213 lines
29 KiB
XML
213 lines
29 KiB
XML
<document id="738B57AF109E8BC5AA20FC90675F4600" ID="10.11646/zootaxa.4236.1.11SLASH10254" ID-CLB-Dataset="36470" ID-DOI="10.11646/zootaxa.4236.1.11" ID-GBIF-Dataset="0fa5d3ee-feb7-48c1-9271-735e5f2738de" ID-ISSN="1175-5326" ID-Zenodo-Dep="321976" ID-ZooBank="36F97777-A034-4E19-A749-912804C0928D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487757199576" checkinUser="plazi" docAuthor="Kamiński, Marcin Jan" docDate="2017" docId="03DC87A2FF8D5A65BE8A7B3FFC547611" docLanguage="en" docName="zootaxa.4236.1.11SLASH10254.pdf" docOrigin="Zootaxa 4236 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atrocrates smithi Kamiński, 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="187" masterDocId="FFE5FFDAFF8E5A61BE1D7C20FFD97104" masterDocTitle="Three new species of trigonopoid Platynotina (Tenebrionidae: Pedinini) from the Baviaanskloof Nature Reserve, South Africa" masterLastPageNumber="193" masterPageNumber="183" pageNumber="186" updateTime="1698437199826" updateUser="ExternalLinkService">
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<mods:titleInfo id="869160C898E0905EBD2B85166DB7E9F7">
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<mods:title id="23C84888A5E223D228C70214AA06BE5B">Three new species of trigonopoid Platynotina (Tenebrionidae: Pedinini) from the Baviaanskloof Nature Reserve, South Africa</mods:title>
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<mods:name id="741E55D781EAA77F500B2C968953CDF6" type="personal">
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<mods:roleTerm id="B0CBFBB2FEA81460B786C5F4DE191F4A">Author</mods:roleTerm>
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<mods:namePart id="CED3D9D8385991645C4AA1EBFC0B3BA5">Kamiński, Marcin Jan</mods:namePart>
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<mods:typeOfResource id="AF5E9778B4A34DB6CB879F53001B4BC5">text</mods:typeOfResource>
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<mods:title id="857A206333AF274980813D7CFA95FBA6">Zootaxa</mods:title>
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<mods:part id="1DE22482877306321AA353AD4EF91A4E">
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<mods:date id="799F8D467B0B47E43AD70F9421C4EE99">2017</mods:date>
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<mods:number id="8D92FEFAA567852150C2B070BC3319C1">4236</mods:number>
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<mods:number id="E6C889B3FA150A7D1342AC60E45ADE59">1</mods:number>
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<mods:identifier id="94B728EA2A08F12DA5184F9026E930AE" type="DOI">10.11646/zootaxa.4236.1.11</mods:identifier>
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<treatment id="03DC87A2FF8D5A65BE8A7B3FFC547611" ID-DOI="http://doi.org/10.5281/zenodo.5695108" ID-GBIF-Taxon="127696721" ID-Zenodo-Dep="5695108" LSID="urn:lsid:plazi:treatment:03DC87A2FF8D5A65BE8A7B3FFC547611" httpUri="http://treatment.plazi.org/id/03DC87A2FF8D5A65BE8A7B3FFC547611" lastPageId="4" lastPageNumber="187" pageId="3" pageNumber="186">
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<subSubSection id="C36F653FFF8D5A62BE8A7B3FFE5E765E" pageId="3" pageNumber="186" type="nomenclature">
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<paragraph id="8BCA36B4FF8D5A62BE8A7B3FFE12763D" blockId="3.[151,459,1823,1883]" box="[151,459,1823,1849]" pageId="3" pageNumber="186">
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<heading id="D08281D8FF8D5A62BE8A7B3FFE12763D" bold="true" box="[151,459,1823,1849]" fontSize="11" level="1" pageId="3" pageNumber="186" reason="1">
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<emphasis id="B901EAA6FF8D5A62BE8A7B3FFE12763D" bold="true" box="[151,459,1823,1849]" pageId="3" pageNumber="186">
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<taxonomicName id="4C754D37FF8D5A62BE8A7B3FFEBE763D" box="[151,359,1823,1849]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="186" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
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<emphasis id="B901EAA6FF8D5A62BE8A7B3FFEBE763D" bold="true" box="[151,359,1823,1849]" italics="true" pageId="3" pageNumber="186">Atrocrates smithi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23257DDFF8D5A62BF737B3FFE12763D" box="[366,459,1823,1849]" pageId="3" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BCA36B4FF8D5A62BE8A7B62FE5E765E" blockId="3.[151,459,1823,1883]" box="[151,391,1858,1883]" pageId="3" pageNumber="186">
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(
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<figureCitation id="134E2A31FF8D5A62BE827B62FF3C765E" box="[159,229,1858,1883]" captionStart="FIGURE 1" captionStartId="2.[151,250,1099,1121]" captionTargetBox="[150,1436,303,1062]" captionTargetId="figure@2.[150,1436,303,1074]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of the newly described species (Geelhoutbos— S. baviaanskloofensis; Doodsklip— A. smithi; Komdomo— A. kandai) and location of the Baviaanskloof Nature Reserve in relation to the surrounding towns in South Africa." httpUri="https://zenodo.org/record/321977/files/figure.png" pageId="3" pageNumber="186">Figs 1</figureCitation>
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,
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<figureCitation id="134E2A31FF8D5A62BEEE7B62FEDA765E" box="[243,259,1858,1882]" captionStart="FIGURE 2" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[275,1326,203,1878]" captionTargetId="figure@5.[261,1326,193,1878]" captionTargetPageId="5" captionText="FIGURE 2. Habitat of Schelodontes baviaanskloofensis, surroundings of the Geelhoutbos campsite (A, B): habitat of Atrocrates smithi, surroundings of the Doodsklip campsite (C, D, E). Two specimens of A. smithi dwelling in the soil (D). Specimen of A. smithi (bottom, left) and Zadenos delalandei (Mulsant et Rey, 1854) (Tenebrionidae: Melambiina) (top, right) found under a dung patch (E)." httpUri="https://zenodo.org/record/321978/files/figure.png" pageId="3" pageNumber="186">2</figureCitation>
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,
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<figureCitation id="134E2A31FF8D5A62BF127B62FEC4765E" box="[271,285,1858,1882]" captionStart="FIGURE 3" captionStartId="6.[151,250,1264,1286]" captionTargetBox="[222,1341,237,1207]" captionTargetId="figure@6.[192,1394,193,1243]" captionTargetPageId="6" captionText="FIGURE 3. Body structure of chosen representatives of Atrocrates (A – C) and Schelodontes (D – G). A. kandai, paratype (A); A. peringueyi (B); A. smithi, paratype (C); S. baviaanskloofensis, paratype (D); S. chevrolati (E); S. oblitus (F); S. omeri (G)." httpUri="https://zenodo.org/record/321979/files/figure.png" pageId="3" pageNumber="186">3</figureCitation>
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C, 4A–F)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F653FFF8D5A65BE8A7BAAFED971FC" lastPageId="4" lastPageNumber="187" pageId="3" pageNumber="186" type="materials_examined">
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<paragraph id="8BCA36B4FF8D5A65BE8A7BAAFED971FC" blockId="3.[151,1437,1930,2027]" lastBlockId="4.[151,1437,151,1813]" lastPageId="4" lastPageNumber="187" pageId="3" pageNumber="186">
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<materialsCitation id="3B1D3CE9FF8D5A62BE8A7BAAFE4176EF" ID-GBIF-Occurrence="1434655613" collectingDate="2015-11-12" country="South Africa" location="Doodsklip" pageId="3" pageNumber="186" specimenCount="2" typeStatus="holotype">
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<emphasis id="B901EAA6FF8D5A62BE8A7BAAFE9376A7" bold="true" box="[151,330,1930,1955]" pageId="3" pageNumber="186">
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<typeStatus id="54CE8816FF8D5A62BE8A7BAAFF0B76A7" box="[151,210,1930,1955]" pageId="3" pageNumber="186">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54CE8816FF8D5A62BF4B7BAAFE1876A7" box="[342,449,1930,1955]" pageId="3" pageNumber="186" type="holotype">Holotype</typeStatus>
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, preserved in 96% ethanol, male (Ditsong National Museum of Natural History in Pretoria): “
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<collectingCountry id="F3627624FF8D5A62BF0C7B8EFDF576C3" box="[273,556,1966,1991]" name="South Africa" pageId="3" pageNumber="186">Republic of South Africa</collectingCountry>
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Baviaanskloof,
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<location id="8EAA606FFF8D5A62BCFC7B8EFC8F76C3" LSID="urn:lsid:plazi:treatment:03DC87A2FF8D5A65BE8A7B3FFC547611:8EAA606FFF8D5A62BCFC7B8EFC8F76C3" box="[737,854,1966,1991]" country="South Africa" name="Doodsklip" pageId="3" pageNumber="186">Doodsklip</location>
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//
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<date id="FFCB1074FF8D5A62BD6F7B8EFC2976C3" box="[882,1008,1966,1991]" pageId="3" pageNumber="186" value="2015-11-12">
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<collectingDate id="EF8FE99CFF8D5A62BD6F7B8EFC2976C3" box="[882,1008,1966,1991]" pageId="3" pageNumber="186" value="2015-11-12">12/11/2015</collectingDate>
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</date>
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// -33.658; 24.432; under stone // Leg. Marcin Jan Kamiński”
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</materialsCitation>
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. Paratypes, pinned, male and female (Museum and Institute of Zoology, Polish Academy of Sciences), preserved in 96% ethanol, 1 males and 2 females (Museum and Institute of Zoology, Polish Academy of Sciences),
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<materialsCitation id="3B1D3CE9FF8A5A65BF087C9CFF2471FC" ID-GBIF-Occurrence="1434655610" collectingDate="2015-11-12" country="South Africa" location="Doodsklip" pageId="4" pageNumber="187" specimenCount="18" specimenCount-female="7" specimenCount-male="11" typeStatus="holotype">
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<specimenCount id="9D73FD3DFF8A5A65BF087C9CFEA771D1" box="[277,382,188,213]" pageId="4" pageNumber="187" type="male">11 males</specimenCount>
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and
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<specimenCount id="9D73FD3DFF8A5A65BFA27C9DFDE871D1" box="[447,561,188,213]" pageId="4" pageNumber="187" type="female">7 females</specimenCount>
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(Ditsong National Museum of Natural History in Pretoria): same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F653FFF8A5A65BEDA7D24FEE173C9" pageId="4" pageNumber="187" type="diagnosis">
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<paragraph id="8BCA36B4FF8A5A65BEDA7D24FF0A7380" blockId="4.[151,1437,151,1813]" pageId="4" pageNumber="187">
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<emphasis id="B901EAA6FF8A5A65BEDA7D24FE987019" bold="true" box="[199,321,260,285]" pageId="4" pageNumber="187">Diagnosis.</emphasis>
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<taxonomicName id="4C754D37FF8A5A65BF547D24FE747019" box="[329,429,260,285]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="smithi">
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<emphasis id="B901EAA6FF8A5A65BF547D24FE747019" box="[329,429,260,285]" italics="true" pageId="4" pageNumber="187">A. smithi</emphasis>
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</taxonomicName>
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resembles
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<taxonomicName id="4C754D37FF8A5A65BC307D24FD407019" box="[557,665,260,285]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="kandai">
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<emphasis id="B901EAA6FF8A5A65BC307D24FD407019" box="[557,665,260,285]" italics="true" pageId="4" pageNumber="187">A. kandai</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C754D37FF8A5A65BCCF7D24FCB07018" box="[722,873,260,284]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="peringueyi">
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<emphasis id="B901EAA6FF8A5A65BCCF7D24FCB07018" box="[722,873,260,284]" italics="true" pageId="4" pageNumber="187">A. peringueyi</emphasis>
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</taxonomicName>
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by sharing the following character combinations: medial lobe of mentum hexagonal, narrowing towards apex; sides of pronotal disc rounded; elytra with punctatesulcate intervals; elytral humeri protruding outwards; and male mesotibiae without denticles. All three abovementioned species differ from most of their congeners by having narrow protarsi in males (widened in all other
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<taxonomicName id="4C754D37FF8A5A65BE8A7DB4FED570AF" box="[151,268,404,427]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="genus">
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<emphasis id="B901EAA6FF8A5A65BE8A7DB4FED570AF" box="[151,268,404,427]" italics="true" pageId="4" pageNumber="187">Atrocrates</emphasis>
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</taxonomicName>
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species except
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<taxonomicName id="4C754D37FF8A5A65BFA47DB4FDF070A9" box="[441,553,404,429]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="galbasi">
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<emphasis id="B901EAA6FF8A5A65BFA47DB4FDF070A9" box="[441,553,404,429]" italics="true" pageId="4" pageNumber="187">A. galbasi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C754D37FF8A5A65BC427DB4FD2970A9" box="[607,752,404,429]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="matthewsi">
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<emphasis id="B901EAA6FF8A5A65BC427DB4FD2970A9" box="[607,752,404,429]" italics="true" pageId="4" pageNumber="187">A. matthewsi</emphasis>
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</taxonomicName>
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).
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<taxonomicName id="4C754D37FF8A5A65BD1B7DB4FCB370A9" box="[774,874,404,429]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="smithi">
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<emphasis id="B901EAA6FF8A5A65BD1B7DB4FCB370A9" box="[774,874,404,429]" italics="true" pageId="4" pageNumber="187">A. smithi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C754D37FF8A5A65BD837DB4FBD070A9" box="[926,1033,404,429]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="kandai">
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<emphasis id="B901EAA6FF8A5A65BD837DB4FBD070A9" box="[926,1033,404,429]" italics="true" pageId="4" pageNumber="187">A. kandai</emphasis>
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</taxonomicName>
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differ from
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<taxonomicName id="4C754D37FF8A5A65BA8C7DB4FAFE70A8" box="[1169,1319,404,428]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="peringueyi">
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<emphasis id="B901EAA6FF8A5A65BA8C7DB4FAFE70A8" box="[1169,1319,404,428]" italics="true" pageId="4" pageNumber="187">A. peringueyi</emphasis>
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</taxonomicName>
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by having more coarse and dense punctation on the pronotal disc (
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<figureCitation id="134E2A31FF8A5A65BD0B7D97FC8070D4" box="[790,857,439,464]" captionStart="FIGURE 3" captionStartId="6.[151,250,1264,1286]" captionTargetBox="[222,1341,237,1207]" captionTargetId="figure@6.[192,1394,193,1243]" captionTargetPageId="6" captionText="FIGURE 3. Body structure of chosen representatives of Atrocrates (A – C) and Schelodontes (D – G). A. kandai, paratype (A); A. peringueyi (B); A. smithi, paratype (C); S. baviaanskloofensis, paratype (D); S. chevrolati (E); S. oblitus (F); S. omeri (G)." httpUri="https://zenodo.org/record/321979/files/figure.png" pageId="4" pageNumber="187">Fig. 3</figureCitation>
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A–C); preapical indentations on the male protibiae (
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<figureCitation id="134E2A31FF8A5A65BE827DFCFF3D70F0" box="[159,228,476,501]" captionStart="FIGURE 4" captionStartId="7.[151,250,1446,1468]" captionTargetBox="[311,1276,366,1412]" captionTargetId="figure@7.[302,1285,359,1424]" captionTargetPageId="7" captionText="FIGURE 4. Morphology of the newly described species: Atrocrates smithi sp. nov., paratype (A – F) and Schelodontes baviaanskloofensis sp. nov., paratype (G – K). Head dorsal (A) and ventral (C) views, pronotum (B, G), elytra (D, F, H), intercoxal process of prosternum (I), 5 th abdominal ventrite, legs: dorsal (E 1) and ventral (E 2) views of male protibia, male metaleg (K), female protibia (paratype)." httpUri="https://zenodo.org/record/321980/files/figure.png" pageId="4" pageNumber="187">Fig. 4</figureCitation>
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E) (protibiae evenly narrowing towards apex in
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<taxonomicName id="4C754D37FF8A5A65BD0D7DFCFC7370F0" box="[784,938,476,500]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="peringueyi">
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<emphasis id="B901EAA6FF8A5A65BD0D7DFCFC7370F0" box="[784,938,476,500]" italics="true" pageId="4" pageNumber="187">A. peringueyi</emphasis>
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</taxonomicName>
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); prominent denticles on the inner sides of metafemora (
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<figureCitation id="134E2A31FF8A5A65BF337DDFFEAA731C" box="[302,371,511,536]" captionStart="FIGURE 4" captionStartId="7.[151,250,1446,1468]" captionTargetBox="[311,1276,366,1412]" captionTargetId="figure@7.[302,1285,359,1424]" captionTargetPageId="7" captionText="FIGURE 4. Morphology of the newly described species: Atrocrates smithi sp. nov., paratype (A – F) and Schelodontes baviaanskloofensis sp. nov., paratype (G – K). Head dorsal (A) and ventral (C) views, pronotum (B, G), elytra (D, F, H), intercoxal process of prosternum (I), 5 th abdominal ventrite, legs: dorsal (E 1) and ventral (E 2) views of male protibia, male metaleg (K), female protibia (paratype)." httpUri="https://zenodo.org/record/321980/files/figure.png" pageId="4" pageNumber="187">Fig. 4</figureCitation>
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E). From
|
||
<taxonomicName id="4C754D37FF8A5A65BFFF7E20FD96731C" box="[482,591,511,536]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="kandai">
|
||
<emphasis id="B901EAA6FF8A5A65BFFF7E20FD96731C" box="[482,591,511,536]" italics="true" pageId="4" pageNumber="187">A. kandai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C754D37FF8A5A65BC427E20FD1C731C" box="[607,709,511,536]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="smithi">
|
||
<emphasis id="B901EAA6FF8A5A65BC427E20FD1C731C" box="[607,709,511,536]" italics="true" pageId="4" pageNumber="187">A. smithi</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by a wider lateral border of the pronotum (width ratio of the lateral margin to the submarginal depression ca. 4.0; in
|
||
<taxonomicName id="4C754D37FF8A5A65BDE67E04FBB17339" box="[1019,1128,548,573]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="kandai">
|
||
<emphasis id="B901EAA6FF8A5A65BDE67E04FBB17339" box="[1019,1128,548,573]" italics="true" pageId="4" pageNumber="187">A. kandai</emphasis>
|
||
</taxonomicName>
|
||
ca. 2.0), obtuse intercoxal process of prosternum (angular in
|
||
<taxonomicName id="4C754D37FF8A5A65BC037E68FD537364" box="[542,650,583,608]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="kandai">
|
||
<emphasis id="B901EAA6FF8A5A65BC037E68FD537364" box="[542,650,583,608]" italics="true" pageId="4" pageNumber="187">A. kandai</emphasis>
|
||
</taxonomicName>
|
||
) and larger body size (
|
||
<taxonomicName id="4C754D37FF8A5A65BD8E7E68FC217364" box="[915,1016,583,608]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="smithi">
|
||
<emphasis id="B901EAA6FF8A5A65BD8E7E68FC217364" box="[915,1016,583,608]" italics="true" pageId="4" pageNumber="187">A. smithi</emphasis>
|
||
</taxonomicName>
|
||
= 10.0 – 11.0 mm;
|
||
<taxonomicName id="4C754D37FF8A5A65BAC17E68FA917364" box="[1244,1352,583,608]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="kandai">
|
||
<emphasis id="B901EAA6FF8A5A65BAC17E68FA917364" box="[1244,1352,583,608]" italics="true" pageId="4" pageNumber="187">A. kandai</emphasis>
|
||
</taxonomicName>
|
||
ca. 8.0 mm).
|
||
</paragraph>
|
||
<paragraph id="8BCA36B4FF8A5A65BEDA7EAFFEE173C9" blockId="4.[151,1437,151,1813]" pageId="4" pageNumber="187">
|
||
According to Iwan’s (2016) infrageneric classification concept,
|
||
<taxonomicName id="4C754D37FF8A5A65BD897EB0FC2173AC" box="[916,1016,655,680]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="smithi">
|
||
<emphasis id="B901EAA6FF8A5A65BD897EB0FC2173AC" box="[916,1016,655,680]" italics="true" pageId="4" pageNumber="187">A. smithi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C754D37FF8A5A65BA327EB0FB4373AC" box="[1071,1178,655,680]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="kandai">
|
||
<emphasis id="B901EAA6FF8A5A65BA327EB0FB4373AC" box="[1071,1178,655,680]" italics="true" pageId="4" pageNumber="187">A. kandai</emphasis>
|
||
</taxonomicName>
|
||
belong to the
|
||
<taxonomicName id="4C754D37FF8A5A65BB217EAFFA4573AC" box="[1340,1436,655,680]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="species" species="bellamyi">
|
||
<emphasis id="B901EAA6FF8A5A65BB217EAFFA4573AC" box="[1340,1436,655,680]" italics="true" pageId="4" pageNumber="187">bellamyi</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F653FFF8A5A65BEDA7EF7FE3F7781" pageId="4" pageNumber="187" type="description">
|
||
<paragraph id="8BCA36B4FF8A5A65BEDA7EF7FE0A758E" blockId="4.[151,1437,151,1813]" pageId="4" pageNumber="187">
|
||
<emphasis id="B901EAA6FF8A5A65BEDA7EF7FE8073F4" bold="true" box="[199,345,727,752]" pageId="4" pageNumber="187">Description.</emphasis>
|
||
Measurements. Body—length = 10.0–11.0 mm; width ratio elytra / pronotum ca. 1.0; length ratio elytra / pronotum ca. 1.8. Clypeus—clypeal emargination width/depth ratio ca. 16.0. Antennae—length ratio antenna / pronotum ca. 0.7; ratio length of antenna / width of 3rd antennomere ca. 10.0; length ratio of antennomere 3rd/2nd ca. 2.0; ratio length / width of 5th antennomere ca. 1.4. Maxillary palp—ratio width of maxillary palp / length of 3rd antennomere ca. 1.3. Pronotum—pronotum length / width ratio ca. 0.8; length ratio pronotum (measured in the middle) / side of pronotum (measured at the level of anterior angle) ca. 0.9. Elytra—elytra length / width ratio = 1.3–1.4. Scutellum—width ratio anterior margin of elytra / scutellum at base ca. 3.2. Metaventrite—length ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae ca. 3.1. Abdomen—width ratio process of 1st abdominal ventrite / process of metaventrite = 1.6. Legs—protibia length / width ratio = 1.9–2.0; profemur length / width ratio ca. 3.0. Tarsi—length ratio metatarsomere 1st / 2nd ca. 1.6. Male genitalia—length of parameres / basal part of aedeagus ca. 0.4; length of clavae / parameres ca. 0.6. Female genitalia—length of paraproct/all coxites ca. 1.1.
|
||
</paragraph>
|
||
<paragraph id="8BCA36B4FF8A5A65BEDA78B5FE3F7781" blockId="4.[151,1437,151,1813]" pageId="4" pageNumber="187">
|
||
Body surface bare and dull (
|
||
<figureCitation id="134E2A31FF8A5A65BC0678B5FDBB75AA" box="[539,610,1173,1198]" captionStart="FIGURE 3" captionStartId="6.[151,250,1264,1286]" captionTargetBox="[222,1341,237,1207]" captionTargetId="figure@6.[192,1394,193,1243]" captionTargetPageId="6" captionText="FIGURE 3. Body structure of chosen representatives of Atrocrates (A – C) and Schelodontes (D – G). A. kandai, paratype (A); A. peringueyi (B); A. smithi, paratype (C); S. baviaanskloofensis, paratype (D); S. chevrolati (E); S. oblitus (F); S. omeri (G)." httpUri="https://zenodo.org/record/321979/files/figure.png" pageId="4" pageNumber="187">Fig. 3</figureCitation>
|
||
C). Head with dense punctures, 1–3 diameters apart. Clypeus scarcely emarginated. Mentum widest at middle; narrowing towards apex. Submentum triangular. Prosternite medially covered with long but sparse setae. Hypomeron glabrous and impunctate. Pronotum covered with fine punctures, 1–3 diameters apart; anterior and basal borders complete; lateral border widened basally, broad (width
|
||
<date id="FFCB1074FF8A5A65BB377922FA9F741E" box="[1322,1350,1282,1306]" pageId="4" pageNumber="187">4x</date>
|
||
greater than submarginal depression width); lateral margins actuate from anterior to posterior ends. Intercoxal process of prosternum obtuse in lateral view. Scutellum triangular, with fine punctures. Elytral striae punctate-sulcate; punctures fine, 2–3 diameters apart. Intervals dull; with visible microsculpture (magnification
|
||
<date id="FFCB1074FF8A5A65BAF5794DFACE7482" box="[1256,1303,1389,1414]" pageId="4" pageNumber="187" value="1950-10">50x</date>
|
||
); punctures scarcely visible, 3–4 diameters apart. Elytral humeri protruding laterally. 5th abdominal ventrite with submarginal sulcus; with irregularly distributed punctures, 1–2 diameters apart. Pro- and mesotarsi in both sexes narrow. Female protibiae evenly widened towards apex. Male protibiae with preapical concavity on the inner side. Female metafemora simple. Inner face of male metafemora with longitudinal concavities ending with large denticles. Other leg segments without modifications. Parameres evenly narrowing towards apex; not fused basally. Ovipositor with coxites subequal in length to paraproct. Valvifers short and wide, other lobes elongate. Gonostyli situated on dorsal part of apical lobe of coxities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F653FFF8A5A65BEDA7AB0FC8177C9" pageId="4" pageNumber="187" type="etymology">
|
||
<paragraph id="8BCA36B4FF8A5A65BEDA7AB0FC8177C9" blockId="4.[151,1437,151,1813]" pageId="4" pageNumber="187">
|
||
<emphasis id="B901EAA6FF8A5A65BEDA7AB0FE9777AD" bold="true" box="[199,334,1680,1705]" pageId="4" pageNumber="187">Etymology.</emphasis>
|
||
I have named this species in honor of my friend Prof. Aaron D. Smith, specialist in
|
||
<taxonomicName id="4C754D37FF8A5A65BAE87AB0FA4177AD" box="[1269,1432,1680,1705]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="187" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
|
||
, to whom I am greatly indebted for his help during my projects.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F653FFF8A5A65BEDA7AF8FC547611" pageId="4" pageNumber="187" type="distribution">
|
||
<paragraph id="8BCA36B4FF8A5A65BEDA7AF8FC547611" blockId="4.[151,1437,151,1813]" pageId="4" pageNumber="187">
|
||
<emphasis id="B901EAA6FF8A5A65BEDA7AF8FEB977F5" bold="true" box="[199,352,1752,1777]" pageId="4" pageNumber="187">Distribution.</emphasis>
|
||
This species has been collected in the eastern part of the Baviaanskloof Nature Reserve, near the Doodsklip campsite (GPS coordinates: -33.658, 24.432) (
|
||
<figureCitation id="134E2A31FF8A5A65BD077ADCFCB87611" box="[794,865,1788,1813]" captionStart="FIGURE 1" captionStartId="2.[151,250,1099,1121]" captionTargetBox="[150,1436,303,1062]" captionTargetId="figure@2.[150,1436,303,1074]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of the newly described species (Geelhoutbos— S. baviaanskloofensis; Doodsklip— A. smithi; Komdomo— A. kandai) and location of the Baviaanskloof Nature Reserve in relation to the surrounding towns in South Africa." httpUri="https://zenodo.org/record/321977/files/figure.png" pageId="4" pageNumber="187">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="134E2A31FF8A5A65BD727ADDFC597611" box="[879,896,1789,1813]" captionStart="FIGURE 2" captionStartId="5.[151,250,1918,1940]" captionTargetBox="[275,1326,203,1878]" captionTargetId="figure@5.[261,1326,193,1878]" captionTargetPageId="5" captionText="FIGURE 2. Habitat of Schelodontes baviaanskloofensis, surroundings of the Geelhoutbos campsite (A, B): habitat of Atrocrates smithi, surroundings of the Doodsklip campsite (C, D, E). Two specimens of A. smithi dwelling in the soil (D). Specimen of A. smithi (bottom, left) and Zadenos delalandei (Mulsant et Rey, 1854) (Tenebrionidae: Melambiina) (top, right) found under a dung patch (E)." httpUri="https://zenodo.org/record/321978/files/figure.png" pageId="4" pageNumber="187">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |