63B" LSID="urn:lsid:plazi:treatment:03DC87B7EF07FF99FE30FE0ECC4CE761:8EAA607AEF06FF9DFDCCFBACCA42E63B" box="[515,889,1074,1101]" country="Brazil" latitude="-24.062777" longLatPrecision="21" longitude="-47.97889" municipality="Sao Miguel Arcanjo" name="Parque Estadual Carlos Botelho" pageId="35" pageNumber="255" stateProvince="Sao Paulo">Parque Estadual Carlos Botelho</location>
; [
<geoCoordinate id="EE415066EF06FF9DFC45FBACCD37E63A" box="[906,1036,1073,1101]" degrees="24" direction="south" minutes="3" orientation="latitude" pageId="35" pageNumber="255" precision="15" seconds="46" value="-24.062777">24°3′46″ S</geoCoordinate>
, 
<geoCoordinate id="EE415066EF06FF9DFBD7FBACCD8EE63A" box="[1048,1205,1073,1101]" degrees="47" direction="west" minutes="58" orientation="longitude" pageId="35" pageNumber="255" precision="15" seconds="44" value="-47.97889">47°58′44″ W</geoCoordinate>
]; 
<quantity id="4C8D9B44EF06FF9DFB07FBACCC2FE63A" box="[1224,1300,1075,1101]" metricMagnitude="2" metricUnit="m" metricValue="8.23" pageId="35" pageNumber="255" unit="m" value="823.0">
<elevation id="0058D192EF06FF9DFB07FBACCC2FE63A" box="[1224,1300,1075,1101]" metricMagnitude="2" metricUnit="m" metricValue="8.23" pageId="35" pageNumber="255" unit="m" value="823.0">823 m</elevation>
</quantity>
; 
<date id="FFCB1061EF06FF9DFAEAFBACC9C0E607" pageId="35" pageNumber="255" value="2012-01-29">
<collectingDate id="EF8FE989EF06FF9DFAEAFBACC9C0E607" pageId="35" pageNumber="255" value="2012-01-29">29 Jan. 2012</collectingDate>
</date>
; 
<collectorName id="26805377EF06FF9DFEC8FBC9C853E607" box="[263,360,1110,1136]" pageId="35" pageNumber="255">E. Bovy</collectorName>
(2); human feces; 
<collectionCode id="ED64AE64EF06FF9DFD8CFBC9CBAFE607" box="[579,660,1110,1136]" pageId="35" pageNumber="255">CEMT</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1D3CFCEF06FF9DFD62FBC9CD44E607" box="[685,1151,1110,1136]" collectionCode="MNHN" country="Brazil" pageId="35" pageNumber="255" specimenCount="2" specimenCount-male="2" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="9D73FD28EF06FF9DFD62FBC9CBC9E607" box="[685,754,1110,1136]" count="2" pageId="35" pageNumber="255" type="male">2 ♂♂</specimenCount>
; [no date]; [anonymous]; 
<collectionCode id="ED64AE64EF06FF9DFBEEFBC9CD44E607" box="[1057,1151,1110,1136]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="255">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C36F652AEF06FF9BFF72FB08CBB3E1C7" lastPageId="37" lastPageNumber="257" pageId="35" pageNumber="255" type="description">
<paragraph id="8BCA36A1EF06FF9DFF72FB08C875E6C6" blockId="35.[189,334,1175,1201]" box="[189,334,1175,1201]" pageId="35" pageNumber="255">
<heading id="D08281CDEF06FF9DFF72FB08C875E6C6" bold="true" box="[189,334,1175,1201]" fontSize="11" level="3" pageId="35" pageNumber="255" reason="3">
<emphasis id="B901EAB3EF06FF9DFF72FB08C875E6C6" bold="true" box="[189,334,1175,1201]" pageId="35" pageNumber="255">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36A1EF06FF9DFF25FB59CBAAE697" blockId="35.[189,657,1222,1284]" box="[234,657,1222,1248]" pageId="35" pageNumber="255">
<emphasis id="B901EAB3EF06FF9DFF25FB59C8A6E697" bold="true" box="[234,413,1222,1248]" pageId="35" pageNumber="255">
Male 
<typeStatus id="54CE8803EF06FF9DFEFEFB59C8A6E697" box="[305,413,1222,1248]" pageId="35" pageNumber="255" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="134E2A24EF06FF9DFE63FB59CB3DE697" box="[428,518,1222,1248]" captionStart="Figs 9–12" captionStartId="11.[189,242,1928,1954]" captionTargetBox="[207,1370,321,1842]" captionTargetId="figure-18@11.[207,1378,265,1880]" captionTargetPageId="11" captionText="Figs 9–12. Dorsal habitus. 9. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 10. Homocopris torulosus, specimen ♀ (repository unknown). 11. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 12. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824974" httpUri="https://zenodo.org/record/10824974/files/figure.png" pageId="35" pageNumber="255">Figs 11</figureCitation>
, 
<figureCitation id="134E2A24EF06FF9DFDDCFB59CB08E697" box="[531,563,1222,1248]" captionStart="Figs 21–24" captionStartId="16.[189,242,1928,1954]" captionTargetBox="[210,1378,289,1875]" captionTargetId="figure-12@16.[210,1378,265,1875]" captionTargetPageId="16" captionText="Figs 21–24. Head and pronotum, lateral view.21. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 22. Homocopris torulosus, specimen ♀ (repository unknown). 23. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 24. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824982" httpUri="https://zenodo.org/record/10824982/files/figure.png" pageId="35" pageNumber="255">23</figureCitation>
, 
<figureCitation id="134E2A24EF06FF9DFDF0FB59CB64E697" box="[575,607,1222,1248]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="35" pageNumber="255">30</figureCitation>
, 
<figureCitation id="134E2A24EF06FF9DFDA4FB59CBB1E697" box="[619,650,1222,1248]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="35" pageNumber="255">36</figureCitation>
)
</paragraph>
<paragraph id="8BCA36A1EF06FF9DFF72FB76CBBEE774" blockId="35.[189,657,1222,1284]" box="[189,645,1257,1284]" pageId="35" pageNumber="255">
<smallCapsWord id="8D2CA07DEF06FF9DFF72FB76C855E776" baselines="1276,1276" box="[189,366,1257,1283]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="35" pageNumber="255">MEASUREMENTS</smallCapsWord>
. Body length 
<quantity id="4C8D9B44EF06FF9DFDDAFB76CBBBE774" box="[533,640,1257,1284]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="35" pageNumber="255" unit="mm" value="15.1">15.1 mm</quantity>
.
</paragraph>
<paragraph id="8BCA36A1EF06FF9DFF72FAB4CA36E7A6" blockId="35.[189,1401,1322,1490]" pageId="35" pageNumber="255">
<smallCapsWord id="8D2CA07DEF06FF9DFF72FAB4C9C7E734" baselines="1342,1342" box="[189,252,1323,1349]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="35" pageNumber="255">HEAD</smallCapsWord>
. Clypeus bidentate with broad, rounded teeth. Anteroventral clypeal tooth broad. Clypeal and genal surface rugose, clypeogenal suture demarcated by shallow notch. Posterior genal angle obtuse. Frontal surface reduced, smooth. Dorsal ocular width subequal to one-sixth interocular distance. Interocular surface medially smooth. Cephalic horn emerging centrally from clypeofrontal surface, tapering apically, gently curved, 
<quantity id="4C8D9B44EF06FF9DFDF2FA28CBACE7A5" box="[573,663,1463,1490]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="35" pageNumber="255" unit="mm" value="4.7">4.7 mm</quantity>
in length.
</paragraph>
<paragraph id="8BCA36A1EF06FF9DFF72FA66CB1CE40B" blockId="35.[188,1399,1529,1661]" pageId="35" pageNumber="255">
<smallCapsWord id="8D2CA07DEF06FF9DFF72FA66C87BE466" baselines="1548,1548" box="[189,320,1529,1555]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Prothorax" pageId="35" pageNumber="255">PROTHORAX</smallCapsWord>
. Anteromedial pronotal edge curvature continuous. Lateral pronotal edge angular on anterior fourth in dorsal view. Anterior inclined pronotal surface with fine, transverse rugosities. Pronotal ridge with two sets of distinct bilaterally paired tubercles. Posterior pronotal surface uniformly punctate. Prosternal apex acutely tipped.
</paragraph>
<paragraph id="8BCA36A1EF06FF9DFF72F93BCB95E572" blockId="35.[189,1399,1700,1797]" pageId="35" pageNumber="255">
<smallCapsWord id="8D2CA07DEF06FF9DFF72F93BC829E4CB" baselines="1719,1719" box="[189,274,1700,1726]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Elytra" pageId="35" pageNumber="255">ELYTRA</smallCapsWord>
. Glossy overall. Striae 1–6 widest anteriorly; narrowing posteriorly. Third and fourth striae posteriorly joined. Eighth stria absent throughout. Ninth stria effaced along anterior quarter of elytral length. Interstriae convex, finely punctate.
</paragraph>
<paragraph id="8BCA36A1EF06FF9DFF72F8B3CDD6E531" blockId="35.[189,1261,1835,1862]" box="[189,1261,1835,1862]" pageId="35" pageNumber="255">
<smallCapsWord id="8D2CA07DEF06FF9DFF72F8B3C814E533" baselines="1855,1855" box="[189,303,1836,1862]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Hindwing" pageId="35" pageNumber="255">HINDWING</smallCapsWord>
. Membrane edge between AP and AA sinuous. AP vein entire, converging with J.
</paragraph>
<paragraph id="8BCA36A1EF06FF9DFF72F8F2CAE1E5B9" blockId="35.[189,1399,1901,1998]" pageId="35" pageNumber="255">
<smallCapsWord id="8D2CA07DEF06FF9DFF72F8F2C80FE5F2" baselines="1920,1920" box="[189,308,1901,1927]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Ventrites" pageId="35" pageNumber="255">VENTRITES</smallCapsWord>
. Meso-metasternal suture straight. Surface of median metasternal lobe with uneven texture and deep, anteriorly divided impression. Lateral metasternal lobe densely setose. Fifth abdominal ventrite with sparse medial punctation. Pygidium coarsely punctate.
</paragraph>
<paragraph id="8BCA36A1EF01FF9AFF72FE96CB92E31E" blockId="36.[189,1399,265,361]" pageId="36" pageNumber="256">
<smallCapsWord id="8D2CA07DEF01FF9AFF72FE96C86FE356" baselines="284,284" box="[189,340,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Prothoracic" pageId="36" pageNumber="256">PROTHORACIC</smallCapsWord>
<smallCapsWord id="8D2CA07DEF01FF9AFEAFFE91C8AFE356" baselines="284" box="[352,404,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="legs" pageId="36" pageNumber="256">LEGS</smallCapsWord>
. Anterior and posterior surfaces of profemur with dense row of long dark setae. Ventral profemoral surface finely punctate, sparsely setose. Protibia with three distinct teeth. Protibial forespur apically tapered, curving inward.
</paragraph>
<paragraph id="8BCA36A1EF01FF9AFF72FE0ECA14E386" blockId="36.[189,1399,401,498]" pageId="36" pageNumber="256">
<smallCapsWord id="8D2CA07DEF01FF9AFF72FE0EC853E3DE" baselines="420,420" box="[189,360,401,427]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesothoracic" pageId="36" pageNumber="256">MESOTHORACIC</smallCapsWord>
<smallCapsWord id="8D2CA07DEF01FF9AFEBAFE09C892E3DE" baselines="420" box="[373,425,406,425]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="legs" pageId="36" pageNumber="256">LEGS</smallCapsWord>
. Posterior surface of mesotrochanter with tuft of long dark setae. Anterior and posterior surfaces of mesofemur with sparse row of long dark setae. Outer surface of mesotibia serrate. Width at mesotibial apex one-third mesotibial length.
</paragraph>
<paragraph id="8BCA36A1EF01FF9AFF72FD86CA1DE00E" blockId="36.[189,1399,537,634]" pageId="36" pageNumber="256">
<smallCapsWord id="8D2CA07DEF01FF9AFF72FD86C85EE046" baselines="556,556" box="[189,357,537,563]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Metathoracic" pageId="36" pageNumber="256">METATHORACIC</smallCapsWord>
<smallCapsWord id="8D2CA07DEF01FF9AFEBDFD81C89DE046" baselines="556" box="[370,422,542,561]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="legs" pageId="36" pageNumber="256">LEGS</smallCapsWord>
. Posterior surface of metatrochanter with tuft of long dark setae. Anterior and posterior surfaces of metafemur with sparse row of long dark setae. Outer surface of metatibia serrate. Width at metatibial apex one-third metatibial length.
</paragraph>
<paragraph id="8BCA36A1EF01FF9AFF72FD3EC9C0E153" blockId="36.[188,1398,672,804]" pageId="36" pageNumber="256">
<smallCapsWord id="8D2CA07DEF01FF9AFF72FD3EC9C4E0CE" baselines="692,692" box="[189,255,673,699]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="36" pageNumber="256">MALE</smallCapsWord>
<smallCapsWord id="8D2CA07DEF01FF9AFEC2FD39C840E0CE" baselines="692" box="[269,379,678,697]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="36" pageNumber="256">GENITALIA</smallCapsWord>
. Aedeagus length: 
<quantity id="4C8D9B44EF01FF9AFDA2FD3FCBEBE0CC" box="[621,720,672,699]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="36" pageNumber="256" unit="mm" value="3.5">3.5 mm</quantity>
. Medially paired sclerites of genital segment elongate, subequal in length to lateral sclerites. Paramere (
<figureCitation id="134E2A24EF01FF9AFD36FD5BCA74E0A9" box="[761,847,708,734]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="36" pageNumber="256">Fig. 30</figureCitation>
) laterally flattened, slightly tapering from base to apex with apex evenly rounded. FLP endophallite (
<figureCitation id="134E2A24EF01FF9AFCE6FD78CA45E176" box="[809,894,743,769]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="36" pageNumber="256">Fig. 36</figureCitation>
) elongate with two indentations along right edge.
</paragraph>
<caption id="DF0A6629EF01FF9AFF72F81DCB6BE595" ID-DOI="http://doi.org/10.5281/zenodo.10824996" ID-Zenodo-Dep="10824996" httpUri="https://zenodo.org/record/10824996/files/figure.png" pageId="36" pageNumber="256" startId="36.[189,232,1922,1948]" targetBox="[189,1398,975,1873]" targetPageId="36">
<paragraph id="8BCA36A1EF01FF9AFF72F81DCB6BE595" blockId="36.[189,1399,1922,2019]" pageId="36" pageNumber="256">
<emphasis id="B901EAB3EF01FF9AFF72F81DC827E5EB" bold="true" box="[189,284,1922,1948]" pageId="36" pageNumber="256">Fig. 65</emphasis>
. 
<taxonomicName id="4C754D22EF01FF9AFEFBF81DCA2EE5EB" authority="(Harold, 1867)" authorityName="Génier &amp; Darling" authorityYear="2024" baseAuthorityName="Harold" baseAuthorityYear="1867" box="[308,789,1922,1949]" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="256" phylum="Arthropoda" rank="species" species="achamas" status="gen. et comb. nov.">
<emphasis id="B901EAB3EF01FF9AFEFBF81DCB70E5EB" box="[308,587,1922,1948]" italics="true" pageId="36" pageNumber="256">Andinocopris achamas</emphasis>
(
<bibRefCitation id="EFE44B50EF01FF9AFDABF81DCA30E5EB" author="Harold E. von" box="[612,779,1922,1949]" pageId="36" pageNumber="256" pagination="95 - 104" refId="ref26084" refString="Harold E. von 1867. Diagnosen neuer Coprophagen. Coleopterologische Hefte 2: 95 - 104. Available from https: // www. biodiversitylibrary. org / page / 16069770 [accessed 6 Feb. 2024]." type="journal article" year="1867">Harold, 1867</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23257C8EF01FF9AFCE9F81CCD21E5EA" box="[806,1050,1922,1949]" pageId="36" pageNumber="256" rank="species">gen. et comb. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C754D22EF01FF9AFBA9F81DC89EE5C8" authority="(Waterhouse, 1891)" authorityName="Génier &amp; Darling" authorityYear="2024" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="256" phylum="Arthropoda" rank="species" species="buckleyi" status="gen. et comb. nov.">
<emphasis id="B901EAB3EF01FF9AFBA9F81DCC4CE5EB" box="[1126,1399,1922,1948]" italics="true" pageId="36" pageNumber="256">Andinocopris buckleyi</emphasis>
(
<bibRefCitation id="EFE44B50EF01FF9AFF0AF83AC8A7E5C8" author="Waterhouse C. O." box="[197,412,1957,1983]" pageId="36" pageNumber="256" pagination="348 - 363" refId="ref28366" refString="Waterhouse C. O. 1891. New Scarabaeidae in the British Museum: a fourth contribution. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology. Sixth Series 7 (40): 348 - 363. https: // doi. org / 10.1080 / 00222939109460622" type="journal article" year="1891">Waterhouse, 1891</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23257C8EF01FF9AFE63F839CBBEE5B7" box="[428,645,1957,1984]" pageId="36" pageNumber="256" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, distribution map. Legend and scale bar included in the top and bottom right corners, respectively.
</paragraph>
</caption>
<paragraph id="8BCA36A1EF00FF9BFF25FE96CB75E354" blockId="37.[189,1399,265,396]" box="[234,590,265,291]" pageId="37" pageNumber="257">
<emphasis id="B901EAB3EF00FF9BFF25FE96C88BE354" bold="true" box="[234,432,265,291]" pageId="37" pageNumber="257">
Female 
<typeStatus id="54CE8803EF00FF9BFE82FE96C88BE354" box="[333,432,265,291]" pageId="37" pageNumber="257" type="allotype">allotype</typeStatus>
</emphasis>
(
<figureCitation id="134E2A24EF00FF9BFE0FFE96CB20E354" box="[448,539,265,291]" captionStart="Figs 9–12" captionStartId="11.[189,242,1928,1954]" captionTargetBox="[207,1370,321,1842]" captionTargetId="figure-18@11.[207,1378,265,1880]" captionTargetPageId="11" captionText="Figs 9–12. Dorsal habitus. 9. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 10. Homocopris torulosus, specimen ♀ (repository unknown). 11. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 12. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824974" httpUri="https://zenodo.org/record/10824974/files/figure.png" pageId="37" pageNumber="257">Figs 12</figureCitation>
, 
<figureCitation id="134E2A24EF00FF9BFDE8FE96CB7DE354" box="[551,582,265,291]" captionStart="Figs 21–24" captionStartId="16.[189,242,1928,1954]" captionTargetBox="[210,1378,289,1875]" captionTargetId="figure-12@16.[210,1378,265,1875]" captionTargetPageId="16" captionText="Figs 21–24. Head and pronotum, lateral view.21. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 22. Homocopris torulosus, specimen ♀ (repository unknown). 23. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 24. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824982" httpUri="https://zenodo.org/record/10824982/files/figure.png" pageId="37" pageNumber="257">24</figureCitation>
)
</paragraph>
<paragraph id="8BCA36A1EF00FF9BFF72FEB3C80EE3FB" blockId="37.[189,1399,265,396]" pageId="37" pageNumber="257">
Similar to male with the following exceptions: Body length 
<quantity id="4C8D9B44EF00FF9BFC4AFEB3CACAE330" box="[901,1009,300,327]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="37" pageNumber="257" unit="mm" value="14.3">14.3 mm</quantity>
. Clypeofrontal carina transverse, centrally raised. Interocular surface chagrined. Anterior pronotal ridge low, followed by shallow concavity.
</paragraph>
<paragraph id="8BCA36A1EF00FF9BFF72FE2BC808E3B9" blockId="37.[189,307,436,462]" box="[189,307,436,462]" pageId="37" pageNumber="257">
<heading id="D08281CDEF00FF9BFF72FE2BC808E3B9" bold="true" box="[189,307,436,462]" fontSize="11" level="3" pageId="37" pageNumber="257" reason="3">
<emphasis id="B901EAB3EF00FF9BFF72FE2BC808E3B9" bold="true" box="[189,307,436,462]" pageId="37" pageNumber="257">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36A1EF00FF9BFF72FE7DC8C6E011" blockId="37.[188,1399,482,615]" pageId="37" pageNumber="257">
Body length 
<quantity id="4C8D9B44EF00FF9BFE98FE7CC8E1E38A" box="[343,474,483,509]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="37" pageNumber="257" unit="mm" value="16.0" valueMax="19.0" valueMin="13.0">13–19 mm</quantity>
. Small males with medially projecting clypeofrontal ridge instead of cephalic horn. Anteromedial pronotal edge evenly curved more often than indented. Pronotal ridge in small males low, medially notched. Elytral interstriae finely punctate (visible at 10 ×) more often than minutely punctate (invisible at 10 ×).
</paragraph>
<paragraph id="8BCA36A1EF00FF9BFF72FD12C8FFE0D0" blockId="37.[189,452,653,679]" box="[189,452,653,679]" pageId="37" pageNumber="257">
<emphasis id="B901EAB3EF00FF9BFF72FD12C86DE0D0" bold="true" box="[189,342,653,679]" pageId="37" pageNumber="257">Distribution</emphasis>
(
<figureCitation id="134E2A24EF00FF9BFEA9FD12C887E0D0" box="[358,444,653,679]" captionStart="Fig" captionStartId="37.[189,232,1952,1978]" captionTargetBox="[189,1398,998,1904]" captionTargetId="figure-178@37.[189,1398,997,1904]" captionTargetPageId="37" captionText="Fig. 66. Homocopris grossiorum Darling &amp; Génier sp. nov. and Homocopris williami Darling &amp; Génier sp. nov., distribution map. Legend and scale bar included in the top and bottom right corners, respectively." figureDoi="http://doi.org/10.5281/zenodo.10825000" httpUri="https://zenodo.org/record/10825000/files/figure.png" pageId="37" pageNumber="257">Fig. 66</figureCitation>
)
</paragraph>
<paragraph id="8BCA36A1EF00FF9BFF72FD23CD5FE08D" blockId="37.[189,1398,700,762]" pageId="37" pageNumber="257">
Serra do Mar and Serra Geral Mountains in the Brazilian states of 
<collectingRegion id="49B1F843EF00FF9BFC25FD23CDA7E0A1" box="[1002,1180,700,726]" country="Brazil" name="Rio de Janeiro" pageId="37" pageNumber="257">Rio de Janeiro</collectingRegion>
, 
<collectingRegion id="49B1F843EF00FF9BFB64FD23CC1FE0A1" box="[1195,1316,700,726]" country="Brazil" name="Sao Paulo" pageId="37" pageNumber="257">São Paulo</collectingRegion>
, 
<collectingRegion id="49B1F843EF00FF9BFAFAFD23C818E08E" country="Brazil" name="Santa Catarina" pageId="37" pageNumber="257">Santa Catarina</collectingRegion>
, and 
<collectingRegion id="49B1F843EF00FF9BFEAEFD40CB04E08E" box="[353,575,735,761]" country="Brazil" name="Rio Grande do Sul" pageId="37" pageNumber="257">Rio Grande do Sul</collectingRegion>
, with recorded elevations from 
<quantity id="4C8D9B44EF00FF9BFC7CFD40CD5BE08D" box="[947,1120,735,762]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="37" pageNumber="257" unit="m" value="1200.0" valueMax="1600.0" valueMin="800.0">800 to 1600 m</quantity>
.
</paragraph>
<paragraph id="8BCA36A1EF00FF9BFF72FCBEC845E14D" blockId="37.[189,382,800,827]" box="[189,382,800,827]" pageId="37" pageNumber="257">
<heading id="D08281CDEF00FF9BFF72FCBEC845E14D" bold="true" box="[189,382,800,827]" fontSize="11" level="3" pageId="37" pageNumber="257" reason="3">
<emphasis id="B901EAB3EF00FF9BFF72FCBEC845E14D" bold="true" box="[189,382,800,827]" pageId="37" pageNumber="257">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36A1EF00FF9BFF72FCD0CBB3E1C7" blockId="37.[189,1399,847,944]" pageId="37" pageNumber="257">
Specimens with data were collected in cloud and sub-montane Atlantic Forest. Some specimens came to carrion and dung traps, pitfall traps baited with human faeces and faeces of 
<taxonomicName id="4C754D22EF00FF9BFB98FCECCC61E1FB" authority="Brisson, 1762" authorityName="Brisson" authorityYear="1762" box="[1111,1370,882,908]" class="Mammalia" family="Tapiridae" genus="Tapirus" kingdom="Animalia" order="Perissodactyla" pageId="37" pageNumber="257" phylum="Chordata" rank="genus">
<emphasis id="B901EAB3EF00FF9BFB98FCECCD8BE1FB" box="[1111,1200,883,908]" italics="true" pageId="37" pageNumber="257">Tapirus</emphasis>
Brisson, 1762
</taxonomicName>
at elevations between 
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and 
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.
</paragraph>
</subSubSection>
<caption id="DF0A6629EF00FF9BFF72F83FCC4CE5A9" ID-DOI="http://doi.org/10.5281/zenodo.10825000" ID-Zenodo-Dep="10825000" httpUri="https://zenodo.org/record/10825000/files/figure.png" pageId="37" pageNumber="257" startId="37.[189,232,1952,1978]" targetBox="[189,1398,998,1904]" targetPageId="37">
<paragraph id="8BCA36A1EF00FF9BFF72F83FCC4CE5A9" blockId="37.[188,1399,1952,2014]" pageId="37" pageNumber="257">
<emphasis id="B901EAB3EF00FF9BFF72F83FC82AE5CD" bold="true" box="[189,273,1952,1978]" pageId="37" pageNumber="257">Fig. 66</emphasis>
. 
<taxonomicName id="4C754D22EF00FF9BFEEFF83ECA2AE5CD" authority="Darling &amp; Genier" authorityName="Darling &amp; Genier" authorityYear="2024" box="[288,785,1952,1979]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="257" phylum="Arthropoda" rank="species" species="grossiorum" status="sp. nov.">
<emphasis id="B901EAB3EF00FF9BFEEFF83ECB06E5CD" box="[288,573,1953,1978]" italics="true" pageId="37" pageNumber="257">Homocopris grossiorum</emphasis>
Darling &amp; Génier
</taxonomicName>
<taxonomicNameLabel id="A23257C8EF00FF9BFCD7F83ECA49E5CC" box="[792,882,1953,1979]" pageId="37" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C754D22EF00FF9BFC65F83ECC4DE5CD" authority="Darling &amp; Genier" authorityName="Darling &amp; Genier" authorityYear="2024" box="[938,1398,1952,1979]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="257" phylum="Arthropoda" rank="species" species="williami" status="sp. nov.">
<emphasis id="B901EAB3EF00FF9BFC65F83ECD99E5CD" box="[938,1186,1952,1978]" italics="true" pageId="37" pageNumber="257">Homocopris williami</emphasis>
Darling &amp; Génier
</taxonomicName>
<taxonomicNameLabel id="A23257C8EF00FF9BFF73F85BC82EE5A9" box="[188,277,1988,2014]" pageId="37" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
, distribution map. Legend and scale bar included in the top and bottom right corners, respectively.
</paragraph>
</caption>
<subSubSection id="C36F652AEF03FF99FF72FE92CC4CE761" lastPageId="39" lastPageNumber="259" pageId="38" pageNumber="258" type="key">
<paragraph id="8BCA36A1EF03FF98FF72FE92CC69E35D" blockId="38.[189,1362,269,298]" box="[189,1362,269,298]" pageId="38" pageNumber="258">
<emphasis id="B901EAB3EF03FF98FF72FE92CC69E35D" bold="true" box="[189,1362,269,298]" italics="true" pageId="38" pageNumber="258">
Identification key to species of 
<taxonomicName id="4C754D22EF03FF98FD9CFE91CAE4E35D" authority="Burmeister, 1846" authorityName="Burmeister" authorityYear="1846" box="[595,991,270,298]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="258" phylum="Arthropoda" rank="genus">
Homocopris 
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</taxonomicName>
and 
<taxonomicName id="4C754D22EF03FF98FBEEFE92CDE8E35D" authority="Génier &amp; Darling, 2024" authorityName="Génier &amp; Darling" authorityYear="2024" box="[1057,1235,269,298]" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="258" phylum="Arthropoda" rank="genus" status="gen. nov.">Andinocopris</taxonomicName>
<taxonomicNameLabel id="A23257C8EF03FF98FB14FE91CC69E35D" box="[1243,1362,270,298]" pageId="38" pageNumber="258" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<key id="FFE46190EF03FF99FF72FEDDCC4CE761" lastPageId="39" lastPageNumber="259" pageId="38" pageNumber="258">
<keyStep id="30812E04EF03FF98FF72FEDDCC4CE0A8" pageId="38" pageNumber="258">
<paragraph id="8BCA36A1EF03FF98FF72FEDDCC4CE07B" blockId="38.[189,1399,322,736]" pageId="38" pageNumber="258">
<keyLead id="30849594EF03FF98FF72FEDDCC4CE07B" pageId="38" pageNumber="258">
1 Dorsal ocular width greater than one quarter of interocular distance; lateral pronotal carina and pronotal edge joined anteriorly and posteriorly, forming a closed ellipse in lateral view (
<figureCitation id="134E2A24EF03FF98FAC2FEFAC831E3D5" captionStart="Figs13–16" captionStartId="14.[189,242,1891,1917]" captionTargetBox="[221,1346,305,1841]" captionTargetId="figure-12@14.[221,1366,263,1841]" captionTargetPageId="14" captionText="Figs13–16.Headandpronotum,lateralview.13. Andinocoprisachamas(Harold,1867)gen.etcomb.nov., lectotype, ♂ (MNHN). 14. Andinocopris achamas, paralectotype, ♀ (MNHN). 15. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 16. Andinocopris buckleyi, specimen ♀ (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.10824978" httpUri="https://zenodo.org/record/10824978/files/figure.png" pageId="38" pageNumber="258">Figs 13– 16</figureCitation>
); meso-metasternal suture posteriorly arcuate between mesocoxae (
<figureCitation id="134E2A24EF03FF98FBDAFE17CD51E3D5" box="[1045,1130,392,418]" captionStart="Figs 58–64. 58" captionStartId="31.[189,242,1857,1883]" captionTargetBox="[234,1353,267,1780]" captionTargetId="figure-17@31.[232,1356,265,1815]" captionTargetPageId="31" captionText="Figs 58–64. 58. Copris victorini Boheman, 1857, pronotum oblique view. 59. Sulcophanaeus carnifex (Linnaeus, 1758), pronotum oblique view. 60. Andinocopris achamas (Harold, 1867) gen. et comb. nov., meso-metasternal suture, ventral view.61. Homocopris torulosus(Eschscholtz, 1822), mesometasternal suture, ventral view. 62. A. achamas, left elytron, oblique view. 63. H. torulosus, elytron underside. 64. Sulcophanaeus carnifex, hindwing." figureDoi="http://doi.org/10.5281/zenodo.10824994" httpUri="https://zenodo.org/record/10824994/files/figure.png" pageId="38" pageNumber="258">Fig. 60</figureCitation>
). Metasternum broadly flat (
<figureCitation id="134E2A24EF03FF98FED2FE33C84DE3B1" box="[285,374,428,454]" captionStart="Figs 42–45" captionStartId="25.[189,242,1822,1848]" captionTargetBox="[232,1356,285,1784]" captionTargetId="figure-18@25.[232,1356,265,1787]" captionTargetPageId="25" captionText="Figs 42–45. Metasternum, ventral view. In specimens with a metasternal depression, the lateral depression is bordered with white dotted lines. For Homocopris grossiorum Darling &amp; Génier sp. nov., a red dotted line indicates where the metasternal depression is interrupted. 42. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♀ (MNHN) 43. Homocopris grossiorum, holotype ♂ (MZSP). 44. Homocopris punctatissimus (Curtis, 1845), specimen ♀ (NMPC). 45. Homocopris torulosus (Eschscholtz, 1822), specimen ♂ (CMNC)." figureDoi="http://doi.org/10.5281/zenodo.10824990" httpUri="https://zenodo.org/record/10824990/files/figure.png" pageId="38" pageNumber="258">Fig. 42</figureCitation>
), some individuals with a narrow and shallow longitudinal sulcus. 
<specimenCount id="9D73FD28EF03FF98FBB2FE33CDA8E3B1" box="[1149,1171,428,454]" count="1" pageId="38" pageNumber="258" type="male">♂</specimenCount>
pronotal armament forming an overhanging ridge or bifurcating projection in large individuals (
<figureCitation id="134E2A24EF03FF98FBAEFE50CDDDE39D" box="[1121,1254,463,490]" captionStart="Figs13–16" captionStartId="14.[189,242,1891,1917]" captionTargetBox="[221,1346,305,1841]" captionTargetId="figure-12@14.[221,1366,263,1841]" captionTargetPageId="14" captionText="Figs13–16.Headandpronotum,lateralview.13. Andinocoprisachamas(Harold,1867)gen.etcomb.nov., lectotype, ♂ (MNHN). 14. Andinocopris achamas, paralectotype, ♀ (MNHN). 15. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 16. Andinocopris buckleyi, specimen ♀ (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.10824978" httpUri="https://zenodo.org/record/10824978/files/figure.png" pageId="38" pageNumber="258">Figs 13, 15</figureCitation>
). 
<collectingCountry id="F3627631EF03FF98FB34FE50CC49E39E" box="[1275,1394,463,489]" name="Colombia" pageId="38" pageNumber="258">Colombia</collectingCountry>
, 
<collectingCountry id="F3627631EF03FF98FF25FE6DC875E07B" box="[234,334,498,524]" name="Ecuador" pageId="38" pageNumber="258">Ecuador</collectingCountry>
, 
<collectingCountry id="F3627631EF03FF98FE96FE6CC8B4E07A" box="[345,399,499,525]" name="Peru" pageId="38" pageNumber="258">Peru</collectingCountry>
.............................................................................................. 
<taxonomicName id="4C754D22EF03FF98FB83FE6ECDD4E07B" authority="Génier &amp; Darling, 2024" authorityName="Génier &amp; Darling" authorityYear="2024" box="[1100,1263,497,524]" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="258" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B901EAB3EF03FF98FB83FE6ECDD4E07B" bold="true" box="[1100,1263,497,524]" italics="true" pageId="38" pageNumber="258">Andinocopris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257C8EF03FF98FB39FE6CCC5AE07A" box="[1270,1377,499,525]" pageId="38" pageNumber="258" rank="genus">gen. nov.</taxonomicNameLabel>
2
</keyLead>
</paragraph>
<paragraph id="8BCA36A1EF03FF98FF0EFD89CC4CE0A8" blockId="38.[189,1399,322,736]" pageId="38" pageNumber="258">
<keyLead id="30849594EF03FF98FF0EFD89CC4CE0A8" pageId="38" pageNumber="258">
– Dorsal ocular width approximately one sixth of interocular distance; lateral pronotal carina and pronotal edge only joined anteriorly, forming an open ellipse in lateral view (
<figureCitation id="134E2A24EF03FF98FB43FDA7CC23E024" box="[1164,1304,568,595]" captionStart-0="Figs 17–20" captionStart-1="Figs 21–24" captionStartId-0="15.[189,242,1928,1954]" captionStartId-1="16.[189,242,1928,1954]" captionTargetBox-0="[208,1379,304,1881]" captionTargetBox-1="[210,1378,289,1875]" captionTargetId-0="figure-18@15.[208,1380,265,1881]" captionTargetId-1="figure-12@16.[210,1378,265,1875]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Figs 17–20. Head and pronotum, lateral view. 17. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 18. Homocopris grossiorum, allotype, ♀ (CEMT). 19. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 20. Homocopris punctatissimus, specimen ♀ (BNPC)." captionText-1="Figs 21–24. Head and pronotum, lateral view.21. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 22. Homocopris torulosus, specimen ♀ (repository unknown). 23. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 24. Homocopris williami, allotype, ♀ (MZSP)." figureDoi-0="http://doi.org/10.5281/zenodo.10824980" figureDoi-1="http://doi.org/10.5281/zenodo.10824982" httpUri-0="https://zenodo.org/record/10824980/files/figure.png" httpUri-1="https://zenodo.org/record/10824982/files/figure.png" pageId="38" pageNumber="258">Figs 17–24</figureCitation>
); meso-metasternal suture approximately straight between median coxae (
<figureCitation id="134E2A24EF03FF98FBDEFDC3CD57E002" box="[1041,1132,603,630]" captionStart="Figs 58–64. 58" captionStartId="31.[189,242,1857,1883]" captionTargetBox="[234,1353,267,1780]" captionTargetId="figure-17@31.[232,1356,265,1815]" captionTargetPageId="31" captionText="Figs 58–64. 58. Copris victorini Boheman, 1857, pronotum oblique view. 59. Sulcophanaeus carnifex (Linnaeus, 1758), pronotum oblique view. 60. Andinocopris achamas (Harold, 1867) gen. et comb. nov., meso-metasternal suture, ventral view.61. Homocopris torulosus(Eschscholtz, 1822), mesometasternal suture, ventral view. 62. A. achamas, left elytron, oblique view. 63. H. torulosus, elytron underside. 64. Sulcophanaeus carnifex, hindwing." figureDoi="http://doi.org/10.5281/zenodo.10824994" httpUri="https://zenodo.org/record/10824994/files/figure.png" pageId="38" pageNumber="258">Fig. 61</figureCitation>
); metasternum with a large median depression (
<figureCitation id="134E2A24EF03FF98FDD0FDE0CB97E0ED" box="[543,684,639,666]" captionStart="Figs 42–45" captionStartId="25.[189,242,1822,1848]" captionTargetBox="[232,1356,285,1784]" captionTargetId="figure-18@25.[232,1356,265,1787]" captionTargetPageId="25" captionText="Figs 42–45. Metasternum, ventral view. In specimens with a metasternal depression, the lateral depression is bordered with white dotted lines. For Homocopris grossiorum Darling &amp; Génier sp. nov., a red dotted line indicates where the metasternal depression is interrupted. 42. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♀ (MNHN) 43. Homocopris grossiorum, holotype ♂ (MZSP). 44. Homocopris punctatissimus (Curtis, 1845), specimen ♀ (NMPC). 45. Homocopris torulosus (Eschscholtz, 1822), specimen ♂ (CMNC)." figureDoi="http://doi.org/10.5281/zenodo.10824990" httpUri="https://zenodo.org/record/10824990/files/figure.png" pageId="38" pageNumber="258">Figs 43–45</figureCitation>
). 
<specimenCount id="9D73FD28EF03FF98FD0CFDE0CBE2E0EE" box="[707,729,639,665]" count="1" pageId="38" pageNumber="258" type="male">♂</specimenCount>
pronotal armament forming bilaterally paired tubercles along an inclined ridge in large individuals (
<figureCitation id="134E2A24EF03FF98FD3EFD3DCA60E0CA" box="[753,859,674,701]" captionStart="Figs 5–8" captionStartId="10.[189,242,1928,1954]" captionTargetBox="[215,1383,296,1868]" captionTargetId="figure-12@10.[205,1383,265,1889]" captionTargetPageId="10" captionText="Figs 5–8. Dorsal habitus. 5. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 6. Homocopris grossiorum, allotype, ♀ (CEMT). 7. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 8. Homocopris punctatissimus, specimen ♀ (NMPC)." figureDoi="http://doi.org/10.5281/zenodo.10824972" httpUri="https://zenodo.org/record/10824972/files/figure.png" pageId="38" pageNumber="258">Figs 5, 7</figureCitation>
, 
<figureCitation id="134E2A24EF03FF98FCA8FD3DCA98E0CB" box="[871,931,674,700]" captionStart="Figs 9–12" captionStartId="11.[189,242,1928,1954]" captionTargetBox="[207,1370,321,1842]" captionTargetId="figure-18@11.[207,1378,265,1880]" captionTargetPageId="11" captionText="Figs 9–12. Dorsal habitus. 9. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 10. Homocopris torulosus, specimen ♀ (repository unknown). 11. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 12. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824974" httpUri="https://zenodo.org/record/10824974/files/figure.png" pageId="38" pageNumber="258">9, 11</figureCitation>
). 
<collectingCountry id="F3627631EF03FF98FC76FD3DCAC2E0CB" box="[953,1017,674,700]" name="Chile" pageId="38" pageNumber="258">Chile</collectingCountry>
, 
<collectingCountry id="F3627631EF03FF98FBCAFD3DCD46E0CB" box="[1029,1149,674,700]" name="Argentina" pageId="38" pageNumber="258">Argentina</collectingCountry>
, 
<collectingCountry id="F3627631EF03FF98FB46FD3DCDEAE0CB" box="[1161,1233,674,700]" name="Brazil" pageId="38" pageNumber="258">Brazil</collectingCountry>
<emphasis id="B901EAB3EF03FF98FB12FD3CCAC2E0A8" italics="true" pageId="38" pageNumber="258">.................... ..........................................................................................................</emphasis>
<taxonomicName id="4C754D22EF03FF98FC35FD5ACC5AE0A8" ID-CoL="62V95" authority="Burmeister, 1846" authorityName="Burmeister" authorityYear="1846" box="[1018,1377,709,736]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B901EAB3EF03FF98FC35FD5ACDB5E0A8" bold="true" box="[1018,1166,709,735]" italics="true" pageId="38" pageNumber="258">Homocopris</emphasis>
<bibRefCitation id="EFE44B50EF03FF98FB59FD5ACC5AE0A8" author="Burmeister H." box="[1174,1377,709,736]" pageId="38" pageNumber="258" refId="ref24974" refString="Burmeister H. 1846. Genera quaedam insectorum, Vol. 1 (10). A. Burmeister, Berlin [Berolini]. https: // doi. org / 10.5962 / bhl. title. 8144" type="book" year="1846">Burmeister, 1846</bibRefCitation>
</taxonomicName>
3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812E04EF03FF98FF72FC98CC4CE1FD" pageId="38" pageNumber="258">
<paragraph id="8BCA36A1EF03FF98FF72FC98CC4CE133" blockId="38.[189,1399,775,907]" pageId="38" pageNumber="258">
<keyLead id="30849594EF03FF98FF72FC98CC4CE133" pageId="38" pageNumber="258">
2. Elytral interstriae flat (
<figureCitation id="134E2A24EF03FF98FE32FC98CB63E156" box="[509,600,775,801]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling &amp; Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="38" pageNumber="258">Fig. 46</figureCitation>
). Fifth abdominal ventrite with dense setigerous punctation. Body length: 
<quantity id="4C8D9B44EF03FF98FE8CFCB5C8FEE133" box="[323,453,810,836]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.4" metricValueMin="2.0" pageId="38" pageNumber="258" unit="mm" value="27.0" valueMax="34.0" valueMin="20.0">20–34 mm</quantity>
. 
<collectingCountry id="F3627631EF03FF98FE1FFCB5CB7CE133" box="[464,583,810,836]" name="Colombia" pageId="38" pageNumber="258">Colombia</collectingCountry>
, 
<collectingCountry id="F3627631EF03FF98FD9CFCB5CB8EE133" box="[595,693,810,836]" name="Ecuador" pageId="38" pageNumber="258">Ecuador</collectingCountry>
(
<collectingRegion id="49B1F843EF03FF98FD0BFCB5CA2DE133" box="[708,790,810,836]" country="Ecuador" name="Carchi" pageId="38" pageNumber="258">Carchi</collectingRegion>
) .................................... 
<taxonomicName id="4C754D22EF03FF98FBFCFCB5CC4CE133" authority="(Harold, 1867)" authorityName="Génier &amp; Darling" authorityYear="2024" baseAuthorityName="Harold" baseAuthorityYear="1867" box="[1075,1399,809,836]" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="258" phylum="Arthropoda" rank="species" species="achamas">
<emphasis id="B901EAB3EF03FF98FBFCFCB5CDFAE133" bold="true" box="[1075,1217,809,836]" italics="true" pageId="38" pageNumber="258">A. achamas</emphasis>
(
<bibRefCitation id="EFE44B50EF03FF98FB1FFCB5CC56E133" author="Harold E. von" box="[1232,1389,810,836]" pageId="38" pageNumber="258" pagination="95 - 104" refId="ref26084" refString="Harold E. von 1867. Diagnosen neuer Coprophagen. Coleopterologische Hefte 2: 95 - 104. Available from https: // www. biodiversitylibrary. org / page / 16069770 [accessed 6 Feb. 2024]." type="journal article" year="1867">Harold, 1867</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA36A1EF03FF98FF0EFCD1CC4CE1FD" blockId="38.[189,1399,775,907]" pageId="38" pageNumber="258">
<keyLead id="30849594EF03FF98FF0EFCD1CC4CE1FD" pageId="38" pageNumber="258">
– Elytral interstriae slightly convex. Fifth abdominal ventrite smooth. Body length: 
<quantity id="4C8D9B44EF03FF98FB25FCD2CC49E11F" box="[1258,1394,845,872]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="38" pageNumber="258" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
. 
<collectingCountry id="F3627631EF03FF98FF25FCEFC875E1FD" box="[234,334,880,906]" name="Ecuador" pageId="38" pageNumber="258">Ecuador</collectingCountry>
, 
<collectingCountry id="F3627631EF03FF98FE95FCEEC8ABE1FC" box="[346,400,881,907]" name="Peru" pageId="38" pageNumber="258">Peru</collectingCountry>
.................................................................................... 
<taxonomicName id="4C754D22EF03FF98FBCEFCEFCC4CE1FD" authority="(Waterhouse, 1891)" authorityName="Génier &amp; Darling" authorityYear="2024" baseAuthorityName="Waterhouse" baseAuthorityYear="1891" box="[1025,1399,880,907]" class="Insecta" family="Scarabaeidae" genus="Andinocopris" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="258" phylum="Arthropoda" rank="species" species="buckleyi">
<emphasis id="B901EAB3EF03FF98FBCEFCEFCDB3E1FC" bold="true" box="[1025,1160,880,907]" italics="true" pageId="38" pageNumber="258">A. buckleyi</emphasis>
(
<bibRefCitation id="EFE44B50EF03FF98FB58FCEFCC56E1FD" author="Waterhouse C. O." box="[1175,1389,880,907]" pageId="38" pageNumber="258" pagination="348 - 363" refId="ref28366" refString="Waterhouse C. O. 1891. New Scarabaeidae in the British Museum: a fourth contribution. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology. Sixth Series 7 (40): 348 - 363. https: // doi. org / 10.1080 / 00222939109460622" type="journal article" year="1891">Waterhouse, 1891</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF0A6629EF03FF98FF72F834CC08E59F" ID-DOI="http://doi.org/10.5281/zenodo.10825002" ID-Zenodo-Dep="10825002" httpUri="https://zenodo.org/record/10825002/files/figure.png" pageId="38" pageNumber="258" startId="38.[189,232,1963,1989]" targetBox="[189,1398,1007,1914]" targetPageId="38">
<paragraph id="8BCA36A1EF03FF98FF72F834CC08E59F" blockId="38.[189,1398,1963,2025]" pageId="38" pageNumber="258">
<emphasis id="B901EAB3EF03FF98FF72F834C82DE5B2" bold="true" box="[189,278,1963,1989]" pageId="38" pageNumber="258">Fig. 67</emphasis>
. 
<taxonomicName id="4C754D22EF03FF98FEE7F833CA12E5B1" authority="(Curtis, 1845)" baseAuthorityName="Curtis" baseAuthorityYear="1845" box="[296,809,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="258" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis id="B901EAB3EF03FF98FEE7F833CB49E5B2" box="[296,626,1964,1989]" italics="true" pageId="38" pageNumber="258">Homocopris punctatissimus</emphasis>
(
<bibRefCitation id="EFE44B50EF03FF98FD49F834CA1BE5B1" author="Curtis J." box="[646,800,1963,1990]" pageId="38" pageNumber="258" pagination="441 - 475" refId="ref25304" refString="Curtis J. 1845. Descriptions etc. of the insects collected by Captain P. P. King, R. N. F. R. S. &amp; L. S. in the survey of the Straits of Magellan. The Transactions of the Linnean Society of London 19: 441 - 475. https: // doi. org / 10.1111 / j. 1096 - 3642.1842. tb 00371. x" type="journal article" year="1845">Curtis, 1845</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="4C754D22EF03FF98FCA1F833CC54E5B2" authority="(Eschscholtz, 1822)" baseAuthorityName="Eschscholtz" baseAuthorityYear="1822" box="[878,1391,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="258" phylum="Arthropoda" rank="species" species="torulosus">
<emphasis id="B901EAB3EF03FF98FCA1F833CD4CE5B2" box="[878,1143,1963,1989]" italics="true" pageId="38" pageNumber="258">Homocopris torulosus</emphasis>
(
<bibRefCitation id="EFE44B50EF03FF98FB45F834CC5CE5B2" author="Eschscholtz J. F. G. von" box="[1162,1383,1963,1989]" pageId="38" pageNumber="258" refId="ref25539" refString="Eschscholtz J. F. G. von 1822. Beschreibungen neuer Insectens. Entomographien. Erster Lieferung, G. Reimer, Berlin. https: // doi. org / 10.1515 / 9783112513941" type="book" year="1822">Eschscholtz, 1822</bibRefCitation>
)
</taxonomicName>
, distribution map. Legend and scale bar included in the top and bottom right corners, respectively.
</paragraph>
</caption>
<keyStep id="30812E04EF02FF99FF72FE96CC4CE06D" pageId="39" pageNumber="259">
<paragraph id="8BCA36A1EF02FF99FF72FE96CC4CE3FA" blockId="39.[189,1399,265,538]" pageId="39" pageNumber="259">
<keyLead id="30849594EF02FF99FF72FE96CC4CE3FA" pageId="39" pageNumber="259">
3. Posterior pronotal surface uniformly punctate. Metasternal impression divided anteriorly (
<figureCitation id="134E2A24EF02FF99FADFFE96CC52E354" box="[1296,1385,265,291]" captionStart="Figs 42–45" captionStartId="25.[189,242,1822,1848]" captionTargetBox="[232,1356,285,1784]" captionTargetId="figure-18@25.[232,1356,265,1787]" captionTargetPageId="25" captionText="Figs 42–45. Metasternum, ventral view. In specimens with a metasternal depression, the lateral depression is bordered with white dotted lines. For Homocopris grossiorum Darling &amp; Génier sp. nov., a red dotted line indicates where the metasternal depression is interrupted. 42. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♀ (MNHN) 43. Homocopris grossiorum, holotype ♂ (MZSP). 44. Homocopris punctatissimus (Curtis, 1845), specimen ♀ (NMPC). 45. Homocopris torulosus (Eschscholtz, 1822), specimen ♂ (CMNC)." figureDoi="http://doi.org/10.5281/zenodo.10824990" httpUri="https://zenodo.org/record/10824990/files/figure.png" pageId="39" pageNumber="259">Fig. 43</figureCitation>
). 
<specimenCount id="9D73FD28EF02FF99FF24FEB2C83AE330" box="[235,257,301,327]" count="1" pageId="39" pageNumber="259" type="male">♂</specimenCount>
anterior inclined pronotal surface with fine, transverse rugosities (
<figureCitation id="134E2A24EF02FF99FBE8FEB3CDBAE331" box="[1063,1153,300,326]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling &amp; Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="39" pageNumber="259">Fig. 48</figureCitation>
); one or two sets of bilaterally paired tubercles transversely spread along pronotal ridge in large individuals (
<figureCitation id="134E2A24EF02FF99FB3EFED0CC07E31D" box="[1265,1340,335,362]" captionStart="Figs 5–8" captionStartId="10.[189,242,1928,1954]" captionTargetBox="[215,1383,296,1868]" captionTargetId="figure-12@10.[205,1383,265,1889]" captionTargetPageId="10" captionText="Figs 5–8. Dorsal habitus. 5. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 6. Homocopris grossiorum, allotype, ♀ (CEMT). 7. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 8. Homocopris punctatissimus, specimen ♀ (NMPC)." figureDoi="http://doi.org/10.5281/zenodo.10824972" httpUri="https://zenodo.org/record/10824972/files/figure.png" pageId="39" pageNumber="259">Figs 5</figureCitation>
, 
<figureCitation id="134E2A24EF02FF99FA87FED0CC5CE31E" box="[1352,1383,335,361]" captionStart="Figs 9–12" captionStartId="11.[189,242,1928,1954]" captionTargetBox="[207,1370,321,1842]" captionTargetId="figure-18@11.[207,1378,265,1880]" captionTargetPageId="11" captionText="Figs 9–12. Dorsal habitus. 9. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 10. Homocopris torulosus, specimen ♀ (repository unknown). 11. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 12. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824974" httpUri="https://zenodo.org/record/10824974/files/figure.png" pageId="39" pageNumber="259">11</figureCitation>
); paramere apex broad, rounded (
<figureCitation id="134E2A24EF02FF99FD91FEEDCBDDE3FA" box="[606,742,370,397]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="39" pageNumber="259">Figs 27, 30</figureCitation>
). 
<collectingCountry id="F3627631EF02FF99FD34FEEDCA78E3FB" box="[763,835,370,396]" name="Brazil" pageId="39" pageNumber="259">Brazil</collectingCountry>
........................................................................ 4
</keyLead>
</paragraph>
<paragraph id="8BCA36A1EF02FF99FF0EFE09CC4CE06D" blockId="39.[189,1399,265,538]" pageId="39" pageNumber="259">
<keyLead id="30849594EF02FF99FF0EFE09CC4CE06D" pageId="39" pageNumber="259">
– Posterior pronotal surface unevenly punctate-rugulate (
<figureCitation id="134E2A24EF02FF99FC5DFE09CAD4E3C7" box="[914,1007,406,432]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling &amp; Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="39" pageNumber="259">Fig. 49</figureCitation>
). Metasternal impression entire (
<figureCitation id="134E2A24EF02FF99FF3CFE26C845E3A3" box="[243,382,441,468]" captionStart="Figs 42–45" captionStartId="25.[189,242,1822,1848]" captionTargetBox="[232,1356,285,1784]" captionTargetId="figure-18@25.[232,1356,265,1787]" captionTargetPageId="25" captionText="Figs 42–45. Metasternum, ventral view. In specimens with a metasternal depression, the lateral depression is bordered with white dotted lines. For Homocopris grossiorum Darling &amp; Génier sp. nov., a red dotted line indicates where the metasternal depression is interrupted. 42. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♀ (MNHN) 43. Homocopris grossiorum, holotype ♂ (MZSP). 44. Homocopris punctatissimus (Curtis, 1845), specimen ♀ (NMPC). 45. Homocopris torulosus (Eschscholtz, 1822), specimen ♂ (CMNC)." figureDoi="http://doi.org/10.5281/zenodo.10824990" httpUri="https://zenodo.org/record/10824990/files/figure.png" pageId="39" pageNumber="259">Figs 44–45</figureCitation>
). 
<specimenCount id="9D73FD28EF02FF99FE5BFE26C891E3A4" box="[404,426,441,467]" count="1" pageId="39" pageNumber="259" type="male">♂</specimenCount>
anterior inclined pronotal surface with irregular, fused rugosities (
<figureCitation id="134E2A24EF02FF99FB05FE26CC1FE3A4" box="[1226,1316,441,467]" captionStart="Figs 46–57. 46–49" captionStartId="29.[189,242,1787,1813]" captionTargetBox="[278,1306,298,1712]" captionTargetId="figure-17@29.[258,1330,265,1743]" captionTargetPageId="29" captionText="Figs 46–57. 46–49. Elytral striae, posterolateral view. 46. Andinocopris achamas (Harold, 1867) gen. et comb. nov., lectotype, ♂ (MNHN). 47. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). Anterior pronotal surface. 48. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 49. Homocopris torulosus, neotype, ♂ (CMNC). 50–57. Type specimen labels. 50. Pinotus achamas Harold, 1867. 51. Pinotus buckleyi (Waterhouse, 1891). 52. Homocopris grossiorum. 53. Copris punctatissimus (Curtis, 1845). 54. Copris torulosus Eschscholtz, 1822. 55. Homocopris williami Darling &amp; Génier sp. nov. 56. Pinotus simulator Luederwaldt, 1936. 57. Pinotus dahli Landin, 1955." figureDoi="http://doi.org/10.5281/zenodo.10824992" httpUri="https://zenodo.org/record/10824992/files/figure.png" pageId="39" pageNumber="259">Fig. 49</figureCitation>
); three sets of bilaterally paired tubercles transversely spread along pronotal ridge in large individuals (
<figureCitation id="134E2A24EF02FF99FF3CFE60C804E06D" box="[243,319,511,538]" captionStart="Figs 5–8" captionStartId="10.[189,242,1928,1954]" captionTargetBox="[215,1383,296,1868]" captionTargetId="figure-12@10.[205,1383,265,1889]" captionTargetPageId="10" captionText="Figs 5–8. Dorsal habitus. 5. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 6. Homocopris grossiorum, allotype, ♀ (CEMT). 7. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 8. Homocopris punctatissimus, specimen ♀ (NMPC)." figureDoi="http://doi.org/10.5281/zenodo.10824972" httpUri="https://zenodo.org/record/10824972/files/figure.png" pageId="39" pageNumber="259">Figs 7</figureCitation>
, 
<figureCitation id="134E2A24EF02FF99FE84FE60C867E06E" box="[331,348,511,537]" captionStart="Figs 9–12" captionStartId="11.[189,242,1928,1954]" captionTargetBox="[207,1370,321,1842]" captionTargetId="figure-18@11.[207,1378,265,1880]" captionTargetPageId="11" captionText="Figs 9–12. Dorsal habitus. 9. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 10. Homocopris torulosus, specimen ♀ (repository unknown). 11. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 12. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824974" httpUri="https://zenodo.org/record/10824974/files/figure.png" pageId="39" pageNumber="259">9</figureCitation>
); paramere apex acutely tipped (
<figureCitation id="134E2A24EF02FF99FD14FE60CA5EE06E" box="[731,869,511,537]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="39" pageNumber="259">Figs 28–29</figureCitation>
). 
<collectingCountry id="F3627631EF02FF99FCB8FE60CAD7E06E" box="[887,1004,511,537]" name="Argentina" pageId="39" pageNumber="259">Argentina</collectingCountry>
(Río Negro), 
<collectingCountry id="F3627631EF02FF99FB5FFE60CDEBE06E" box="[1168,1232,511,537]" name="Chile" pageId="39" pageNumber="259">Chile</collectingCountry>
.................. 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812E04EF02FF99FF72FDDECC4CE102" pageId="39" pageNumber="259">
<paragraph id="8BCA36A1EF02FF99FF72FDDECC4CE09F" blockId="39.[189,1399,577,885]" pageId="39" pageNumber="259">
<keyLead id="30849594EF02FF99FF72FDDECC4CE09F" pageId="39" pageNumber="259">
4. Anterior pronotal edge tends to be medially indented. Elytral interstriae punctation tends not to be visible at 10× magnification. 
<specimenCount id="9D73FD28EF02FF99FD8CFDFACB62E008" box="[579,601,613,639]" count="1" pageId="39" pageNumber="259" type="male">♂</specimenCount>
pronotal ridge with a single medial pair of distinct tubercles in large individuals (
<figureCitation id="134E2A24EF02FF99FEB5FD18C885E0D5" box="[378,446,647,674]" captionStart="Figs 5–8" captionStartId="10.[189,242,1928,1954]" captionTargetBox="[215,1383,296,1868]" captionTargetId="figure-12@10.[205,1383,265,1889]" captionTargetPageId="10" captionText="Figs 5–8. Dorsal habitus. 5. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 6. Homocopris grossiorum, allotype, ♀ (CEMT). 7. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 8. Homocopris punctatissimus, specimen ♀ (NMPC)." figureDoi="http://doi.org/10.5281/zenodo.10824972" httpUri="https://zenodo.org/record/10824972/files/figure.png" pageId="39" pageNumber="259">Fig. 5</figureCitation>
). Paramere apex unevenly rounded, projecting dorsally (
<figureCitation id="134E2A24EF02FF99FB87FD18CDA0E0D6" box="[1096,1179,647,673]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="39" pageNumber="259">Fig. 27</figureCitation>
). FLP endophallite reduced (
<figureCitation id="134E2A24EF02FF99FE9AFD35C896E0B2" box="[341,429,682,709]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="39" pageNumber="259">Fig. 33</figureCitation>
). 
<collectingCountry id="F3627631EF02FF99FE0EFD35CB32E0B3" box="[449,521,682,708]" name="Brazil" pageId="39" pageNumber="259">Brazil</collectingCountry>
(
<collectingRegion id="49B1F843EF02FF99FDD7FD35CB80E0B3" box="[536,699,682,708]" country="Brazil" name="Minas Gerais" pageId="39" pageNumber="259">Minas Gerais</collectingRegion>
, 
<collectingRegion id="49B1F843EF02FF99FD09FD35CA48E0B3" box="[710,883,682,708]" country="Brazil" name="Rio de Janeiro" pageId="39" pageNumber="259">Rio de Janeiro</collectingRegion>
, 
<collectingRegion id="49B1F843EF02FF99FCB1FD34CACDE0B3" box="[894,1014,682,709]" country="Brazil" name="Sao Paulo" pageId="39" pageNumber="259">São Paulo</collectingRegion>
) ................................................. ........................................................................................... 
<taxonomicName id="4C754D22EF02FF99FC5FFD51CC2EE09F" authority="Darling &amp; Genier" authorityName="Darling &amp; Genier" authorityYear="2024" box="[912,1301,717,744]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="259" phylum="Arthropoda" rank="species" species="grossiorum" status="sp. nov.">
<emphasis id="B901EAB3EF02FF99FC5FFD51CD04E090" bold="true" box="[912,1087,717,744]" italics="true" pageId="39" pageNumber="259">H. grossiorum</emphasis>
Darling &amp; Génier
</taxonomicName>
<taxonomicNameLabel id="A23257C8EF02FF99FAD3FD51CC4CE09F" box="[1308,1399,718,744]" pageId="39" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BCA36A1EF02FF99FF0EFD6ECC4CE102" blockId="39.[189,1399,577,885]" pageId="39" pageNumber="259">
<keyLead id="30849594EF02FF99FF0EFD6ECC4CE102" pageId="39" pageNumber="259">
– Anterior pronotal edge evenly curved. Elytral interstriae punctation tends to be visible at 10 × magnification. 
<specimenCount id="9D73FD28EF02FF99FE23FC8ACB39E158" box="[492,514,789,815]" count="1" pageId="39" pageNumber="259" type="male">♂</specimenCount>
pronotal ridge with two pairs of distinct tubercles in large individuals (
<figureCitation id="134E2A24EF02FF99FF3CFCA8C87CE126" box="[243,327,823,849]" captionStart="Figs 9–12" captionStartId="11.[189,242,1928,1954]" captionTargetBox="[207,1370,321,1842]" captionTargetId="figure-18@11.[207,1378,265,1880]" captionTargetPageId="11" captionText="Figs 9–12. Dorsal habitus. 9. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 10. Homocopris torulosus, specimen ♀ (repository unknown). 11. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 12. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824974" httpUri="https://zenodo.org/record/10824974/files/figure.png" pageId="39" pageNumber="259">Fig. 11</figureCitation>
). Paramere apex evenly rounded (
<figureCitation id="134E2A24EF02FF99FD17FCA8CA16E126" box="[728,813,823,849]" captionStart="Figs25–30" captionStartId="19.[189,242,1857,1883]" captionTargetBox="[239,1354,353,1767]" captionTargetId="figure-17@19.[234,1354,265,1809]" captionTargetPageId="19" captionText="Figs25–30. Paramere,lateralview.25. Andinocoprisachamas(Harold,1867)gen.etcomb.nov.,specimen ♂ (CMNC). 26. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂ (MNHN). 27. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 28. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 29. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 30. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824984" httpUri="https://zenodo.org/record/10824984/files/figure.png" pageId="39" pageNumber="259">Fig. 30</figureCitation>
). FLP endophallite elongate (
<figureCitation id="134E2A24EF02FF99FB48FCA8CDE4E126" box="[1159,1247,823,849]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="39" pageNumber="259">Fig. 36</figureCitation>
). 
<collectingCountry id="F3627631EF02FF99FB3CFCA8CC00E126" box="[1267,1339,823,849]" name="Brazil" pageId="39" pageNumber="259">Brazil</collectingCountry>
(
<collectingRegion id="49B1F843EF02FF99FA83FCA8C85EE103" country="Brazil" name="Rio de Janeiro" pageId="39" pageNumber="259">Rio de Janeiro</collectingRegion>
, 
<collectingRegion id="49B1F843EF02FF99FEBEFCC4C8D1E103" box="[369,490,858,885]" country="Brazil" name="Sao Paulo" pageId="39" pageNumber="259">São Paulo</collectingRegion>
, 
<collectingRegion id="49B1F843EF02FF99FE39FCC4CB9FE103" box="[502,676,858,885]" country="Brazil" name="Santa Catarina" pageId="39" pageNumber="259">Santa Catarina</collectingRegion>
, 
<collectingRegion id="49B1F843EF02FF99FD7FFCC5CAB6E103" box="[688,909,858,884]" country="Brazil" name="Rio Grande do Sul" pageId="39" pageNumber="259">Rio Grande do Sul</collectingRegion>
) ... 
<taxonomicName id="4C754D22EF02FF99FC78FCC4CC2EE103" authority="Darling &amp; Genier" authorityName="Darling &amp; Genier" authorityYear="2024" box="[951,1301,858,885]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="259" phylum="Arthropoda" rank="species" species="williami" status="sp. nov.">
<emphasis id="B901EAB3EF02FF99FC78FCC4CD04E102" bold="true" box="[951,1087,858,885]" italics="true" pageId="39" pageNumber="259">H. williami</emphasis>
Darling &amp; Génier
</taxonomicName>
<taxonomicNameLabel id="A23257C8EF02FF99FAD3FCC4CC4CE102" box="[1308,1399,859,885]" pageId="39" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812E04EF02FF99FF72FC03CC4CE761" pageId="39" pageNumber="259">
<paragraph id="8BCA36A1EF02FF99FF72FC03CC4CE611" blockId="39.[189,1401,924,1303]" pageId="39" pageNumber="259">
<keyLead id="30849594EF02FF99FF72FC03CC4CE611" pageId="39" pageNumber="259">
5. Lateral pronotal edge angular on anterior fourth in dorsal view (
<figureCitation id="134E2A24EF02FF99FC70FC03CD0DE1C1" box="[959,1078,924,950]" captionStart="Figs 9–12" captionStartId="11.[189,242,1928,1954]" captionTargetBox="[207,1370,321,1842]" captionTargetId="figure-18@11.[207,1378,265,1880]" captionTargetPageId="11" captionText="Figs 9–12. Dorsal habitus. 9. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 10. Homocopris torulosus, specimen ♀ (repository unknown). 11. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 12. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824974" httpUri="https://zenodo.org/record/10824974/files/figure.png" pageId="39" pageNumber="259">Figs 9–10</figureCitation>
); width of ellipse formed by lateral pronotal carina and edge approximately one third the elliptical length (
<figureCitation id="134E2A24EF02FF99FB9DFC20CDEDE1AE" box="[1106,1238,959,985]" captionStart="Figs 21–24" captionStartId="16.[189,242,1928,1954]" captionTargetBox="[210,1378,289,1875]" captionTargetId="figure-12@16.[210,1378,265,1875]" captionTargetPageId="16" captionText="Figs 21–24. Head and pronotum, lateral view.21. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 22. Homocopris torulosus, specimen ♀ (repository unknown). 23. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP). 24. Homocopris williami, allotype, ♀ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824982" httpUri="https://zenodo.org/record/10824982/files/figure.png" pageId="39" pageNumber="259">Figs 21–22</figureCitation>
). Metasternal depression extending into anterior portion of median metasternal lobe (
<figureCitation id="134E2A24EF02FF99FB8CFC7DCDA5E18A" box="[1091,1182,994,1021]" captionStart="Figs 42–45" captionStartId="25.[189,242,1822,1848]" captionTargetBox="[232,1356,285,1784]" captionTargetId="figure-18@25.[232,1356,265,1787]" captionTargetPageId="25" captionText="Figs 42–45. Metasternum, ventral view. In specimens with a metasternal depression, the lateral depression is bordered with white dotted lines. For Homocopris grossiorum Darling &amp; Génier sp. nov., a red dotted line indicates where the metasternal depression is interrupted. 42. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♀ (MNHN) 43. Homocopris grossiorum, holotype ♂ (MZSP). 44. Homocopris punctatissimus (Curtis, 1845), specimen ♀ (NMPC). 45. Homocopris torulosus (Eschscholtz, 1822), specimen ♂ (CMNC)." figureDoi="http://doi.org/10.5281/zenodo.10824990" httpUri="https://zenodo.org/record/10824990/files/figure.png" pageId="39" pageNumber="259">Fig. 45</figureCitation>
). 
<specimenCount id="9D73FD28EF02FF99FB7AFC7CCDF0E18A" box="[1205,1227,995,1021]" count="1" pageId="39" pageNumber="259" type="male">♂</specimenCount>
cephalic horn length never exceeding interocular distance. FLP endophallite elongate, bent, with internal angle adjacent to sclerotized extension (
<figureCitation id="134E2A24EF02FF99FDB7FBB6CBEBE634" box="[632,720,1064,1091]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="39" pageNumber="259">Fig. 35</figureCitation>
). 
<collectingCountry id="F3627631EF02FF99FD2CFBB6CA63E634" box="[739,856,1065,1091]" name="Argentina" pageId="39" pageNumber="259">Argentina</collectingCountry>
(Río Negro), 
<collectingCountry id="F3627631EF02FF99FC32FBB6CD06E634" box="[1021,1085,1065,1091]" name="Chile" pageId="39" pageNumber="259">Chile</collectingCountry>
........................................ ........................................................................................................ 
<taxonomicName id="4C754D22EF02FF99FC3FFBD3CC4CE611" ID-CoL="3MFLK" authority="(Eschscholtz, 1822)" baseAuthorityName="Eschscholtz" baseAuthorityYear="1822" box="[1008,1399,1099,1126]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="259" phylum="Arthropoda" rank="species" species="torulosus">
<emphasis id="B901EAB3EF02FF99FC3FFBD3CDBDE611" bold="true" box="[1008,1158,1099,1126]" italics="true" pageId="39" pageNumber="259">H. torulosus</emphasis>
(
<bibRefCitation id="EFE44B50EF02FF99FB5AFBD3CC56E611" author="Eschscholtz J. F. G. von" box="[1173,1389,1100,1126]" pageId="39" pageNumber="259" refId="ref25539" refString="Eschscholtz J. F. G. von 1822. Beschreibungen neuer Insectens. Entomographien. Erster Lieferung, G. Reimer, Berlin. https: // doi. org / 10.1515 / 9783112513941" type="book" year="1822">Eschscholtz, 1822</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA36A1EF02FF99FF0EFBEFCC4CE761" blockId="39.[189,1401,924,1303]" pageId="39" pageNumber="259">
<keyLead id="30849594EF02FF99FF0EFBEFCC4CE761" pageId="39" pageNumber="259">
– Lateral pronotal edge broadly arcuate on anterior half in dorsal view (
<figureCitation id="134E2A24EF02FF99FBF9FBF0CD98E6FE" box="[1078,1187,1135,1161]" captionStart="Figs 5–8" captionStartId="10.[189,242,1928,1954]" captionTargetBox="[215,1383,296,1868]" captionTargetId="figure-12@10.[205,1383,265,1889]" captionTargetPageId="10" captionText="Figs 5–8. Dorsal habitus. 5. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 6. Homocopris grossiorum, allotype, ♀ (CEMT). 7. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 8. Homocopris punctatissimus, specimen ♀ (NMPC)." figureDoi="http://doi.org/10.5281/zenodo.10824972" httpUri="https://zenodo.org/record/10824972/files/figure.png" pageId="39" pageNumber="259">Figs 7–8</figureCitation>
); width of ellipse formed by lateral pronotal carina and edge approximately one-half the elliptical length (
<figureCitation id="134E2A24EF02FF99FAC0FB0DC832E6A7" captionStart="Figs 17–20" captionStartId="15.[189,242,1928,1954]" captionTargetBox="[208,1379,304,1881]" captionTargetId="figure-18@15.[208,1380,265,1881]" captionTargetPageId="15" captionText="Figs 17–20. Head and pronotum, lateral view. 17. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 18. Homocopris grossiorum, allotype, ♀ (CEMT). 19. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 20. Homocopris punctatissimus, specimen ♀ (BNPC)." figureDoi="http://doi.org/10.5281/zenodo.10824980" httpUri="https://zenodo.org/record/10824980/files/figure.png" pageId="39" pageNumber="259">Figs 19– 20</figureCitation>
). Metasternal depression limited to posterior portion of median metasternal lobe(
<figureCitation id="134E2A24EF02FF99FB57FB2ACDD6E6A7" box="[1176,1261,1205,1232]" captionStart="Figs 42–45" captionStartId="25.[189,242,1822,1848]" captionTargetBox="[232,1356,285,1784]" captionTargetId="figure-18@25.[232,1356,265,1787]" captionTargetPageId="25" captionText="Figs 42–45. Metasternum, ventral view. In specimens with a metasternal depression, the lateral depression is bordered with white dotted lines. For Homocopris grossiorum Darling &amp; Génier sp. nov., a red dotted line indicates where the metasternal depression is interrupted. 42. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♀ (MNHN) 43. Homocopris grossiorum, holotype ♂ (MZSP). 44. Homocopris punctatissimus (Curtis, 1845), specimen ♀ (NMPC). 45. Homocopris torulosus (Eschscholtz, 1822), specimen ♂ (CMNC)." figureDoi="http://doi.org/10.5281/zenodo.10824990" httpUri="https://zenodo.org/record/10824990/files/figure.png" pageId="39" pageNumber="259">Fig.44</figureCitation>
). 
<specimenCount id="9D73FD28EF02FF99FB33FB29CC29E6A7" box="[1276,1298,1206,1232]" count="1" pageId="39" pageNumber="259" type="male">♂</specimenCount>
cephalic horn length up to twice the interocular distance in large individuals; FLP endophallite crescent-shaped, with one end rounded and the other toothed (
<figureCitation id="134E2A24EF02FF99FD39FB63CA75E761" box="[758,846,1276,1302]" captionStart="Figs 31–36" captionStartId="21.[189,242,1857,1883]" captionTargetBox="[263,1329,282,1811]" captionTargetId="figure-18@21.[233,1354,265,1811]" captionTargetPageId="21" captionText="Figs 31–36. FLP endophallite, dorsal view. 31. Andinocopris achamas (Harold, 1867) gen. et comb. nov., specimen ♂. 32. Andinocopris buckleyi (Waterhouse, 1891) gen. et comb. nov., specimen ♂. 33. Homocopris grossiorum Darling &amp; Génier sp. nov., holotype, ♂ (CEMT). 34. Homocopris punctatissimus (Curtis, 1845), specimen ♂ (CMNC). 35. Homocopris torulosus (Eschscholtz, 1822), neotype, ♂ (CMNC). 36. Homocopris williami Darling &amp; Génier sp. nov., holotype, ♂ (MZSP)." figureDoi="http://doi.org/10.5281/zenodo.10824986" httpUri="https://zenodo.org/record/10824986/files/figure.png" pageId="39" pageNumber="259">Fig. 34</figureCitation>
). 
<collectingCountry id="F3627631EF02FF99FCACFB63CA98E761" box="[867,931,1276,1302]" name="Chile" pageId="39" pageNumber="259">Chile</collectingCountry>
........ 
<taxonomicName id="4C754D22EF02FF99FC3FFB63CC4CE761" ID-CoL="BDQ5Q" authority="(Curtis, 1845)" baseAuthorityName="Curtis" baseAuthorityYear="1845" box="[1008,1399,1276,1303]" class="Insecta" family="Scarabaeidae" genus="Homocopris" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="259" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis id="B901EAB3EF02FF99FC3FFB63CDF0E761" bold="true" box="[1008,1227,1276,1302]" italics="true" pageId="39" pageNumber="259">H. punctatissimus</emphasis>
(
<bibRefCitation id="EFE44B50EF02FF99FB15FB63CC56E760" author="Curtis J." box="[1242,1389,1276,1303]" pageId="39" pageNumber="259" pagination="441 - 475" refId="ref25304" refString="Curtis J. 1845. Descriptions etc. of the insects collected by Captain P. P. King, R. N. F. R. S. &amp; L. S. in the survey of the Straits of Magellan. The Transactions of the Linnean Society of London 19: 441 - 475. https: // doi. org / 10.1111 / j. 1096 - 3642.1842. tb 00371. x" type="journal article" year="1845">Curtis, 1845</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>